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</paragraph>
</subSubSection>
<subSubSection id="C3213CD7FC12FFE8FFE00C33FF35AB44" lastPageId="21" lastPageNumber="40" pageId="19" pageNumber="38" type="discussion">
<paragraph id="8B846F5CFC12FFEEFFE00C33FE26ADE2" blockId="19.[121,749,1798,1998]" pageId="19" pageNumber="38">
<emphasis id="B94FB34EFC12FFEEFFE00C33FF1FAD32" bold="true" box="[121,225,1798,1822]" pageId="19" pageNumber="38">Remarks</emphasis>
. The description of
<taxonomicName id="4C3B14DFFC12FFEEFE5C0C33FDA1AD31" authorityName="Skuse" authorityYear="1890" box="[453,607,1798,1821]" class="Insecta" family="Limoniidae" genus="Thrypticomyia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="aureipennis">
<emphasis id="B94FB34EFC12FFEEFE5C0C33FDA1AD31" box="[453,607,1798,1821]" italics="true" pageId="19" pageNumber="38">T. aureipennis</emphasis>
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provided by
<bibRefCitation id="EFAA12ADFC12FFEEFFE00C16FEFBAD17" author="Skuse, F. A. A." box="[121,261,1827,1851]" pageId="19" pageNumber="38" pagination="757 - 892" refId="ref12726" refString="Skuse, F. A. A. 1890. Diptera of Australia. Part VII. The Tipulidae Brevipalpi. Proceedings of the Linnean Society of New South Wales 4: 757 - 892. https: // doi. org / 10.5962 / bhl. part. 15067" type="journal article" year="1890">Skuse (1890)</bibRefCitation>
did not include a detailed description or figure of the hypopygium, making comparison with other species in the genus difficult. The availability of fresh material of
<taxonomicName id="4C3B14DFFC12FFEEFFE00C49FEEEADBF" authorityName="Skuse" authorityYear="1890" box="[121,272,1916,1939]" class="Insecta" family="Limoniidae" genus="Thrypticomyia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="aureipennis">
<emphasis id="B94FB34EFC12FFEEFFE00C49FEEEADBF" box="[121,272,1916,1939]" italics="true" pageId="19" pageNumber="38">T. aureipennis</emphasis>
</taxonomicName>
now allows for a thorough description of the hypopygium, a description of the ovipositor and provision of figures to aid in identification.
</paragraph>
<paragraph id="8B846F5CFC12FFEEFCA109C5FB1DA96F" blockId="19.[794,1422,752,835]" pageId="19" pageNumber="38">
Photographs are included to show the wing (
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), and a lateral view of the ovipositor (
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). A dorsal view of the hypopygium (
<figureCitation id="130073D9FC12FFEEFC4F081EFBDAA96F" box="[982,1060,811,835]" captionStart="Figure 45" captionStartId="20.[167,235,499,521]" captionTargetBox="[166,1459,159,481]" captionTargetId="figure-10@20.[166,1466,151,485]" captionTargetPageId="20" captionText="Figure 45. Thrypticomyia aureipennis male, wing." figureDoi="http://doi.org/10.5281/zenodo.7177964" httpUri="https://zenodo.org/record/7177964/files/figure.png" pageId="19" pageNumber="38">Fig. 45</figureCitation>
) is also provided.
</paragraph>
<paragraph id="8B846F5CFC12FFEEFC83086DFB59AF27" blockId="19.[794,1422,856,1291]" pageId="19" pageNumber="38">
<emphasis id="B94FB34EFC12FFEEFC83086DFCABA95C" bold="true" box="[794,853,856,880]" pageId="19" pageNumber="38">Male</emphasis>
. Hypopygium with tergite and sternite yellowish brown, gonocoxites darker greyish brown, lobe of inner gonostylus greyish white. Distal margin of T9 moderately bilobed, with broad U-shaped concavity. Ventral lobe of gonocoxite elongate, tip blunt, subequal to length of gonocoxite. Inner gonostylus 1½2 times the length of gonocoxite, lobe swollen, rounded, with an ovoid sclerite on dorsal face, two spines set on the medial edge of ovoid sclerite, each arising from a prominent tubercle, one at the base of the sclerite, one higher, rostral prolongation short, arched, tapering to a narrow tip. Outer gonostylus an evenly arched rod, narrowing gradually to a sharp tip. Aedeagus exceeding length of gonocoxite, parameres about ¾ the length of the aedeagus, lateral lobe low and rounded, mesal lobe gradually tapering to a blunt tip.
</paragraph>
<paragraph id="8B846F5CFC12FFEEFC830E15FB58AFBC" blockId="19.[794,1422,1312,1424]" pageId="19" pageNumber="38">
<emphasis id="B94FB34EFC12FFEEFC830E15FC93AF14" bold="true" box="[794,877,1312,1336]" pageId="19" pageNumber="38">Female</emphasis>
. Size and colouration much the same as for male. Ovipositor with tergite and sternite largely brownish. Cercus of moderate length, lightly curved. Tip of hypogynial valve reaching to about ¾ length of cercus.
</paragraph>
<paragraph id="8B846F5CFC12FFEEFC830E90FBB2ADE2" blockId="19.[793,1422,1445,1998]" pageId="19" pageNumber="38">
<emphasis id="B94FB34EFC12FFEEFC830E90FC6CAF91" bold="true" box="[794,914,1445,1469]" pageId="19" pageNumber="38">Discussion</emphasis>
. Five species of
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<emphasis id="B94FB34EFC12FFEEFBDA0E93FB21AF92" box="[1091,1247,1446,1470]" italics="true" pageId="19" pageNumber="38">Thrypticomyia</emphasis>
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are known from
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, predominantly from the Cairns area of northern
<collectingRegion id="49FFA1BEFC12FFEEFC830ED5FC65AFD4" box="[794,923,1504,1528]" country="Australia" name="Queensland" pageId="19" pageNumber="38">Queensland</collectingRegion>
, with
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<emphasis id="B94FB34EFC12FFEEFC410ED4FB90AFD4" box="[984,1134,1505,1528]" italics="true" pageId="19" pageNumber="38">T. aureipennis</emphasis>
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seemingly the only species extending south beyond
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. The Australian species are readily distinguished by features of the wing:
<taxonomicName id="4C3B14DFFC12FFEEFA820D29FC2CAC7C" authority="Alexander, 1956" authorityName="Alexander" authorityYear="1956" class="Insecta" family="Limoniidae" genus="Thrypticomyia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="marksae">
<emphasis id="B94FB34EFC12FFEEFA820D29FA73AC1F" box="[1307,1421,1563,1587]" italics="true" pageId="19" pageNumber="38">T. marksae</emphasis>
Alexander, 1956
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, having the discal medial cell open, all other species with discal medial cell closed;
<taxonomicName id="4C3B14DFFC12FFEEFB7C0D63FC36ACA7" authority="Alexander, 1921" authorityName="Alexander" authorityYear="1921" class="Insecta" family="Limoniidae" genus="Thrypticomyia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="fumidapicalis">
<emphasis id="B94FB34EFC12FFEEFB7C0D63FA73AC42" box="[1253,1421,1622,1646]" italics="true" pageId="19" pageNumber="38">T. fumidapicalis</emphasis>
Alexander, 1921
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, with clouded dark pigmentation in apical ⅕ of wing, remaining species may have greyish suffusion to the wing or darkened areas but not with the wing tip prominently darkened;
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<emphasis id="B94FB34EFC12FFEEFC130DF9FBD6ACCF" box="[906,1064,1740,1763]" italics="true" pageId="19" pageNumber="38">T. microstigma</emphasis>
Alexander, 1921
</taxonomicName>
, with Sc ending well before origin of Rs and stigma greatly reduced, in other species Sc close to origin of Rs and stigma more prominent;
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<emphasis id="B94FB34EFC12FFEEFC830C11FC8CAD17" box="[794,882,1827,1851]" italics="true" pageId="19" pageNumber="38">T. doddi</emphasis>
Alexander, 1921
</taxonomicName>
, and
<taxonomicName id="4C3B14DFFC12FFEEFBFF0C11FB01AD17" authorityName="Skuse" authorityYear="1890" box="[1126,1279,1828,1851]" class="Insecta" family="Limoniidae" genus="Thrypticomyia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="aureipennis">
<emphasis id="B94FB34EFC12FFEEFBFF0C11FB01AD17" box="[1126,1279,1828,1851]" italics="true" pageId="19" pageNumber="38">T. aureipennis</emphasis>
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very similar, differing in placement of a cross-vein between the costa and R, this cross-vein considered as the tip of R
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by
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, being only a short distance before R
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in
</quantity>
<taxonomicName id="4C3B14DFFC12FFEEFAB40C49FA79ADBF" authorityName="Alexander" authorityYear="1921" box="[1325,1415,1915,1939]" class="Insecta" family="Limoniidae" genus="Thrypticomyia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="doddi">
<emphasis id="B94FB34EFC12FFEEFAB40C49FA79ADBF" box="[1325,1415,1915,1939]" italics="true" pageId="19" pageNumber="38">T. doddi</emphasis>
</taxonomicName>
, while in
<taxonomicName id="4C3B14DFFC12FFEEFCEF0CACFBF2AD9C" authorityName="Skuse" authorityYear="1890" box="[886,1036,1945,1968]" class="Insecta" family="Limoniidae" genus="Thrypticomyia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="aureipennis">
<emphasis id="B94FB34EFC12FFEEFCEF0CACFBF2AD9C" box="[886,1036,1945,1968]" italics="true" pageId="19" pageNumber="38">T. aureipennis</emphasis>
</taxonomicName>
it is set further from R
<subScript id="17BF6D19FC12FFEEFB640C90FAFBAD9F" attach="left" box="[1277,1285,1957,1971]" fontSize="6" pageId="19" pageNumber="38">2</subScript>
, towards the proximal edge of the stigma.
</paragraph>
<caption id="DF443FD4FC15FFE9FF3E0AC6FD70A825" ID-DOI="http://doi.org/10.5281/zenodo.7177964" ID-Zenodo-Dep="7177964" box="[167,654,499,521]" httpUri="https://zenodo.org/record/7177964/files/figure.png" pageId="20" pageNumber="39" startId="20.[167,235,499,521]" targetBox="[166,1459,159,481]" targetPageId="20">
<paragraph id="8B846F5CFC15FFE9FF3E0AC6FD70A825" blockId="20.[167,654,499,521]" box="[167,654,499,521]" pageId="20" pageNumber="39">
<emphasis id="B94FB34EFC15FFE9FF3E0AC6FEF6A824" bold="true" box="[167,264,499,521]" pageId="20" pageNumber="39">Figure 45</emphasis>
.
<taxonomicName id="4C3B14DFFC15FFE9FE8D0AC6FDE8A825" authorityName="Skuse" authorityYear="1890" box="[276,534,499,521]" class="Insecta" family="Limoniidae" genus="Thrypticomyia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="39" phylum="Arthropoda" rank="species" species="aureipennis">
<emphasis id="B94FB34EFC15FFE9FE8D0AC6FDE8A825" box="[276,534,499,521]" italics="true" pageId="20" pageNumber="39">Thrypticomyia aureipennis</emphasis>
</taxonomicName>
male, wing.
</paragraph>
</caption>
<paragraph id="8B846F5CFC15FFE9FF5C097FFA78A93E" blockId="20.[166,794,586,786]" lastBlockId="20.[839,1467,586,786]" pageId="20" pageNumber="39">
Beyond
<collectingCountry id="F32C2FCCFC15FFE9FEBC097FFE72A84E" box="[293,396,586,610]" name="Australia" pageId="20" pageNumber="39">Australia</collectingCountry>
<taxonomicName id="4C3B14DFFC15FFE9FE01097EFDC5A84E" authorityName="Skuse" authorityYear="1890" box="[408,571,587,610]" class="Insecta" family="Limoniidae" genus="Thrypticomyia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="39" phylum="Arthropoda" rank="species" species="aureipennis">
<emphasis id="B94FB34EFC15FFE9FE01097EFDC5A84E" box="[408,571,587,610]" italics="true" pageId="20" pageNumber="39">T. aureipennis</emphasis>
</taxonomicName>
is also know from
<collectingCountry id="F32C2FCCFC15FFE9FF3E095DFF00A853" box="[167,254,616,639]" name="Vanuatu" pageId="20" pageNumber="39">Vanuatu</collectingCountry>
and
<collectingCountry id="F32C2FCCFC15FFE9FEAB095DFE83A8AC" box="[306,381,616,640]" name="Samoa" pageId="20" pageNumber="39">Samoa</collectingCountry>
, with minor variations across the three populations, prompting
<bibRefCitation id="EFAA12ADFC15FFE9FE2C09B0FD97A8B1" author="Edwards, F. W." box="[437,617,645,669]" pageId="20" pageNumber="39" pagination="25 - 102" refId="ref12639" refString="Edwards, F. W. 1928. Nematocera. In Insects of Samoa and other Samoan Terrestrial Arthropoda - Part VI, Diptera, Fasc. 2, ed. F. W. Edwards and H. F. Barnes, pp. 25 - 102. London: British Museum (Natural History)." type="book chapter" year="1928">Edwards (1928)</bibRefCitation>
to suggest that individuals from
<collectingCountry id="F32C2FCCFC15FFE9FEF50996FE38A896" box="[364,454,675,698]" name="Vanuatu" pageId="20" pageNumber="39">Vanuatu</collectingCountry>
and
<collectingCountry id="F32C2FCCFC15FFE9FD9A0996FDB3A897" box="[515,589,675,699]" name="Samoa" pageId="20" pageNumber="39">Samoa</collectingCountry>
represent distinct varieties of the species. The Vanuatuan individuals having the rostral spines of the inner gonostylus closer together and on lower, less distinct, tubercles than in Australian individuals (
<bibRefCitation id="EFAA12ADFC15FFE9FCD7097FFC06A84F" author="Edwards, F. W." box="[846,1016,586,611]" pageId="20" pageNumber="39" pagination="230 - 236" refId="ref12591" refString="Edwards, F. W. 1927. Crane-flies collected by Dr. P. A. Buxton in the New Hebrides. Annals and Magazine of Natural History 9 (20): 230 - 236. https: // doi. org / 10.1080 / 00222932708655589" type="journal article" year="1927">Edwards, 1927</bibRefCitation>
). While the Samoan
<taxonomicName id="4C3B14DFFC15FFE9FB7D097FFA82A84E" box="[1252,1404,586,610]" pageId="20" pageNumber="39" rank="variety" variety="differs">variety differs</taxonomicName>
from the Australian form in having somewhat dusky wings with a less prominent stigma (
<bibRefCitation id="EFAA12ADFC15FFE9FBFE09B0FAEAA8B1" author="Edwards, F. W." box="[1127,1300,645,669]" pageId="20" pageNumber="39" pagination="25 - 102" refId="ref12639" refString="Edwards, F. W. 1928. Nematocera. In Insects of Samoa and other Samoan Terrestrial Arthropoda - Part VI, Diptera, Fasc. 2, ed. F. W. Edwards and H. F. Barnes, pp. 25 - 102. London: British Museum (Natural History)." type="book chapter" year="1928">Edwards, 1928</bibRefCitation>
). Furthermore, both Vanuatuan and Samoan varieties have the ventral lobe of the gonocoxite shorter and broader than in Australian individuals (
<bibRefCitation id="EFAA12ADFC15FFE9FC4D09E8FB7FA8D9" author="Edwards, F. W." box="[980,1153,733,757]" pageId="20" pageNumber="39" pagination="25 - 102" refId="ref12639" refString="Edwards, F. W. 1928. Nematocera. In Insects of Samoa and other Samoan Terrestrial Arthropoda - Part VI, Diptera, Fasc. 2, ed. F. W. Edwards and H. F. Barnes, pp. 25 - 102. London: British Museum (Natural History)." type="book chapter" year="1928">Edwards, 1928</bibRefCitation>
), where it is comparatively elongate, subequal in length to that of the gonocoxite.
</paragraph>
<caption id="DF443FD4FC15FFE9FF3F0C1BFB1AAD69" ID-DOI="http://doi.org/10.5281/zenodo.7177966" ID-Zenodo-Dep="7177966" box="[166,1252,1838,1861]" httpUri="https://zenodo.org/record/7177966/files/figure.png" pageId="20" pageNumber="39" startId="20.[166,243,1838,1860]" targetBox="[237,1291,842,1810]" targetPageId="20">
<paragraph id="8B846F5CFC15FFE9FF3F0C1BFB1AAD69" blockId="20.[166,1252,1838,1861]" box="[166,1252,1838,1861]" pageId="20" pageNumber="39">
<emphasis id="B94FB34EFC15FFE9FF3F0C1BFECBAD6F" bold="true" box="[166,309,1838,1860]" pageId="20" pageNumber="39">Figures 4647</emphasis>
.
<taxonomicName id="4C3B14DFFC15FFE9FED80C1AFDBDAD68" authorityName="Skuse" authorityYear="1890" box="[321,579,1839,1860]" class="Insecta" family="Limoniidae" genus="Thrypticomyia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="39" phylum="Arthropoda" rank="species" species="aureipennis">
<emphasis id="B94FB34EFC15FFE9FED80C1AFDBDAD68" box="[321,579,1839,1860]" italics="true" pageId="20" pageNumber="39">Thrypticomyia aureipennis</emphasis>
</taxonomicName>
. (
<emphasis id="B94FB34EFC15FFE9FDCE0C1AFD91AD68" box="[599,623,1839,1860]" italics="true" pageId="20" pageNumber="39">46</emphasis>
) male, hypopygium, dorsal view; (
<emphasis id="B94FB34EFC15FFE9FC580C1AFC27AD68" box="[961,985,1839,1860]" italics="true" pageId="20" pageNumber="39">47</emphasis>
) female, ovipositor, lateral.
</paragraph>
</caption>
<paragraph id="8B846F5CFC14FFE8FFE00BADFF35AB44" blockId="21.[121,749,151,360]" pageId="21" pageNumber="40">
ACKNOWLEDGEMENTS. Specimens collected from protected areas were done so under NSW National Parks &amp; Wildlife Service scientific licence number SL101930 and Forestry NSW Research Permit number HF55651. Russell Cox and Natalie Tees of the Australian Museum are thanked for their assistance with the
<typeStatus id="5480D1FEFC14FFE8FD5A0A36FD12AB34" box="[707,748,259,280]" pageId="21" pageNumber="40">type</typeStatus>
material of
<taxonomicName id="4C3B14DFFC14FFE8FF760A28FD9AAB1E" authorityName="Skuse" authorityYear="1890" box="[239,612,285,306]" class="Insecta" family="Limoniidae" genus="Teucholabis" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="40" phylum="Arthropoda" rank="species" species="meridiana" subGenus="Teucholabis">
<emphasis id="B94FB34EFC14FFE8FF760A28FD9AAB1E" box="[239,612,285,306]" italics="true" pageId="21" pageNumber="40">Teucholabis (Teucholabis) meridiana</emphasis>
</taxonomicName>
. John Martin (Australian Museum) is thanked for photographing
<taxonomicName id="4C3B14DFFC14FFE8FDF70A0DFF3BAB4B" authorityName="Billingham &amp; Theischinger" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Dicranomyia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="40" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="B94FB34EFC14FFE8FDF70A0DFF3BAB4B" italics="true" pageId="21" pageNumber="40">Dicranomyia globosa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>