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<document id="C5E8203357C9021F781AA05672DAE6FB" ID-DOI="10.1649/611" ID-ISSN="1938-4394" ID-Zenodo-Dep="10104359" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630168753109" checkinUser="tatiana" docAuthor="Sokolov, Igor M., Carlton, Christopher &amp; Cornell, James F." docDate="2004" docId="845C87831260FE205E0EFE0CFDA5FDF6" docLanguage="en" docName="611.pdf.imf" docOrigin="The Coleopterists Bulletin 58 (2)" docSource="http://dx.doi.org/10.1649/611" docStyle="DocumentStyle:07711681861DB98305E87147CAB6DAD5.4:TheColeopBull.2003-2009.journal_article.0cover" docStyleId="07711681861DB98305E87147CAB6DAD5" docStyleName="TheColeopBull.2003-2009.journal_article.0cover" docStyleVersion="4" docTitle="Anillinus lescheni Sokolov and Carlton, new species" docType="treatment" docVersion="3" lastPageNumber="218" masterDocId="7865FFFB1240FE015F43FFFDFFABFFE0" masterDocTitle="Review of Anillinus, with Descriptions of 17 New Species and a Key to Soil and Litter Species (Coleoptera: Carabidae: Trechinae: Bembidiini)" masterLastPageNumber="233" masterPageNumber="185" pageNumber="217" updateTime="1699629423378" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D90D1D727267C00AC0964824A5586CF5">Review of Anillinus, with Descriptions of 17 New Species and a Key to Soil and Litter Species (Coleoptera: Carabidae: Trechinae: Bembidiini)</mods:title>
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<mods:namePart id="775653F722949DA9398C76EE909F2057">Sokolov, Igor M.</mods:namePart>
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<mods:namePart id="58C10D0B7A3E4B686BB39EDE55DD8DDB">Carlton, Christopher</mods:namePart>
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<mods:namePart id="3495A3593420637DCC4250514148CB8E">Cornell, James F.</mods:namePart>
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<mods:title id="AB0A09CA937C126A9B308C84AA147EC6">The Coleopterists Bulletin</mods:title>
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<mods:date id="1EC098015BC86454C51370B7CF59BA7B">2004</mods:date>
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<heading id="570281F91260FE215E0EFE0CFCE5FDE6" box="[333,846,497,518]" centered="true" fontSize="9" level="2" pageId="32" pageNumber="217" reason="2">
<taxonomicName id="CBF54D161260FE215E0EFE0CFD60FDE6" ID-CoL="5V28D" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" box="[333,715,497,518]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="217" phylum="Arthropoda" rank="species" species="lescheni" status="sp. nov.">
<emphasis id="3E81EA871260FE215E0EFE0CFE53FDE6" box="[333,504,497,518]" italics="true" pageId="32" pageNumber="217">Anillinus lescheni</emphasis>
Sokolov and Carlton
</taxonomicName>
,
<emphasis id="3E81EA871260FE215D94FE0CFCE5FDE6" bold="true" box="[727,846,497,518]" pageId="32" pageNumber="217">
<taxonomicNameLabel id="25B257FC1260FE215D94FE0CFCE5FDE6" box="[727,846,497,518]" pageId="32" pageNumber="217" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="44EF651E1260FE215EFAFDF1FD48FDC1" box="[441,739,524,545]" pageId="32" pageNumber="217" type="description">
<paragraph id="0C4A36951260FE215EFAFDF1FD48FDC1" blockId="32.[333,846,497,545]" box="[441,739,524,545]" pageId="32" pageNumber="217">
(
<figureCitation id="94CE2A101260FE215E83FDF1FDBAFDC1" box="[448,529,524,545]" captionStart="Figs" captionStartId="7.[169,208,1334,1353]" captionText="Figs. 1013. Anillinus spp., right metafemora, setae omitted. 10) A. elongatus Jeannel;" pageId="32" pageNumber="217">Figs. 12</figureCitation>
,
<figureCitation id="94CE2A101260FE215D5EFDF1FD9CFDC1" box="[541,567,524,545]" captionStart="Figs" captionStartId="7.[169,208,1404,1423]" captionTargetBox="[160,966,196,1313]" captionTargetId="figure-1@7.[159,966,194,1313]" captionTargetPageId="7" captionText="Figs. 1417. Anillinus spp., right metatibiae, setae and spurs omitted. 14) A. elongatus Jeannel; 15) A. lescheni n. sp.; 16) A. fortis (Horn); 17) A. stephani n. sp." figureDoi="http://doi.org/10.5281/zenodo.10104366" httpUri="https://zenodo.org/record/10104366/files/figure.png" pageId="32" pageNumber="217">15</figureCitation>
,
<figureCitation id="94CE2A101260FE215D00FDF1FDF6FDC1" box="[579,605,524,545]" captionStart="Figs" captionStartId="9.[169,208,1196,1215]" captionTargetBox="[154,974,196,1174]" captionTargetId="figure-309@9.[151,975,194,1175]" captionTargetPageId="9" captionText="Figs. 2531. Aedeagi of Anillinus spp., right lateral aspects, scale ¼ 0.1 mm. 25) A. docwatsoni n. sp.; 26) A. erwini n.sp.; 27) A. folkertsi n.sp.; 28) A. kovariki n.sp.; 29) Anillinus langdoni n.sp.; 30) A. lescheni n. sp.; 31) A. loweae n.sp." figureDoi="http://doi.org/10.5281/zenodo.10104370" httpUri="https://zenodo.org/record/10104370/files/figure.png" pageId="32" pageNumber="217">30</figureCitation>
, Map fig. 40)
</paragraph>
</subSubSection>
<subSubSection id="44EF651E1260FE215F84FDCEFF5EFC12" pageId="32" pageNumber="217" type="materials_examined">
<paragraph id="0C4A36951260FE215F84FDCEFC9AFD83" blockId="32.[175,1005,563,1010]" pageId="32" pageNumber="217">
<materialsCitation id="BC9D3CC81260FE215F84FDCEFC4EFDA9" box="[199,997,563,585]" collectionCode="USNM" collectorName="K. Stephan" county="Latimer Co." location="Male" pageId="32" pageNumber="217" specimenCount="1" stateProvince="Oklahoma" typeStatus="holotype">
<emphasis id="3E81EA871260FE215F84FDCEFE8FFDA8" bold="true" box="[199,292,563,584]" pageId="32" pageNumber="217">
<typeStatus id="D34E88371260FE215F84FDCEFE8FFDA8" box="[199,292,563,584]" pageId="32" pageNumber="217" type="holotype">Holotype</typeStatus>
</emphasis>
(
<collectionCode id="6AE4AE501260FE215E7AFDC9FE2BFDA9" box="[313,384,564,585]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="32" pageNumber="217" type="Museum">USNM</collectionCode>
).
<location id="092A604E1260FE215EDAFDC9FE61FDA9" LSID="urn:lsid:plazi:treatment:845C87831260FE205E0EFE0CFDA5FDF6:092A604E1260FE215EDAFDC9FE61FDA9" box="[409,458,564,585]" county="Latimer Co." name="Male" pageId="32" pageNumber="217" stateProvince="Oklahoma">Male</location>
labeled /
<collectingRegion id="CE31F8771260FE215D72FDC9FD3DFDA9" box="[561,662,564,585]" country="United States of America" name="Oklahoma" pageId="32" pageNumber="217">Oklahoma</collectingRegion>
,
<collectingCounty id="E52B4E191260FE215DE4FDC9FC8FFDA9" box="[679,804,564,585]" pageId="32" pageNumber="217">Latimer Co.</collectingCounty>
, IV-85,
<collectorName id="A10053431260FE215CC1FDC9FC4EFDA9" box="[898,997,564,585]" pageId="32" pageNumber="217">K.Stephan</collectorName>
</materialsCitation>
/
<typeStatus id="D34E88371260FE215FECFDB3FE84FD83" box="[175,303,590,611]" pageId="32" pageNumber="217" type="holotype">HOLOTYPE</typeStatus>
,
<taxonomicName id="CBF54D161260FE215E79FDB3FD11FD83" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" box="[314,698,590,611]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="217" phylum="Arthropoda" rank="species" species="lescheni">Anillinus lescheni Sokolov and Carlton</taxonomicName>
, des. 2004/.
</paragraph>
<paragraph id="0C4A36951260FE215F84FD94FC46FC68" blockId="32.[175,1005,563,1010]" pageId="32" pageNumber="217">
<emphasis id="3E81EA871260FE215F84FD94FE86FD9E" bold="true" box="[199,301,617,638]" pageId="32" pageNumber="217">
<typeStatus id="D34E88371260FE215F84FD94FE86FD9E" box="[199,301,617,638]" pageId="32" pageNumber="217" type="paratype">Paratypes</typeStatus>
</emphasis>
(21). Four males and
<materialsCitation id="BC9D3CC81260FE215D4AFD94FEACFD53" collectingDate="1982-04" collectionCode="TAMU" collectorName="Karl Stephan" county="Latimer Co." location="One" pageId="32" pageNumber="217" specimenCount="2" specimenCount-female="2" stateProvince="Oklahoma" typeStatus="paratype">
<specimenCount id="1AF3FD1C1260FE215D4AFD94FDCEFD9E" box="[521,613,617,638]" pageId="32" pageNumber="217" type="female">2 females</specimenCount>
with same data as holotype.
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female labeled /
<collectingRegion id="CE31F8771260FE215E44FD7EFEC7FD78" box="[263,364,643,664]" country="United States of America" name="Oklahoma" pageId="32" pageNumber="217">Oklahoma</collectingRegion>
,
<collectingCounty id="E52B4E191260FE215E3FFD7EFE5CFD78" box="[380,503,643,664]" pageId="32" pageNumber="217">Latimer Co.</collectingCounty>
,
<date id="784B10551260FE215D44FD7EFDD3FD78" box="[519,632,643,664]" pageId="32" pageNumber="217" value="1982-04">
<collectingDate id="680FE9BD1260FE215D44FD7EFDD3FD78" box="[519,632,643,664]" pageId="32" pageNumber="217" value="1982-04">April 1982</collectingDate>
</date>
,
<collectorName id="A10053431260FE215DCBFD7EFCA1FD78" box="[648,778,643,664]" pageId="32" pageNumber="217">Karl Stephan</collectorName>
/ collected under rock (
<collectionCode id="6AE4AE501260FE215FF5FD63FEABFD53" box="[182,256,670,691]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="32" pageNumber="217" type="Herbarium">TAMU</collectionCode>
)
</materialsCitation>
;/
<materialsCitation id="BC9D3CC81260FE215E59FD63FDBAFD2E" collectionCode="TAMU, JFC" collectorName="Karl Stephan" county="Latimer Co." location="Latimer Co." pageId="32" pageNumber="217" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Oklahoma" typeStatus="paratype">
<specimenCount id="1AF3FD1C1260FE215E59FD63FEC3FD53" box="[282,360,670,691]" pageId="32" pageNumber="217" type="male">2 males</specimenCount>
,
<specimenCount id="1AF3FD1C1260FE215E37FD63FE63FD53" box="[372,456,670,691]" pageId="32" pageNumber="217" type="female">1 female</specimenCount>
labeled /
<collectingRegion id="CE31F8771260FE215D66FD63FD21FD53" box="[549,650,670,691]" country="United States of America" name="Oklahoma" pageId="32" pageNumber="217">Oklahoma</collectingRegion>
,
<collectingCounty id="E52B4E191260FE215DD6FD63FCA6FD53" box="[661,781,670,691]" pageId="32" pageNumber="217">Latimer Co.</collectingCounty>
, -IV-83,
<collectorName id="A10053431260FE215C24FD63FC4EFD53" box="[871,997,670,691]" pageId="32" pageNumber="217">Karl Stephan</collectorName>
/ collected under rock/ (
<collectionCode id="6AE4AE501260FE215EC9FD44FE7FFD2E" box="[394,468,697,718]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="32" pageNumber="217" type="Herbarium">TAMU</collectionCode>
,
<collectionCode id="6AE4AE501260FE215EA3FD44FDA1FD2E" box="[480,522,697,718]" pageId="32" pageNumber="217">JFC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="BC9D3CC81260FE215D5EFD44FD95FD08" collectionCode="JFC" collectorName="Karl Stephan" county="Latimer Co." location="Latimer Co." pageId="32" pageNumber="217" specimenCount="1" specimenCount-male="1" stateProvince="Oklahoma" typeStatus="paratype">
<specimenCount id="1AF3FD1C1260FE215D5EFD44FDF4FD2E" box="[541,607,697,718]" pageId="32" pageNumber="217" type="male">1 male</specimenCount>
labeled /
<collectingRegion id="CE31F8771260FE215DF8FD44FC8BFD2E" box="[699,800,697,718]" country="United States of America" name="Oklahoma" pageId="32" pageNumber="217">Oklahoma</collectingRegion>
,
<collectingCounty id="E52B4E191260FE215C68FD44FC09FD2E" box="[811,930,697,718]" pageId="32" pageNumber="217">Latimer Co.</collectingCounty>
, IV-85,
<collectorName id="A10053431260FE215FECFD2EFE80FD08" box="[175,299,723,744]" pageId="32" pageNumber="217">Karl Stephan</collectorName>
/ collected under rock/ (
<collectionCode id="6AE4AE501260FE215D4EFD2EFD93FD08" box="[525,568,723,744]" pageId="32" pageNumber="217">JFC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="BC9D3CC81260FE215D0BFD2EFE17FCE3" collectingDate="1985-06" collectionCode="TAMU" collectorName="K. Stephan" county="Latimer Co." location="Latimer Co." pageId="32" pageNumber="217" specimenCount="1" specimenCount-male="1" stateProvince="Oklahoma" typeStatus="paratype">
<specimenCount id="1AF3FD1C1260FE215D0BFD2EFD23FD08" box="[584,648,723,744]" pageId="32" pageNumber="217" type="male">1 male</specimenCount>
labeled /
<collectingRegion id="CE31F8771260FE215D9CFD2EFCEFFD08" box="[735,836,723,744]" country="United States of America" name="Oklahoma" pageId="32" pageNumber="217">Oklahoma</collectingRegion>
,
<collectingCounty id="E52B4E191260FE215C0EFD2EFC69FD08" box="[845,962,723,744]" pageId="32" pageNumber="217">Latimer Co.</collectingCounty>
,
<date id="784B10551260FE215C8FFD2EFF4AFCE3" pageId="32" pageNumber="217" value="1985-06">
<collectingDate id="680FE9BD1260FE215C8FFD2EFF4AFCE3" pageId="32" pageNumber="217" value="1985-06">VI- 1985</collectingDate>
</date>
,
<collectorName id="A10053431260FE215FACFD13FEF9FCE3" box="[239,338,750,771]" pageId="32" pageNumber="217">K.Stephan</collectorName>
/ (
<collectionCode id="6AE4AE501260FE215E29FD13FE1EFCE3" box="[362,437,750,771]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="32" pageNumber="217" type="Herbarium">TAMU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="BC9D3CC81260FE215E8AFD13FEEBFCFD" collectingDate="1987-03" collectionCode="JFC" collectorName="Karl Stephan" county="Latimer Co." location="Latimer Co." pageId="32" pageNumber="217" specimenCount="1" specimenCount-male="1" stateProvince="Oklahoma" typeStatus="paratype">
<specimenCount id="1AF3FD1C1260FE215E8AFD13FDA6FCE3" box="[457,525,750,771]" pageId="32" pageNumber="217" type="male">1 male</specimenCount>
labeled /
<collectingRegion id="CE31F8771260FE215D2FFD13FD7AFCE3" box="[620,721,750,771]" country="United States of America" name="Oklahoma" pageId="32" pageNumber="217">Oklahoma</collectingRegion>
,
<collectingCounty id="E52B4E191260FE215D9DFD13FCFDFCE3" box="[734,854,750,771]" pageId="32" pageNumber="217">Latimer Co.</collectingCounty>
,
<date id="784B10551260FE215C27FD13FC1EFCE3" box="[868,949,750,771]" pageId="32" pageNumber="217" value="1987-03">
<collectingDate id="680FE9BD1260FE215C27FD13FC1EFCE3" box="[868,949,750,771]" pageId="32" pageNumber="217" value="1987-03">III-1987</collectingDate>
</date>
,
<collectorName id="A10053431260FE215C81FD13FF50FCFD" pageId="32" pageNumber="217">Karl Stephan</collectorName>
/ (
<collectionCode id="6AE4AE501260FE215E4CFCF5FE91FCFD" box="[271,314,776,797]" pageId="32" pageNumber="217">JFC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="BC9D3CC81260FE215E09FCF5FC43FCFD" box="[330,1000,776,797]" collectingDate="1987-04" collectionCode="JFC" collectorName="Karl Stephan" county="Latimer Co." location="Latimer Co." pageId="32" pageNumber="217" specimenCount="1" specimenCount-male="1" stateProvince="Oklahoma" typeStatus="paratype">
<specimenCount id="1AF3FD1C1260FE215E09FCF5FE21FCFD" box="[330,394,776,797]" pageId="32" pageNumber="217" type="male">1 male</specimenCount>
labeled /
<collectingRegion id="CE31F8771260FE215EA3FCF5FDEEFCFD" box="[480,581,776,797]" country="United States of America" name="Oklahoma" pageId="32" pageNumber="217">Oklahoma</collectingRegion>
,
<collectingCounty id="E52B4E191260FE215D0DFCF5FD68FCFD" box="[590,707,776,797]" pageId="32" pageNumber="217">Latimer Co.</collectingCounty>
,
<date id="784B10551260FE215D8FFCF5FCB5FCFD" box="[716,798,776,797]" pageId="32" pageNumber="217" value="1987-04">
<collectingDate id="680FE9BD1260FE215D8FFCF5FCB5FCFD" box="[716,798,776,797]" pageId="32" pageNumber="217" value="1987-04">IV-1987</collectingDate>
</date>
,
<collectorName id="A10053431260FE215C6BFCF5FC08FCFD" box="[808,931,776,797]" pageId="32" pageNumber="217">Karl Stephan</collectorName>
/ (
<collectionCode id="6AE4AE501260FE215CF4FCF5FC49FCFD" box="[951,994,776,797]" pageId="32" pageNumber="217">JFC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="BC9D3CC81260FE215FECFCDEFCD7FCD8" box="[175,892,803,824]" collectingDate="1989-03" collectionCode="JFC" collectorName="Karl Stephan" county="Latimer Co." location="Latimer Co." pageId="32" pageNumber="217" specimenCount="1" specimenCount-male="1" stateProvince="Oklahoma" typeStatus="paratype">
<specimenCount id="1AF3FD1C1260FE215FECFCDEFF5FFCD8" box="[175,244,803,824]" pageId="32" pageNumber="217" type="male">1 male</specimenCount>
labeled /
<collectingRegion id="CE31F8771260FE215E15FCDEFE10FCD8" box="[342,443,803,824]" country="United States of America" name="Oklahoma" pageId="32" pageNumber="217">Oklahoma</collectingRegion>
,
<collectingCounty id="E52B4E191260FE215E8AFCDEFDE8FCD8" box="[457,579,803,824]" pageId="32" pageNumber="217">Latimer Co.</collectingCounty>
,
<date id="784B10551260FE215D12FCDEFD09FCD8" box="[593,674,803,824]" pageId="32" pageNumber="217" value="1989-03">
<collectingDate id="680FE9BD1260FE215D12FCDEFD09FCD8" box="[593,674,803,824]" pageId="32" pageNumber="217" value="1989-03">III-1989</collectingDate>
</date>
,
<collectorName id="A10053431260FE215DF2FCDEFC9AFCD8" box="[689,817,803,824]" pageId="32" pageNumber="217">Karl Stephan</collectorName>
/ (
<collectionCode id="6AE4AE501260FE215C09FCDEFCDEFCD8" box="[842,885,803,824]" pageId="32" pageNumber="217">JFC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="BC9D3CC81260FE215CC9FCDEFCDBFCB2" collectingDate="1991-04" collectionCode="TAMU" collectorName="Karl Stephan" county="Latimer Co." location="Latimer Co." pageId="32" pageNumber="217" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Oklahoma" typeStatus="paratype">
<specimenCount id="1AF3FD1C1260FE215CC9FCDEFC7AFCD8" box="[906,977,803,824]" pageId="32" pageNumber="217" type="male">1 male</specimenCount>
,
<specimenCount id="1AF3FD1C1260FE215CA3FCDEFF44FCB2" pageId="32" pageNumber="217" type="female">1 female</specimenCount>
labeled /
<collectingRegion id="CE31F8771260FE215E06FCC0FE01FCB2" box="[325,426,829,850]" country="United States of America" name="Oklahoma" pageId="32" pageNumber="217">Oklahoma</collectingRegion>
,
<collectingCounty id="E52B4E191260FE215EF1FCC0FD8EFCB2" box="[434,549,829,850]" pageId="32" pageNumber="217">Latimer Co.</collectingCounty>
, -
<date id="784B10551260FE215D77FCC0FD23FCB2" box="[564,648,829,850]" pageId="32" pageNumber="217" value="1991-04">
<collectingDate id="680FE9BD1260FE215D77FCC0FD23FCB2" box="[564,648,829,850]" pageId="32" pageNumber="217" value="1991-04">IV-1991</collectingDate>
</date>
,
<collectorName id="A10053431260FE215DD2FCC0FCA0FCB2" box="[657,779,829,850]" pageId="32" pageNumber="217">Karl Stephan</collectorName>
/ (
<collectionCode id="6AE4AE501260FE215C5DFCC0FCC2FCB2" box="[798,873,829,850]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="32" pageNumber="217" type="Herbarium">TAMU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="BC9D3CC81260FE215C3BFCC0FC2AFC8D" collectingDate="1992-03" collectionCode="TAMU" collectorName="Karl Stephan" county="Latimer Co." location="Latimer Co." pageId="32" pageNumber="217" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Oklahoma" typeStatus="paratype">
<specimenCount id="1AF3FD1C1260FE215C3BFCC0FC12FCB2" box="[888,953,829,850]" pageId="32" pageNumber="217" type="male">1 male</specimenCount>
,
<specimenCount id="1AF3FD1C1260FE215C82FCC0FF76FC8D" pageId="32" pageNumber="217" type="female">1 female</specimenCount>
labeled /
<collectingRegion id="CE31F8771260FE215E7FFCA5FE0AFC8D" box="[316,417,856,877]" country="United States of America" name="Oklahoma" pageId="32" pageNumber="217">Oklahoma</collectingRegion>
,
<collectingCounty id="E52B4E191260FE215EEEFCA5FD8DFC8D" box="[429,550,856,877]" pageId="32" pageNumber="217">Latimer Co.</collectingCounty>
, -
<date id="784B10551260FE215D7AFCA5FD20FC8D" box="[569,651,856,877]" pageId="32" pageNumber="217" value="1992-03">
<collectingDate id="680FE9BD1260FE215D7AFCA5FD20FC8D" box="[569,651,856,877]" pageId="32" pageNumber="217" value="1992-03">III-1992</collectingDate>
</date>
,
<collectorName id="A10053431260FE215DDBFCA5FCB3FC8D" box="[664,792,856,877]" pageId="32" pageNumber="217">Karl Stephan</collectorName>
/ (
<collectionCode id="6AE4AE501260FE215C6CFCA5FCD1FC8D" box="[815,890,856,877]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="32" pageNumber="217" type="Herbarium">TAMU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="BC9D3CC81260FE215CCEFCA5FC41FC68" collectingDate="1992-03" collectionCode="TAMU" collectorName="Karl Stephan" county="Latimer Co." location="Latimer Co." pageId="32" pageNumber="217" specimenCount="2" specimenCount-female="2" stateProvince="Oklahoma" typeStatus="paratype">
<specimenCount id="1AF3FD1C1260FE215CCEFCA5FC47FC8D" box="[909,1004,856,877]" pageId="32" pageNumber="217" type="female">2 females</specimenCount>
labeled /
<collectingRegion id="CE31F8771260FE215E42FC8EFECDFC68" box="[257,358,883,904]" country="United States of America" name="Oklahoma" pageId="32" pageNumber="217">Oklahoma</collectingRegion>
,
<collectingCounty id="E52B4E191260FE215E33FC8EFE4EFC68" box="[368,485,883,904]" pageId="32" pageNumber="217">Latimer Co.</collectingCounty>
, -
<date id="784B10551260FE215EB6FC8EFDECFC68" box="[501,583,883,904]" pageId="32" pageNumber="217" value="1992-03">
<collectingDate id="680FE9BD1260FE215EB6FC8EFDECFC68" box="[501,583,883,904]" pageId="32" pageNumber="217" value="1992-03">III-1992</collectingDate>
</date>
,
<collectorName id="A10053431260FE215D12FC8EFD66FC68" box="[593,717,883,904]" pageId="32" pageNumber="217">Karl Stephan</collectorName>
/ ex gopher burrow/ (
<collectionCode id="6AE4AE501260FE215CD4FC8EFC49FC68" box="[919,994,883,904]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="32" pageNumber="217" type="Herbarium">TAMU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="0C4A36951260FE215F84FC70FF5EFC12" blockId="32.[175,1005,563,1010]" pageId="32" pageNumber="217">
<emphasis id="3E81EA871260FE215F84FC70FEC4FC42" bold="true" box="[199,367,909,930]" pageId="32" pageNumber="217">Specific Epithet.</emphasis>
This species is named for Richard Leschen (Landcare Research,
<collectingRegion id="CE31F8771260FE215FECFC55FEA5FC5D" box="[175,270,936,957]" country="New Zealand" name="Auckland" pageId="32" pageNumber="217">Auckland</collectingRegion>
,
<collectingCountry id="74E276051260FE215E59FC55FE36FC5D" box="[282,413,936,957]" name="New Zealand" pageId="32" pageNumber="217">New Zealand</collectingCountry>
) in honor of his contributions to
<taxonomicName id="CBF54D161260FE215DA6FC55FCE6FC5D" authorityName="Linnaeus" authorityYear="1758" box="[741,845,936,957]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="217" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
systematics and in recognition of Karl Stephans effectiveness in training coleopterists how to collect.
</paragraph>
</subSubSection>
<subSubSection id="44EF651E1260FE215F86FBFBFCB2FA00" pageId="32" pageNumber="217" type="description">
<paragraph id="0C4A36951260FE215F86FBFBFE0AFB71" blockId="32.[175,1006,1030,1504]" pageId="32" pageNumber="217">
<emphasis id="3E81EA871260FE215F86FBFBFE98FBF9" bold="true" box="[197,307,1030,1049]" pageId="32" pageNumber="217">Description.</emphasis>
Large-sized species (ABL range,
<quantity id="CB0D9B701260FE215D2DFBFBFD5BFBF9" box="[622,752,1030,1049]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="32" pageNumber="217" unit="mm" value="2.35" valueMax="2.5" valueMin="2.2">2.202.50 mm</quantity>
; mean,
<quantity id="CB0D9B701260FE215C7EFBFBFC3BFBF9" box="[829,912,1030,1049]" metricMagnitude="-3" metricUnit="m" metricValue="2.38" pageId="32" pageNumber="217" unit="mm" value="2.38">2.38 mm</quantity>
, n
<emphasis id="3E81EA871260FE215CF1FBFBFC6BFBF9" bold="true" box="[946,960,1030,1049]" pageId="32" pageNumber="217">¼</emphasis>
10). Habitus elongate, parallel-sided (WE/ABL, 0.29), head relatively broad (WH/WPm, 0.81), pronotum wide (WPm/WE, 0.97). Color of body various, from rufotestaceous to amber testaceous, appendages testaceous. Dorsal microsculpture distinct, covering pronotum and head except for small areas with indistinct microsculpture at anterior parts of frons between frontal carinae and frontal grooves.
</paragraph>
<paragraph id="0C4A36951260FE215F86FB6BFD60FB11" blockId="32.[175,1006,1030,1504]" pageId="32" pageNumber="217">Pronotum moderately convex, broad (WPm/LP, 1.46), with margins markedly constricted posteriad (WPm/WPp, 1.52) and slightly sinuate before posterior angles. Anterior angles evident, slightly prominent. Posterior angles nearly rectangular (951008). Width between anterior angles much greater than between posterior angles (WPa/WPp, 1.05).</paragraph>
<paragraph id="0C4A36951260FE215F86FB0BFE79FAD8" blockId="32.[175,1006,1030,1504]" pageId="32" pageNumber="217">Elytra slightly convex, widely depressed along suture, relatively short (LE/ABL, 0.57), with traces 34 interneurs. Humeri prominent, only slightly rounded. Vestiture of elytra short (less than one-fourth length of discal setae).</paragraph>
<paragraph id="0C4A36951260FE215F86FAC0FEE4FA78" blockId="32.[175,1006,1030,1504]" pageId="32" pageNumber="217">
Males with mesotibiae bearing a posterior row of approximately six, evenly spaced teeth. Metafemora (
<figureCitation id="94CE2A101260FE215E67FAA8FECFFA88" box="[292,356,1365,1384]" captionStart="Figs" captionStartId="7.[169,208,1334,1353]" captionText="Figs. 1013. Anillinus spp., right metafemora, setae omitted. 10) A. elongatus Jeannel;" pageId="32" pageNumber="217">Fig. 12</figureCitation>
) swollen, broadest in apical one-third and bearing a posterior row of low tubercles in middle one-third. Metatibiae (
<figureCitation id="94CE2A101260FE215D5FFA90FDF0FA60" box="[540,603,1389,1408]" captionStart="Figs" captionStartId="7.[169,208,1404,1423]" captionTargetBox="[160,966,196,1313]" captionTargetId="figure-1@7.[159,966,194,1313]" captionTargetPageId="7" captionText="Figs. 1417. Anillinus spp., right metatibiae, setae and spurs omitted. 14) A. elongatus Jeannel; 15) A. lescheni n. sp.; 16) A. fortis (Horn); 17) A. stephani n. sp." figureDoi="http://doi.org/10.5281/zenodo.10104366" httpUri="https://zenodo.org/record/10104366/files/figure.png" pageId="32" pageNumber="217">Fig. 15</figureCitation>
) bearing a posterior row of irregular rounded, granular tubercles.
</paragraph>
<paragraph id="0C4A36951260FE215F86FA60FCB2FA00" blockId="32.[175,1006,1030,1504]" pageId="32" pageNumber="217">
Median lobe (
<figureCitation id="94CE2A101260FE215E7DFA60FED5FA50" box="[318,382,1437,1456]" captionStart="Figs" captionStartId="9.[169,208,1196,1215]" captionTargetBox="[154,974,196,1174]" captionTargetId="figure-309@9.[151,975,194,1175]" captionTargetPageId="9" captionText="Figs. 2531. Aedeagi of Anillinus spp., right lateral aspects, scale ¼ 0.1 mm. 25) A. docwatsoni n. sp.; 26) A. erwini n.sp.; 27) A. folkertsi n.sp.; 28) A. kovariki n.sp.; 29) Anillinus langdoni n.sp.; 30) A. lescheni n. sp.; 31) A. loweae n.sp." figureDoi="http://doi.org/10.5281/zenodo.10104370" httpUri="https://zenodo.org/record/10104370/files/figure.png" pageId="32" pageNumber="217">Fig. 30</figureCitation>
) large, slightly arcuate, with an elongate ventrolateral lobe. Internal sac with large curved copulatory sclerite. Internal sac bearing approximately ten sharp, conical spines. Ventral sclerite absent, but two dense clusters of ventral spines present.
</paragraph>
</subSubSection>
<subSubSection id="44EF651E1260FE205F84FA0EFE67FE97" lastPageId="33" lastPageNumber="218" pageId="32" pageNumber="217" type="distribution">
<paragraph id="0C4A36951260FE215F84FA0EFC41F9C2" blockId="32.[175,1004,1523,1570]" pageId="32" pageNumber="217">
<emphasis id="3E81EA871260FE215F84FA0EFEE1F9E8" bold="true" box="[199,330,1523,1544]" pageId="32" pageNumber="217">Distribution.</emphasis>
Known from Latimer Co.,
<collectingRegion id="CE31F8771260FE215D1DFA0EFD6AF9E8" box="[606,705,1523,1544]" country="United States of America" name="Oklahoma" pageId="32" pageNumber="217">Oklahoma</collectingRegion>
(
<figureCitation id="94CE2A101260FE215D93FA0EFCB2F9E8" box="[720,793,1523,1544]" captionStart="Fig" captionStartId="19.[169,200,849,868]" captionText="Fig. 40. Locality records for Anillinus spp." pageId="32" pageNumber="217">Fig. 40</figureCitation>
). This species and its sympatric congener
<taxonomicName id="CBF54D161260FE215E30F9F0FE74F9C2" box="[371,479,1549,1570]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="217" phylum="Arthropoda" rank="species" species="stephani">
<emphasis id="3E81EA871260FE215E30F9F0FE74F9C2" box="[371,479,1549,1570]" italics="true" pageId="32" pageNumber="217">A. stephani</emphasis>
</taxonomicName>
represent the western-most occurrences of the genus.
</paragraph>
<paragraph id="0C4A36951261FE205FEFFF3FFE67FE97" blockId="33.[148,978,194,534]" pageId="33" pageNumber="218">
<emphasis id="3E81EA871261FE205FEFFF3FFEABFF37" bold="true" box="[172,256,194,215]" pageId="33" pageNumber="218">Habitat.</emphasis>
Although Karl Stephans collecting efforts encompass the entire area of Latimer County, two of his favorite collecting localities are his house near Red Oak and Robbers Cave State Park. Habitat where specimens were collected at the state park is rocky upland oak/hickory/shortleaf pine forest. Specimens were taken from beneath large rocks during rainy weather (K. Stephan pers. comm.). This locality is also the
<typeStatus id="D34E88371261FE205CEBFED0FC7AFEA2" box="[936,977,301,322]" pageId="33" pageNumber="218">type</typeStatus>
and only known locality for the pselaphine staphylinid
<taxonomicName id="CBF54D161261FE205DE9FEBAFE63FE97" authority="Carlton and Allen (Carlton and Allen 1989)" authorityName="Carlton and Allen (Carlton and Allen" authorityYear="1989" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="218" phylum="Arthropoda" rank="species" species="stephani">
<emphasis id="3E81EA871261FE205DE9FEBAFCF3FEBC" box="[682,856,327,348]" italics="true" pageId="33" pageNumber="218">Arianops stephani</emphasis>
Carlton and Allen (Carlton and Allen 1989)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="44EF651E1261FE205FEFFE81FDA5FDF6" pageId="33" pageNumber="218" type="diagnosis">
<paragraph id="0C4A36951261FE205FEFFE81FDA5FDF6" blockId="33.[148,978,194,534]" pageId="33" pageNumber="218">
<emphasis id="3E81EA871261FE205FEFFE81FE3CFE71" bold="true" box="[172,407,380,401]" pageId="33" pageNumber="218">Differential Diagnosis.</emphasis>
<taxonomicName id="CBF54D161261FE205EE0FE81FDF9FE71" box="[419,594,380,401]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="218" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="3E81EA871261FE205EE0FE81FDF9FE71" box="[419,594,380,401]" italics="true" pageId="33" pageNumber="218">Anillinus lescheni</emphasis>
</taxonomicName>
is parallel-sided with distinct microsculpture on the forebody (
<tableCitation id="4177032E1261FE205EE6FE6AFE5FFE4C" box="[421,500,407,428]" captionStart="Table 2" captionStartId="43.[167,219,627,646]" captionText="Table 2. Grouping of Anillinus species in accordance with their microsculpture patterns." pageId="33" pageNumber="218">Table 2</tableCitation>
, group II of endogean species). This species is sympatric with
<taxonomicName id="CBF54D161261FE205E02FE4FFE13FE27" authorityName="Carlton and Allen (Carlton and Allen" authorityYear="1989" box="[321,440,434,455]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="218" phylum="Arthropoda" rank="species" species="stephani">
<emphasis id="3E81EA871261FE205E02FE4FFE13FE27" box="[321,440,434,455]" italics="true" pageId="33" pageNumber="218">A. stephani</emphasis>
</taxonomicName>
, which shares the elongate habitus and distinct microsculpture. The unusual tubercles of the metatibiae and large ventrolateral process of the aedeagus of males distinguish this species from
<taxonomicName id="CBF54D161261FE205DDAFE1AFCA8FE1C" authorityName="Carlton and Allen (Carlton and Allen" authorityYear="1989" box="[665,771,487,508]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="218" phylum="Arthropoda" rank="species" species="stephani">
<emphasis id="3E81EA871261FE205DDAFE1AFCA8FE1C" box="[665,771,487,508]" italics="true" pageId="33" pageNumber="218">A. stephani</emphasis>
</taxonomicName>
and all other species. It is also much larger than
<taxonomicName id="CBF54D161261FE205EDFFDFCFDA3FDF6" authorityName="Carlton and Allen (Carlton and Allen" authorityYear="1989" box="[412,520,513,534]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="218" phylum="Arthropoda" rank="species" species="stephani">
<emphasis id="3E81EA871261FE205EDFFDFCFDA3FDF6" box="[412,520,513,534]" italics="true" pageId="33" pageNumber="218">A. stephani</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>