treatments-xml/data/17/46/87/174687DE4D20637F62F0E15AFD1A2782.xml

<document id="02E1D653DB0A6D98DD423030191E560A" ID-CLB-Dataset="36898" ID-DOI="10.11646/zootaxa.3765.1.5" ID-GBIF-Dataset="09a8a185-0090-4ee2-9094-0bd1beab38a3" ID-ISSN="1175-5326" ID-Zenodo-Dep="285583" ID-ZooBank="9672C31D-6D1D-4CAF-A69E-BE45AE9AB635" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460384326662" checkinUser="plazi" docAuthor="Vicente, Carlos San, Frutos, Inmaculada &amp; Cartes, Joan E." docDate="2014" docId="174687DE4D20637F62F0E15AFD1A2782" docLanguage="en" docName="zt03765p091.pdf" docOrigin="Zootaxa 3765 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Petalophthalmus papilloculatus Vicente, Frutos &amp; Cartes, 2014, sp. nov." docType="treatment" docVersion="7" lastPageNumber="88" masterDocId="EB7FFFA64D2163746267E614FF922427" masterDocTitle="Petalophthalmus papilloculatus sp. nov. (Crustacea: Mysida: Petalophthalmidae), a new bathyal suprabenthic mysid from the Galicia Bank (NE Atlantic Ocean)" masterLastPageNumber="91" masterPageNumber="77" pageNumber="78" updateTime="1698297283302" updateUser="ExternalLinkService">
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<mods:title id="064BD4D3A77073EC12C2C6AC23E96755">Petalophthalmus papilloculatus sp. nov. (Crustacea: Mysida: Petalophthalmidae), a new bathyal suprabenthic mysid from the Galicia Bank (NE Atlantic Ocean)</mods:title>
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<mods:namePart id="7097FF878E8D28C0481606ECC1B6C9C6">Vicente, Carlos San</mods:namePart>
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<mods:namePart id="D40B33B4E2B2111D36524277A6927A5B">Frutos, Inmaculada</mods:namePart>
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<taxonomicName id="58EF4D4B4D20637562F0E15AFD972340" box="[151,517,1870,1895]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="papilloculatus" status="sp. nov.">
<emphasis id="AD9BEADA4D20637562F0E15AFD972340" bold="true" box="[151,517,1870,1895]" italics="true" pageId="1" pageNumber="78">Petalophthalmus papilloculatus</emphasis>
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(
<figureCitation id="07D42A4D4D20637562F8E166FE9B23AC" box="[159,265,1906,1931]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="4.[151,250,1936,1958]" captionStartId-1="5.[151,250,1917,1939]" captionStartId-2="6.[151,250,1985,2007]" captionStartId-3="7.[151,250,1866,1888]" captionStartId-4="8.[151,250,1901,1923]" captionStartId-5="9.[151,250,1916,1938]" captionTargetBox-0="[186,1394,206,1897]" captionTargetBox-1="[206,1365,203,1881]" captionTargetBox-2="[219,1374,199,1956]" captionTargetBox-3="[215,1383,204,1835]" captionTargetBox-4="[184,1375,203,1870]" captionTargetBox-5="[193,1393,200,1901]" captionTargetId-0="figure@4.[174,1403,193,1905]" captionTargetId-1="figure@5.[190,1395,193,1898]" captionTargetId-2="figure@6.[197,1387,193,1963]" captionTargetId-3="figure@7.[196,1391,193,1845]" captionTargetId-4="figure@8.[179,1404,193,1883]" captionTargetId-5="figure@9.[179,1408,193,1911]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionText-0="FIGURE 2. Petalophthalmus papilloculatus sp. nov.: (A, B, D) holotype female MNCN 20.04 / 9201; (C, E) allotype male MNCN 20.04 / 9202. (A) habitus in lateral view, (B) anterior part of carapace with eyes and right antennule in dorsal view, (C) antennule in dorsal view, (D) antenna in ventral view, (E) antenna in ventral view." captionText-1="FIGURE 3. Petalophthalmus papilloculatus sp. nov.: (A – I) holotype female MNCN 20.04 / 9201. (A) labrum, (B) maxillule, (C) cuspidate-serrate seta on maxillule basis, (D) maxilla, (E) left mandible, (F) cuspidate seta with supracuticular pocket of the left mandible first article, (G) incisor to molar process of left mandible, (H) right mandible, (I) incisor to molar process of right mandible." captionText-2="FIGURE 4. Petalophthalmus papilloculatus sp. nov.: (A – C) holotype female MNCN 20.04 / 9201; (D, E) allotype male MNCN 20.04 / 9202. (A) first thoracopod, (B) second thoracopod, (C) third thoracopod, (D) third thoracopod, (E) fourth thoracopod." captionText-3="FIGURE 5. Petalophthalmus papilloculatus sp. nov.: (A, B) holotype female MNCN 20.04 / 9201; (C, D, F, G) allotype male MNCN 20.04 / 9202; (E) paratype female MNCN 20.04 / 9203. (A) fifth thoracopod, (B) distal article of endopod of fifth thoracopod, (C) sixth thoracopod, (D) distal article of endopod of sixth thoracopod, (E) seventh thoracopod, (F) eighth thoracopod, (G) distal article of endopod of eighth thoracopod." captionText-4="FIGURE 6. Petalophthalmus papilloculatus sp. nov.: (A – E) holotype female MNCN 20.04 / 9201; (F – I) allotype male MNCN 20.04 / 9202. (A) first pleopod, (B) second pleopod, (C) third pleopod, (D) fourth pleopod, (E) fifth pleopod, (F) first pleopod, (G) distal article of endopod of first pleopod, (H) second pleopod, (I) distal end of first article of exopod of second pleopod, (J) third pleopod, (K) fourth pleopod, (L) fifth pleopod." captionText-5="FIGURE 7. Petalophthalmus papilloculatus sp. nov.: (A – D) holotype female MNCN 20.04 / 9201; (E – G) allotype male MNCN 20.04 / 9202. (A) telson in dorsal view, (B) telson apex in dorsal view, (C) medial pair of serrate setae and three pairs of ventral small teeth on telson apex, (D) right uropod in ventral view, (E) telson in dorsal view, (F) telson apex in dorsal view, (G) medial serrate seta and pairs of ventral small teeth on telson apex." httpUri-0="https://zenodo.org/record/285585/files/figure.png" httpUri-1="https://zenodo.org/record/285586/files/figure.png" httpUri-2="https://zenodo.org/record/285587/files/figure.png" httpUri-3="https://zenodo.org/record/285588/files/figure.png" httpUri-4="https://zenodo.org/record/285589/files/figure.png" httpUri-5="https://zenodo.org/record/285590/files/figure.png" pageId="1" pageNumber="78">Figs. 2–7</figureCitation>
)
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Material examined. 
<typeStatus id="4054886A4D20637563E1E1AEFE6723F4" box="[390,501,1978,2003]" pageId="1" pageNumber="78" type="holotype">Holotype</typeStatus>
.
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1 brooding (empty marsupium) female (
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), 
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20.04/9201, Galicia Bank, RV “Miguel Oliver”, INDEMARES BANGAL0711 cruise, 
<date id="EB5110084D23637661E7E68CFBB52497" box="[896,1063,152,176]" pageId="2" pageNumber="79" value="2011-08-08">8 August 2011</date>
, GOC73 bottom trawl, haul G10, 
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, 
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depth, very fine sand bottom, temperature: 6.36ºC, salinity: 35.38; dissected, one vial.
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<emphasis id="AD9BEADA4D23637662A0E710FEA1253A" bold="true" box="[199,307,260,285]" pageId="2" pageNumber="79">Allotype.</emphasis>
1 mature male (34.0 mm 
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), 
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20.04/9202, Galicia Bank, RV “Miguel Oliver”, INDEMARES BANGAL0711 cruise, 
<date id="EB5110084D2363766024E733FD7A2567" box="[579,744,295,320]" pageId="2" pageNumber="79" value="2011-08-06">6 August 2011</date>
, beam trawl, haul 
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V8, 
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º38.48’N 
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, 
<quantity id="58179B2D4D236376669BE733FAC32518" box="[1276,1361,295,320]" metricMagnitude="3" metricUnit="m" metricValue="1.565" pageId="2" pageNumber="79" unit="m" value="1565.0">1565 m</quantity>
depth, fine sand bottom, temperature: 6.36ºC, salinity: 35.38; dissected, one vial.
</paragraph>
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<emphasis id="AD9BEADA4D23637662F0E0DDFE8522F8" bold="true" box="[151,279,1737,1759]" pageId="2" pageNumber="79">FIGURE 1.</emphasis>
Location of sampling stations where the mysid 
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<emphasis id="AD9BEADA4D236376617DE0DDFBF222F8" box="[794,1120,1737,1759]" italics="true" pageId="2" pageNumber="79">Petalophthalmus papilloculatus</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="B6A857A14D236376660DE0DDFB2C22F8" box="[1130,1214,1737,1759]" pageId="2" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
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was collected in the Galicia Bank (NE Atlantic Ocean) with various bottom trawling devices during INDEMARES BANGAL 0 711 cruise. G: GOC73 otter trawl, TS: suprabenthic sled, V: beam trawl. Isobaths expressed in metres.
</paragraph>
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<typeStatus id="4054886A4D23637662A0E150FED3237A" box="[199,321,1860,1885]" pageId="2" pageNumber="79" type="paratype">Paratypes</typeStatus>
.
</emphasis>
1 immature female (
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), 
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20.04/9203, partially dissected, one vial. 1 immature female (
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<collectionCode id="F9FEAE0D4D2363766309E17DFE0623A6" box="[366,404,1897,1921]" pageId="2" pageNumber="79">TL</collectionCode>
), 
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20.04/9204, not dissected, one vial. Both from 
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locality. 2 immature females (
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CL, 
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), 
<collectionCode id="F9FEAE0D4D2363766030E199FD392382" LSID="urn:lsid:biocol.org:col:35275" box="[599,683,1933,1957]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="2" pageNumber="79">MNCN</collectionCode>
20.04/9205, Galicia Bank, RV “Miguel Oliver”, INDEMARES BANGAL0711 cruise, 
<date id="EB5110084D23637663C7E1A5FDDE23EE" box="[416,588,1969,1993]" pageId="2" pageNumber="79" value="2011-08-08">8 August 2011</date>
, beam trawl, haul 
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V10, 
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</date>
º41.88’N 
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, 
<quantity id="58179B2D4D23637666E2E1A5FB4E23EF" box="[1157,1244,1969,1993]" metricMagnitude="3" metricUnit="m" metricValue="1.72" pageId="2" pageNumber="79" unit="m" value="1720.0">1720 m</quantity>
depth, very fine sand bottom, temperature: 6.36ºC, salinity: 35.38; not dissected, one vial. 2 immature females (
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, 
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) 
<collectionCode id="F9FEAE0D4D22637762B4E68CFF6B2488" box="[211,249,152,175]" pageId="3" pageNumber="80">TL</collectionCode>
; 
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20.04/9206, not dissected, one vial. 1 brooding female (empty marsupium) (
<quantity id="58179B2D4D22637766D7E683FA952488" box="[1200,1287,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="80" unit="mm" value="5.5">5.5 mm</quantity>
CL), 
<collectionCode id="F9FEAE0D4D226377672FE68CFA0E2497" LSID="urn:lsid:biocol.org:col:35275" box="[1352,1436,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="3" pageNumber="80">MNCN</collectionCode>
20.04/9207, not dissected, one vial. 1 mature male (
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<collectionCode id="F9FEAE0D4D2263776123E6A8FCF824F3" box="[836,874,188,212]" pageId="3" pageNumber="80">TL</collectionCode>
), 
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20.04/9208, not dissected one vial; from allotype locality. 
<specimenCount id="89E9FD414D226377633FE6F4FE5A24DF" box="[344,456,223,248]" pageId="3" pageNumber="80" type="juvenile">1 juvenile</specimenCount>
(
<quantity id="58179B2D4D22637763B0E6CBFDAE24D0" box="[471,572,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="3" pageNumber="80" unit="mm" value="14.6">14.6 mm</quantity>
<collectionCode id="F9FEAE0D4D2263776024E6F4FDFB24DF" box="[579,617,224,248]" pageId="3" pageNumber="80">TL</collectionCode>
), 
<collectionCode id="F9FEAE0D4D226377601AE6F4FD4324DF" LSID="urn:lsid:biocol.org:col:35275" box="[637,721,224,248]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="3" pageNumber="80">MNCN</collectionCode>
20.04/9209, Galicia Bank, RV “Miguel Oliver”, INDEMARES BANGAL0711 cruise, 
<date id="EB5110084D22637763FBE711FDD4253B" box="[412,582,260,284]" pageId="3" pageNumber="80" value="2011-08-07">7 August 2011</date>
, suprabenthic sled, haul TS9, 
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<geoCoordinate id="FADB500F4D2263776642E710FB3E253A" box="[1061,1196,260,285]" direction="west" orientation="longitude" pageId="3" pageNumber="80" precision="9" value="-11.989833">11º59.39’W</geoCoordinate>
, 
<quantity id="58179B2D4D22637766DFE710FA9D253B" box="[1208,1295,260,285]" metricMagnitude="3" metricUnit="m" metricValue="1.669" pageId="3" pageNumber="80" unit="m" value="1669.0">1669 m</quantity>
depth, 
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near-bottom water layer, fine sand bottom, temperature 5.65ºC, salinity: 35.27; not dissected, one vial. 
<specimenCount id="89E9FD414D22637767E9E73CFF632542" pageId="3" pageNumber="80" type="juvenile">1 juvenile</specimenCount>
(
<quantity id="58179B2D4D2263776364E758FEFE2543" box="[259,364,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="3" pageNumber="80" unit="mm" value="10.4">10.4 mm</quantity>
<collectionCode id="F9FEAE0D4D2263776312E758FE092543" box="[373,411,332,356]" pageId="3" pageNumber="80">TL</collectionCode>
), 
<collectionCode id="F9FEAE0D4D22637763D5E758FD942543" LSID="urn:lsid:biocol.org:col:35275" box="[434,518,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="3" pageNumber="80">MNCN</collectionCode>
20.04/9210, Galicia Bank, RV “Miguel Oliver”, INDEMARES BANGAL0711 cruise, 
<date id="EB5110084D226377628EE764FEEC25AF" box="[233,382,367,392]" pageId="3" pageNumber="80" value="2011-07-29">29 July 2011</date>
, suprabenthic sled, haul TS2, 
<geoCoordinate id="FADB500F4D22637760B4E77BFCC625AF" box="[723,852,367,392]" direction="north" orientation="latitude" pageId="3" pageNumber="80" precision="9" value="42.966167">42º57.97’N</geoCoordinate>
<geoCoordinate id="FADB500F4D226377613BE77BFC7125AF" box="[860,995,367,392]" direction="west" orientation="longitude" pageId="3" pageNumber="80" precision="9" value="-12.008667">12º00.52’W</geoCoordinate>
, 
<quantity id="58179B2D4D2263776197E77BFBD425A0" box="[1008,1094,367,392]" metricMagnitude="3" metricUnit="m" metricValue="1.7259999999999998" pageId="3" pageNumber="80" unit="m" value="1726.0">1726 m</quantity>
depth, 
<quantity id="58179B2D4D22637766FCE77BFA9925A0" box="[1179,1291,367,392]" metricMagnitude="-1" metricUnit="m" metricValue="7.75" metricValueMax="9.0" metricValueMin="6.5" pageId="3" pageNumber="80" unit="cm" value="77.5" valueMax="90.0" valueMin="65.0">65–90 cm</quantity>
near-bottom water layer, fine sand bottom, temperature: 5.65ºC, salinity: 35.27; not dissected, one vial.
</paragraph>
</subSubSection>
<subSubSection id="D7F565434D22637762A0E7A3FA0E25F7" box="[199,1436,439,464]" pageId="3" pageNumber="80" type="etymology">
<paragraph id="9F5036C84D22637762A0E7A3FA0E25F7" blockId="3.[151,1436,151,2037]" box="[199,1436,439,464]" pageId="3" pageNumber="80">
<emphasis id="AD9BEADA4D22637762A0E7A3FEDC25F7" bold="true" box="[199,334,439,464]" pageId="3" pageNumber="80">Etymology.</emphasis>
This species is named for the peculiar papilla located on the inner dorsomedial surface of the eyes.
</paragraph>
</subSubSection>
<subSubSection id="D7F565434D22637762A0E7C8FDE8268F" pageId="3" pageNumber="80" type="diagnosis">
<paragraph id="9F5036C84D22637762A0E7C8FDE8268F" blockId="3.[151,1436,151,2037]" pageId="3" pageNumber="80">
<emphasis id="AD9BEADA4D22637762A0E7C8FED325D2" bold="true" box="[199,321,476,501]" pageId="3" pageNumber="80">Diagnosis.</emphasis>
Anterior margin of the carapace almost truncate without rostrum. Eyes flattened and leaf-like with an ocular papilla on the dorsomedial inner surface. Antennal scale with an apical lobe. First article of mandibular palp with a small lobe on the inner distal margin. Distal outer margin of the basal segment of the uropod exopod with three cuspidate setae that increase in length from outer to inner one. Lateral margin of telson with 20–36 cuspidate setae. Telson apex armed with 3–4 pairs of serrate setae decreasing in length form the outer to the inner ones, and 3–8 pairs of ventral small spines.
</paragraph>
</subSubSection>
<subSubSection id="D7F565434D22637E62A0E4A0FD63222F" lastPageId="10" lastPageNumber="87" pageId="3" pageNumber="80" type="description">
<paragraph id="9F5036C84D22637762A0E4A0FB5B26EA" blockId="3.[151,1436,151,2037]" box="[199,1225,692,717]" pageId="3" pageNumber="80">
<emphasis id="AD9BEADA4D22637762A0E4A0FECB26EA" bold="true" box="[199,345,692,717]" pageId="3" pageNumber="80">Description.</emphasis>
The morphological characteristic refers to both sexes, unless otherwise stated.
</paragraph>
<paragraph id="9F5036C84D22637762A0E4CCFF58277B" blockId="3.[151,1436,151,2037]" pageId="3" pageNumber="80">
<emphasis id="AD9BEADA4D22637762A0E4CCFEA726C8" box="[199,309,728,751]" italics="true" pageId="3" pageNumber="80">Carapace</emphasis>
short and membranous, leaving the last three or four thoracic somites uncovered dorsally (
<figureCitation id="07D42A4D4D2263776753E4C3FAEA26D7" box="[1332,1400,727,752]" captionStart="FIGURE 2" captionStartId="4.[151,250,1936,1958]" captionTargetBox="[186,1394,206,1897]" captionTargetId="figure@4.[174,1403,193,1905]" captionTargetPageId="4" captionText="FIGURE 2. Petalophthalmus papilloculatus sp. nov.: (A, B, D) holotype female MNCN 20.04 / 9201; (C, E) allotype male MNCN 20.04 / 9202. (A) habitus in lateral view, (B) anterior part of carapace with eyes and right antennule in dorsal view, (C) antennule in dorsal view, (D) antenna in ventral view, (E) antenna in ventral view." httpUri="https://zenodo.org/record/285585/files/figure.png" pageId="3" pageNumber="80">Fig. 2</figureCitation>
A); anterior margin rectilinear and without rostral projection, lateral corners anteriorly rounded; one median acute tooth anterior to cervical sulcus; posterior margin not emarginated dorsally and without posterolateral lobes (
<figureCitation id="07D42A4D4D2263776709E50BFF34277B" captionStart="FIGURE 2" captionStartId="4.[151,250,1936,1958]" captionTargetBox="[186,1394,206,1897]" captionTargetId="figure@4.[174,1403,193,1905]" captionTargetPageId="4" captionText="FIGURE 2. Petalophthalmus papilloculatus sp. nov.: (A, B, D) holotype female MNCN 20.04 / 9201; (C, E) allotype male MNCN 20.04 / 9202. (A) habitus in lateral view, (B) anterior part of carapace with eyes and right antennule in dorsal view, (C) antennule in dorsal view, (D) antenna in ventral view, (E) antenna in ventral view." httpUri="https://zenodo.org/record/285585/files/figure.png" pageId="3" pageNumber="80">Fig. 2</figureCitation>
A).
</paragraph>
<paragraph id="9F5036C84D22637762A0E57DFE382783" blockId="3.[151,1436,151,2037]" pageId="3" pageNumber="80">
<emphasis id="AD9BEADA4D22637762A0E57DFF6E27A7" box="[199,252,873,896]" italics="true" pageId="3" pageNumber="80">Eyes</emphasis>
flattened and leaf-like (
<figureCitation id="07D42A4D4D226377606AE573FDC727A7" box="[525,597,871,896]" captionStart="FIGURE 2" captionStartId="4.[151,250,1936,1958]" captionTargetBox="[186,1394,206,1897]" captionTargetId="figure@4.[174,1403,193,1905]" captionTargetPageId="4" captionText="FIGURE 2. Petalophthalmus papilloculatus sp. nov.: (A, B, D) holotype female MNCN 20.04 / 9201; (C, E) allotype male MNCN 20.04 / 9202. (A) habitus in lateral view, (B) anterior part of carapace with eyes and right antennule in dorsal view, (C) antennule in dorsal view, (D) antenna in ventral view, (E) antenna in ventral view." httpUri="https://zenodo.org/record/285585/files/figure.png" pageId="3" pageNumber="80">Figs 2</figureCitation>
A, B), without functional visual element, with a definite ocular papilla on mesial margin (
<figureCitation id="07D42A4D4D2263776322E598FE1B2783" box="[325,393,908,933]" captionStart="FIGURE 2" captionStartId="4.[151,250,1936,1958]" captionTargetBox="[186,1394,206,1897]" captionTargetId="figure@4.[174,1403,193,1905]" captionTargetPageId="4" captionText="FIGURE 2. Petalophthalmus papilloculatus sp. nov.: (A, B, D) holotype female MNCN 20.04 / 9201; (C, E) allotype male MNCN 20.04 / 9202. (A) habitus in lateral view, (B) anterior part of carapace with eyes and right antennule in dorsal view, (C) antennule in dorsal view, (D) antenna in ventral view, (E) antenna in ventral view." httpUri="https://zenodo.org/record/285585/files/figure.png" pageId="3" pageNumber="80">Fig. 2</figureCitation>
B).
</paragraph>
<paragraph id="9F5036C84D22637762A0E5BBFDEC205A" blockId="3.[151,1436,151,2037]" pageId="3" pageNumber="80">
<emphasis id="AD9BEADA4D22637762A0E5BBFEAB27EF" box="[199,313,943,968]" italics="true" pageId="3" pageNumber="80">Antennule</emphasis>
peduncle very long and slender, longer than carapace. First article almost as long as the remaining two combined, armed on distal dorsal margin with one cuspidate seta and 2–4 pappose setae, rounded lobe on the outer proximal margin armed with two pappose setae; second article armed with 2–3 simple setae and one cuspidate seta on distal outer margin; third article shorter than second, armed with two–three dorsal pappose setae on distal margin (
<figureCitation id="07D42A4D4D226377630EE22BFE23207F" box="[361,433,1087,1112]" captionStart="FIGURE 2" captionStartId="4.[151,250,1936,1958]" captionTargetBox="[186,1394,206,1897]" captionTargetId="figure@4.[174,1403,193,1905]" captionTargetPageId="4" captionText="FIGURE 2. Petalophthalmus papilloculatus sp. nov.: (A, B, D) holotype female MNCN 20.04 / 9201; (C, E) allotype male MNCN 20.04 / 9202. (A) habitus in lateral view, (B) anterior part of carapace with eyes and right antennule in dorsal view, (C) antennule in dorsal view, (D) antenna in ventral view, (E) antenna in ventral view." httpUri="https://zenodo.org/record/285585/files/figure.png" pageId="3" pageNumber="80">Fig. 2</figureCitation>
B–C). Outer flagellum thinner than inner in females, basal portion of outer flagellum considerably thickened than inner in males.
</paragraph>
<paragraph id="9F5036C84D22637762A0E29CFB0D2175" blockId="3.[151,1436,151,2037]" pageId="3" pageNumber="80">
<emphasis id="AD9BEADA4D22637762A0E29CFEB720B8" box="[199,293,1160,1183]" italics="true" pageId="3" pageNumber="80">Antenna</emphasis>
peduncle 3-articulate, extending to 4/5 scale length (
<figureCitation id="07D42A4D4D22637761E7E293FC562087" box="[896,964,1159,1184]" captionStart="FIGURE 2" captionStartId="4.[151,250,1936,1958]" captionTargetBox="[186,1394,206,1897]" captionTargetId="figure@4.[174,1403,193,1905]" captionTargetPageId="4" captionText="FIGURE 2. Petalophthalmus papilloculatus sp. nov.: (A, B, D) holotype female MNCN 20.04 / 9201; (C, E) allotype male MNCN 20.04 / 9202. (A) habitus in lateral view, (B) anterior part of carapace with eyes and right antennule in dorsal view, (C) antennule in dorsal view, (D) antenna in ventral view, (E) antenna in ventral view." httpUri="https://zenodo.org/record/285585/files/figure.png" pageId="3" pageNumber="80">Fig. 2</figureCitation>
D–E); first article short, as long as broad, inner margin produced medially into triangular lobe; second article five times as long as broad, distal margin armed with 2–3 simple setae; third article longer than second, distal margin armed with 4 simple setae; flagellum 5- articulate in females, with more than 60 articles and longer than peduncle in males, most articles short and scarcely as long as broad. Antennal scale about 6–7 times as long as maximum width, extending slightly beyond first article of antennular peduncle; margins setose all round; apical lobe short, nearly 1/15 scale length.
</paragraph>
<paragraph id="9F5036C84D22637762A0E348FA1D2152" blockId="3.[151,1436,151,2037]" box="[199,1423,1372,1397]" pageId="3" pageNumber="80">
<emphasis id="AD9BEADA4D22637762A0E348FEB22152" box="[199,288,1372,1397]" italics="true" pageId="3" pageNumber="80">Labrum</emphasis>
more or less symmetrical, distal margin with short irregularly distributed thin simple setae (
<figureCitation id="07D42A4D4D2263776740E348FAF82152" box="[1319,1386,1372,1397]" captionStart="FIGURE 3" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[206,1365,203,1881]" captionTargetId="figure@5.[190,1395,193,1898]" captionTargetPageId="5" captionText="FIGURE 3. Petalophthalmus papilloculatus sp. nov.: (A – I) holotype female MNCN 20.04 / 9201. (A) labrum, (B) maxillule, (C) cuspidate-serrate seta on maxillule basis, (D) maxilla, (E) left mandible, (F) cuspidate seta with supracuticular pocket of the left mandible first article, (G) incisor to molar process of left mandible, (H) right mandible, (I) incisor to molar process of right mandible." httpUri="https://zenodo.org/record/285586/files/figure.png" pageId="3" pageNumber="80">Fig. 3</figureCitation>
A).
</paragraph>
<paragraph id="9F5036C84D22637762A0E394FEE72293" blockId="3.[151,1436,151,2037]" pageId="3" pageNumber="80">
<emphasis id="AD9BEADA4D22637762A0E394FEAC21BE" box="[199,318,1408,1433]" italics="true" pageId="3" pageNumber="80">Mandibles</emphasis>
with elongated and 3-articulate palp (
<figureCitation id="07D42A4D4D2263776081E394FCB921BF" box="[742,811,1408,1433]" captionStart="FIGURE 3" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[206,1365,203,1881]" captionTargetId="figure@5.[190,1395,193,1898]" captionTargetPageId="5" captionText="FIGURE 3. Petalophthalmus papilloculatus sp. nov.: (A – I) holotype female MNCN 20.04 / 9201. (A) labrum, (B) maxillule, (C) cuspidate-serrate seta on maxillule basis, (D) maxilla, (E) left mandible, (F) cuspidate seta with supracuticular pocket of the left mandible first article, (G) incisor to molar process of left mandible, (H) right mandible, (I) incisor to molar process of right mandible." httpUri="https://zenodo.org/record/285586/files/figure.png" pageId="3" pageNumber="80">Figs 3</figureCitation>
E, H); first article very small, armed with one cuspidate seta (with supracuticular pocket) on distal outer margin (
<figureCitation id="07D42A4D4D2263776148E3B7FCE6219B" box="[815,884,1443,1468]" captionStart="FIGURE 3" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[206,1365,203,1881]" captionTargetId="figure@5.[190,1395,193,1898]" captionTargetPageId="5" captionText="FIGURE 3. Petalophthalmus papilloculatus sp. nov.: (A – I) holotype female MNCN 20.04 / 9201. (A) labrum, (B) maxillule, (C) cuspidate-serrate seta on maxillule basis, (D) maxilla, (E) left mandible, (F) cuspidate seta with supracuticular pocket of the left mandible first article, (G) incisor to molar process of left mandible, (H) right mandible, (I) incisor to molar process of right mandible." httpUri="https://zenodo.org/record/285586/files/figure.png" pageId="3" pageNumber="80">Fig. 3</figureCitation>
F) and a small lobe on the inner distal margin; second article about twice as long as third with 8–9 strong simple setae and about 11 pappose short setae on inner distal margin; third article armed on inner margin with five strong simple setae on inner margin and three large conspicuous simple setae on distal margin. 
<emphasis id="AD9BEADA4D22637760EAE01AFCAE2200" box="[653,828,1550,1575]" italics="true" pageId="3" pageNumber="80">Right mandible</emphasis>
with incisor process composed of a single chitinous ridge with one broad terminal tooth and a smaller one behind it; lacinia mobilis reduced, setal row reduced to single spine, molar process with one chitinous ridge (
<figureCitation id="07D42A4D4D22637760A3E041FC9E224A" box="[708,780,1621,1646]" captionStart="FIGURE 3" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[206,1365,203,1881]" captionTargetId="figure@5.[190,1395,193,1898]" captionTargetPageId="5" captionText="FIGURE 3. Petalophthalmus papilloculatus sp. nov.: (A – I) holotype female MNCN 20.04 / 9201. (A) labrum, (B) maxillule, (C) cuspidate-serrate seta on maxillule basis, (D) maxilla, (E) left mandible, (F) cuspidate seta with supracuticular pocket of the left mandible first article, (G) incisor to molar process of left mandible, (H) right mandible, (I) incisor to molar process of right mandible." httpUri="https://zenodo.org/record/285586/files/figure.png" pageId="3" pageNumber="80">
Fig. 
<date id="EB5110084D226377609AE041FC9E224A" box="[765,780,1621,1645]" pageId="3" pageNumber="80">3</date>
</figureCitation>
I). 
<emphasis id="AD9BEADA4D2263776157E041FC422249" box="[816,976,1621,1646]" italics="true" pageId="3" pageNumber="80">Left mandible</emphasis>
with incisor process composed of two chitinous ridges with rounded distal end; lacinia mobilis and setal row absent, molar process similar to that of right mandible (
<figureCitation id="07D42A4D4D2263776369E088FEC32293" box="[270,337,1692,1717]" captionStart="FIGURE 3" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[206,1365,203,1881]" captionTargetId="figure@5.[190,1395,193,1898]" captionTargetPageId="5" captionText="FIGURE 3. Petalophthalmus papilloculatus sp. nov.: (A – I) holotype female MNCN 20.04 / 9201. (A) labrum, (B) maxillule, (C) cuspidate-serrate seta on maxillule basis, (D) maxilla, (E) left mandible, (F) cuspidate seta with supracuticular pocket of the left mandible first article, (G) incisor to molar process of left mandible, (H) right mandible, (I) incisor to molar process of right mandible." httpUri="https://zenodo.org/record/285586/files/figure.png" pageId="3" pageNumber="80">Fig. 3</figureCitation>
G).
</paragraph>
<paragraph id="9F5036C84D22637762A0E0ABFBA122DC" blockId="3.[151,1436,151,2037]" pageId="3" pageNumber="80">
<emphasis id="AD9BEADA4D22637762A0E0ABFEA122FF" box="[199,307,1727,1752]" italics="true" pageId="3" pageNumber="80">Maxillule</emphasis>
comparatively small (
<figureCitation id="07D42A4D4D2263776056E0ABFDE722FF" box="[561,629,1727,1752]" captionStart="FIGURE 3" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[206,1365,203,1881]" captionTargetId="figure@5.[190,1395,193,1898]" captionTargetPageId="5" captionText="FIGURE 3. Petalophthalmus papilloculatus sp. nov.: (A – I) holotype female MNCN 20.04 / 9201. (A) labrum, (B) maxillule, (C) cuspidate-serrate seta on maxillule basis, (D) maxilla, (E) left mandible, (F) cuspidate seta with supracuticular pocket of the left mandible first article, (G) incisor to molar process of left mandible, (H) right mandible, (I) incisor to molar process of right mandible." httpUri="https://zenodo.org/record/285586/files/figure.png" pageId="3" pageNumber="80">Fig. 3</figureCitation>
B), outer lobe distally armed with seven strong cuspidate-serrate setae, each one with one row of denticles (
<figureCitation id="07D42A4D4D226377604BE0F7FDFD22DC" box="[556,623,1763,1788]" captionStart="FIGURE 3" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[206,1365,203,1881]" captionTargetId="figure@5.[190,1395,193,1898]" captionTargetPageId="5" captionText="FIGURE 3. Petalophthalmus papilloculatus sp. nov.: (A – I) holotype female MNCN 20.04 / 9201. (A) labrum, (B) maxillule, (C) cuspidate-serrate seta on maxillule basis, (D) maxilla, (E) left mandible, (F) cuspidate seta with supracuticular pocket of the left mandible first article, (G) incisor to molar process of left mandible, (H) right mandible, (I) incisor to molar process of right mandible." httpUri="https://zenodo.org/record/285586/files/figure.png" pageId="3" pageNumber="80">Fig. 3</figureCitation>
C); inner lobe with seven pappose setae.
</paragraph>
<paragraph id="9F5036C84D22637762A0E112FBC023AF" blockId="3.[151,1436,151,2037]" pageId="3" pageNumber="80">
<emphasis id="AD9BEADA4D22637762A0E112FE8C2338" box="[199,286,1798,1823]" italics="true" pageId="3" pageNumber="80">Maxilla</emphasis>
with elongate and narrow exopod, extending to half length of endopod distal article, outer margin armed with long pappose setae, inner margin with seven distal pappose setae (
<figureCitation id="07D42A4D4D226377667EE13DFBCC2365" box="[1049,1118,1833,1858]" captionStart="FIGURE 3" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[206,1365,203,1881]" captionTargetId="figure@5.[190,1395,193,1898]" captionTargetPageId="5" captionText="FIGURE 3. Petalophthalmus papilloculatus sp. nov.: (A – I) holotype female MNCN 20.04 / 9201. (A) labrum, (B) maxillule, (C) cuspidate-serrate seta on maxillule basis, (D) maxilla, (E) left mandible, (F) cuspidate seta with supracuticular pocket of the left mandible first article, (G) incisor to molar process of left mandible, (H) right mandible, (I) incisor to molar process of right mandible." httpUri="https://zenodo.org/record/285586/files/figure.png" pageId="3" pageNumber="80">Fig. 3</figureCitation>
D). Endopod with the distal article long and narrow, about 2.6 times as long as broad, densely setose on inner margin. Two coxal endites armed with pappose setae on inner margins, basal endite with proximal longer pappose seta.
</paragraph>
<paragraph id="9F5036C84D22637762A0E180FE4A23D2" blockId="3.[151,1436,151,2037]" pageId="3" pageNumber="80">
<emphasis id="AD9BEADA4D22637762A0E180FEC9238A" box="[199,347,1940,1965]" italics="true" pageId="3" pageNumber="80">Thoracopods</emphasis>
classified into four different shape groups: first and second thoracopods represented as robust appendages (maxillipeds), third and fourth with peculiar endopods, fifth with apically setose endopod and sixth to eighth with clawed dactylus.
</paragraph>
<caption id="CB9066404D25637062F0E184FC4623C4" httpUri="https://zenodo.org/record/285585/files/figure.png" pageId="4" pageNumber="81" targetBox="[186,1394,206,1897]" targetPageId="4">
<paragraph id="9F5036C84D25637062F0E184FC4623C4" blockId="4.[151,1436,1936,2019]" pageId="4" pageNumber="81">
<emphasis id="AD9BEADA4D25637062F0E184FE842381" bold="true" box="[151,278,1936,1958]" pageId="4" pageNumber="81">FIGURE 2.</emphasis>
<taxonomicName id="58EF4D4B4D2563706379E184FDF62381" box="[286,612,1936,1958]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="4" pageNumber="81" phylum="Arthropoda" rank="species" species="papilloculatus" status="sp. nov.">
<emphasis id="AD9BEADA4D2563706379E184FDF62381" box="[286,612,1936,1958]" italics="true" pageId="4" pageNumber="81">Petalophthalmus papilloculatus</emphasis>
</taxonomicName>
<emphasis id="AD9BEADA4D256370600AE184FD522381" bold="true" box="[621,704,1936,1958]" pageId="4" pageNumber="81">
<taxonomicNameLabel id="B6A857A14D256370600AE184FD522381" box="[621,704,1936,1958]" pageId="4" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A, B, D) holotype female MNCN 20.04/9201; (C, E) allotype male MNCN 20.04/9202. (A) habitus in lateral view, (B) anterior part of carapace with eyes and right antennule in dorsal view, (C) antennule in dorsal view, (D) antenna in ventral view, (E) antenna in ventral view.
</paragraph>
</caption>
<caption id="CB9066404D24637162F0E169FF6F23C8" httpUri="https://zenodo.org/record/285586/files/figure.png" pageId="5" pageNumber="82" targetBox="[206,1365,203,1881]" targetPageId="5">
<paragraph id="9F5036C84D24637162F0E169FF6F23C8" blockId="5.[151,1436,1917,2031]" pageId="5" pageNumber="82">
<emphasis id="AD9BEADA4D24637162F0E169FE8623B4" bold="true" box="[151,276,1917,1939]" pageId="5" pageNumber="82">FIGURE 3.</emphasis>
<taxonomicName id="58EF4D4B4D246371637BE169FDF223B4" box="[284,608,1917,1939]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="5" pageNumber="82" phylum="Arthropoda" rank="species" species="papilloculatus" status="sp. nov.">
<emphasis id="AD9BEADA4D246371637BE169FDF223B4" box="[284,608,1917,1939]" italics="true" pageId="5" pageNumber="82">Petalophthalmus papilloculatus</emphasis>
</taxonomicName>
<emphasis id="AD9BEADA4D246371600FE169FD2823B4" bold="true" box="[616,698,1917,1939]" pageId="5" pageNumber="82">
<taxonomicNameLabel id="B6A857A14D246371600FE169FD2823B4" box="[616,698,1917,1939]" pageId="5" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A–I) holotype female MNCN 20.04/9201. (A) labrum, (B) maxillule, (C) cuspidate-serrate seta on maxillule basis, (D) maxilla, (E) left mandible, (F) cuspidate seta with supracuticular pocket of the left mandible first article, (G) incisor to molar process of left mandible, (H) right mandible, (I) incisor to molar process of right mandible.
</paragraph>
</caption>
<caption id="CB9066404D27637262F0E1D5FAED23D1" httpUri="https://zenodo.org/record/285587/files/figure.png" pageId="6" pageNumber="83" targetBox="[219,1374,199,1956]" targetPageId="6">
<paragraph id="9F5036C84D27637262F0E1D5FAED23D1" blockId="6.[151,1436,1985,2038]" pageId="6" pageNumber="83">
<emphasis id="AD9BEADA4D27637262F0E1D5FE8023F0" bold="true" box="[151,274,1985,2007]" pageId="6" pageNumber="83">FIGURE 4.</emphasis>
<taxonomicName id="58EF4D4B4D276372637FE1D5FDC823F0" box="[280,602,1985,2007]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="6" pageNumber="83" phylum="Arthropoda" rank="species" species="papilloculatus" status="sp. nov.">
<emphasis id="AD9BEADA4D276372637FE1D5FDC823F0" box="[280,602,1985,2007]" italics="true" pageId="6" pageNumber="83">Petalophthalmus papilloculatus</emphasis>
</taxonomicName>
<emphasis id="AD9BEADA4D2763726007E1D5FD3D23F0" bold="true" box="[608,687,1985,2007]" pageId="6" pageNumber="83">
<taxonomicNameLabel id="B6A857A14D2763726007E1D5FD3D23F0" box="[608,687,1985,2007]" pageId="6" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A–C) holotype female MNCN 20.04/9201; (D, E) allotype male MNCN 20.04/9202. (A) first thoracopod, (B) second thoracopod, (C) third thoracopod, (D) third thoracopod, (E) fourth thoracopod.
</paragraph>
</caption>
<caption id="CB9066404D26637362F0E15EFC87239B" httpUri="https://zenodo.org/record/285588/files/figure.png" pageId="7" pageNumber="84" targetBox="[215,1383,204,1835]" targetPageId="7">
<paragraph id="9F5036C84D26637362F0E15EFC87239B" blockId="7.[151,1436,1866,1980]" pageId="7" pageNumber="84">
<emphasis id="AD9BEADA4D26637362F0E15EFE862347" bold="true" box="[151,276,1866,1888]" pageId="7" pageNumber="84">FIGURE 5.</emphasis>
<taxonomicName id="58EF4D4B4D266373637CE15EFDCD2347" box="[283,607,1866,1888]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="papilloculatus" status="sp. nov.">
<emphasis id="AD9BEADA4D266373637CE15EFDCD2347" box="[283,607,1866,1888]" italics="true" pageId="7" pageNumber="84">Petalophthalmus papilloculatus</emphasis>
</taxonomicName>
<emphasis id="AD9BEADA4D2663736001E15EFD252347" bold="true" box="[614,695,1866,1888]" pageId="7" pageNumber="84">
<taxonomicNameLabel id="B6A857A14D2663736001E15EFD252347" box="[614,695,1866,1888]" pageId="7" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A, B) holotype female MNCN 20.04/9201; (C, D, F, G) allotype male MNCN 20.04/9202; (E) paratype female MNCN 20.04/9203. (A) fifth thoracopod, (B) distal article of endopod of fifth thoracopod, (C) sixth thoracopod, (D) distal article of endopod of sixth thoracopod, (E) seventh thoracopod, (F) eighth thoracopod, (G) distal article of endopod of eighth thoracopod.
</paragraph>
</caption>
<caption id="CB9066404D29637C62F0E179FD3A23F8" httpUri="https://zenodo.org/record/285589/files/figure.png" pageId="8" pageNumber="85" targetBox="[184,1375,203,1870]" targetPageId="8">
<paragraph id="9F5036C84D29637C62F0E179FD3A23F8" blockId="8.[151,1436,1901,2015]" pageId="8" pageNumber="85">
<emphasis id="AD9BEADA4D29637C62F0E179FE8123A4" bold="true" box="[151,275,1901,1923]" pageId="8" pageNumber="85">FIGURE 6.</emphasis>
<taxonomicName id="58EF4D4B4D29637C637EE179FDCE23A4" box="[281,604,1901,1923]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="papilloculatus" status="sp. nov.">
<emphasis id="AD9BEADA4D29637C637EE179FDCE23A4" box="[281,604,1901,1923]" italics="true" pageId="8" pageNumber="85">Petalophthalmus papilloculatus</emphasis>
</taxonomicName>
<emphasis id="AD9BEADA4D29637C6004E179FD2623A4" bold="true" box="[611,692,1901,1923]" pageId="8" pageNumber="85">
<taxonomicNameLabel id="B6A857A14D29637C6004E179FD2623A4" box="[611,692,1901,1923]" pageId="8" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A–E) holotype female MNCN 20.04/9201; (F–I) allotype male MNCN 20.04/9202. (A) first pleopod, (B) second pleopod, (C) third pleopod, (D) fourth pleopod, (E) fifth pleopod, (F) first pleopod, (G) distal article of endopod of first pleopod, (H) second pleopod, (I) distal end of first article of exopod of second pleopod, (J) third pleopod, (K) fourth pleopod, (L) fifth pleopod.
</paragraph>
</caption>
<caption id="CB9066404D28637D62F0E168FCA023C9" httpUri="https://zenodo.org/record/285590/files/figure.png" pageId="9" pageNumber="86" targetBox="[193,1393,200,1901]" targetPageId="9">
<paragraph id="9F5036C84D28637D62F0E168FCA023C9" blockId="9.[151,1436,1916,2030]" pageId="9" pageNumber="86">
<emphasis id="AD9BEADA4D28637D62F0E168FE8A23B5" bold="true" box="[151,280,1916,1938]" pageId="9" pageNumber="86">FIGURE 7.</emphasis>
<taxonomicName id="58EF4D4B4D28637D6343E168FDFE23B5" box="[292,620,1916,1938]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="papilloculatus" status="sp. nov.">
<emphasis id="AD9BEADA4D28637D6343E168FDFE23B5" box="[292,620,1916,1938]" italics="true" pageId="9" pageNumber="86">Petalophthalmus papilloculatus</emphasis>
</taxonomicName>
<emphasis id="AD9BEADA4D28637D601FE168FD5C23B5" bold="true" box="[632,718,1916,1938]" pageId="9" pageNumber="86">
<taxonomicNameLabel id="B6A857A14D28637D601FE168FD5C23B5" box="[632,718,1916,1938]" pageId="9" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A–D) holotype female MNCN 20.04/9201; (E–G) allotype male MNCN 20.04/9202. (A) telson in dorsal view, (B) telson apex in dorsal view, (C) medial pair of serrate setae and three pairs of ventral small teeth on telson apex, (D) right uropod in ventral view, (E) telson in dorsal view, (F) telson apex in dorsal view, (G) medial serrate seta and pairs of ventral small teeth on telson apex.
</paragraph>
</caption>
<paragraph id="9F5036C84D2B637E62A0E68DFC99253B" blockId="10.[151,1436,151,2013]" pageId="10" pageNumber="87">
<emphasis id="AD9BEADA4D2B637E62A0E68DFE142497" box="[199,390,151,176]" italics="true" pageId="10" pageNumber="87">First thoracopod</emphasis>
with long narrow epipodite, without exopod (
<figureCitation id="07D42A4D4D2B637E61F5E683FC452497" box="[914,983,151,176]" captionStart="FIGURE 4" captionStartId="6.[151,250,1985,2007]" captionTargetBox="[219,1374,199,1956]" captionTargetId="figure@6.[197,1387,193,1963]" captionTargetPageId="6" captionText="FIGURE 4. Petalophthalmus papilloculatus sp. nov.: (A – C) holotype female MNCN 20.04 / 9201; (D, E) allotype male MNCN 20.04 / 9202. (A) first thoracopod, (B) second thoracopod, (C) third thoracopod, (D) third thoracopod, (E) fourth thoracopod." httpUri="https://zenodo.org/record/285587/files/figure.png" pageId="10" pageNumber="87">Fig. 4</figureCitation>
A). Endopod powerful and robust; basis without lobe; preischium very short; ischium produced into inner triangular lobe, tipped with four simple long seta; carpopropodus about twice as long as its greatest width, tapering somewhat distally; dactylus fused with nail to form a long curved claw armed with a few simple setae.
</paragraph>
<paragraph id="9F5036C84D2B637E62A0E733FE342647" blockId="10.[151,1436,151,2013]" pageId="10" pageNumber="87">
<emphasis id="AD9BEADA4D2B637E62A0E733FE372567" box="[199,421,295,320]" italics="true" pageId="10" pageNumber="87">Second thoracopod</emphasis>
longer and broader than first thoracopod, without epipodite and exopodite (
<figureCitation id="07D42A4D4D2B637E6756E733FAE92567" box="[1329,1403,295,320]" captionStart="FIGURE 4" captionStartId="6.[151,250,1985,2007]" captionTargetBox="[219,1374,199,1956]" captionTargetId="figure@6.[197,1387,193,1963]" captionTargetPageId="6" captionText="FIGURE 4. Petalophthalmus papilloculatus sp. nov.: (A – C) holotype female MNCN 20.04 / 9201; (D, E) allotype male MNCN 20.04 / 9202. (A) first thoracopod, (B) second thoracopod, (C) third thoracopod, (D) third thoracopod, (E) fourth thoracopod." httpUri="https://zenodo.org/record/285587/files/figure.png" pageId="10" pageNumber="87">Fig. 4</figureCitation>
B). Endopod with preischium unarmed; ischium inner margin produced into a large quadrangular lobe overreaching merus distal margin, about 4 times as long as broad and bearing short cuspidate setae on inner and outer margins and five long simple setae on apex; merus longest, 3.4 times as long as broad, inner margin armed with row of simple long and short cuspidate setae; carpopropodus twice as long as greatest width, tapering somewhat distally, outer margin armed with two cuspidate setae and one long subdistal simple seta, inner margin armed with an irregularly row of simple long setae, short cuspidate setae and apical pappose setae; dactylus fused with nail to form long curved claw, armed with one long and robust simple seta on inner proximal margin and one simple seta on middle inner margin.
</paragraph>
<paragraph id="9F5036C84D2B637E62A0E478FE95268F" blockId="10.[151,1436,151,2013]" pageId="10" pageNumber="87">
<emphasis id="AD9BEADA4D2B637E62A0E478FE9626A2" box="[199,260,620,645]" italics="true" pageId="10" pageNumber="87">Third</emphasis>
and 
<emphasis id="AD9BEADA4D2B637E635DE478FD8326A2" box="[314,529,620,645]" italics="true" pageId="10" pageNumber="87">fourth thoracopods</emphasis>
(
<figureCitation id="07D42A4D4D2B637E6078E478FDF426A3" box="[543,614,620,645]" captionStart="FIGURE 4" captionStartId="6.[151,250,1985,2007]" captionTargetBox="[219,1374,199,1956]" captionTargetId="figure@6.[197,1387,193,1963]" captionTargetPageId="6" captionText="FIGURE 4. Petalophthalmus papilloculatus sp. nov.: (A – C) holotype female MNCN 20.04 / 9201; (D, E) allotype male MNCN 20.04 / 9202. (A) first thoracopod, (B) second thoracopod, (C) third thoracopod, (D) third thoracopod, (E) fourth thoracopod." httpUri="https://zenodo.org/record/285587/files/figure.png" pageId="10" pageNumber="87">Figs 4</figureCitation>
C–E). Endopod reduced to 1 or 2-articulate naked articles. Exopod 19/20- articulate.
</paragraph>
<paragraph id="9F5036C84D2B637E62A0E4A0FA1D271F" blockId="10.[151,1436,151,2013]" pageId="10" pageNumber="87">
<emphasis id="AD9BEADA4D2B637E62A0E4A0FE1A26EA" box="[199,392,692,717]" italics="true" pageId="10" pageNumber="87">Fifth thoracopod</emphasis>
slender (
<figureCitation id="07D42A4D4D2B637E6393E4A0FDA826EA" box="[500,570,692,717]" captionStart="FIGURE 5" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[215,1383,204,1835]" captionTargetId="figure@7.[196,1391,193,1845]" captionTargetPageId="7" captionText="FIGURE 5. Petalophthalmus papilloculatus sp. nov.: (A, B) holotype female MNCN 20.04 / 9201; (C, D, F, G) allotype male MNCN 20.04 / 9202; (E) paratype female MNCN 20.04 / 9203. (A) fifth thoracopod, (B) distal article of endopod of fifth thoracopod, (C) sixth thoracopod, (D) distal article of endopod of sixth thoracopod, (E) seventh thoracopod, (F) eighth thoracopod, (G) distal article of endopod of eighth thoracopod." httpUri="https://zenodo.org/record/285588/files/figure.png" pageId="10" pageNumber="87">Fig. 5</figureCitation>
A). Endopod with only distal setae; ischium, merus and carpus subequal in length; propodus divided in 2 segments, the distal one about twice as long as the proximal one and armed on its inner distal margin with two rows of simple setae; dactylus very small and densely setose, armed with two proximal serrate setae and one distal hook-like serrate seta (
<figureCitation id="07D42A4D4D2B637E6152E50BFCEA271F" box="[821,888,799,824]" captionStart="FIGURE 5" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[215,1383,204,1835]" captionTargetId="figure@7.[196,1391,193,1845]" captionTargetPageId="7" captionText="FIGURE 5. Petalophthalmus papilloculatus sp. nov.: (A, B) holotype female MNCN 20.04 / 9201; (C, D, F, G) allotype male MNCN 20.04 / 9202; (E) paratype female MNCN 20.04 / 9203. (A) fifth thoracopod, (B) distal article of endopod of fifth thoracopod, (C) sixth thoracopod, (D) distal article of endopod of sixth thoracopod, (E) seventh thoracopod, (F) eighth thoracopod, (G) distal article of endopod of eighth thoracopod." httpUri="https://zenodo.org/record/285588/files/figure.png" pageId="10" pageNumber="87">Fig. 5</figureCitation>
B). Exopod shorter than endopod, 18-articulate.
</paragraph>
<paragraph id="9F5036C84D2B637E62A0E550FD5F27CA" blockId="10.[151,1436,151,2013]" pageId="10" pageNumber="87">
<emphasis id="AD9BEADA4D2B637E62A0E550FF6D277A" box="[199,255,836,861]" italics="true" pageId="10" pageNumber="87">Sixth</emphasis>
to 
<emphasis id="AD9BEADA4D2B637E6344E550FE6E277A" box="[291,508,836,861]" italics="true" pageId="10" pageNumber="87">eighth thoracopods</emphasis>
(
<figureCitation id="07D42A4D4D2B637E606DE550FDC3277A" box="[522,593,836,861]" captionStart="FIGURE 5" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[215,1383,204,1835]" captionTargetId="figure@7.[196,1391,193,1845]" captionTargetPageId="7" captionText="FIGURE 5. Petalophthalmus papilloculatus sp. nov.: (A, B) holotype female MNCN 20.04 / 9201; (C, D, F, G) allotype male MNCN 20.04 / 9202; (E) paratype female MNCN 20.04 / 9203. (A) fifth thoracopod, (B) distal article of endopod of fifth thoracopod, (C) sixth thoracopod, (D) distal article of endopod of sixth thoracopod, (E) seventh thoracopod, (F) eighth thoracopod, (G) distal article of endopod of eighth thoracopod." httpUri="https://zenodo.org/record/285588/files/figure.png" pageId="10" pageNumber="87">Figs 5</figureCitation>
C, E–F). Endopod bearing a few plumose setae; preischium slightly shorter than ischium, merus about twice as long as carpopropodus, carpopropodus undivided; dactylus short, terminating in one simple seta, dactylus apparently fused to distal curved nail (
<figureCitation id="07D42A4D4D2B637E61EEE598FC432782" box="[905,977,908,933]" captionStart="FIGURE 5" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[215,1383,204,1835]" captionTargetId="figure@7.[196,1391,193,1845]" captionTargetPageId="7" captionText="FIGURE 5. Petalophthalmus papilloculatus sp. nov.: (A, B) holotype female MNCN 20.04 / 9201; (C, D, F, G) allotype male MNCN 20.04 / 9202; (E) paratype female MNCN 20.04 / 9203. (A) fifth thoracopod, (B) distal article of endopod of fifth thoracopod, (C) sixth thoracopod, (D) distal article of endopod of sixth thoracopod, (E) seventh thoracopod, (F) eighth thoracopod, (G) distal article of endopod of eighth thoracopod." httpUri="https://zenodo.org/record/285588/files/figure.png" pageId="10" pageNumber="87">Figs 5</figureCitation>
D, G). Exopod longer than endopod, 17- articulate. Penis elongate, cylindrical, ending in hemispherical apex, without setae (
<figureCitation id="07D42A4D4D2B637E66FCE5BBFB7427EF" box="[1179,1254,943,968]" captionStart="FIGURE 5" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[215,1383,204,1835]" captionTargetId="figure@7.[196,1391,193,1845]" captionTargetPageId="7" captionText="FIGURE 5. Petalophthalmus papilloculatus sp. nov.: (A, B) holotype female MNCN 20.04 / 9201; (C, D, F, G) allotype male MNCN 20.04 / 9202; (E) paratype female MNCN 20.04 / 9203. (A) fifth thoracopod, (B) distal article of endopod of fifth thoracopod, (C) sixth thoracopod, (D) distal article of endopod of sixth thoracopod, (E) seventh thoracopod, (F) eighth thoracopod, (G) distal article of endopod of eighth thoracopod." httpUri="https://zenodo.org/record/285588/files/figure.png" pageId="10" pageNumber="87">Fig. 5</figureCitation>
F). In females, marsupium composed of seven pairs of oostegites.
</paragraph>
<paragraph id="9F5036C84D2B637E62A0E5E3FF6920E2" blockId="10.[151,1436,151,2013]" pageId="10" pageNumber="87">
<emphasis id="AD9BEADA4D2B637E62A0E5E3FEBD2037" box="[199,303,1015,1040]" italics="true" pageId="10" pageNumber="87">Pleopods</emphasis>
uniramous in female, increasing in length towards posterior pairs; first pleopod 2-articulate, remaining pleopods 3-articulate (
<figureCitation id="07D42A4D4D2B637E6068E208FDC52012" box="[527,599,1052,1077]" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1923]" captionTargetBox="[184,1375,203,1870]" captionTargetId="figure@8.[179,1404,193,1883]" captionTargetPageId="8" captionText="FIGURE 6. Petalophthalmus papilloculatus sp. nov.: (A – E) holotype female MNCN 20.04 / 9201; (F – I) allotype male MNCN 20.04 / 9202. (A) first pleopod, (B) second pleopod, (C) third pleopod, (D) fourth pleopod, (E) fifth pleopod, (F) first pleopod, (G) distal article of endopod of first pleopod, (H) second pleopod, (I) distal end of first article of exopod of second pleopod, (J) third pleopod, (K) fourth pleopod, (L) fifth pleopod." httpUri="https://zenodo.org/record/285589/files/figure.png" pageId="10" pageNumber="87">Figs 6</figureCitation>
A–E). Male pleopods biramous. First pair with endopod 2-articulate (
<figureCitation id="07D42A4D4D2B637E6709E208FF34207F" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1923]" captionTargetBox="[184,1375,203,1870]" captionTargetId="figure@8.[179,1404,193,1883]" captionTargetPageId="8" captionText="FIGURE 6. Petalophthalmus papilloculatus sp. nov.: (A – E) holotype female MNCN 20.04 / 9201; (F – I) allotype male MNCN 20.04 / 9202. (A) first pleopod, (B) second pleopod, (C) third pleopod, (D) fourth pleopod, (E) fifth pleopod, (F) first pleopod, (G) distal article of endopod of first pleopod, (H) second pleopod, (I) distal end of first article of exopod of second pleopod, (J) third pleopod, (K) fourth pleopod, (L) fifth pleopod." httpUri="https://zenodo.org/record/285589/files/figure.png" pageId="10" pageNumber="87">Fig. 6</figureCitation>
F), distal article finger-like shaped, ending in two small simple setae (
<figureCitation id="07D42A4D4D2B637E61A5E22BFB95207F" box="[962,1031,1087,1112]" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1923]" captionTargetBox="[184,1375,203,1870]" captionTargetId="figure@8.[179,1404,193,1883]" captionTargetPageId="8" captionText="FIGURE 6. Petalophthalmus papilloculatus sp. nov.: (A – E) holotype female MNCN 20.04 / 9201; (F – I) allotype male MNCN 20.04 / 9202. (A) first pleopod, (B) second pleopod, (C) third pleopod, (D) fourth pleopod, (E) fifth pleopod, (F) first pleopod, (G) distal article of endopod of first pleopod, (H) second pleopod, (I) distal end of first article of exopod of second pleopod, (J) third pleopod, (K) fourth pleopod, (L) fifth pleopod." httpUri="https://zenodo.org/record/285589/files/figure.png" pageId="10" pageNumber="87">Fig. 6</figureCitation>
G); exopod 16-articulate. Second to fifth pleopods with exopod slightly longer than endopod; exopods 18/19-articulate, endopods 11-articulate (
<figureCitation id="07D42A4D4D2B637E670DE270FF372087" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1923]" captionTargetBox="[184,1375,203,1870]" captionTargetId="figure@8.[179,1404,193,1883]" captionTargetPageId="8" captionText="FIGURE 6. Petalophthalmus papilloculatus sp. nov.: (A – E) holotype female MNCN 20.04 / 9201; (F – I) allotype male MNCN 20.04 / 9202. (A) first pleopod, (B) second pleopod, (C) third pleopod, (D) fourth pleopod, (E) fifth pleopod, (F) first pleopod, (G) distal article of endopod of first pleopod, (H) second pleopod, (I) distal end of first article of exopod of second pleopod, (J) third pleopod, (K) fourth pleopod, (L) fifth pleopod." httpUri="https://zenodo.org/record/285589/files/figure.png" pageId="10" pageNumber="87">Figs 6</figureCitation>
H, J–L). Exopod basal article of second pleopod bearing rounded distal lobe armed with two strong simple setae (
<figureCitation id="07D42A4D4D2B637E62F8E2B8FF7620E2" box="[159,228,1196,1221]" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1923]" captionTargetBox="[184,1375,203,1870]" captionTargetId="figure@8.[179,1404,193,1883]" captionTargetPageId="8" captionText="FIGURE 6. Petalophthalmus papilloculatus sp. nov.: (A – E) holotype female MNCN 20.04 / 9201; (F – I) allotype male MNCN 20.04 / 9202. (A) first pleopod, (B) second pleopod, (C) third pleopod, (D) fourth pleopod, (E) fifth pleopod, (F) first pleopod, (G) distal article of endopod of first pleopod, (H) second pleopod, (I) distal end of first article of exopod of second pleopod, (J) third pleopod, (K) fourth pleopod, (L) fifth pleopod." httpUri="https://zenodo.org/record/285589/files/figure.png" pageId="10" pageNumber="87">
Fig. 
<date id="EB5110084D2B637E62B3E2B8FF7620E2" box="[212,228,1196,1221]" pageId="10" pageNumber="87">6</date>
</figureCitation>
I).
</paragraph>
<paragraph id="9F5036C84D2B637E62A0E2DBFC8E2173" blockId="10.[151,1436,151,2013]" pageId="10" pageNumber="87">
<emphasis id="AD9BEADA4D2B637E62A0E2DBFE8F20CF" box="[199,285,1231,1256]" italics="true" pageId="10" pageNumber="87">Uropod</emphasis>
endopod slender, without statocyst, not extending beyond telson apex, fully setose, distal inner margin concave (
<figureCitation id="07D42A4D4D2B637E6365E2E0FED6212B" box="[258,324,1268,1293]" captionStart="FIGURE 7" captionStartId="9.[151,250,1916,1938]" captionTargetBox="[193,1393,200,1901]" captionTargetId="figure@9.[179,1408,193,1911]" captionTargetPageId="9" captionText="FIGURE 7. Petalophthalmus papilloculatus sp. nov.: (A – D) holotype female MNCN 20.04 / 9201; (E – G) allotype male MNCN 20.04 / 9202. (A) telson in dorsal view, (B) telson apex in dorsal view, (C) medial pair of serrate setae and three pairs of ventral small teeth on telson apex, (D) right uropod in ventral view, (E) telson in dorsal view, (F) telson apex in dorsal view, (G) medial serrate seta and pairs of ventral small teeth on telson apex." httpUri="https://zenodo.org/record/285590/files/figure.png" pageId="10" pageNumber="87">Fig. 7</figureCitation>
D). Uropod exopod 2-articulate, longer and broader than endopod, extending slightly beyond telson apex; distal article about one third of the basal article length; outer margin of the basal article entire, ending with three cuspidate setae lengthening from outer to inner one.
</paragraph>
<paragraph id="9F5036C84D2B637E62A0E34BFD63222F" blockId="10.[151,1436,151,2013]" pageId="10" pageNumber="87">
<emphasis id="AD9BEADA4D2B637E62A0E34BFE9D215F" box="[199,271,1375,1400]" italics="true" pageId="10" pageNumber="87">Telson</emphasis>
quadrangular, 2.8 times as long as broad, nearly as long as the sixth abdominal somite; 2/3 of posterior lateral margins armed with 25–37 cuspidate setae increasing distally in size (
<figureCitation id="07D42A4D4D2B637E666CE390FBC621BB" box="[1035,1108,1412,1437]" captionStart="FIGURE 7" captionStartId="9.[151,250,1916,1938]" captionTargetBox="[193,1393,200,1901]" captionTargetId="figure@9.[179,1408,193,1911]" captionTargetPageId="9" captionText="FIGURE 7. Petalophthalmus papilloculatus sp. nov.: (A – D) holotype female MNCN 20.04 / 9201; (E – G) allotype male MNCN 20.04 / 9202. (A) telson in dorsal view, (B) telson apex in dorsal view, (C) medial pair of serrate setae and three pairs of ventral small teeth on telson apex, (D) right uropod in ventral view, (E) telson in dorsal view, (F) telson apex in dorsal view, (G) medial serrate seta and pairs of ventral small teeth on telson apex." httpUri="https://zenodo.org/record/285590/files/figure.png" pageId="10" pageNumber="87">Figs 7</figureCitation>
A, E). In females, apex with four pairs of serrate setae decreasing in length towards medial point and three pairs of ventral small teeth (
<figureCitation id="07D42A4D4D2B637E675BE3B3FA1121E7" box="[1340,1411,1447,1472]" captionStart="FIGURE 7" captionStartId="9.[151,250,1916,1938]" captionTargetBox="[193,1393,200,1901]" captionTargetId="figure@9.[179,1408,193,1911]" captionTargetPageId="9" captionText="FIGURE 7. Petalophthalmus papilloculatus sp. nov.: (A – D) holotype female MNCN 20.04 / 9201; (E – G) allotype male MNCN 20.04 / 9202. (A) telson in dorsal view, (B) telson apex in dorsal view, (C) medial pair of serrate setae and three pairs of ventral small teeth on telson apex, (D) right uropod in ventral view, (E) telson in dorsal view, (F) telson apex in dorsal view, (G) medial serrate seta and pairs of ventral small teeth on telson apex." httpUri="https://zenodo.org/record/285590/files/figure.png" pageId="10" pageNumber="87">Figs 7</figureCitation>
B, C). In males, apex with three pairs of serrate setae decreasing in length towards medial point, one medial serrate seta, and 7–8 pairs of ventral small teeth (
<figureCitation id="07D42A4D4D2B637E600DE3FBFD22222F" box="[618,688,1519,1544]" captionStart="FIGURE 7" captionStartId="9.[151,250,1916,1938]" captionTargetBox="[193,1393,200,1901]" captionTargetId="figure@9.[179,1408,193,1911]" captionTargetPageId="9" captionText="FIGURE 7. Petalophthalmus papilloculatus sp. nov.: (A – D) holotype female MNCN 20.04 / 9201; (E – G) allotype male MNCN 20.04 / 9202. (A) telson in dorsal view, (B) telson apex in dorsal view, (C) medial pair of serrate setae and three pairs of ventral small teeth on telson apex, (D) right uropod in ventral view, (E) telson in dorsal view, (F) telson apex in dorsal view, (G) medial serrate seta and pairs of ventral small teeth on telson apex." httpUri="https://zenodo.org/record/285590/files/figure.png" pageId="10" pageNumber="87">Figs 7</figureCitation>
F, G).
</paragraph>
</subSubSection>
<subSubSection id="D7F565434D2B637E62A0E000FC122277" pageId="10" pageNumber="87" type="distribution">
<paragraph id="9F5036C84D2B637E62A0E000FC122277" blockId="10.[151,1436,151,2013]" pageId="10" pageNumber="87">
<emphasis id="AD9BEADA4D2B637E62A0E000FEF2220A" bold="true" box="[199,352,1556,1581]" pageId="10" pageNumber="87">Distribution.</emphasis>
The known distributional area of the new 
<taxonomicName id="58EF4D4B4D2B637E612FE000FB94220A" box="[840,1030,1556,1581]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="10" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="AD9BEADA4D2B637E612FE000FB94220A" box="[840,1030,1556,1581]" italics="true" pageId="10" pageNumber="87">Petalophthalmus</emphasis>
</taxonomicName>
species is at the moment limited to the Galicia Bank (NE Atlantic), between 
<date id="EB5110084D2B637E6002E023FC9E2277" box="[613,780,1591,1616]" pageId="10" pageNumber="87" value="1536" valueMax="1809">
1536 and 
<quantity id="58179B2D4D2B637E60B3E02CFC9E2277" box="[724,780,1592,1616]" metricMagnitude="3" metricUnit="m" metricValue="1.809" pageId="10" pageNumber="87" unit="m" value="1809.0">1809</quantity>
</date>
m depths.
</paragraph>
</subSubSection>
<subSubSection id="D7F565434D2B637E62A0E048FADE23B2" pageId="10" pageNumber="87" type="discussion">
<paragraph id="9F5036C84D2B637E62A0E048FEC9230F" blockId="10.[151,1436,151,2013]" pageId="10" pageNumber="87">
<emphasis id="AD9BEADA4D2B637E62A0E048FEA92252" bold="true" box="[199,315,1628,1653]" pageId="10" pageNumber="87">Remarks.</emphasis>
According to 
<bibRefCitation id="FB7E4B394D2B637E63B8E048FD092252" author="Tattersall" box="[479,667,1628,1653]" pageId="10" pageNumber="87" refString="Tattersall, O. S. (1968) A new species of Petalophthalmus (Mysidacea) based on specimens from off Puerto Rico, hitherto referred to Petalophthalmus oculatus. Journal of Zoology, 155, 271 - 282. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1968. tb 03052. x" type="journal article" year="1968">Tattersall (1968)</bibRefCitation>
, the main diagnostic features of the genus 
<taxonomicName id="58EF4D4B4D2B637E66E5E048FAAD2252" box="[1154,1343,1628,1653]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="10" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="AD9BEADA4D2B637E66E5E048FAAD2252" box="[1154,1343,1628,1653]" italics="true" pageId="10" pageNumber="87">Petalophthalmus</emphasis>
</taxonomicName>
are: the long and slender antennular peduncle; the powerful, long and prehensile mandibular palp; the prominent lobe on the ischium of thoracopods 1–2; the 2-articulated uropodal exopod with outer proximal margin entire, ending with 3 cuspidate setae at distal angle; the quadrangular shape of the telson with slightly emarginate apex and armed with serrate setae. The specimens herein described share all these diagnostic features and are therefore ascribed to genus 
<taxonomicName id="58EF4D4B4D2B637E62F0E11BFEC7230F" box="[151,341,1807,1832]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="10" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="AD9BEADA4D2B637E62F0E11BFEC7230F" box="[151,341,1807,1832]" italics="true" pageId="10" pageNumber="87">Petalophthalmus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="9F5036C84D2B637E62A0E121FADE23B2" blockId="10.[151,1436,151,2013]" pageId="10" pageNumber="87">
<taxonomicName id="58EF4D4B4D2B637E62A0E121FE14236A" box="[199,390,1844,1869]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="papilloculatus" status="sp. nov.">
<emphasis id="AD9BEADA4D2B637E62A0E121FE14236A" box="[199,390,1844,1869]" italics="true" pageId="10" pageNumber="87">P. papilloculatus</emphasis>
</taxonomicName>
<emphasis id="AD9BEADA4D2B637E63F6E120FE7D236A" bold="true" box="[401,495,1844,1869]" pageId="10" pageNumber="87">
<taxonomicNameLabel id="B6A857A14D2B637E63F6E120FE7D236A" box="[401,495,1844,1869]" pageId="10" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the sixth species to be discovered in genus 
<taxonomicName id="58EF4D4B4D2B637E667FE120FB44236A" box="[1048,1238,1844,1869]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="10" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="AD9BEADA4D2B637E667FE120FB44236A" box="[1048,1238,1844,1869]" italics="true" pageId="10" pageNumber="87">Petalophthalmus</emphasis>
</taxonomicName>
. It can be easily distinguished from 
<taxonomicName id="58EF4D4B4D2B637E6311E14DFE612357" box="[374,499,1879,1904]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="oculatus">
<emphasis id="AD9BEADA4D2B637E6311E14DFE612357" box="[374,499,1879,1904]" italics="true" pageId="10" pageNumber="87">P. oculatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="58EF4D4B4D2B637E6063E14DFD3C2357" box="[516,686,1879,1904]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="caribbeanus">
<emphasis id="AD9BEADA4D2B637E6063E14DFD3C2357" box="[516,686,1879,1904]" italics="true" pageId="10" pageNumber="87">P. caribbeanus</emphasis>
</taxonomicName>
, 
<taxonomicName id="58EF4D4B4D2B637E60D9E14DFCAE2357" box="[702,828,1881,1904]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="macrops">
<emphasis id="AD9BEADA4D2B637E60D9E14DFCAE2357" box="[702,828,1881,1904]" italics="true" pageId="10" pageNumber="87">P. macrops</emphasis>
</taxonomicName>
and 
<emphasis id="AD9BEADA4D2B637E611FE14DFC2A2357" box="[888,952,1879,1904]" italics="true" pageId="10" pageNumber="87">P. l i u i</emphasis>
by the structure of its eyes, leaf-like and without visual elements (
<emphasis id="AD9BEADA4D2B637E63D6E169FE6B23B3" box="[433,505,1917,1940]" italics="true" pageId="10" pageNumber="87">versus</emphasis>
eyes being more or less globular in shape, with functional visual elements).
</paragraph>
</subSubSection>
<subSubSection id="D7F565434D2B637F62A0E1B5FD1A2782" lastPageId="11" lastPageNumber="88" pageId="10" pageNumber="87" type="reference_group">
<paragraph id="9F5036C84D2B637E62A0E1B5FE6523FA" blockId="10.[151,1436,151,2013]" pageId="10" pageNumber="87">
<taxonomicName id="58EF4D4B4D2B637E62A0E1B5FE10239E" box="[199,386,1952,1977]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="papilloculatus" status="sp. nov.">
<emphasis id="AD9BEADA4D2B637E62A0E1B5FE10239E" box="[199,386,1952,1977]" italics="true" pageId="10" pageNumber="87">P. papilloculatus</emphasis>
</taxonomicName>
<emphasis id="AD9BEADA4D2B637E63EFE1B4FE73239E" bold="true" box="[392,481,1952,1977]" pageId="10" pageNumber="87">
<taxonomicNameLabel id="B6A857A14D2B637E63EFE1B4FE73239E" box="[392,481,1952,1977]" pageId="10" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shows some similarity to 
<taxonomicName id="58EF4D4B4D2B637E6163E1B5FB1A239E" ID-CoL="4FCNW" authority="Willemoes-Suhm, 1875" authorityName="Willemoes-Suhm" authorityYear="1875" box="[772,1160,1952,1977]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="armiger">
<emphasis id="AD9BEADA4D2B637E6163E1B5FCEB239F" box="[772,889,1953,1976]" italics="true" pageId="10" pageNumber="87">P. armiger</emphasis>
<bibRefCitation id="FB7E4B394D2B637E6118E1B4FB1A239E" author="Willemoes-Suhm" box="[895,1160,1952,1977]" pageId="10" pageNumber="87" refString="Willemoes-Suhm, R. von. (1875) On some Atlantic Crustacea from the ' Challenger' Expedition (communicated by W. Thomson). Transactions of the Linnean Society of London. Second Series. Zoology, 1 (1), 23 - 59." type="journal article" year="1875">Willemoes-Suhm, 1875</bibRefCitation>
</taxonomicName>
but can be distinguished by the main following features:
</paragraph>
<paragraph id="9F5036C84D2A637F62A0E68CFE9324F3" blockId="11.[151,1436,151,933]" pageId="11" pageNumber="88">
(1) Anterior margin of the carapace truncate, without rostrum (
<emphasis id="AD9BEADA4D2A637F61A7E68DFB952497" box="[960,1031,153,176]" italics="true" pageId="11" pageNumber="88">versus</emphasis>
an acute triangular rostrum in 
<taxonomicName id="58EF4D4B4D2A637F67E0E68DFF6324F3" ID-CoL="4FCNW" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="armiger">
<emphasis id="AD9BEADA4D2A637F67E0E68DFF6324F3" italics="true" pageId="11" pageNumber="88">P. armiger</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="9F5036C84D2A637F62A0E6F4FCF024DF" blockId="11.[151,1436,151,933]" box="[199,866,223,248]" pageId="11" pageNumber="88">
(2) An ocular papilla (
<emphasis id="AD9BEADA4D2A637F63A6E6F5FD9A24DF" box="[449,520,225,248]" italics="true" pageId="11" pageNumber="88">versus</emphasis>
without papilla in 
<taxonomicName id="58EF4D4B4D2A637F60BBE6F5FCC024DF" ID-CoL="4FCNW" box="[732,850,225,248]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="armiger">
<emphasis id="AD9BEADA4D2A637F60BBE6F5FCC024DF" box="[732,850,225,248]" italics="true" pageId="11" pageNumber="88">P. armiger</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="9F5036C84D2A637F62A0E710FB76253B" blockId="11.[151,1436,151,933]" box="[199,1252,260,285]" pageId="11" pageNumber="88">
(3) Antennal scale relatively longer and with an apical lobe that is not described in 
<emphasis id="AD9BEADA4D2A637F660EE711FB76253B" box="[1129,1252,261,284]" italics="true" pageId="11" pageNumber="88">
<taxonomicName id="58EF4D4B4D2A637F660EE711FB72253B" ID-CoL="4FCNW" box="[1129,1248,261,284]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="armiger">P. armiger</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="9F5036C84D2A637F62A0E73CFE9325AF" blockId="11.[151,1436,151,933]" pageId="11" pageNumber="88">
(4) Molar process of mandible with only one chitinous ridge (
<emphasis id="AD9BEADA4D2A637F611BE73DFC512567" box="[892,963,297,320]" italics="true" pageId="11" pageNumber="88">versus</emphasis>
three ridges separated by deep grooves in 
<taxonomicName id="58EF4D4B4D2A637F62F0E759FE822543" ID-CoL="4FCNW" box="[151,272,333,356]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="armiger">
<emphasis id="AD9BEADA4D2A637F62F0E759FE822543" box="[151,272,333,356]" italics="true" pageId="11" pageNumber="88">P. armiger</emphasis>
</taxonomicName>
. First article of mandibular palp with a small lobe on its inner distal margin (
<emphasis id="AD9BEADA4D2A637F66D9E759FA972543" box="[1214,1285,333,356]" italics="true" pageId="11" pageNumber="88">versus</emphasis>
absent in 
<taxonomicName id="58EF4D4B4D2A637F67E0E759FF6325AF" ID-CoL="4FCNW" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="armiger">
<emphasis id="AD9BEADA4D2A637F67E0E759FF6325AF" italics="true" pageId="11" pageNumber="88">P. armiger</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="9F5036C84D2A637F62A0E780FD1425F7" blockId="11.[151,1436,151,933]" pageId="11" pageNumber="88">
(5) Male second pleopod with a rounded distal lobe on exopod first article, bearing two strong simple setae (
<emphasis id="AD9BEADA4D2A637F62C7E7ADFF7A25F7" box="[160,232,441,464]" italics="true" pageId="11" pageNumber="88">versus</emphasis>
lobe and setae absent in 
<taxonomicName id="58EF4D4B4D2A637F6398E7ADFDE725F7" ID-CoL="4FCNW" box="[511,629,441,464]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="armiger">
<emphasis id="AD9BEADA4D2A637F6398E7ADFDE725F7" box="[511,629,441,464]" italics="true" pageId="11" pageNumber="88">P. armiger</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="9F5036C84D2A637F62A0E7C8FA98263F" blockId="11.[151,1436,151,933]" pageId="11" pageNumber="88">
(6) Basal article of uropodal exopod with three distal cuspidate setae on outer distal corner, lengthening from outer to inner one (versus two or three cuspidate setae, lengthening from inner to outer in 
<taxonomicName id="58EF4D4B4D2A637F66E3E415FB68263F" ID-CoL="4FCNW" box="[1156,1274,513,536]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="armiger">
<emphasis id="AD9BEADA4D2A637F66E3E415FB68263F" box="[1156,1274,513,536]" italics="true" pageId="11" pageNumber="88">P. armiger</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="9F5036C84D2A637F62A0E430FAFF261B" blockId="11.[151,1436,151,933]" box="[199,1389,548,573]" pageId="11" pageNumber="88">
(7) Telson of mature specimens with 25–37 setae along lateral margins (
<emphasis id="AD9BEADA4D2A637F6188E431FBA4261B" box="[1007,1078,549,572]" italics="true" pageId="11" pageNumber="88">versus</emphasis>
40–50 setae in 
<taxonomicName id="58EF4D4B4D2A637F6681E431FACE261B" ID-CoL="4FCNW" box="[1254,1372,549,572]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="armiger">
<emphasis id="AD9BEADA4D2A637F6681E431FACE261B" box="[1254,1372,549,572]" italics="true" pageId="11" pageNumber="88">P. armiger</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="9F5036C84D2A637F62A0E45CFEE5268F" blockId="11.[151,1436,151,933]" pageId="11" pageNumber="88">
(8) Telson apex armed with 3–4 pairs of serrate setae decreasing in length towards the medial point and 3–8 pairs of small ventral teeth (
<emphasis id="AD9BEADA4D2A637F63B1E479FD8F26A3" box="[470,541,621,644]" italics="true" pageId="11" pageNumber="88">versus</emphasis>
5 pairs of serrate setae, a single median serrate seta and 5 pair of small ventral teeth in 
<taxonomicName id="58EF4D4B4D2A637F6296E485FEF4268F" ID-CoL="4FCNW" box="[241,358,657,680]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="armiger">
<emphasis id="AD9BEADA4D2A637F6296E485FEF4268F" box="[241,358,657,680]" italics="true" pageId="11" pageNumber="88">P. armiger</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="9F5036C84D2A637F62A0E4A0FD1A2782" blockId="11.[151,1436,151,933]" pageId="11" pageNumber="88">
The endopod of the third and fourth thoracopods of 
<taxonomicName id="58EF4D4B4D2A637F6170E4A1FC4626EA" box="[791,980,692,717]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="papilloculatus" status="sp. nov.">
<emphasis id="AD9BEADA4D2A637F6170E4A1FC4626EA" box="[791,980,692,717]" italics="true" pageId="11" pageNumber="88">P. papilloculatus</emphasis>
</taxonomicName>
<emphasis id="AD9BEADA4D2A637F61BBE4A0FBA526EA" bold="true" box="[988,1079,692,717]" pageId="11" pageNumber="88">
<taxonomicNameLabel id="B6A857A14D2A637F61BBE4A0FBA526EA" box="[988,1079,692,717]" pageId="11" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shows a rudimentary structure, reduced to 1–2 articles. Such a peculiarity was observed in all the specimens herein examined and was previously mentioned for 
<taxonomicName id="58EF4D4B4D2A637F635CE4E9FCF12733" ID-CoL="4FCNZ" authority="Tchindonova and Vereshchaka 1991" authorityName="Tchindonova and Vereshchaka" authorityYear="1991" box="[315,867,764,789]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="macrops">
<emphasis id="AD9BEADA4D2A637F635CE4E9FE272733" box="[315,437,765,788]" italics="true" pageId="11" pageNumber="88">P. macrops</emphasis>
(
<bibRefCitation id="FB7E4B394D2A637F63A3E4E8FCC92733" author="Tchindonova" box="[452,859,764,789]" pageId="11" pageNumber="88" refString="Tchindonova, Y. G. &amp; Vereshchaka, A. L. (1991) New mysid species Petalophthalmus macrops sp. n. from the west part of the Indian Ocean. Zoologicheskii Zhurnal, 70 (1), 135 - 138." type="journal article" year="1991">Tchindonova and Vereshchaka 1991</bibRefCitation>
)
</taxonomicName>
and for 
<taxonomicName id="58EF4D4B4D2A637F61A7E4E9FB0E2733" ID-CoL="4FCNY" authority="Wang 1998" authorityName="Wang" authorityYear="1998" box="[960,1180,764,789]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="liui">
<emphasis id="AD9BEADA4D2A637F61A7E4E9FB932732" box="[960,1025,764,789]" italics="true" pageId="11" pageNumber="88">P. liui</emphasis>
(
<bibRefCitation id="FB7E4B394D2A637F6668E4E8FB062733" author="Wang" box="[1039,1172,764,788]" pageId="11" pageNumber="88" refString="Wang, S. (1998) On new and rare species of Mysidacea (Crustacea) from the Northern South China Sea. Studia Marina Sinica = Hai Yang Ke Xue Ji Kan, 40, 199 - 244." type="journal article" year="1998">Wang 1998</bibRefCitation>
)
</taxonomicName>
. On the contrary, these thoracopod endopods are not rudimentary in 
<taxonomicName id="58EF4D4B4D2A637F60F7E535FCA4271F" ID-CoL="4FCNX" box="[656,822,799,824]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="caribbeanus">
<emphasis id="AD9BEADA4D2A637F60F7E535FCA4271F" box="[656,822,799,824]" italics="true" pageId="11" pageNumber="88">P. caribbeanus</emphasis>
</taxonomicName>
, as shown by 
<bibRefCitation id="FB7E4B394D2A637F61B0E50BFB02271F" author="Tattersall" box="[983,1168,799,824]" pageId="11" pageNumber="88" refString="Tattersall, O. S. (1968) A new species of Petalophthalmus (Mysidacea) based on specimens from off Puerto Rico, hitherto referred to Petalophthalmus oculatus. Journal of Zoology, 155, 271 - 282. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1968. tb 03052. x" type="journal article" year="1968">Tattersall (1968)</bibRefCitation>
and were not described in the case of 
<taxonomicName id="58EF4D4B4D2A637F6333E551FE40277B" ID-CoL="4FCNW" box="[340,466,837,860]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="armiger">
<emphasis id="AD9BEADA4D2A637F6333E551FE40277B" box="[340,466,837,860]" italics="true" pageId="11" pageNumber="88">P. armiger</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="FB7E4B394D2A637F6078E550FCBF277A" author="Willemoes-Suhm" box="[543,813,836,861]" pageId="11" pageNumber="88" refString="Willemoes-Suhm, R. von. (1875) On some Atlantic Crustacea from the ' Challenger' Expedition (communicated by W. Thomson). Transactions of the Linnean Society of London. Second Series. Zoology, 1 (1), 23 - 59." type="journal article" year="1875">Willemoes-Suhm 1875</bibRefCitation>
; 
<bibRefCitation id="FB7E4B394D2A637F6126E550FC2E277A" author="Sars" box="[833,956,836,861]" pageId="11" pageNumber="88" refString="Sars, G. O. (1885) Report on the Schizopoda collected by H. M. S. &quot; Challenger &quot; during the years 1873 - 1876. Reports of the scientific Results of the Voyage of H. M. S. Challenger Zoology, 13 (37), 1 - 128." type="journal article" year="1885">Sars 1885</bibRefCitation>
; Tatttersall and 
<bibRefCitation id="FB7E4B394D2A637F66E0E550FAA8277A" author="Tattersall" box="[1159,1338,836,861]" pageId="11" pageNumber="88" refString="Tattersall, W. M. &amp; Tattersall, O. S. (1951) The British Mysidacea. Ray Society, London, 1 - 460." type="book chapter" year="1951">Tattersall 1951</bibRefCitation>
; broken appendages) and 
<taxonomicName id="58EF4D4B4D2A637F6301E57DFE7527A7" ID-CoL="4FCP2" box="[358,487,871,896]" class="Malacostraca" family="Petalophthalmidae" genus="Petalophthalmus" kingdom="Animalia" order="Mysida" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="oculatus">
<emphasis id="AD9BEADA4D2A637F6301E57DFE7527A7" box="[358,487,871,896]" italics="true" pageId="11" pageNumber="88">P. oculatus</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="FB7E4B394D2A637F6048E573FD2127A7" author="Pillai" box="[559,691,871,896]" pageId="11" pageNumber="88" refString="Pillai, N. K. (1968) Redescription of Petalophthalmus oculatus (Crustacea: Mysidacea) based on an adult female collected from near the type locality. Journal of Zoology, 155, 283 - 291. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1968. tb 03053. x" type="journal article" year="1968">Pillai 1968</bibRefCitation>
; the figured ‘fourth thoracopod 4’ is in fact the fifth one, as demonstrated by the habitus of this species).
</paragraph>
</subSubSection>
</treatment>
</document>