treatments-xml/data/03/92/32/0392327B7E756318FF37FC8DFBA44C04.xml

<document id="F0AACEB9127D5184DCE16A47F53EFC3E" ID-DOI="10.11646/zootaxa.5395.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10438327" ID-ZooBank="CCA38E8E-26E5-47D4-A27D-305554F6097F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1703753817594" checkinUser="plazi" docAuthor="Feng, Zegang, Zhang, Feng &amp; Chen, Jun" docDate="2023" docId="0392327B7E756318FF37FC8DFBA44C04" docLanguage="en" docName="zootaxa.5395.1.1.pdf" docOrigin="Zootaxa 5395 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5395.1.1/52550" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.16:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="16" docTitle="Bisetocreagris brevis Feng &amp; Zhang &amp; Chen 2023, sp. nov." docType="treatment" docUuid="2264FA33-7C38-4EBB-8A1C-E60AC84B3F23" docUuidSource="ZooBank" docVersion="3" lastPageNumber="22" masterDocId="FFAB4A037E7A630DFFA0FA66FFBA4815" masterDocTitle="Cave-dwelling Neobisiidae in South China Karst, with descriptions of twelve new species of Bisetocreagris (Pseudoscorpiones, Neobisiidae) from Guizhou Province" masterLastPageNumber="77" masterPageNumber="1" pageNumber="16" updateTime="1704289532487" updateUser="julia">
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<mods:title id="DA740A9B659B6901F0B1E065A160F802">Cave-dwelling Neobisiidae in South China Karst, with descriptions of twelve new species of Bisetocreagris (Pseudoscorpiones, Neobisiidae) from Guizhou Province</mods:title>
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<mods:namePart id="38E4C61B4325461D9B663983F14B5B90">Feng, Zegang</mods:namePart>
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<mods:affiliation id="1BF98AD0E74B92B44E1D4AFE10659100">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, P. R. China https: // orcid. org / 0000 - 0002 - 5809 - 5984 &amp; College of Life Sciences, University of Chinese Academy of Sciences, Beijing, 100049, P. R. China chenj @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 0325 - 2532 &amp; Key Laboratory of Zoological Systematics and Application, Hebei Basic Science Center for Biotic Interaction, College of Life sciences, Hebei University, Baoding, Hebei 071002, P. R. China dudu 06042001 @ 163. com; https: // orcid. org / 0000 - 0002 - 3347 - 1031 * Corresponding author</mods:affiliation>
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<mods:namePart id="16C300C6E0F50D7E5EE7367086B55DAD">Zhang, Feng</mods:namePart>
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<mods:affiliation id="A523355590301F9B448744769E434CB8">Key Laboratory of Zoological Systematics and Application, Hebei Basic Science Center for Biotic Interaction, College of Life sciences, Hebei University, Baoding, Hebei 071002, P. R. China dudu 06042001 @ 163. com; https: // orcid. org / 0000 - 0002 - 3347 - 1031 * Corresponding author</mods:affiliation>
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<mods:namePart id="9941E16C48269ECF17E928C1F5349431">Chen, Jun</mods:namePart>
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<mods:affiliation id="913938FE4A255DB19937499F017C6222">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, P. R. China https: // orcid. org / 0000 - 0002 - 5809 - 5984 &amp; College of Life Sciences, University of Chinese Academy of Sciences, Beijing, 100049, P. R. China chenj @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 0325 - 2532</mods:affiliation>
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<taxonomicName id="4C3BF8EE7E756302FF37FC8DFE294F10" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[151,403,1770,1797]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="B94F5F7F7E756302FF37FC8DFE294F10" bold="true" box="[151,403,1770,1797]" italics="true" pageId="15" pageNumber="16">Bisetocreagris brevis</emphasis>
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(Ħae双ŁÞ伪AE)
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<figureCitation id="13009FE87E756302FF37FD55FEAE4F58" box="[151,276,1843,1869]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStart-2="FIGURE 14" captionStart-3="FIGURE 15" captionStart-4="FIGURE 16" captionStartId-0="16.[151,250,1174,1198]" captionStartId-1="17.[151,250,1062,1086]" captionStartId-2="17.[151,250,1851,1875]" captionStartId-3="18.[151,250,1943,1967]" captionStartId-4="19.[151,250,1874,1898]" captionTargetBox-0="[151,1435,181,1150]" captionTargetBox-1="[151,1436,181,1038]" captionTargetBox-2="[151,1435,1110,1827]" captionTargetBox-3="[178,1410,181,1919]" captionTargetBox-4="[206,1382,186,1846]" captionTargetId-0="figure-21@16.[151,1435,181,1150]" captionTargetId-1="figure-17@17.[151,1436,181,1038]" captionTargetId-2="figure-33@17.[151,1435,1110,1827]" captionTargetId-3="figure-21@18.[178,1410,181,1919]" captionTargetId-4="figure-17@19.[202,1386,181,1850]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="17" captionTargetPageId-3="18" captionTargetPageId-4="19" captionText-0="FIGURE 12. Yehu Cave, type locality of Bisetocreagris brevis sp. nov., A. Surrounding surface vegetation with cave entrance; B‒C. Entrances (B: Outside the entrance; C: Inside the entrance); D. Area where B. brevis sp. nov. specimens were collected." captionText-1="FIGURE 13. Live individual of Bisetocreagris brevis sp. nov. (male) on the wall." captionText-2="FIGURE 14. Bisetocreagris brevis sp. nov., A. Holotype male, dorsal view; B. Paratype female, dorsal view." captionText-3="FIGURE 15. Bisetocreagris brevis sp. nov., holotype male (A–F, H–I), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." captionText-4="FIGURE 16. Bisetocreagris brevis sp. nov., holotype male (A–G, I–M), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Movable finger of chelicera, showing galea; E. Subterminal tarsal seta; F. Galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (C–E), 0.25 mm (B, G–H), 0.5 mm (A, J), 1 mm (I, K–M)." figureDoi-0="http://doi.org/10.5281/zenodo.10438353" figureDoi-1="http://doi.org/10.5281/zenodo.10438355" figureDoi-2="http://doi.org/10.5281/zenodo.10438357" figureDoi-3="http://doi.org/10.5281/zenodo.10438359" figureDoi-4="http://doi.org/10.5281/zenodo.10438361" httpUri-0="https://zenodo.org/record/10438353/files/figure.png" httpUri-1="https://zenodo.org/record/10438355/files/figure.png" httpUri-2="https://zenodo.org/record/10438357/files/figure.png" httpUri-3="https://zenodo.org/record/10438359/files/figure.png" httpUri-4="https://zenodo.org/record/10438361/files/figure.png" pageId="15" pageNumber="16">Figs 12–16</figureCitation>
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<emphasis id="B94F5F7F7E756302FF37FD1DFEF84F81" bold="true" box="[151,322,1914,1941]" pageId="15" pageNumber="16">Type material.</emphasis>
<materialsCitation id="3B5389307E756302FEE6FD1DFB2C4FC8" collectingDate="2020-08-22" collectorName="Zegang Feng &amp; Hongru Xu &amp; Yanmeng Hou" country="China" county="Yinjiang County" elevation="982" latitude="27.851883" location="Chanxi Town, Zhoujiaao Village, Yehu Cave" longLatPrecision="1" longitude="108.50988" municipality="Tongren City" pageId="15" pageNumber="16" specimenCode="MHBU-GZ20082201" specimenCount="1" stateProvince="Guizhou Province" typeStatus="holotype">
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male (Ps.- 
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): 
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, 
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, 
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, 
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, 
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(
<emphasis id="B94F5F7F7E756302FD6BFDF9FB654FAC" bold="true" box="[715,1247,1951,1977]" pageId="15" pageNumber="16">âṅåȇ仁Ṅ印ũ县dzäDz周ẋ坳ť夜壶ñ</emphasis>
), [
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, 
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], 
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a.s.l.
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<collectingDate id="EFC15C457E756302FE78FDA5FD364FC8" box="[472,652,1987,2013]" pageId="15" pageNumber="16" value="2020-08-22">22 August 2020</collectingDate>
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, 
<collectorName id="26CEE6BB7E756302FD37FDA5FC974FC8" box="[663,813,1987,2013]" pageId="15" pageNumber="16">Zegang Feng</collectorName>
, 
<collectorName id="26CEE6BB7E756302FC98FDA5FC004FC8" box="[824,954,1987,2013]" pageId="15" pageNumber="16">Hongru Xu</collectorName>
, 
<collectorName id="26CEE6BB7E756302FC64FDA5FBDF4FC8" box="[964,1125,1987,2013]" pageId="15" pageNumber="16">Yanmeng Hou</collectorName>
leg.
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<emphasis id="B94F5F7F7E6A631DFF37FEF0FEA54CBB" bold="true" box="[151,287,1174,1198]" pageId="16" pageNumber="17">FIGURE 12.</emphasis>
Yehu Cave, type locality of 
<taxonomicName id="4C3BF8EE7E6A631DFD9EFEF1FCAE4CBB" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[574,788,1174,1198]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="B94F5F7F7E6A631DFD9EFEF1FCAE4CBB" box="[574,788,1174,1198]" italics="true" pageId="16" pageNumber="17">Bisetocreagris brevis</emphasis>
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<taxonomicNameLabel id="A27CE2047E6A631DFCBAFEF0FCD04CBB" box="[794,874,1174,1198]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
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, A. Surrounding surface vegetation with cave entrance; B‒C. Entrances (B: Outside the entrance; C: Inside the entrance); D. Area where 
<taxonomicName id="4C3BF8EE7E6A631DFC6EFEDDFB9D4CC7" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[974,1063,1210,1234]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="B94F5F7F7E6A631DFC6EFEDDFB9D4CC7" box="[974,1063,1210,1234]" italics="true" pageId="16" pageNumber="17">B. brevis</emphasis>
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<taxonomicNameLabel id="A27CE2047E6A631DFB8DFEDCFBC74CC7" box="[1069,1149,1210,1234]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
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specimens were collected.
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: 
<specimenCount id="9D3D48E47E6A631DFEE3FF43FE234D2B" box="[323,409,1316,1343]" count="3" pageId="16" pageNumber="17" type="male">3 males</specimenCount>
(Ps.- 
<specimenCode id="DB9D2B167E6A631DFE6EFF43FD4C4D2A" box="[462,758,1317,1343]" collectionCode="MHBU-GZ" pageId="16" pageNumber="17">MHBU-GZ20082202‒04</specimenCode>
)
</materialsCitation>
, 
<materialsCitation id="3B5389307E6A631DFCA9FF42FF474D77" collectingDate="2020-08-22" collectorName="Zegang Feng &amp; Hongru Xu &amp; Yanmeng Hou" country="China" county="Yinjiang County" elevation="982" latitude="27.851883" location="Town" longLatPrecision="1" longitude="108.50988" municipality="Tongren City" pageId="16" pageNumber="17" specimenCode="MHBU-GZ20082205-10" specimenCount="6" specimenCount-female="6" stateProvince="Guizhou" typeStatus="paratype">
<specimenCount id="9D3D48E47E6A631DFCA9FF42FCCF4D2B" box="[777,885,1316,1342]" count="6" pageId="16" pageNumber="17" type="female">6 females</specimenCount>
(Ps.- 
<specimenCode id="DB9D2B167E6A631DFC0AFF42FB684D2A" box="[938,1234,1316,1343]" collectionCode="MHBU-GZ" pageId="16" pageNumber="17">MHBU-GZ20082205‒10</specimenCode>
), same data as the holotype
</materialsCitation>
; 
<materialsCitation id="3B5389307E6A631DFEA8FF2FFA2A4D77" box="[264,1424,1352,1379]" collectingDate="2014-10-21" collectorName="Xuankong Jiang" country="China" county="Yinjiang County" elevation="982" latitude="27.851883" location="Town" longLatPrecision="1" longitude="108.50988" municipality="Tongren City" pageId="16" pageNumber="17" specimenCode="MHBU-GZ14102101" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou" typeStatus="paratype">
<specimenCount id="9D3D48E47E6A631DFEA8FF2FFED34D77" box="[264,361,1352,1379]" count="1" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
(Ps.- 
<specimenCode id="DB9D2B167E6A631DFE3EFF2FFD264D76" box="[414,668,1353,1379]" collectionCode="MHBU-GZ" pageId="16" pageNumber="17">MHBU-GZ14102101</specimenCode>
), same location as holotype, 
<date id="FF85A5AD7E6A631DFC7EFF2FFB244D76" box="[990,1182,1352,1379]" pageId="16" pageNumber="17" value="2014-10-21">
<collectingDate id="EFC15C457E6A631DFC7EFF2FFB244D76" box="[990,1182,1352,1379]" pageId="16" pageNumber="17" value="2014-10-21">21 October 2014</collectingDate>
</date>
, 
<collectorName id="26CEE6BB7E6A631DFB09FF2EFAE54D77" box="[1193,1375,1352,1378]" pageId="16" pageNumber="17">Xuankong Jiang</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C321D0E67E6A631DFF67FF0BFD2E4DBF" pageId="16" pageNumber="17" type="etymology">
<paragraph id="8B84836D7E6A631DFF67FF0BFD2E4DBF" blockId="16.[151,1437,1316,2027]" pageId="16" pageNumber="17">
<emphasis id="B94F5F7F7E6A631DFF67FF0BFEF44D92" bold="true" box="[199,334,1389,1415]" pageId="16" pageNumber="17">Etymology.</emphasis>
The species name is derived from the Latin word 
<taxonomicName id="4C3BF8EE7E6A631DFC29FF0BFC624D93" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[905,984,1388,1415]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="brevis">
“ 
<emphasis id="B94F5F7F7E6A631DFC35FF0AFC624D93" box="[917,984,1388,1414]" italics="true" pageId="16" pageNumber="17">brevis</emphasis>
</taxonomicName>
”, meaning short and little, and refers to the shape of the pedicel of the pedipalp hand.
</paragraph>
</subSubSection>
<subSubSection id="C321D0E67E6A631DFF67FFD2FAC54E2E" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8B84836D7E6A631DFF67FFD2FAC54E2E" blockId="16.[151,1437,1316,2027]" pageId="16" pageNumber="17">
<emphasis id="B94F5F7F7E6A631DFF67FFD2FEDB4DDB" bold="true" box="[199,353,1460,1486]" pageId="16" pageNumber="17">Distribution.</emphasis>
This new species is known only from the 
<typeStatus id="54803DCF7E6A631DFCE5FFD3FCCF4DDA" box="[837,885,1461,1487]" pageId="16" pageNumber="17">type</typeStatus>
locality Yehu Cave (
<figureCitation id="13009FE87E6A631DFBC5FFD2FB784DDA" box="[1125,1218,1460,1487]" captionStart="FIGURE 1" captionStartId="4.[151,250,1063,1087]" captionTargetBox="[156,1436,186,1039]" captionTargetId="figure-21@4.[151,1436,181,1039]" captionTargetPageId="4" captionText="FIGURE 1. Distribution map of the type locality of troglomorphic Bisetocreagris species in China. A. B. maomaotou Gao, Wynne et Zhang, 2018 (Maomaotou Cave); B. B. que sp. nov. (Yanggong Cave); C. B. longidigitata sp. nov. (Mao’er Cave); D. B. brevis sp. nov. (Yehu Cave); E. B. aspera sp. nov. (Tu Cave); F. B. jiangi sp. nov. (Anjialin Cave); G. B. tenuipalma sp. nov. (Sanjie Cave); H. B. changbing sp. nov. (Sanyuan and Guanyin caves); I. B. ampla sp. nov. (Hei Cave); J. B. gracilenta Gao et Zhang, 2017 (Shanmu Cave); K. B. guangshanensis Gao et Zhang, 2017 (Guangshan Cave); L. B. chumi sp. nov. (Tufei Cave); M. B. sanmen sp. nov. (Sanmen Cave); N. B. xishui sp. nov. (Dongkoushang Cave); O. B. baozinensis Mahnert et Li, 2016 (Baozi Cave); P. B. chuanensis Mahnert et Li, 2016 (Chuan Cave); Q. B. grandis sp. nov. (Wulibeidadong Cave); R. B. scaura (Mahnert, 2003) (Dahei Cave); S. B. martii (Mahnert, 2003) and B. titania (Mahnert, 2003) (Guoquan Cave); T. B. xiaoensis Li et Liu, 2017 (Xiao Cave); U. B. juanxuae Mahnert et Li, 2016 (Dao Cave); V. B. cavernarum Mahnert et Li, 2016 (Ert Long Cave); W. B. chinacavernicola (Schawaller, 1995) (Xian Nin and Chao-Tian caves)." figureDoi="http://doi.org/10.5281/zenodo.10438331" httpUri="https://zenodo.org/record/10438331/files/figure.png" pageId="16" pageNumber="17">Figs 1D</figureCitation>
, 
<figureCitation id="13009FE87E6A631DFB70FFD3FB544DDA" box="[1232,1262,1461,1487]" captionStart="FIGURE 12" captionStartId="16.[151,250,1174,1198]" captionTargetBox="[151,1435,181,1150]" captionTargetId="figure-21@16.[151,1435,181,1150]" captionTargetPageId="16" captionText="FIGURE 12. Yehu Cave, type locality of Bisetocreagris brevis sp. nov., A. Surrounding surface vegetation with cave entrance; B‒C. Entrances (B: Outside the entrance; C: Inside the entrance); D. Area where B. brevis sp. nov. specimens were collected." figureDoi="http://doi.org/10.5281/zenodo.10438353" httpUri="https://zenodo.org/record/10438353/files/figure.png" pageId="16" pageNumber="17">12</figureCitation>
), located about ~ 
<quantity id="4CC32E887E6A631DFF05FFBEFF4C4DE7" box="[165,246,1496,1523]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="16" pageNumber="17" unit="km" value="4.5">4.5 km</quantity>
to the east of 
<collectingRegion id="49FF4D8F7E6A631DFE3DFFBEFE544DE7" box="[413,494,1496,1522]" country="China" name="Shaanxi" pageId="16" pageNumber="17">Chanxi</collectingRegion>
Town, Tongren City. The entrance is irregular oval (~ 
<quantity id="4CC32E887E6A631DFBFAFFBEFB264DE6" box="[1114,1180,1496,1523]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="17" unit="m" value="1.5">1.5 m</quantity>
high and ~ 
<quantity id="4CC32E887E6A631DFABFFFBFFAF74DE6" box="[1311,1357,1497,1523]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="16" pageNumber="17" unit="m" value="4.0">4 m</quantity>
wide), sheltered in a forest about 
<quantity id="4CC32E887E6A631DFE60FF9AFE434E02" box="[448,505,1532,1559]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="17" unit="m" value="15.0">15 m</quantity>
below the man-made diversion canal and surrounded by weeds. The cave is moist, about 
<quantity id="4CC32E887E6A631DFF7CFC47FE994E2E" box="[220,291,1569,1595]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="17" unit="m" value="100.0">100 m</quantity>
long, extending horizontally. All specimens were collected on the wall (
<figureCitation id="13009FE87E6A631DFBEDFC46FB094E2E" box="[1101,1203,1568,1595]" captionStart="FIGURE 12" captionStartId="16.[151,250,1174,1198]" captionTargetBox="[151,1435,181,1150]" captionTargetId="figure-21@16.[151,1435,181,1150]" captionTargetPageId="16" captionText="FIGURE 12. Yehu Cave, type locality of Bisetocreagris brevis sp. nov., A. Surrounding surface vegetation with cave entrance; B‒C. Entrances (B: Outside the entrance; C: Inside the entrance); D. Area where B. brevis sp. nov. specimens were collected." figureDoi="http://doi.org/10.5281/zenodo.10438353" httpUri="https://zenodo.org/record/10438353/files/figure.png" pageId="16" pageNumber="17">Fig. 12D</figureCitation>
) of the dark zone.
</paragraph>
</subSubSection>
<subSubSection id="C321D0E67E6A631DFF67FC22FC314F23" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8B84836D7E6A631DFF67FC22FC314F23" blockId="16.[151,1437,1316,2027]" pageId="16" pageNumber="17">
<emphasis id="B94F5F7F7E6A631DFF67FC22FEEA4E4B" bold="true" box="[199,336,1604,1630]" pageId="16" pageNumber="17">Diagnosis. (</emphasis>
<specimenCount id="9D3D48E47E6A631DFEF1FC23FED04E4A" box="[337,362,1605,1631]" count="1" pageId="16" pageNumber="17" type="male">♂</specimenCount>
<specimenCount id="9D3D48E47E6A631DFEC9FC23FEC04E4A" box="[361,378,1605,1631]" count="1" pageId="16" pageNumber="17" type="female">♀</specimenCount>
<emphasis id="B94F5F7F7E6A631DFEDBFC22FE3E4E4B" bold="true" box="[379,388,1604,1630]" pageId="16" pageNumber="17">)</emphasis>
. Troglomorphic habitus; carapace without eyes or eyespots; epistome triangular, with rounded tip; carapace with 4–6 setae on posterior margin; pedipalp smooth and slender, both chelal fingers with 120–139 teeth; femur 8.14–8.81 times (length 2.19–2.44), patella 6.84–7.40 times (length 2.05–2.27) longer than wide, pedicel about one-third of the total length of patella. Hand with pedicel 3.00–3.42 times (length 1.54–1.72), chela with pedicel 7.69–8.02 (
<specimenCount id="9D3D48E47E6A631DFE05FCB3FE064EFA" box="[421,444,1749,1775]" count="1" pageId="16" pageNumber="17" type="male">♂</specimenCount>
), 7.13–7.79 (
<specimenCount id="9D3D48E47E6A631DFDF1FCB3FDD94EFA" box="[593,611,1749,1775]" count="1" pageId="16" pageNumber="17" type="female">♀</specimenCount>
) times (length 3.61–3.92) longer than wide, finger 1.41–1.52 times longer than hand with pedicel. Chelicera: rallum with 10–14 pinnate setae, distal one with a slightly expanded base, distal-most 1–3 blades slightly separated from others, proximal one short.
</paragraph>
</subSubSection>
<subSubSection id="C321D0E67E6A6318FF67FD26FD714A0C" lastPageId="21" lastPageNumber="22" pageId="16" pageNumber="17" type="description">
<paragraph id="8B84836D7E6A631DFF67FD26FD374F6B" blockId="16.[151,1437,1316,2027]" pageId="16" pageNumber="17">
<emphasis id="B94F5F7F7E6A631DFF67FD26FEE34F4F" bold="true" box="[199,345,1856,1882]" pageId="16" pageNumber="17">Description.</emphasis>
Male (
<typeStatus id="54803DCF7E6A631DFE03FD26FDB04F4F" box="[419,522,1856,1882]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
; 
<typeStatus id="54803DCF7E6A631DFDB3FD27FDC44F4E" box="[531,638,1857,1883]" pageId="16" pageNumber="17" type="paratype">paratypes</typeStatus>
males in parentheses; 
<figureCitation id="13009FE87E6A631DFCD2FD26FC7F4F4E" box="[882,965,1856,1883]" captionStart="FIGURE 13" captionStartId="17.[151,250,1062,1086]" captionTargetBox="[151,1436,181,1038]" captionTargetId="figure-17@17.[151,1436,181,1038]" captionTargetPageId="17" captionText="FIGURE 13. Live individual of Bisetocreagris brevis sp. nov. (male) on the wall." figureDoi="http://doi.org/10.5281/zenodo.10438355" httpUri="https://zenodo.org/record/10438355/files/figure.png" pageId="16" pageNumber="17">Figs 13</figureCitation>
, 
<figureCitation id="13009FE87E6A631DFC6EFD27FBB84F4E" box="[974,1026,1857,1883]" captionStart="FIGURE 14" captionStartId="17.[151,250,1851,1875]" captionTargetBox="[151,1435,1110,1827]" captionTargetId="figure-33@17.[151,1435,1110,1827]" captionTargetPageId="17" captionText="FIGURE 14. Bisetocreagris brevis sp. nov., A. Holotype male, dorsal view; B. Paratype female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10438357" httpUri="https://zenodo.org/record/10438357/files/figure.png" pageId="16" pageNumber="17">14A</figureCitation>
). Carapace, chelicerae, and pedipalps reddish brown; abdomen and legs yellowish.
</paragraph>
<paragraph id="8B84836D7E6A631DFF67FDEFFB894FFF" blockId="16.[151,1437,1316,2027]" pageId="16" pageNumber="17">
Carapace (
<figureCitation id="13009FE87E6A631DFEE1FDEEFE6B4FB6" box="[321,465,1928,1955]" captionStart="FIGURE 15" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[178,1410,181,1919]" captionTargetId="figure-21@18.[178,1410,181,1919]" captionTargetPageId="18" captionText="FIGURE 15. Bisetocreagris brevis sp. nov., holotype male (A–F, H–I), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438359" httpUri="https://zenodo.org/record/10438359/files/figure.png" pageId="16" pageNumber="17">Figs 15A–B</figureCitation>
, 
<figureCitation id="13009FE87E6A631DFE7FFDEEFDAE4FB7" box="[479,532,1928,1954]" captionStart="FIGURE 16" captionStartId="19.[151,250,1874,1898]" captionTargetBox="[206,1382,186,1846]" captionTargetId="figure-17@19.[202,1386,181,1850]" captionTargetPageId="19" captionText="FIGURE 16. Bisetocreagris brevis sp. nov., holotype male (A–G, I–M), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Movable finger of chelicera, showing galea; E. Subterminal tarsal seta; F. Galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (C–E), 0.25 mm (B, G–H), 0.5 mm (A, J), 1 mm (I, K–M)." figureDoi="http://doi.org/10.5281/zenodo.10438361" httpUri="https://zenodo.org/record/10438361/files/figure.png" pageId="16" pageNumber="17">16A</figureCitation>
): smooth, 1.18 (1.17–1.23) times longer than wide, with a total of 27 (24–26) setae, including 4 (3–4) on anterior margin, 6 (4–4) on posterior margin, and 2–3 (2–3) on each side of antero-lateral margin; without eyes or eyespots (
<figureCitation id="13009FE87E6A631DFDB8FDB6FDCD4FFE" box="[536,631,2000,2027]" captionStart="FIGURE 15" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[178,1410,181,1919]" captionTargetId="figure-21@18.[178,1410,181,1919]" captionTargetPageId="18" captionText="FIGURE 15. Bisetocreagris brevis sp. nov., holotype male (A–F, H–I), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438359" httpUri="https://zenodo.org/record/10438359/files/figure.png" pageId="16" pageNumber="17">Fig 15B</figureCitation>
); epistome triangular, with rounded tip.
</paragraph>
<caption id="DF44D3E57E6B631CFF37FE40FC6D4C2B" ID-DOI="http://doi.org/10.5281/zenodo.10438355" ID-Zenodo-Dep="10438355" box="[151,983,1062,1086]" httpUri="https://zenodo.org/record/10438355/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,1062,1086]" targetBox="[151,1436,181,1038]" targetPageId="17" targetType="figure">
<paragraph id="8B84836D7E6B631CFF37FE40FC6D4C2B" blockId="17.[151,983,1062,1086]" box="[151,983,1062,1086]" pageId="17" pageNumber="18">
<emphasis id="B94F5F7F7E6B631CFF37FE40FE9A4C2B" bold="true" box="[151,288,1062,1086]" pageId="17" pageNumber="18">FIGURE 13.</emphasis>
Live individual of 
<taxonomicName id="4C3BF8EE7E6B631CFE43FE41FD034C2B" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[483,697,1062,1086]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="B94F5F7F7E6B631CFE43FE41FD034C2B" box="[483,697,1062,1086]" italics="true" pageId="17" pageNumber="18">Bisetocreagris brevis</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E6B631CFD60FE40FCAA4C2B" bold="true" box="[704,784,1062,1086]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A27CE2047E6B631CFD60FE40FCAA4C2B" box="[704,784,1062,1086]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(male) on the wall.
</paragraph>
</caption>
<caption id="DF44D3E57E6B631CFF37FD5DFB564F46" ID-DOI="http://doi.org/10.5281/zenodo.10438357" ID-Zenodo-Dep="10438357" box="[151,1260,1851,1876]" httpUri="https://zenodo.org/record/10438357/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,1851,1875]" targetBox="[151,1435,1110,1827]" targetPageId="17" targetType="figure">
<paragraph id="8B84836D7E6B631CFF37FD5DFB564F46" blockId="17.[151,1260,1851,1876]" box="[151,1260,1851,1876]" pageId="17" pageNumber="18">
<emphasis id="B94F5F7F7E6B631CFF37FD5DFE9A4F46" bold="true" box="[151,288,1851,1875]" pageId="17" pageNumber="18">FIGURE 14.</emphasis>
<taxonomicName id="4C3BF8EE7E6B631CFE86FD5AFE464F46" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[294,508,1851,1875]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="B94F5F7F7E6B631CFE86FD5AFE464F46" box="[294,508,1851,1875]" italics="true" pageId="17" pageNumber="18">Bisetocreagris brevis</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E6B631CFDA2FD5AFDE94F41" bold="true" box="[514,595,1852,1876]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A27CE2047E6B631CFDA2FD5AFDE94F41" box="[514,595,1852,1876]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Holotype male, dorsal view; B. Paratype female, dorsal view.
</paragraph>
</caption>
<caption id="DF44D3E57E68631FFF37FDF1FC394FED" ID-DOI="http://doi.org/10.5281/zenodo.10438359" ID-Zenodo-Dep="10438359" httpUri="https://zenodo.org/record/10438359/files/figure.png" pageId="18" pageNumber="19" startId="18.[151,250,1943,1967]" targetBox="[178,1410,181,1919]" targetPageId="18" targetType="figure">
<paragraph id="8B84836D7E68631FFF37FDF1FC394FED" blockId="18.[151,1437,1943,2040]" pageId="18" pageNumber="19">
<emphasis id="B94F5F7F7E68631FFF37FDF1FE984FBA" bold="true" box="[151,290,1943,1967]" pageId="18" pageNumber="19">FIGURE 15.</emphasis>
<taxonomicName id="4C3BF8EE7E68631FFE8AFDFEFDB94FA5" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[298,515,1944,1968]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="B94F5F7F7E68631FFE8AFDFEFDB94FA5" box="[298,515,1944,1968]" italics="true" pageId="18" pageNumber="19">Bisetocreagris brevis</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E68631FFDABFDFEFDE44FA5" bold="true" box="[523,606,1944,1968]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="A27CE2047E68631FFDABFDFEFDE44FA5" box="[523,606,1944,1968]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (A–F, H–I), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view.
</paragraph>
</caption>
<caption id="DF44D3E57E69631EFF37FD34FD1B4FEF" ID-DOI="http://doi.org/10.5281/zenodo.10438361" ID-Zenodo-Dep="10438361" httpUri="https://zenodo.org/record/10438361/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,1874,1898]" targetBox="[206,1382,186,1846]" targetPageId="19" targetType="figure">
<paragraph id="8B84836D7E69631EFF37FD34FD1B4FEF" blockId="19.[151,1436,1874,2042]" pageId="19" pageNumber="20">
<emphasis id="B94F5F7F7E69631EFF37FD34FE9E4F7F" bold="true" box="[151,292,1874,1898]" pageId="19" pageNumber="20">FIGURE 16.</emphasis>
<taxonomicName id="4C3BF8EE7E69631EFE8EFD35FDB24F7F" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[302,520,1874,1898]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="B94F5F7F7E69631EFE8EFD35FDB24F7F" box="[302,520,1874,1898]" italics="true" pageId="19" pageNumber="20">Bisetocreagris brevis</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E69631EFDB2FD34FDDC4F7F" bold="true" box="[530,614,1874,1898]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="A27CE2047E69631EFDB2FD34FDDC4F7F" box="[530,614,1874,1898]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (A–G, I–M), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Movable finger of chelicera, showing galea; E. Subterminal tarsal seta; F. Galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (C–E), 0.25 mm (B, G–H), 0.5 mm (A, J), 1 mm (I, K–M).
</paragraph>
</caption>
<paragraph id="8B84836D7E6E6319FF67FAF1FD744954" blockId="20.[151,1437,150,2013]" pageId="20" pageNumber="21">
Chelicera (
<figureCitation id="13009FE87E6E6319FEE0FAF1FE1348A4" box="[320,425,150,177]" captionStart="FIGURE 15" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[178,1410,181,1919]" captionTargetId="figure-21@18.[178,1410,181,1919]" captionTargetPageId="18" captionText="FIGURE 15. Bisetocreagris brevis sp. nov., holotype male (A–F, H–I), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438359" httpUri="https://zenodo.org/record/10438359/files/figure.png" pageId="20" pageNumber="21">Figs 15C</figureCitation>
, 
<figureCitation id="13009FE87E6E6319FE13FAF1FD9E48A4" box="[435,548,151,177]" captionStart="FIGURE 16" captionStartId="19.[151,250,1874,1898]" captionTargetBox="[206,1382,186,1846]" captionTargetId="figure-17@19.[202,1386,181,1850]" captionTargetPageId="19" captionText="FIGURE 16. Bisetocreagris brevis sp. nov., holotype male (A–G, I–M), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Movable finger of chelicera, showing galea; E. Subterminal tarsal seta; F. Galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (C–E), 0.25 mm (B, G–H), 0.5 mm (A, J), 1 mm (I, K–M)." figureDoi="http://doi.org/10.5281/zenodo.10438361" httpUri="https://zenodo.org/record/10438361/files/figure.png" pageId="20" pageNumber="21">16B–D, F</figureCitation>
): hand with 6 (6–7) setae, movable finger with 1 (1) submedial seta; fixed finger with 14–15 (13–19) teeth; movable finger with 12–14 (13–16) teeth; serrula exterior with 35 (33) lamellae; serrula interior with 25 (23) lamellae. Galea (
<figureCitation id="13009FE87E6E6319FDE4FAB9FD7448EC" box="[580,718,223,249]" captionStart="FIGURE 16" captionStartId="19.[151,250,1874,1898]" captionTargetBox="[206,1382,186,1846]" captionTargetId="figure-17@19.[202,1386,181,1850]" captionTargetPageId="19" captionText="FIGURE 16. Bisetocreagris brevis sp. nov., holotype male (A–G, I–M), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Movable finger of chelicera, showing galea; E. Subterminal tarsal seta; F. Galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (C–E), 0.25 mm (B, G–H), 0.5 mm (A, J), 1 mm (I, K–M)." figureDoi="http://doi.org/10.5281/zenodo.10438361" httpUri="https://zenodo.org/record/10438361/files/figure.png" pageId="20" pageNumber="21">Figs 16D, F</figureCitation>
) elongated with only one main branch, apically forked. Rallum (
<figureCitation id="13009FE87E6E6319FF3FFB65FEBF4908" box="[159,261,259,285]" captionStart="FIGURE 16" captionStartId="19.[151,250,1874,1898]" captionTargetBox="[206,1382,186,1846]" captionTargetId="figure-17@19.[202,1386,181,1850]" captionTargetPageId="19" captionText="FIGURE 16. Bisetocreagris brevis sp. nov., holotype male (A–G, I–M), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Movable finger of chelicera, showing galea; E. Subterminal tarsal seta; F. Galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (C–E), 0.25 mm (B, G–H), 0.5 mm (A, J), 1 mm (I, K–M)." figureDoi="http://doi.org/10.5281/zenodo.10438361" httpUri="https://zenodo.org/record/10438361/files/figure.png" pageId="20" pageNumber="21">Fig. 16C</figureCitation>
) with 11 (10–14) pinnate blade, distal-most blade with a slightly expanded base, distal-most 1–2 blades slightly separated from others, proximal one short.
</paragraph>
<paragraph id="8B84836D7E6E6319FF67FB2DFB274AD8" blockId="20.[151,1437,150,2013]" pageId="20" pageNumber="21">
Pedipalps (
<figureCitation id="13009FE87E6E6319FEE3FB2DFE744970" box="[323,462,330,357]" captionStart="FIGURE 15" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[178,1410,181,1919]" captionTargetId="figure-21@18.[178,1410,181,1919]" captionTargetPageId="18" captionText="FIGURE 15. Bisetocreagris brevis sp. nov., holotype male (A–F, H–I), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438359" httpUri="https://zenodo.org/record/10438359/files/figure.png" pageId="20" pageNumber="21">Figs 15D–E</figureCitation>
, 
<figureCitation id="13009FE87E6E6319FE79FB2DFD9C4970" box="[473,550,331,357]" captionStart="FIGURE 16" captionStartId="19.[151,250,1874,1898]" captionTargetBox="[206,1382,186,1846]" captionTargetId="figure-17@19.[202,1386,181,1850]" captionTargetPageId="19" captionText="FIGURE 16. Bisetocreagris brevis sp. nov., holotype male (A–G, I–M), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Movable finger of chelicera, showing galea; E. Subterminal tarsal seta; F. Galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (C–E), 0.25 mm (B, G–H), 0.5 mm (A, J), 1 mm (I, K–M)." figureDoi="http://doi.org/10.5281/zenodo.10438361" httpUri="https://zenodo.org/record/10438361/files/figure.png" pageId="20" pageNumber="21">16I–K</figureCitation>
): apex of coxae rounded, with 5 (5) setae on each side, pedipalpal coxae with 6 (6–8) setae. Pedipalp slender and smooth. Trochanter 3.26 (3.20–3.32) times longer than wide, femur 8.36 (8.42– 8.70), patella 7.28 (6.84–7.40) times longer than wide, pedicel about one-third of the total length of patella, chela (with pedicel) 7.98 (7.69–8.02), chela (without pedicel) 7.59 (7.24–7.60) times longer than wide, movable finger 1.61 (1.64–1.72) times longer than hand (without pedicel). Fixed chelal finger with 8 trichobothria, movable finger with 4, 
<emphasis id="B94F5F7F7E6E6319FF51FB99FEB14A0C" box="[241,267,511,537]" italics="true" pageId="20" pageNumber="21">eb</emphasis>
and 
<emphasis id="B94F5F7F7E6E6319FEE8FB99FED74A0C" box="[328,365,511,537]" italics="true" pageId="20" pageNumber="21">esb</emphasis>
on lateral margin of hand; 
<emphasis id="B94F5F7F7E6E6319FD0EFB99FD7E4A0C" box="[686,708,511,537]" italics="true" pageId="20" pageNumber="21">ib</emphasis>
, 
<emphasis id="B94F5F7F7E6E6319FD74FB99FD544A0D" box="[724,750,511,536]" italics="true" pageId="20" pageNumber="21">ist</emphasis>
and 
<emphasis id="B94F5F7F7E6E6319FC8AFB99FCF14A0C" box="[810,843,511,537]" italics="true" pageId="20" pageNumber="21">isb</emphasis>
closely grouped at the base of the fixed finger; 
<emphasis id="B94F5F7F7E6E6319FADDFB99FA264A0D" box="[1405,1436,511,536]" italics="true" pageId="20" pageNumber="21">est</emphasis>
proximal of finger middle; 
<emphasis id="B94F5F7F7E6E6319FE6DF845FE404A29" box="[461,506,547,573]" italics="true" pageId="20" pageNumber="21">it -et</emphasis>
at same level near fingertip; on movable finger 
<emphasis id="B94F5F7F7E6E6319FB82F845FB8F4A29" box="[1058,1077,547,572]" italics="true" pageId="20" pageNumber="21">st</emphasis>
nearer to 
<emphasis id="B94F5F7F7E6E6319FB09F845FB0B4A29" box="[1193,1201,547,572]" italics="true" pageId="20" pageNumber="21">t</emphasis>
than to 
<emphasis id="B94F5F7F7E6E6319FAB0F845FA934A28" box="[1296,1321,547,573]" italics="true" pageId="20" pageNumber="21">sb</emphasis>
, the latter nearer 
<emphasis id="B94F5F7F7E6E6319FF45F821FF494A74" box="[229,243,583,609]" italics="true" pageId="20" pageNumber="21">b</emphasis>
than to 
<emphasis id="B94F5F7F7E6E6319FEF2F821FEDF4A75" box="[338,357,583,608]" italics="true" pageId="20" pageNumber="21">st</emphasis>
, the distance between 
<emphasis id="B94F5F7F7E6E6319FDC8F821FD3B4A74" box="[616,641,583,609]" italics="true" pageId="20" pageNumber="21">sb</emphasis>
and 
<emphasis id="B94F5F7F7E6E6319FD1AF821FD724A74" box="[698,712,583,609]" italics="true" pageId="20" pageNumber="21">b</emphasis>
is somewhat equal to that of 
<emphasis id="B94F5F7F7E6E6319FBBAF821FB974A75" box="[1050,1069,583,608]" italics="true" pageId="20" pageNumber="21">st</emphasis>
and 
<emphasis id="B94F5F7F7E6E6319FBC5F821FBD74A75" box="[1125,1133,583,608]" italics="true" pageId="20" pageNumber="21">t</emphasis>
(
<figureCitation id="13009FE87E6E6319FBDDF821FB504A74" box="[1149,1258,582,609]" captionStart="FIGURE 15" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[178,1410,181,1919]" captionTargetId="figure-21@18.[178,1410,181,1919]" captionTargetPageId="18" captionText="FIGURE 15. Bisetocreagris brevis sp. nov., holotype male (A–F, H–I), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438359" httpUri="https://zenodo.org/record/10438359/files/figure.png" pageId="20" pageNumber="21">Figs 15D</figureCitation>
, 
<figureCitation id="13009FE87E6E6319FB57F821FA834A74" box="[1271,1337,583,609]" captionStart="FIGURE 16" captionStartId="19.[151,250,1874,1898]" captionTargetBox="[206,1382,186,1846]" captionTargetId="figure-17@19.[202,1386,181,1850]" captionTargetPageId="19" captionText="FIGURE 16. Bisetocreagris brevis sp. nov., holotype male (A–G, I–M), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Movable finger of chelicera, showing galea; E. Subterminal tarsal seta; F. Galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (C–E), 0.25 mm (B, G–H), 0.5 mm (A, J), 1 mm (I, K–M)." figureDoi="http://doi.org/10.5281/zenodo.10438361" httpUri="https://zenodo.org/record/10438361/files/figure.png" pageId="20" pageNumber="21">16I–J</figureCitation>
). Venom apparatus present only in fixed chelal finger, venom duct short, not extending past half of the distance to 
<emphasis id="B94F5F7F7E6E6319FA9AF80DFAF44A91" box="[1338,1358,619,644]" italics="true" pageId="20" pageNumber="21">et</emphasis>
. Fixed chelal finger with 127 (131–139) pointed and retrorse teeth; movable finger with 128 (129–134) teeth, which are pointed and retrorse in distal, blunt and round in middle and basal part; all teeth contiguous.
</paragraph>
<paragraph id="8B84836D7E6E6319FF67F8B1FD824B94" blockId="20.[151,1437,150,2013]" pageId="20" pageNumber="21">
Abdomen: pleural membrane granulated. Tergal chaetotaxy (I–XI): 6:6:6:6:6:6:6:6:8:7:8 (5–6:3–6:6:6:6:5– 6:6:6–7:6–8:6–8:7);sternal chaetotaxy(V–XI):10:15:14:13:12:13:13:4(10–12:11–14:11–13:12:12–13:12–14:12:4); stigmata with 5–6 (3–7) setae; anal cone with 2 (2) dorsal and 2 (2) ventral setae. Male genital area (
<figureCitation id="13009FE87E6E6319FB4CF979FAEA4B2C" box="[1260,1360,798,825]" captionStart="FIGURE 15" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[178,1410,181,1919]" captionTargetId="figure-21@18.[178,1410,181,1919]" captionTargetPageId="18" captionText="FIGURE 15. Bisetocreagris brevis sp. nov., holotype male (A–F, H–I), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438359" httpUri="https://zenodo.org/record/10438359/files/figure.png" pageId="20" pageNumber="21">Figs 15F</figureCitation>
, 
<figureCitation id="13009FE87E6E6319FAFBF979FA354B2C" box="[1371,1423,799,825]" captionStart="FIGURE 16" captionStartId="19.[151,250,1874,1898]" captionTargetBox="[206,1382,186,1846]" captionTargetId="figure-17@19.[202,1386,181,1850]" captionTargetPageId="19" captionText="FIGURE 16. Bisetocreagris brevis sp. nov., holotype male (A–G, I–M), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Movable finger of chelicera, showing galea; E. Subterminal tarsal seta; F. Galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (C–E), 0.25 mm (B, G–H), 0.5 mm (A, J), 1 mm (I, K–M)." figureDoi="http://doi.org/10.5281/zenodo.10438361" httpUri="https://zenodo.org/record/10438361/files/figure.png" pageId="20" pageNumber="21">16G</figureCitation>
): sternite II with 42 (39–44) scattered setae; sternite III with anteromedian groove flanked by one small seta on each side, with 31 (27–34) posterior setae.
</paragraph>
<paragraph id="8B84836D7E6E6319FF67F9EDFEB74C04" blockId="20.[151,1437,150,2013]" pageId="20" pageNumber="21">
Legs: coxae chaetotaxy (I–IV): 9:7:4:8–9 (7–11:6–9:3–5:8–10). Leg I (
<figureCitation id="13009FE87E6E6319FBA2F9EDFBC84BB0" box="[1026,1138,906,933]" captionStart="FIGURE 15" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[178,1410,181,1919]" captionTargetId="figure-21@18.[178,1410,181,1919]" captionTargetPageId="18" captionText="FIGURE 15. Bisetocreagris brevis sp. nov., holotype male (A–F, H–I), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438359" httpUri="https://zenodo.org/record/10438359/files/figure.png" pageId="20" pageNumber="21">Figs 15H</figureCitation>
, 
<figureCitation id="13009FE87E6E6319FB21F9EDFB094BB0" box="[1153,1203,907,933]" captionStart="FIGURE 16" captionStartId="19.[151,250,1874,1898]" captionTargetBox="[206,1382,186,1846]" captionTargetId="figure-17@19.[202,1386,181,1850]" captionTargetPageId="19" captionText="FIGURE 16. Bisetocreagris brevis sp. nov., holotype male (A–G, I–M), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Movable finger of chelicera, showing galea; E. Subterminal tarsal seta; F. Galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (C–E), 0.25 mm (B, G–H), 0.5 mm (A, J), 1 mm (I, K–M)." figureDoi="http://doi.org/10.5281/zenodo.10438361" httpUri="https://zenodo.org/record/10438361/files/figure.png" pageId="20" pageNumber="21">16L</figureCitation>
): femur 7.05 (6.38– 7.17), patella 4.94 (4.53–4.94), tibia 8.54 (8.15–9.38), basitarsus 5.27 (5.09–5.64), telotarsus 7.00 (7.27–7.50) times longer than wide, femur 1.60 (1.54–1.68) times longer than patella, telotarsus 1.33 (1.27–1.34) times longer than basitarsus.
</paragraph>
<paragraph id="8B84836D7E6E6319FF67FE7DFD754CD0" blockId="20.[151,1437,150,2013]" pageId="20" pageNumber="21">
Leg IV (
<figureCitation id="13009FE87E6E6319FE84FE7DFE3E4C20" box="[292,388,1050,1077]" captionStart="FIGURE 15" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[178,1410,181,1919]" captionTargetId="figure-21@18.[178,1410,181,1919]" captionTargetPageId="18" captionText="FIGURE 15. Bisetocreagris brevis sp. nov., holotype male (A–F, H–I), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438359" httpUri="https://zenodo.org/record/10438359/files/figure.png" pageId="20" pageNumber="21">Figs 15I</figureCitation>
, 
<figureCitation id="13009FE87E6E6319FE2FFE7DFE5D4C20" box="[399,487,1051,1077]" captionStart="FIGURE 16" captionStartId="19.[151,250,1874,1898]" captionTargetBox="[206,1382,186,1846]" captionTargetId="figure-17@19.[202,1386,181,1850]" captionTargetPageId="19" captionText="FIGURE 16. Bisetocreagris brevis sp. nov., holotype male (A–G, I–M), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Movable finger of chelicera, showing galea; E. Subterminal tarsal seta; F. Galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (C–E), 0.25 mm (B, G–H), 0.5 mm (A, J), 1 mm (I, K–M)." figureDoi="http://doi.org/10.5281/zenodo.10438361" httpUri="https://zenodo.org/record/10438361/files/figure.png" pageId="20" pageNumber="21">16E, M</figureCitation>
): femur + patella 8.36 (7.96–8.58) times longer than wide, femur shorter than patella, tibia 11.76 (11.76–12.80), basitarsus 5.77 (5.20–5.54), telotarsus 7.25 (7.31–8.27) times longer than wide, telotarsus 1.16 (1.19–1.26) times longer than basitarsus; basitarsus with one tactile setae (TS=0.13–0.14), telotarsus with one tactile setae (TS=0.59–0.62); subterminal tarsal seta (
<figureCitation id="13009FE87E6E6319FC81FEE1FC3F4CB4" box="[801,901,1159,1185]" captionStart="FIGURE 16" captionStartId="19.[151,250,1874,1898]" captionTargetBox="[206,1382,186,1846]" captionTargetId="figure-17@19.[202,1386,181,1850]" captionTargetPageId="19" captionText="FIGURE 16. Bisetocreagris brevis sp. nov., holotype male (A–G, I–M), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Movable finger of chelicera, showing galea; E. Subterminal tarsal seta; F. Galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (C–E), 0.25 mm (B, G–H), 0.5 mm (A, J), 1 mm (I, K–M)." figureDoi="http://doi.org/10.5281/zenodo.10438361" httpUri="https://zenodo.org/record/10438361/files/figure.png" pageId="20" pageNumber="21">Fig. 16E</figureCitation>
) bifurcated, both branches dentate; arolium not divided, shorter than the slender and simple claws.
</paragraph>
<paragraph id="8B84836D7E6E6319FF67FEA9FB954CFC" blockId="20.[151,1437,150,2013]" box="[199,1071,1231,1257]" pageId="20" pageNumber="21">
Females (
<typeStatus id="54803DCF7E6E6319FE93FEA9FE1E4CFC" box="[307,420,1231,1257]" pageId="20" pageNumber="21" type="paratype">paratypes</typeStatus>
; 
<figureCitation id="13009FE87E6E6319FE0FFEA9FDAF4CFC" box="[431,533,1231,1257]" captionStart="FIGURE 14" captionStartId="17.[151,250,1851,1875]" captionTargetBox="[151,1435,1110,1827]" captionTargetId="figure-33@17.[151,1435,1110,1827]" captionTargetPageId="17" captionText="FIGURE 14. Bisetocreagris brevis sp. nov., A. Holotype male, dorsal view; B. Paratype female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10438357" httpUri="https://zenodo.org/record/10438357/files/figure.png" pageId="20" pageNumber="21">Fig. 14B</figureCitation>
): mostly same as 
<typeStatus id="54803DCF7E6E6319FD7BFEA9FC874CFC" box="[731,829,1231,1257]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
and 
<typeStatus id="54803DCF7E6E6319FCD4FEA9FC654CFC" box="[884,991,1231,1257]" pageId="20" pageNumber="21" type="paratype">paratypes</typeStatus>
males.
</paragraph>
<paragraph id="8B84836D7E6E6319FF67FE95FBE04D40" blockId="20.[151,1437,150,2013]" pageId="20" pageNumber="21">
Chelicera. Hand with 6–7 setae, movable finger with 1 submedial seta; fixed finger with 14–18 teeth; movable finger with 12–16 teeth; serrula exterior with 35–38 lamellae; serrula interior with 27–34 lamellae. Galea with only one main branch and apically forked; rallum of 11–14 blades, and similar to 
<typeStatus id="54803DCF7E6E6319FC51FF5DFBED4D40" box="[1009,1111,1339,1365]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B84836D7E6E6319FF67FF39FCDA4DF0" blockId="20.[151,1437,150,2013]" pageId="20" pageNumber="21">Pedipalps. Apex of coxae rounded, with 5 setae on each side. Pedipalpal coxae with 5–8 setae. Trochanter 3.13–3.28, femur 8.14–8.81, patella 6.84–7.40, chela (with pedicel) 7.13–7.79, chela (without pedicel) 6.70–7.35 times longer than wide, movable finger 1.58–1.65 times longer than hand (without pedicel). Fixed chelal finger with about 121–137 teeth, movable finger with about 120–135 teeth.</paragraph>
<paragraph id="8B84836D7E6E6319FF67FF89FB984E44" blockId="20.[151,1437,150,2013]" pageId="20" pageNumber="21">
Abdomen. Tergal chaetotaxy (I–XI): 5–6:3–6:5–6:4–6:6:5–7:6–7:6–7:6–8:7–8:7–8; sternal chaetotaxy (IV– XI): 9–12:12–15:12–16:12–13:12–14:11–14:12–13:2–4. Female genital area (
<figureCitation id="13009FE87E6E6319FBA3FC75FBCA4E38" box="[1027,1136,1554,1581]" captionStart="FIGURE 15" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[178,1410,181,1919]" captionTargetId="figure-21@18.[178,1410,181,1919]" captionTargetPageId="18" captionText="FIGURE 15. Bisetocreagris brevis sp. nov., holotype male (A–F, H–I), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438359" httpUri="https://zenodo.org/record/10438359/files/figure.png" pageId="20" pageNumber="21">Figs 15G</figureCitation>
, 
<figureCitation id="13009FE87E6E6319FBDCFC75FB0A4E38" box="[1148,1200,1555,1581]" captionStart="FIGURE 16" captionStartId="19.[151,250,1874,1898]" captionTargetBox="[206,1382,186,1846]" captionTargetId="figure-17@19.[202,1386,181,1850]" captionTargetPageId="19" captionText="FIGURE 16. Bisetocreagris brevis sp. nov., holotype male (A–G, I–M), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Movable finger of chelicera, showing galea; E. Subterminal tarsal seta; F. Galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (C–E), 0.25 mm (B, G–H), 0.5 mm (A, J), 1 mm (I, K–M)." figureDoi="http://doi.org/10.5281/zenodo.10438361" httpUri="https://zenodo.org/record/10438361/files/figure.png" pageId="20" pageNumber="21">16H</figureCitation>
): sternite II with 4–9 setae on each side; sternite III with a row of 16–21 setae on the posterior margin.
</paragraph>
<paragraph id="8B84836D7E6E6318FF67FC3DFA9648C0" blockId="20.[151,1437,150,2013]" lastBlockId="21.[151,1437,150,1041]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
Measurements: (length/breadth or depth in mm; ratios for most characters in parentheses). 
<typeStatus id="54803DCF7E6E6319FB52FC3DFAE04E60" box="[1266,1370,1627,1653]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
male (
<typeStatus id="54803DCF7E6E6319FF3FFC19FEB64E8C" box="[159,268,1663,1689]" pageId="20" pageNumber="21" type="paratype">paratypes</typeStatus>
males in square brackets). Body length 4.52 [4.73–5.02]. Carapace 1.13/0.96 (1.18) [1.04–1.16/0.89– 0.96 (1.17–1.23)]. Pedipalpal trochanter 1.01/0.31 (3.26) [0.96–1.03/0.30–0.32 (3.20–3.32)], femur 2.34/0.28 (8.36) [2.19–2.38/0.26–0.28 (8.42–8.70)], patella 2.11/0.29 (7.28) [2.05–2.22/0.29–0.31 (6.84–7.40)], chela (with pedicel) 3.67/0.46 (7.98) [3.61–3.88/0.45–0.49 (7.69–8.02)], chela (without pedicel) 3.49/0.46 (7.59) [3.42–3.67/0.45–0.49 (7.24–7.60)], hand length (without pedicel) 1.38 [1.37–1.44], movable finger length 2.22 (1.61 times longer than hand without pedicel) [2.24–2.41 (1.64–1.72 times longer than hand without pedicel)]. Leg I: trochanter 0.39/0.23 (1.70) [0.37–0.40/0.22–0.25 (1.60–1.68)], femur 1.34/0.19 (7.05) [1.29–1.37/0.18–0.21 (6.38–7.17)], patella 0.84/0.17 (4.94) [0.77–0.89/0.17–0.18 (4.53–4.94)], tibia 1.11/0.13 (8.54) [1.06–1.22/0.13 (8.15–9.38)], basitarsus 0.58/0.11 (5.27) [0.56–0.63/0.11–0.12 (5.09–5.64)], telotarsus 0.77/0.11 (7.00) [0.75–0.80/0.10–0.11 (7.27–7.50)]. Leg IV: trochanter 0.63/0.25 (2.52) [0.57–0.68/0.22–0.27 (2.52–2.62)], femur + patella 2.09/0.25 (8.36) [2.05– 2.23/0.24–0.27 (7.96–8.58)], tibia 2.00/0.17 (11.76) [1.92–2.07/0.15–0.17 (11.76–12.80)], basitarsus 0.75/0.13 (5.77) [0.72–0.78/0.13–0.15 (5.20–5.54)], telotarsus 0.87/0.12 (7.25) [0.91–0.95/0.11–0.13 (7.31–8.27)].
</paragraph>
<paragraph id="8B84836D7E6F6318FF67FAB9FD714A0C" blockId="21.[151,1437,150,1041]" pageId="21" pageNumber="22">
Females (
<typeStatus id="54803DCF7E6F6318FE93FAB9FE1F48EC" box="[307,421,223,249]" pageId="21" pageNumber="22" type="paratype">paratypes</typeStatus>
). Body length 3.41–5.53. Carapace 1.15–1.19/0.91–1.07 (1.11–1.29). Pedipalpal trochanter 1.00–1.09/0.32–0.34 (3.13–3.28), femur 2.25–2.44/0.27–0.29 (8.14–8.81), patella 2.15–2.27/0.30–0.32 (6.84–7.40), chela (with pedicel) 3.68–3.92/0.48–0.53 (7.13–7.79), chela (without pedicel) 3.48–3.68/0.48–0.53 (6.70–7.35), hand length (without pedicel) 1.41–1.52, movable finger length 2.28–2.49 (1.58–1.65 times longer than hand without pedicel). Leg I: trochanter 0.37–0.44/0.24–0.26 (1.54–1.69), femur 1.33–1.41/0.20–0.22 (6.32–7.00), patella 0.82– 0.90/0.17–0.19 (4.32–4.94), tibia 1.11–1.21/0.13–0.14 (8.54–9.00), basitarsus 0.57–0.63/0.11–0.13 (4.46–5.73), telotarsus 0.74–0.82/0.11–0.12 (6.17–7.45). Leg IV: trochanter 0.65–0.71/0.25–0.28 (2.54–2.65), femur + patella 2.12–2.27/0.25–0.29 (7.31–8.88), tibia 1.95–2.11/0.17–0.20 (10.30–11.72), basitarsus 0.75–0.80/0.14–0.16 (4.69– 5.71), telotarsus 0.90–0.99/0.12–0.15 (6.20–8.00).
</paragraph>
</subSubSection>
<subSubSection id="C321D0E67E6F6318FF67F845FBA44C04" pageId="21" pageNumber="22" type="discussion">
<paragraph id="8B84836D7E6F6318FF67F845FDA54B00" blockId="21.[151,1437,150,1041]" pageId="21" pageNumber="22">
<emphasis id="B94F5F7F7E6F6318FF67F845FE814A28" bold="true" box="[199,315,547,573]" pageId="21" pageNumber="22">Remarks.</emphasis>
<taxonomicName id="4C3BF8EE7E6F6318FEE6F845FD8C4A28" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[326,566,547,573]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="B94F5F7F7E6F6318FEE6F845FD8C4A28" box="[326,566,547,573]" italics="true" pageId="21" pageNumber="22">Bisetocreagris brevis</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E6F6318FDE1F845FD244A28" bold="true" box="[577,670,547,573]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="A27CE2047E6F6318FDE1F845FD244A28" box="[577,670,547,573]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles 
<taxonomicName id="4C3BF8EE7E6F6318FC83F845FB1F4A28" authority="Mahnert et Li, 2016" authorityName="Mahnert et Li" authorityYear="2016" box="[803,1189,547,573]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="juanxuae">
<emphasis id="B94F5F7F7E6F6318FC83F845FC174A29" box="[803,941,547,572]" italics="true" pageId="21" pageNumber="22">B. juanxuae</emphasis>
<bibRefCitation id="EFAAFE9C7E6F6318FC17F845FB1F4A28" author="Mahnert, V. &amp; Li, Y. C." box="[951,1189,547,573]" pageId="21" pageNumber="22" pagination="259 - 268" refId="ref48128" refString="Mahnert, V. &amp; Li, Y. C. (2016) Cave-inhabiting Neobisiidae (Arachnida: Pseudoscorpiones) from China, with description of four new species of Bisetocreagris Curcic. Revue suisse de Zoologie, 123, 259 - 268. https: // doi. org / 10.5281 / zenodo. 155299" type="journal article" year="2016">Mahnert et Li, 2016</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C3BF8EE7E6F6318FB13F845FF634A74" authority="(Mahnert, 2003)" authorityName="Dahei Cave" baseAuthorityName="Mahnert" baseAuthorityYear="2003" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="scaura">
<emphasis id="B94F5F7F7E6F6318FB13F845FA984A29" box="[1203,1314,547,572]" italics="true" pageId="21" pageNumber="22">B. scaura</emphasis>
(
<bibRefCitation id="EFAAFE9C7E6F6318FA95F845FF6B4A74" author="Mahnert, V." pageId="21" pageNumber="22" pagination="739 - 748" refId="ref48074" refString="Mahnert, V. (2003) Four new species of pseudoscorpions (Arachnida, Pseudoscorpiones: Neobisiidae, Chernetidae) from caves in Yunnan Province, China. Revue suisse de Zoologie, 110, 739 - 748. https: // doi. org / 10.5962 / bhl. part. 80209" type="journal article" year="2003">Mahnert, 2003</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C3BF8EE7E6F6318FEAFF821FD8C4A74" authority="(Mahnert, 2003)" authorityName="Guoquan Cave" baseAuthorityName="Mahnert" baseAuthorityYear="2003" box="[271,566,583,609]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="titania">
<emphasis id="B94F5F7F7E6F6318FEAFF821FECD4A75" box="[271,375,583,608]" italics="true" pageId="21" pageNumber="22">B. titania</emphasis>
(
<bibRefCitation id="EFAAFE9C7E6F6318FE25F821FD944A74" author="Mahnert, V." box="[389,558,583,609]" pageId="21" pageNumber="22" pagination="739 - 748" refId="ref48074" refString="Mahnert, V. (2003) Four new species of pseudoscorpions (Arachnida, Pseudoscorpiones: Neobisiidae, Chernetidae) from caves in Yunnan Province, China. Revue suisse de Zoologie, 110, 739 - 748. https: // doi. org / 10.5962 / bhl. part. 80209" type="journal article" year="2003">Mahnert, 2003</bibRefCitation>
)
</taxonomicName>
in having a similar slender pedipalpal and the pedipalpal pedicel that is about half the entire length of pedipalpal patella. The new species differs from 
<taxonomicName id="4C3BF8EE7E6F6318FC44F80DFBD64A91" authorityName="Mahnert et Li" authorityYear="2016" box="[996,1132,619,644]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="juanxuae">
<emphasis id="B94F5F7F7E6F6318FC44F80DFBD64A91" box="[996,1132,619,644]" italics="true" pageId="21" pageNumber="22">B. juanxuae</emphasis>
</taxonomicName>
by the absence of eyes or eyespots (
<emphasis id="B94F5F7F7E6F6318FEAAF8E9FE004ABD" box="[266,442,655,680]" italics="true" pageId="21" pageNumber="22">
vs. 
<taxonomicName id="4C3BF8EE7E6F6318FE92F8E9FE004ABD" authorityName="Mahnert et Li" authorityYear="2016" box="[306,442,655,680]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="juanxuae">B. juanxuae</taxonomicName>
</emphasis>
with 2 indistinct small eyes), the pedipalpal patella 6.84–7.40 (
<emphasis id="B94F5F7F7E6F6318FB39F8F6FB794AB9" bold="true" box="[1177,1219,656,684]" pageId="21" pageNumber="22">
<specimenCount id="9D3D48E47E6F6318FB39F8F6FB144AB9" box="[1177,1198,656,684]" count="1" pageId="21" pageNumber="22" type="male">♂</specimenCount>
<specimenCount id="9D3D48E47E6F6318FB0EF8F6FB794AB9" box="[1198,1219,656,684]" count="1" pageId="21" pageNumber="22" type="female">♀</specimenCount>
</emphasis>
) times longer than wide (
<emphasis id="B94F5F7F7E6F6318FF7EF8D5FF464AD9" box="[222,252,691,716]" italics="true" pageId="21" pageNumber="22">vs.</emphasis>
9.6 (
<emphasis id="B94F5F7F7E6F6318FE96F8D2FEF14AC5" bold="true" box="[310,331,692,720]" pageId="21" pageNumber="22">
<specimenCount id="9D3D48E47E6F6318FE96F8D2FEF14AC5" box="[310,331,692,720]" count="1" pageId="21" pageNumber="22" type="male">♂</specimenCount>
</emphasis>
), 9.1(
<emphasis id="B94F5F7F7E6F6318FE2FF8D2FE1E4AC5" bold="true" box="[399,420,692,720]" pageId="21" pageNumber="22">
<specimenCount id="9D3D48E47E6F6318FE2FF8D2FE1E4AC5" box="[399,420,692,720]" count="1" pageId="21" pageNumber="22" type="female">♀</specimenCount>
</emphasis>
) times in 
<taxonomicName id="4C3BF8EE7E6F6318FDB4F8D5FD204AD9" authorityName="Mahnert et Li" authorityYear="2016" box="[532,666,691,716]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="juanxuae">
<emphasis id="B94F5F7F7E6F6318FDB4F8D5FD204AD9" box="[532,666,691,716]" italics="true" pageId="21" pageNumber="22">B. juanxuae</emphasis>
</taxonomicName>
), the chela (with pedicel) 7.69–8.02 (
<emphasis id="B94F5F7F7E6F6318FB9FF8D2FBEE4AC5" bold="true" box="[1087,1108,692,720]" pageId="21" pageNumber="22">
<specimenCount id="9D3D48E47E6F6318FB9FF8D2FBEE4AC5" box="[1087,1108,692,720]" count="1" pageId="21" pageNumber="22" type="male">♂</specimenCount>
</emphasis>
), 7.13–7.79 (
<emphasis id="B94F5F7F7E6F6318FB4CF8D2FABB4AC5" bold="true" box="[1260,1281,692,720]" pageId="21" pageNumber="22">
<specimenCount id="9D3D48E47E6F6318FB4CF8D2FABB4AC5" box="[1260,1281,692,720]" count="1" pageId="21" pageNumber="22" type="female">♀</specimenCount>
</emphasis>
) times longer than wide (
<emphasis id="B94F5F7F7E6F6318FEB7F8B1FE8C4AE5" box="[279,310,727,752]" italics="true" pageId="21" pageNumber="22">vs.</emphasis>
10.8 (
<emphasis id="B94F5F7F7E6F6318FE20F8BEFE2F4AE1" bold="true" box="[384,405,728,756]" pageId="21" pageNumber="22">
<specimenCount id="9D3D48E47E6F6318FE20F8BEFE2F4AE1" box="[384,405,728,756]" count="1" pageId="21" pageNumber="22" type="male">♂</specimenCount>
</emphasis>
), 10.0 (
<emphasis id="B94F5F7F7E6F6318FE50F8BEFDBF4AE1" bold="true" box="[496,517,728,756]" pageId="21" pageNumber="22">
<specimenCount id="9D3D48E47E6F6318FE50F8BEFDBF4AE1" box="[496,517,728,756]" count="1" pageId="21" pageNumber="22" type="female">♀</specimenCount>
</emphasis>
) times in 
<taxonomicName id="4C3BF8EE7E6F6318FDD9F8B1FCBA4AE5" authorityName="Mahnert et Li" authorityYear="2016" box="[633,768,727,752]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="juanxuae">
<emphasis id="B94F5F7F7E6F6318FDD9F8B1FCBA4AE5" box="[633,768,727,752]" italics="true" pageId="21" pageNumber="22">B. juanxuae</emphasis>
</taxonomicName>
); and the presence of 127–139 teeth on both chelal fingers (
<emphasis id="B94F5F7F7E6F6318FF01F89DFF054B01" box="[161,191,763,788]" italics="true" pageId="21" pageNumber="22">vs.</emphasis>
146‒152 teeth in 
<taxonomicName id="4C3BF8EE7E6F6318FE29F89DFDB54B01" authorityName="Mahnert et Li" authorityYear="2016" box="[393,527,763,788]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="juanxuae">
<emphasis id="B94F5F7F7E6F6318FE29F89DFDB54B01" box="[393,527,763,788]" italics="true" pageId="21" pageNumber="22">B. juanxuae</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B84836D7E6F6318FF67F979FDD14BB0" blockId="21.[151,1437,150,1041]" pageId="21" pageNumber="22">
<taxonomicName id="4C3BF8EE7E6F6318FF67F979FE0C4B2C" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[199,438,799,825]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="B94F5F7F7E6F6318FF67F979FE0C4B2C" box="[199,438,799,825]" italics="true" pageId="21" pageNumber="22">Bisetocreagris brevis</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E6F6318FE60F979FDA64B2C" bold="true" box="[448,540,799,825]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="A27CE2047E6F6318FE60F979FDA64B2C" box="[448,540,799,825]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="4C3BF8EE7E6F6318FD17F979FC9F4B2D" authorityName="Dahei Cave" baseAuthorityName="Mahnert" baseAuthorityYear="2003" box="[695,805,799,824]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="scaura">
<emphasis id="B94F5F7F7E6F6318FD17F979FC9F4B2D" box="[695,805,799,824]" italics="true" pageId="21" pageNumber="22">B. scaura</emphasis>
</taxonomicName>
by the pedipalpal femur 8.36–8.70 (
<emphasis id="B94F5F7F7E6F6318FB6CF946FB5B4B29" bold="true" box="[1228,1249,800,828]" pageId="21" pageNumber="22">
<specimenCount id="9D3D48E47E6F6318FB6CF946FB5B4B29" box="[1228,1249,800,828]" count="1" pageId="21" pageNumber="22" type="male">♂</specimenCount>
</emphasis>
), 8.14–8.81 (
<emphasis id="B94F5F7F7E6F6318FADEF946FA294B29" bold="true" box="[1406,1427,800,828]" pageId="21" pageNumber="22">
<specimenCount id="9D3D48E47E6F6318FADEF946FA294B29" box="[1406,1427,800,828]" count="1" pageId="21" pageNumber="22" type="female">♀</specimenCount>
</emphasis>
) times longer than wide (
<emphasis id="B94F5F7F7E6F6318FE04F925FE794B49" box="[420,451,835,860]" italics="true" pageId="21" pageNumber="22">vs.</emphasis>
7.52 (tritonymph) times in 
<taxonomicName id="4C3BF8EE7E6F6318FD54F925FCE54B49" authorityName="Dahei Cave" baseAuthorityName="Mahnert" baseAuthorityYear="2003" box="[756,863,835,860]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="scaura">
<emphasis id="B94F5F7F7E6F6318FD54F925FCE54B49" box="[756,863,835,860]" italics="true" pageId="21" pageNumber="22">B. scaura</emphasis>
</taxonomicName>
), the chela (with pedicel) 7.69–8.02 (
<emphasis id="B94F5F7F7E6F6318FAA1F922FAAC4B75" bold="true" box="[1281,1302,836,864]" pageId="21" pageNumber="22">
<specimenCount id="9D3D48E47E6F6318FAA1F922FAAC4B75" box="[1281,1302,836,864]" count="1" pageId="21" pageNumber="22" type="male">♂</specimenCount>
</emphasis>
), 7.13–7.79 (
<emphasis id="B94F5F7F7E6F6318FF01F90EFF0C4B91" bold="true" box="[161,182,872,900]" pageId="21" pageNumber="22">
<specimenCount id="9D3D48E47E6F6318FF01F90EFF0C4B91" box="[161,182,872,900]" count="1" pageId="21" pageNumber="22" type="female">♀</specimenCount>
</emphasis>
) times longer than wide (
<emphasis id="B94F5F7F7E6F6318FE7BF901FE404B95" box="[475,506,871,896]" italics="true" pageId="21" pageNumber="22">vs.</emphasis>
8.65 (tritonymph) times in 
<taxonomicName id="4C3BF8EE7E6F6318FC95F901FC1B4B95" authorityName="Dahei Cave" baseAuthorityName="Mahnert" baseAuthorityYear="2003" box="[821,929,871,896]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="scaura">
<emphasis id="B94F5F7F7E6F6318FC95F901FC1B4B95" box="[821,929,871,896]" italics="true" pageId="21" pageNumber="22">B. scaura</emphasis>
</taxonomicName>
), and the presence of 127–139 teeth on both chelal fingers (
<emphasis id="B94F5F7F7E6F6318FE9FF9EDFEE44BB1" box="[319,350,907,932]" italics="true" pageId="21" pageNumber="22">vs.</emphasis>
105 teeth in 
<taxonomicName id="4C3BF8EE7E6F6318FE50F9EDFDE14BB1" authorityName="Dahei Cave" baseAuthorityName="Mahnert" baseAuthorityYear="2003" box="[496,603,907,932]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="scaura">
<emphasis id="B94F5F7F7E6F6318FE50F9EDFDE14BB1" box="[496,603,907,932]" italics="true" pageId="21" pageNumber="22">B. scaura</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B84836D7E6F6318FF67F9C9FBA44C04" blockId="21.[151,1437,150,1041]" pageId="21" pageNumber="22">
This new species differs from 
<taxonomicName id="4C3BF8EE7E6F6318FDBBF9C9FD394BDD" authorityName="Guoquan Cave" baseAuthorityName="Mahnert" baseAuthorityYear="2003" box="[539,643,943,968]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="titania">
<emphasis id="B94F5F7F7E6F6318FDBBF9C9FD394BDD" box="[539,643,943,968]" italics="true" pageId="21" pageNumber="22">B. titania</emphasis>
</taxonomicName>
by the pedipalpal femur 8.36–8.70 (
<emphasis id="B94F5F7F7E6F6318FBBEF9D6FB894BD9" bold="true" box="[1054,1075,944,972]" pageId="21" pageNumber="22">
<specimenCount id="9D3D48E47E6F6318FBBEF9D6FB894BD9" box="[1054,1075,944,972]" count="1" pageId="21" pageNumber="22" type="male">♂</specimenCount>
</emphasis>
) times longer than wide (
<emphasis id="B94F5F7F7E6F6318FAF4F9C9FAC84BDD" box="[1364,1394,943,968]" italics="true" pageId="21" pageNumber="22">vs.</emphasis>
8.9 (
<emphasis id="B94F5F7F7E6F6318FF01F9B2FF0C4BE5" bold="true" box="[161,182,980,1008]" pageId="21" pageNumber="22">
<specimenCount id="9D3D48E47E6F6318FF01F9B2FF0C4BE5" box="[161,182,980,1008]" count="1" pageId="21" pageNumber="22" type="male">♂</specimenCount>
</emphasis>
) times in 
<taxonomicName id="4C3BF8EE7E6F6318FE89F9B5FE294BF9" authorityName="Guoquan Cave" baseAuthorityName="Mahnert" baseAuthorityYear="2003" box="[297,403,979,1004]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="titania">
<emphasis id="B94F5F7F7E6F6318FE89F9B5FE294BF9" box="[297,403,979,1004]" italics="true" pageId="21" pageNumber="22">B. titania</emphasis>
</taxonomicName>
), the patella 6.84–7.40 (
<emphasis id="B94F5F7F7E6F6318FD09F9B2FD044BE5" bold="true" box="[681,702,980,1008]" pageId="21" pageNumber="22">
<specimenCount id="9D3D48E47E6F6318FD09F9B2FD044BE5" box="[681,702,980,1008]" count="1" pageId="21" pageNumber="22" type="male">♂</specimenCount>
</emphasis>
) times longer than wide (
<emphasis id="B94F5F7F7E6F6318FC44F9B5FBB94BF9" box="[996,1027,979,1004]" italics="true" pageId="21" pageNumber="22">vs.</emphasis>
8.0 (
<emphasis id="B94F5F7F7E6F6318FB9FF9B2FBEE4BE5" bold="true" box="[1087,1108,980,1008]" pageId="21" pageNumber="22">
<specimenCount id="9D3D48E47E6F6318FB9FF9B2FBEE4BE5" box="[1087,1108,980,1008]" count="1" pageId="21" pageNumber="22" type="male">♂</specimenCount>
</emphasis>
) times in 
<taxonomicName id="4C3BF8EE7E6F6318FB68F9B5FA8B4BF9" authorityName="Guoquan Cave" baseAuthorityName="Mahnert" baseAuthorityYear="2003" box="[1224,1329,979,1004]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="titania">
<emphasis id="B94F5F7F7E6F6318FB68F9B5FA8B4BF9" box="[1224,1329,979,1004]" italics="true" pageId="21" pageNumber="22">B. titania</emphasis>
</taxonomicName>
), and the presence of 127–139 teeth on both chelal finger (
<emphasis id="B94F5F7F7E6F6318FD1EF991FD664C05" box="[702,732,1015,1040]" italics="true" pageId="21" pageNumber="22">vs.</emphasis>
103‒110 teeth in 
<taxonomicName id="4C3BF8EE7E6F6318FC05F991FBB74C05" authorityName="Guoquan Cave" baseAuthorityName="Mahnert" baseAuthorityYear="2003" box="[933,1037,1015,1040]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="titania">
<emphasis id="B94F5F7F7E6F6318FC05F991FBB74C05" box="[933,1037,1015,1040]" italics="true" pageId="21" pageNumber="22">B. titania</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>