treatments-xml/data/5E/46/9A/5E469A23F5D9578AA3BBEF4F046E82DF.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit &amp; Jochum, Adrienne" docDate="2023" docId="5E469A23F5D9578AA3BBEF4F046E82DF" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila tweediei Pall-Gergely &amp; Hunyadi 2023, sp. nov." docType="treatment" docUuid="9D51E177-4754-417E-8F0E-5A139A01C84B" docUuidSource="ZooBank" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
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<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunyadi, Andras</mods:namePart>
<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Alexander</mods:namePart>
<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oerstan, Aydin</mods:namePart>
<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fekete, Judit</mods:namePart>
<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary &amp; Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jochum, Adrienne</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland &amp; Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
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<mods:date>2023</mods:date>
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<mods:number>2023-02-13</mods:number>
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<mods:detail type="volume">
<mods:number>1147</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:9D51E177-4754-417E-8F0E-5A139A01C84B" httpUri="http://treatment.plazi.org/id/5E469A23F5D9578AA3BBEF4F046E82DF" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/9D51E177-4754-417E-8F0E-5A139A01C84B" authority="Páll-Gergely &amp; Hunyadi" authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila tweediei" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tweediei" status="sp. nov.">
Angustopila tweediei 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 83" captionStartId="F83" captionText="Figure 83. Angustopila tweediei Pall-Gergely &amp; Hunyadi, sp. nov. A paratype B-I holotype (HNHM 105314). Apertural (A, B), lateral (C), apical (D) and ventral (E) sides of the shell; sculpture on the protoconch (F); aperture (G); ventral (H) and frontal (I) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure83" httpUri="https://binary.pensoft.net/fig/808892" pageId="0" pageNumber="1">Fig. 83</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: Vietnam • 1 empty shell (H: 0.9 mm, D: 0.81 mm); 
<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
La Province, 10 km from centre of Mộc 
<normalizedToken originalValue="Châu">Chau</normalizedToken>
towards 
<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
La, right side of road no. 6, south of Tất Ngoằng (locality code: 2020/5); 
<geoCoordinate degrees="20" direction="north" minutes="52.58" orientation="latitude" precision="9" value="20.876333">20°52.58'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="35.34" orientation="longitude" precision="9" value="104.589">104°35.34'E</geoCoordinate>
; 715 m a.s.l.; 5 Feb. 2020; A. Hunyadi leg.; HNHM 105314.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: Vietnam • 13 adult shells; same data as for holotype; coll. HA.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
• 1 j/b shell; same data as for holotype; coll. HA.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
A medium-sized 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species with a conical (very slightly conical-globular) shell, a strong parietal tooth reaching the peristome edge, and an upper palatal tooth of comparable height.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell of normal size for the genus, higher than wide; off-white, translucent, conical to very slightly conical-globular; body whorl widest in standard apertural view; protoconch consists of 1.5 whorls with rather weak spiral striation preceding the first teleoconch whorl; teleoconch with very fine irregularly spaced, weak radial growth lines crossed by much stronger irregularly and sparsely-spaced spiral striae (ca. 11-14 on body whorl from apertural view); on both ventral and dorsal surfaces of body whorl spiral lines dominant or spiral and radial lines are of comparable strength; whorls 4-4.25, rounded or very slightly shouldered; aperture slightly oblique to shell axis from lateral view; umbilicus moderately wide; aperture reniform, sinulus relatively narrow and isolated due to the pronounced parietal and palatal teeth; peristome expanded, not reflected; parietal callus protruding, detached from penultimate whorl (aperture not adnate); parietal tooth elevated, strong, but short, reaches parietal callus, perpendicular to parietal wall; palatal tooth almost as high as parietal one, situated near peristome edge, at middle of palatal region, just opposite of parietal tooth.</paragraph>
<caption doi="10.3897/zookeys.1147.93824.figure83" httpUri="https://binary.pensoft.net/fig/808892" pageId="0" pageNumber="1" start="Figure 83" startId="F83">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 83.</emphasis>
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila tweediei" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tweediei">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila tweediei</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi, sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
paratype 
<emphasis bold="true" pageId="0" pageNumber="1">B-I</emphasis>
holotype (HNHM 105314). Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), apical (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
) and ventral (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
) sides of the shell; sculpture on the protoconch (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
); aperture (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
); ventral (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
<paragraph pageId="0" pageNumber="1">
H = 0.86-0.9; D = 0.81-0.85, H/D*100 = 102.4-111.1 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 4), RUD = 26.5-28.4 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 2).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila tweediei" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tweediei">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila tweediei</emphasis>
</taxonomicName>
sp. nov. is most similar to 
<taxonomicName lsidName="A. bathyodon" pageId="0" pageNumber="1" rank="species" species="bathyodon">
<emphasis italics="true" pageId="0" pageNumber="1">A. bathyodon</emphasis>
</taxonomicName>
sp. nov. in terms of shell size and shape, and the position of apertural barriers. However, the spire of the latter species is lower, and both parietal and palatal teeth are situated much deeper inside the aperture. 
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila reticulata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="reticulata">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila reticulata</emphasis>
</taxonomicName>
sp. nov. is similar in general shell shape and size and the formation of the apertural barriers. However, it has a thicker peristome, a protruding palatal lip anterior to the palatal tooth, a rather reticulated superficial sculpture (i.e., the radial lines are strong), and slightly wider umbilicus. The apertural denticles of 
<taxonomicName lsidName="A. huoyani" pageId="0" pageNumber="1" rank="species" species="huoyani">
<emphasis italics="true" pageId="0" pageNumber="1">A. huoyani</emphasis>
</taxonomicName>
are similar, but that species is much larger, has more whorls, and a more horizontally oriented aperture than this new species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
This new species is dedicated to and named after Michael W.F. Tweedie (1907-1993), English naturalist and malacologist, who pioneered collecting Southeast Asian land snails via sediment sampling methods (
<bibRefCitation author="Tweedie, MWF" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="1" pagination="49 - 65" refId="B61" refString="Tweedie, MWF, 1961. On certain Mollusca of the Malayan limestone hills. Bulletin of the Raffles Museum 26: 49 - 65" title="On certain Mollusca of the Malayan limestone hills." volume="26" year="1961">Tweedie 1961</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is known from the type locality only (Fig. 
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Distribution of Angustopila species." figureDoi="10.3897/zookeys.1147.93824.figure27" httpUri="https://binary.pensoft.net/fig/808836" pageId="0" pageNumber="1">27</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>