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<document id="0B11AF213787E7A3516DD48093663CBB" ID-DOI="10.11646/zootaxa.5336.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8272417" ID-ZooBank="7BD6357E-BB67-44E9-B028-4CA7A2769660" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1692697841468" checkinUser="plazi" docAuthor="Uiblein, Franz &amp; Nielsen, Jørgen G." docDate="2023" docId="039287FEFC32FF9476BAEFFEFA2C93D7" docLanguage="en" docName="Zootaxa.5336.2.2.pdf" docOrigin="Zootaxa 5336 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5336.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neobythites steatiticus Alcock 1894" docType="key" docVersion="1" lastPageNumber="202" masterDocId="FFABFF86FC25FF83762DEE25FFB09649" masterDocTitle="Five new ocellus-bearing species of the cusk-eel genus Neobythites (Ophidiidae, Ophidiiformes) from the West Pacific, with establishment of three new species groups" masterLastPageNumber="205" masterPageNumber="179" pageNumber="202" updateTime="1692794457292" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="108782550EEC19BBD44A3E42BD46359F">Five new ocellus-bearing species of the cusk-eel genus Neobythites (Ophidiidae, Ophidiiformes) from the West Pacific, with establishment of three new species groups</mods:title>
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<mods:namePart id="B2EA688C83A395DA81A48C2A48A1A4E4">Uiblein, Franz</mods:namePart>
<mods:affiliation id="68D42D8D364742A1EBBAA57793ABB14D">Institute of Marine Research, P. O. Box 1870 Nordnes, N- 5817 Bergen, Norway. Corresponding author; &amp; National Research Foundation - South African Institute for Aquatic Biodiversity, Makhanda / Grahamstown, South Africa &amp; Vietnam National Museum of Nature, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="F71A767FBBC2BDB77486EB86337296BF">Nielsen, Jørgen G.</mods:namePart>
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<subSubSection id="C3216563FC32FF9476BAEFFEFD8497BC" box="[151,564,474,501]" pageId="23" pageNumber="202" type="nomenclature">
<paragraph id="8B8436E8FC32FF9476BAEFFEFD8497BC" blockId="23.[151,1437,474,1439]" box="[151,564,474,501]" pageId="23" pageNumber="202">
<heading id="D0CC8184FC32FF9476BAEFFEFD8497BC" bold="true" box="[151,564,474,501]" fontSize="11" level="1" pageId="23" pageNumber="202" reason="1">
<emphasis id="B94FEAFAFC32FF9476BAEFFEFD8497BC" bold="true" box="[151,564,474,501]" pageId="23" pageNumber="202">
Key to the
<taxonomicName id="4C3B4D6BFC32FF947734EFFEFE3C97BC" ID-CoL="46C4M" ID-ENA="1649253" authorityName="Alcock" authorityYear="1894" box="[281,396,475,501]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="steatiticus">
<emphasis id="B94FEAFAFC32FF947734EFFEFE3C97BC" bold="true" box="[281,396,475,501]" italics="true" pageId="23" pageNumber="202">steatiticus</emphasis>
</taxonomicName>
species group
</emphasis>
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</paragraph>
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<emphasis id="B94FEAFAFC32FF9476BAEFDEFF139459" bold="true" box="[151,163,507,528]" pageId="23" pageNumber="202">1</emphasis>
Anterior gill arch with 67 long rakers; anal fin without dark margin or band..................................... 2
</keyLead>
</paragraph>
<paragraph id="8B8436E8FC32FF9476BAEC3CFA2C9467" blockId="23.[151,1437,474,1439]" box="[151,1436,537,559]" pageId="23" pageNumber="202">
<keyLead id="30CA95DDFC32FF9476BAEC3CFA2C9467" box="[151,1436,537,559]" pageId="23" pageNumber="202">
<emphasis id="B94FEAFAFC32FF9476BAEC3CFF2F9467" bold="true" box="[151,159,537,558]" pageId="23" pageNumber="202">-</emphasis>
Anterior gill arch with 815 long rakers; anal fin with or without dark margin or band.............................. 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CF2E4DFC32FF9476BAEC13FA2C94EA" pageId="23" pageNumber="202">
<paragraph id="8B8436E8FC32FF9476BAEC13FA2C9420" blockId="23.[151,1437,474,1439]" pageId="23" pageNumber="202">
<keyLead id="30CA95DDFC32FF9476BAEC13FA2C9420" pageId="23" pageNumber="202">
<emphasis id="B94FEAFAFC32FF9476BAEC13FF139402" bold="true" box="[151,163,566,587]" pageId="23" pageNumber="202">2</emphasis>
Ocellus short, placed above anus with ocellus spot covering 9 dorsal-fin rays; longest gill filament 13.0% HL.......................................................................
<taxonomicName id="4C3B4D6BFC32FF947597EC71FBAF9420" authorityName="Nielsen" authorityYear="1995" box="[954,1055,596,617]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="meteori">
<emphasis id="B94FEAFAFC32FF947597EC71FBAF9420" box="[954,1055,596,617]" italics="true" pageId="23" pageNumber="202">N. meteori</emphasis>
</taxonomicName>
(off Sokotra Island, NW Indian Ocean)
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</paragraph>
<paragraph id="8B8436E8FC32FF9476BAEC54FA2C94EA" blockId="23.[151,1437,474,1439]" pageId="23" pageNumber="202">
<keyLead id="30CA95DDFC32FF9476BAEC54FA2C94EA" pageId="23" pageNumber="202">
<emphasis id="B94FEAFAFC32FF9476BAEC54FF2F94CF" bold="true" box="[151,159,625,646]" pageId="23" pageNumber="202">-</emphasis>
Ocellus long, placed above anterior part of anal fin with ocellus spot covering 14 dorsal-fin rays; longest gill filament 9.4% HL............................................................
<taxonomicName id="4C3B4D6BFC32FF9475F2ECAAFBC294EA" authorityName="Uiblein &amp; Nielsen" authorityYear="2018" box="[991,1138,654,676]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="lombokensis">
<emphasis id="B94FEAFAFC32FF9475F2ECAAFBC294EA" box="[991,1138,654,676]" italics="true" pageId="23" pageNumber="202">N. lombokensis</emphasis>
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(off Lombok, E Indian Ocean)
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<keyStep id="30CF2E4DFC32FF9476BAEC8EFA2C95E5" pageId="23" pageNumber="202">
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<keyLead id="30CA95DDFC32FF9476BAEC8EFA2C94B2" pageId="23" pageNumber="202">
<emphasis id="B94FEAFAFC32FF9476BAEC8EFF139489" bold="true" box="[151,163,683,704]" pageId="23" pageNumber="202">4</emphasis>
Vertical, dark bar below ocellus spot; dorsal and anal fins without dark margin or band, dorsal-fin rays 99103, dorsal-fin origin above vertebrae number 23, anal-fin origin below dorsal-fin ray number 2325............................................................................................
<taxonomicName id="4C3B4D6BFC32FF9475A2ECC2FA2C94B2" authority="(Saya de Malha Bank, W Indian Ocean)" baseAuthorityName="Saya de Malha Bank, W Indian Ocean" box="[911,1436,742,764]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="malhaensis">
<emphasis id="B94FEAFAFC32FF9475A2ECC2FBA794B2" box="[911,1047,742,764]" italics="true" pageId="23" pageNumber="202">N. malhaensis</emphasis>
(Saya de Malha Bank, W Indian Ocean)
</taxonomicName>
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</paragraph>
<paragraph id="8B8436E8FC32FF9476BAED21FA2C957F" blockId="23.[151,1437,474,1439]" pageId="23" pageNumber="202">
<keyLead id="30CA95DDFC32FF9476BAED21FA2C957F" pageId="23" pageNumber="202">
<emphasis id="B94FEAFAFC32FF9476BAED21FF2F9550" bold="true" box="[151,159,772,793]" pageId="23" pageNumber="202">-</emphasis>
None to a few indistinct, vertical, dark bars on body; dorsal and/or anal fin with dark margin or band, dorsal-fin rays 8899, dorsal-fin origin above vertebrae number 46, anal-fin origin below dorsal-fin ray number 1721..................... 4
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</paragraph>
<paragraph id="8B8436E8FC32FF9476BAED1BFA2C9538" blockId="23.[151,1437,474,1439]" pageId="23" pageNumber="202">
<keyLead id="30CA95DDFC32FF9476BAED1BFA2C9538" pageId="23" pageNumber="202">
<emphasis id="B94FEAFAFC32FF9476BAED1BFF13951A" bold="true" box="[151,163,830,851]" pageId="23" pageNumber="202">4</emphasis>
Median part of anal fin dark, and distal and proximal parts light; head length 2831% SL; longest gill filament 4.45.9% SL..............................................................
<taxonomicName id="4C3B4D6BFC32FF9475E1ED79FBF99538" authorityName="Alcock" authorityYear="1894" box="[972,1097,860,881]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="steatiticus">
<emphasis id="B94FEAFAFC32FF9475E1ED79FBF99538" box="[972,1097,860,881]" italics="true" pageId="23" pageNumber="202">N. steatiticus</emphasis>
</taxonomicName>
(Bay of Bengal, NE Indian Ocean)
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<keyLead id="30CA95DDFC32FF9476BAED5CFA2C95E5" pageId="23" pageNumber="202">
<emphasis id="B94FEAFAFC32FF9476BAED5CFF2F95C7" bold="true" box="[151,159,889,910]" pageId="23" pageNumber="202">-</emphasis>
Distal part of dorsal and anal fins dark and proximal part light; head length 2128% SL; longest gill filament 1.34.9% SL.. ................................................................................................... 5
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<keyStep id="30CF2E4DFC32FF9476BAED91FA2C9268" pageId="23" pageNumber="202">
<paragraph id="8B8436E8FC32FF9476BAED91FA2C95AE" blockId="23.[151,1437,474,1439]" pageId="23" pageNumber="202">
<keyLead id="30CA95DDFC32FF9476BAED91FA2C95AE" pageId="23" pageNumber="202">
<emphasis id="B94FEAFAFC32FF9476BAED91FF139580" bold="true" box="[151,163,948,969]" pageId="23" pageNumber="202">5</emphasis>
Developed gill rakers 811 (mostly 9 or 10); orbit length 4.25.6% SL and 3746% upper-jaw length; precaudal vertebrae 13 (rarely 12).......................................................................................... 6
</keyLead>
</paragraph>
<paragraph id="8B8436E8FC32FF9476BAEDCBFA2C9268" blockId="23.[151,1437,474,1439]" pageId="23" pageNumber="202">
<keyLead id="30CA95DDFC32FF9476BAEDCBFA2C9268" pageId="23" pageNumber="202">
<emphasis id="B94FEAFAFC32FF9476BAEDCBFF2F924A" bold="true" box="[151,159,1006,1027]" pageId="23" pageNumber="202">-</emphasis>
Developed gill rakers 1015 (mostly 1115); orbit length 4.97.0% SL and 4259% times in upper-jaw length; precaudal vertebrae 12 (rarely 13)................................................................................ 7
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</paragraph>
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<keyStep id="30CF2E4DFC32FF9476BAEA0CFA2D92DF" pageId="23" pageNumber="202">
<paragraph id="8B8436E8FC32FF9476BAEA0CFA2C9212" blockId="23.[151,1437,474,1439]" pageId="23" pageNumber="202">
<keyLead id="30CA95DDFC32FF9476BAEA0CFA2C9212" pageId="23" pageNumber="202">
<emphasis id="B94FEAFAFC32FF9476BAEA0CFF139277" bold="true" box="[151,163,1065,1086]" pageId="23" pageNumber="202">6</emphasis>
Preopercular spine indistinct or a flat process, upper-jaw length 4452% HL, ostium height 0.770.93% SL, 2125% sulcus length and 3238% ostium length..................................
<taxonomicName id="4C3B4D6BFC32FF9475FEEA62FBE79212" box="[979,1111,1094,1116]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="malayanus">
<emphasis id="B94FEAFAFC32FF9475FEEA62FBE79212" box="[979,1111,1094,1116]" italics="true" pageId="23" pageNumber="202">N. malayanus</emphasis>
</taxonomicName>
(
<collectingCountry id="F32C7678FC32FF94724EEA63FB749215" box="[1123,1220,1094,1116]" name="Indonesia" pageId="23" pageNumber="202">Indonesia</collectingCountry>
,
<collectingCountry id="F32C7678FC32FF9472E3EA63FA8D9215" box="[1230,1341,1094,1116]" name="Philippines" pageId="23" pageNumber="202">Philippines</collectingCountry>
,
<collectingCountry id="F32C7678FC32FF94736BEA63FA269212" box="[1350,1430,1094,1115]" name="Vanuatu" pageId="23" pageNumber="202">Vanuatu</collectingCountry>
)
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<paragraph id="8B8436E8FC32FF9476BAEA41FA2D92DF" blockId="23.[151,1437,474,1439]" pageId="23" pageNumber="202">
<keyLead id="30CA95DDFC32FF9476BAEA41FA2D92DF" pageId="23" pageNumber="202">
<emphasis id="B94FEAFAFC32FF9476BAEA41FF2F9230" bold="true" box="[151,159,1124,1145]" pageId="23" pageNumber="202">-</emphasis>
Preopercular spine very short but distinct, upper-jaw length 54% HL, ostium height 0.65% SL, 17% sulcus length and 25% ostium length.................................................................
<taxonomicName id="4C3B4D6BFC32FF9472ADEAA7FB7B92DF" authority="Uiblein &amp; Nielsen, 2023" authorityName="Uiblein &amp; Nielsen" authorityYear="2023" box="[1152,1227,1153,1175]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="pako" status="n. sp.">
<emphasis id="B94FEAFAFC32FF9472ADEAA7FB7B92DF" box="[1152,1227,1153,1175]" italics="true" pageId="23" pageNumber="202">N. pako</emphasis>
</taxonomicName>
<emphasis id="B94FEAFAFC32FF9472FCEAA4FAB792DF" bold="true" box="[1233,1287,1153,1174]" pageId="23" pageNumber="202">
<taxonomicNameLabel id="A27C5781FC32FF9472FCEAA4FAB792DF" box="[1233,1287,1153,1174]" pageId="23" pageNumber="202" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(Solomon Sea)
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</paragraph>
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<keyStep id="30CF2E4DFC32FF9476BAEABBFA2C9360" pageId="23" pageNumber="202">
<paragraph id="8B8436E8FC32FF9476BAEABBFA2C92A6" blockId="23.[151,1437,474,1439]" pageId="23" pageNumber="202">
<keyLead id="30CA95DDFC32FF9476BAEABBFA2C92A6" pageId="23" pageNumber="202">
<emphasis id="B94FEAFAFC32FF9476BAEABBFF1392FA" bold="true" box="[151,163,1182,1203]" pageId="23" pageNumber="202">7</emphasis>
Dorsal-fin rays 9399, anal-fin rays 7883, developed gill rakers 1315, head length 2124% SL, postorbital distance 1114% SL, sulcus length 4.04.7% SL and ostium length 5964% sulcus length.....................................................................................
<taxonomicName id="4C3B4D6BFC32FF9474DEEAFFFC3F92A7" authorityName="Nielsen" authorityYear="1999" box="[755,911,1241,1263]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="monocellatus">
<emphasis id="B94FEAFAFC32FF9474DEEAFFFC3F92A7" box="[755,911,1241,1263]" italics="true" pageId="23" pageNumber="202">N. monocellatus</emphasis>
</taxonomicName>
(
<collectingCountry id="F32C7678FC32FF9475B0EAFCFC4A92A6" box="[925,1018,1241,1263]" name="Honduras" pageId="23" pageNumber="202">Honduras</collectingCountry>
to off
<collectingRegion id="49FFF80AFC32FF947216EAFCFBC592A6" box="[1083,1141,1241,1263]" country="Brazil" name="Bahia" pageId="23" pageNumber="202">Bahia</collectingRegion>
, E
<collectingCountry id="F32C7678FC32FF9472B9EAFCFB6192A6" box="[1172,1233,1241,1263]" name="Brazil" pageId="23" pageNumber="202">Brazil</collectingCountry>
, W and SW Atlantic)
</keyLead>
</paragraph>
<paragraph id="8B8436E8FC32FF9476BAEAD3FA2C9360" blockId="23.[151,1437,474,1439]" pageId="23" pageNumber="202">
<keyLead id="30CA95DDFC32FF9476BAEAD3FA2C9360" pageId="23" pageNumber="202">
<emphasis id="B94FEAFAFC32FF9476BAEAD3FF2F9342" bold="true" box="[151,159,1270,1291]" pageId="23" pageNumber="202">-</emphasis>
Dorsal-fin rays 8995, anal-fin rays 7278, developed gill rakers 1013, head length 2328% SL, postorbital distance 1418% SL, sulcus length 3.44.1% SL, and ostium length 6370% sulcus length........................................ 8
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<paragraph id="8B8436E8FC32FF9476BAEB14FA2C932A" blockId="23.[151,1437,474,1439]" pageId="23" pageNumber="202">
<keyLead id="30CA95DDFC32FF9476BAEB14FA2C932A" pageId="23" pageNumber="202">
<emphasis id="B94FEAFAFC32FF9476BAEB14FF13930F" bold="true" box="[151,163,1329,1350]" pageId="23" pageNumber="202">8</emphasis>
Longest gill filaments 1.43.7% SL; dorsal-fin rays in ocellus spot 812..............................................................................................
<taxonomicName id="4C3B4D6BFC32FF947570EB6AFC60932A" box="[861,976,1358,1380]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="stefanovi">
<emphasis id="B94FEAFAFC32FF947570EB6AFC60932A" box="[861,976,1358,1380]" italics="true" pageId="23" pageNumber="202">N. stefanovi</emphasis>
</taxonomicName>
(Red Sea and Gulf of Aden, NW Indian Ocean)
</keyLead>
</paragraph>
<paragraph id="8B8436E8FC32FF9476BAEB49FA2C93D7" blockId="23.[151,1437,474,1439]" pageId="23" pageNumber="202">
<keyLead id="30CA95DDFC32FF9476BAEB49FA2C93D7" pageId="23" pageNumber="202">
<emphasis id="B94FEAFAFC32FF9476BAEB49FF2F93C8" bold="true" box="[151,159,1388,1409]" pageId="23" pageNumber="202">-</emphasis>
Longest gill filaments 3.94.9% SL; dorsal-fin rays in ocellus spot 68.......................................................................................
<taxonomicName id="4C3B4D6BFC32FF9474D2EBAFFCD093D7" authorityName="Uiblein &amp; Nielsen" authorityYear="2018" box="[767,864,1417,1439]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="202" phylum="Chordata" rank="species" species="gloriae">
<emphasis id="B94FEAFAFC32FF9474D2EBAFFCD093D7" box="[767,864,1417,1439]" italics="true" pageId="23" pageNumber="202">N. gloriae</emphasis>
</taxonomicName>
(Arabian Gulf and inner Gulf of
<collectingCountry id="F32C7678FC32FF9472B2EBACFB6C93D7" box="[1183,1244,1417,1438]" name="Oman" pageId="23" pageNumber="202">Oman</collectingCountry>
, NW Indian Ocean)
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>