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<document id="0A2BE28F5251743C6ADC467EADB83AF2" ID-DOI="10.11865/zs.202015" ID-ISSN="2095-6827" ID-Zenodo-Dep="5365982" ID-ZooBank="6D924257-51F2-449B-93E2-813B9914B191" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630256278547" checkinUser="diego" docAuthor="Ho, Wai-Chun George" docDate="2020" docId="03928D0E1432FFEF3EAD8E7503D7FC3E" docLanguage="en" docName="New taxa of Clitumninae from Vietnam (Phasmatodea_ Phasmatidae).pdf" docOrigin="Zoological Systematics 45 (2)" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Neomedaura yokdonensis Ho 2020, sp. nov." docType="treatment" docVersion="4" lastPageNumber="113" masterDocId="FFABF576143AFFE63E258A040654FFA9" masterDocTitle="New taxa of Clitumninae from Vietnam (Phasmatodea: Phasmatidae)" masterLastPageNumber="117" masterPageNumber="104" pageNumber="112" updateTime="1699199193242" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="5D588ACD05310C67361903909CCF692F">
<mods:title id="25DDE563BBA4BC05DA478FC16BE33E72">New taxa of Clitumninae from Vietnam (Phasmatodea: Phasmatidae)</mods:title>
</mods:titleInfo>
<mods:name id="756489A079B487BC7B3531C24915F981" type="personal">
<mods:role id="6D8942F069C58511B89A91525F9B7560">
<mods:roleTerm id="6239786EC75D6B594D480A6DC69AC4EC">Author</mods:roleTerm>
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<mods:namePart id="C9021DDCA781A0A19870471AEB392226">Ho, Wai-Chun George</mods:namePart>
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<mods:title id="291A43AFC424B095C8685A6E589F93EF">Zoological Systematics</mods:title>
</mods:titleInfo>
<mods:part id="12EE996A05866BA9BF6AE4B38A939158">
<mods:date id="53A30EB98845EB73107EE4476C2F79E7">2020</mods:date>
<mods:detail id="9FB5360FEF8A97D5EEA31A32295EA712" type="volume">
<mods:number id="EE9022C40151D3C30315D09A66F8D541">45</mods:number>
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<mods:start id="46291F528F88CF2B11EB0CA5E9DA1E29">104</mods:start>
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<mods:classification id="ED03EEB366844B333B64161DE97661EA">journal article</mods:classification>
<mods:identifier id="5EE431AFDF65CBA865B4B7819D2E181A" type="DOI">10.11865/zs.202015</mods:identifier>
<mods:identifier id="41FC1C486A863DAD258C69553EB42030" type="ISSN">2095-6827</mods:identifier>
<mods:identifier id="37383D8E42D213BF1F999F5D2190BC94" type="Zenodo-Dep">5365982</mods:identifier>
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<treatment id="03928D0E1432FFEF3EAD8E7503D7FC3E" ID-DOI="http://doi.org/10.5281/zenodo.5459539" ID-Zenodo-Dep="5459539" LSID="urn:lsid:plazi:treatment:03928D0E1432FFEF3EAD8E7503D7FC3E" httpUri="http://treatment.plazi.org/id/03928D0E1432FFEF3EAD8E7503D7FC3E" lastPageId="9" lastPageNumber="113" pageId="8" pageNumber="112">
<subSubSection id="C3216F931432FFEE3EAD8E7504A0FB20" box="[136,756,1137,1162]" pageId="8" pageNumber="112" type="nomenclature">
<paragraph id="8B843C181432FFEE3EAD8E7504A0FB20" blockId="8.[136,756,1137,1162]" box="[136,756,1137,1162]" pageId="8" pageNumber="112">
<heading id="D0CC8B741432FFEE3EAD8E7504A0FB20" box="[136,756,1137,1162]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="112" reason="2">
<emphasis id="B94FE00A1432FFEE3EAD8E750454FB23" bold="true" box="[136,512,1137,1162]" pageId="8" pageNumber="112">
<taxonomicName id="4C3B479B1432FFEE3EAD8E7507F0FB20" ID-CoL="8QQZT" authority="Ho, 2020" authorityName="Ho" authorityYear="2020" box="[136,420,1137,1161]" class="Insecta" family="Phasmatidae" genus="Neomedaura" kingdom="Animalia" order="Phasmida" pageId="8" pageNumber="112" phylum="Arthropoda" rank="species" species="yokdonensis" status="sp. nov.">
<emphasis id="B94FE00A1432FFEE3EAD8E7507F0FB20" bold="true" box="[136,420,1137,1161]" italics="true" pageId="8" pageNumber="112">Neomedaura yokdonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C5D711432FFEE3F8E8E760454FB23" box="[427,512,1138,1162]" pageId="8" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="1300209D1432FFEE3C078E7504CAFB23" box="[546,670,1137,1162]" captionStart="Figures 1–26" captionStartId="2.[155,227,1635,1657]" captionTargetBox="[192,1395,379,1579]" captionTargetId="figure-210@2.[189,1398,376,1585]" captionTargetPageId="2" captionText="Figures 1–26. Clitumninae spp. 1–4. Interphasma lineatum sp. nov. 5–8. Medauroidea cattienensis sp. nov. 9–12. Medauroidea cornuta sp. nov. 13–16. Medauroidea fasciata sp. nov. 17–20. Neomedaura yokdonensis gen. &amp; sp. nov. 21–22. Parapachymorpha pseudospinosa sp. nov. 23–26. Prosentoria vietnamensis sp. nov. 1, 7, 11, 15, 19, 25. Male end of abdomen, lateral view. 2, 8, 12, 16, 20, 26. Male end of abdomen, dorsal view. 3, 5, 9, 13, 17, 21, 23. Female end of abdomen, lateral view. 4, 6, 10, 14, 18, 22, 24. Female end of abdomen, dorsal view. Scale bars=5mm." figureDoi="http://doi.org/10.5281/zenodo.5365984" httpUri="https://zenodo.org/record/5365984/files/figure.png" pageId="8" pageNumber="112">Figs 17–20</figureCitation>
,
<figureCitation id="1300209D1432FFEE3C8D8E7504B8FB20" box="[680,748,1137,1161]" captionStart="Figures 61–68" captionStartId="11.[155,227,971,993]" captionTargetBox="[189,1398,420,921]" captionTargetId="figure-227@11.[189,1398,420,921]" captionTargetPageId="11" captionText="Figures 61–68. Neomedaura yokdonensis gen. &amp; sp. nov. 61. Female habitus. 62. Female abdomen, lateral view. 63. Female abdomen, dorsal view. 64. Male habitus. 65. Male abdomen, lateral view. 66. Male abdomen, dorsal view. 67. Female head and thorax, dorsolateral view. 68. Male head and thorax, dorsolateral view. Scale bars=5mm." figureDoi="http://doi.org/10.5281/zenodo.5365994" httpUri="https://zenodo.org/record/5365994/files/figure.png" pageId="8" pageNumber="112">61–68</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3216F931432FFEF3E988EAD03D7FC3E" lastPageId="9" lastPageNumber="113" pageId="8" pageNumber="112" type="diagnosis">
<paragraph id="8B843C181432FFEE3E988EAD0408FB68" blockId="8.[136,1452,1193,2083]" box="[189,604,1193,1217]" pageId="8" pageNumber="112">Diagnosis. As in the generic diagnosis.</paragraph>
<paragraph id="8B843C181432FFEE3E988EC804D9F8DF" blockId="8.[136,1452,1193,2083]" pageId="8" pageNumber="112">Description. Female. Body slender and long, larger and more robust than male. General colouration of body and legs brown. Head densely granulated. Oblong, longer than pronotum, slightly tapering posteriorly behind compound eyes. Vertex obscurely elevated between compound eyes, with a pair of minute horns. Occiput flat, median and lateral longitudinal furrows distinct, posterior margin weakly raised, with four small swellings. Compound eyes small and rounded, its length about five times of genae. Antennae short, with 15 segments, reaching middle area of profemur; scapus flattened, constricted anteriorly and posteriorly, longer than combined length of pedicellus and third segment; and pedicellus longer than third segment. Thorax rough, densely covered with small granules. Pronotum trapezoidal, gently expanded posteriorly, shorter than head, anterior margin strongly incurved, posterior margin truncate, transverse and longitudinal sulci just crossing after middle point. Mesonotum parallel-sided, longer than combined length of metanotum and median segment. Metanotum more than four times longer than median segment. Abdomen cylindrical, rough, covered with obscure granulations. Median segment rectangular, wider than long. Seventh sternum with distinct praeopercular organ posteromedially, horn-like, apex pointed. Eighth tergum longer than ninth tergum, also longer than anal segment. Anal segment longer than ninth tergum, posterior margin with a small V-shaped emargination, posterolateral angles pointed. Supra-anal plate indistinct. Subgenital plate scoop-shaped, laterally carinate, posterior apex pointed and reaching posterior area of anal segment. Cerci short, flattened, apices pointed and not surpassing posterior margin of anal segment. Legs slender and long. Right foreleg lost. Femora thicker than corresponding tibiae. Left profemur curved basally, anterodorsal carina with six small and indistinct serrations subbasally, other carinae unarmed. Anterodorsal and posterodorsal carinae of mesofemora with one to two small and indistinct serrations subbasally. Other carinae of mesofemora and metafemora unarmed. Protibia and metatibiae almost as long as corresponding femora, mesotibiae as long as mesofemora. Posterodorsal carina of metatibiae with one to two small spines subapically, other carinae unarmed.</paragraph>
<paragraph id="8B843C181432FFEF3E988D850288FDF6" blockId="8.[136,1452,1193,2083]" lastBlockId="9.[136,1453,236,919]" lastPageId="9" lastPageNumber="113" pageId="8" pageNumber="112">Male. Body slender and slim, distinctly smaller and slenderer than female. General colouration of body and legs brown. Head obscurely granulated. Oblong, longer than pronotum, gently tapering posteriorly behind compound eyes. Vertex gently elevated. Occiput flat, with distinct median and lateral longitudinal furrows, posterior margin with four small swellings. Compound eyes small and rounded, its length about four times of genae. Antennae short, with 26 segments, apices reaching subapical area of profemora; scapus flattened, constricted anteriorly and posteriorly, as long as combined length of pedicellus and third segment; and third segment longer than pedicellus. Thorax obscurely granulated. Pronotum rectangular, longer than wide, anterior margin incurved, posterior margin truncate, transverse and longitudinal sulci crossing at middle point. Mesonotum elongate, longer than combined length of metanotum and median segment, broadly emarginated medially. Metanotum more than three times longer than median segment. Abdomen slender and cylindrical. Median segment rectangular, longer than wide. Second to seventh tergites parallel-sided. Third to seventh tergites almost equal in length. Eighth tergum expanded posteriorly, as long as ninth tergum. Anal segment as long as ninth tergum, dilated into two semitergites; semi-tergites straight, medially tapering posteriorly, inner margin incurved, with minute dentations, apices pointed. Poculum small, cup-shaped, posterior margin rounded and reaching posterior area of ninth tergum. Cerci cylindrical, short, apices obtuse and not exceeding posterior apices of anal segment. Legs slender and long. Femora thicker than corresponding tibiae, anterodorsal, posterodorsal, anteroventral and posteroventral carinae unarmed, medioventral carina with a very few minute teeth subapically. Profemora curved basally. Tibiae unarmed, longer than corresponding femora.</paragraph>
<paragraph id="8B843C181433FFEF3E98886E04FFFCA5" blockId="9.[136,1453,236,919]" pageId="9" pageNumber="113">
Measurements. Length. Body,
<specimenCount id="9D3DF7911433FFEF3C2B88620476FD28" box="[526,546,614,641]" pageId="9" pageNumber="113" type="female">♀</specimenCount>
75.0 mm,
<specimenCount id="9D3DF7911433FFEF3CBC886304F9FD2B" box="[665,685,615,642]" pageId="9" pageNumber="113" type="male">♂</specimenCount>
68.0 mm; antennae,
<specimenCount id="9D3DF7911433FFEF3DB0886305FDFD2B" box="[917,937,615,642]" pageId="9" pageNumber="113" type="female">♀</specimenCount>
9.0 mm,
<specimenCount id="9D3DF7911433FFEF3A3788630272FD2B" box="[1042,1062,615,642]" pageId="9" pageNumber="113" type="male">♂</specimenCount>
19.0 mm; head,
<specimenCount id="9D3DF7911433FFEF3AC4886302A1FD2B" box="[1249,1269,615,642]" pageId="9" pageNumber="113" type="female">♀</specimenCount>
6.0 mm,
<specimenCount id="9D3DF7911433FFEF3B7A88630327FD2B" box="[1375,1395,615,642]" pageId="9" pageNumber="113" type="male">♂</specimenCount>
3.0–
<quantity id="4CC391FD1433FFEF3EAD8888068EFD0D" box="[136,218,652,676]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="113" unit="mm" value="3.5">3.5 mm</quantity>
; pronotum,
<specimenCount id="9D3DF7911433FFEF3F7C888D0739FD0D" box="[345,365,649,676]" pageId="9" pageNumber="113" type="female">♀</specimenCount>
3.0 mm,
<specimenCount id="9D3DF7911433FFEF3FEA888D07B7FD0D" box="[463,483,649,676]" pageId="9" pageNumber="113" type="male">♂</specimenCount>
2.0 mm; mesonotum,
<specimenCount id="9D3DF7911433FFEF3CEA888D04B7FD0D" box="[719,739,649,676]" pageId="9" pageNumber="113" type="female">♀</specimenCount>
14.0mm,
<specimenCount id="9D3DF7911433FFEF3D77888D0532FD0D" box="[850,870,649,676]" pageId="9" pageNumber="113" type="male">♂</specimenCount>
15.0–16.0 mm; metanotum,
<specimenCount id="9D3DF7911433FFEF3AB2888D02FFFD0D" box="[1175,1195,649,676]" pageId="9" pageNumber="113" type="female">♀</specimenCount>
9.0 mm,
<specimenCount id="9D3DF7911433FFEF3B28888D0375FD0D" box="[1293,1313,649,676]" pageId="9" pageNumber="113" type="male">♂</specimenCount>
9.0–
<quantity id="4CC391FD1433FFEF3B72888803F3FD0D" box="[1367,1447,652,676]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="9" pageNumber="113" unit="mm" value="9.5">9.5 mm</quantity>
; median segment,
<specimenCount id="9D3DF7911433FFEF3F6788A80702FD6E" box="[322,342,684,711]" pageId="9" pageNumber="113" type="female">♀</specimenCount>
2.0 mm,
<specimenCount id="9D3DF7911433FFEF3F9088A8079DFD6E" box="[437,457,684,711]" pageId="9" pageNumber="113" type="male">♂</specimenCount>
3.0 mm; profemora,
<specimenCount id="9D3DF7911433FFEF3C8088A804EDFD6E" box="[677,697,684,711]" pageId="9" pageNumber="113" type="female">♀</specimenCount>
25.0 mm,
<specimenCount id="9D3DF7911433FFEF3D0088A8056DFD6E" box="[805,825,684,711]" pageId="9" pageNumber="113" type="male">♂</specimenCount>
30.0–31.0 mm; mesofemora,
<specimenCount id="9D3DF7911433FFEF3A5688A802D3FD6E" box="[1139,1159,684,711]" pageId="9" pageNumber="113" type="female">♀</specimenCount>
18.0mm,
<specimenCount id="9D3DF7911433FFEF3AD188A8035CFD6E" box="[1268,1288,684,711]" pageId="9" pageNumber="113" type="male">♂</specimenCount>
18.0–19.0 mm; metafemora,
<specimenCount id="9D3DF7911433FFEF3F3C88CA0779FD40" box="[281,301,718,745]" pageId="9" pageNumber="113" type="female">♀</specimenCount>
25.0 mm,
<specimenCount id="9D3DF7911433FFEF3F8288CB07EFFD43" box="[423,443,719,746]" pageId="9" pageNumber="113" type="male">♂</specimenCount>
23.0–24.0 mm; protibiae,
<specimenCount id="9D3DF7911433FFEF3CC688CB04A3FD43" box="[739,759,719,746]" pageId="9" pageNumber="113" type="female">♀</specimenCount>
27.0 mm,
<specimenCount id="9D3DF7911433FFEF3D5488CB05D1FD43" box="[881,901,719,746]" pageId="9" pageNumber="113" type="male">♂</specimenCount>
34.0–38.0 mm; mesotibiae,
<specimenCount id="9D3DF7911433FFEF3AE788CB0282FD43" box="[1218,1238,719,746]" pageId="9" pageNumber="113" type="female">♀</specimenCount>
18.0 mm,
<specimenCount id="9D3DF7911433FFEF3B7588CB0330FD43" box="[1360,1380,719,746]" pageId="9" pageNumber="113" type="male">♂</specimenCount>
20.0– 22.0 mm; metatibiae,
<specimenCount id="9D3DF7911433FFEF3F4B88F507D6FCA5" box="[366,386,753,780]" pageId="9" pageNumber="113" type="female">♀</specimenCount>
27.0mm,
<specimenCount id="9D3DF7911433FFEF3FD488F50451FCA5" box="[497,517,753,780]" pageId="9" pageNumber="113" type="male">♂</specimenCount>
29.0–31.0 mm.
</paragraph>
<paragraph id="8B843C181433FFEF3E98891307A1FCFB" blockId="9.[136,1453,236,919]" pageId="9" pageNumber="113">
<materialsCitation id="3B5336451433FFEF3E98891307A6FCFB" collectingDate="2012-05-08" collectionCode="CAU, HKES" country="Vietnam" location="Yok Don" municipality="Material" pageId="9" pageNumber="113" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Dak Lak" typeStatus="holotype">
<collectingMunicipality id="6BE0A6621433FFEF3E988913074CFC86" box="[189,280,791,815]" pageId="9" pageNumber="113">Material</collectingMunicipality>
examined.
<typeStatus id="548082BA1433FFEF3FB1891307A3FC86" box="[404,503,791,815]" pageId="9" pageNumber="113" type="holotype">Holotype</typeStatus>
<specimenCount id="9D3DF7911433FFEF3FDB89100446FC86" box="[510,530,788,815]" pageId="9" pageNumber="113" type="female">♀</specimenCount>
,
<collectingCountry id="F32C7C881433FFEF3C3B8913042FFC86" box="[542,635,791,815]" name="Vietnam" pageId="9" pageNumber="113">Vietnam</collectingCountry>
,
<collectingRegion id="49FFF2FA1433FFEF3CA3891304B1FC86" box="[646,741,791,815]" country="Vietnam" name="Dak Lak" pageId="9" pageNumber="113">Dak Lak</collectingRegion>
,
<location id="8EE46AC31433FFEF3CCA89130506FC87" LSID="urn:lsid:plazi:treatment:03928D0E1432FFEF3EAD8E7503D7FC3E:8EE46AC31433FFEF3CCA89130506FC87" box="[751,850,791,815]" country="Vietnam" municipality="Material" name="Yok Don" pageId="9" pageNumber="113" stateProvince="Dak Lak">Yok Don</location>
,
<date id="FF851AD81433FFEF3D79891305B4FC86" box="[860,992,791,815]" pageId="9" pageNumber="113" value="2012-05-08">
<collectingDate id="EFC1E3301433FFEF3D79891305B4FC86" box="[860,992,791,815]" pageId="9" pageNumber="113" value="2012-05-08">8 May 2012</collectingDate>
</date>
, Liu Xingyue (
<collectionCode id="ED2AA4DD1433FFEF3AA4891302EBFC86" box="[1153,1215,791,815]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="9" pageNumber="113">CAU</collectionCode>
).
<typeStatus id="548082BA1433FFEF3AF489130368FC86" box="[1233,1340,791,815]" pageId="9" pageNumber="113" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="9D3DF7911433FFEF3B6389130333FC86" box="[1350,1383,788,815]" pageId="9" pageNumber="113" type="male">2♂</specimenCount>
, same data as holotype (
<collectionCode id="ED2AA4DD1433FFEF3F61893E072AFCFB" box="[324,382,826,850]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="9" pageNumber="113">CAU</collectionCode>
&amp;
<collectionCode id="ED2AA4DD1433FFEF3FBA893E07BDFCFB" box="[415,489,826,850]" pageId="9" pageNumber="113">HKES</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B843C181433FFEF3E98895807F9FCDD" blockId="9.[136,1453,236,919]" box="[189,429,860,884]" pageId="9" pageNumber="113">
Distribution.
<collectingCountry id="F32C7C881433FFEF3F6E895807FDFCDD" box="[331,425,860,884]" name="Vietnam" pageId="9" pageNumber="113">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="8B843C181433FFEF3E98897B03D7FC3E" blockId="9.[136,1453,236,919]" box="[189,1411,895,919]" pageId="9" pageNumber="113">
Etymology. The specific epithet of this new species is derived from the type-locality, Yok Don,
<collectingRegion id="49FFF2FA1433FFEF3A9D897B0343FC3E" box="[1208,1303,895,919]" country="Vietnam" name="Dak Lak" pageId="9" pageNumber="113">Dak Lak</collectingRegion>
,
<collectingCountry id="F32C7C881433FFEF3B07897B03D4FC3E" box="[1314,1408,895,919]" name="Vietnam" pageId="9" pageNumber="113">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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