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<document id="8F9D0E840B9D4292ADF419E21628559B" ID-DOI="10.11646/zootaxa.3737.1.3" ID-GBIF-Dataset="d8c057a6-5061-44ea-9d54-61eee7700e1b" ID-ISSN="1175-5326" ID-Zenodo-Dep="285215" ID-ZooBank="CE23AED0-EBA7-422D-AB3C-47DC2D3FB4D" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460699631104" checkinUser="plazi" docAuthor="Ota, Yuzo" docDate="2013" docId="412987E8FFB63311FF3CFE95BFB42E97" docLanguage="en" docName="zt03737p056.pdf" docOrigin="Zootaxa 3737 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gnathia nasuta Nunomura 1992" docType="treatment" docVersion="6" lastPageNumber="47" masterDocId="BD10FF90FFBF331FFFABFFF9BC292B57" masterDocTitle="Redescription of five gnathiid species from Japan (Crustacea: Isopoda)" masterLastPageNumber="56" masterPageNumber="33" pageNumber="42" updateTime="1698354195244" updateUser="plazi">
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<mods:title id="48619432E39A61AC24C1AF2EC810BB07">Redescription of five gnathiid species from Japan (Crustacea: Isopoda)</mods:title>
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<mods:namePart id="78FBF1A1AA6ABDB9FA0F3BFEA4617E29">Ota, Yuzo</mods:namePart>
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<treatment id="412987E8FFB63311FF3CFE95BFB42E97" ID-DOI="http://doi.org/10.5281/zenodo.6160799" ID-GBIF-Taxon="127666416" ID-Zenodo-Dep="6160799" LSID="urn:lsid:plazi:treatment:412987E8FFB63311FF3CFE95BFB42E97" httpUri="http://treatment.plazi.org/id/412987E8FFB63311FF3CFE95BFB42E97" lastPageId="14" lastPageNumber="47" pageId="9" pageNumber="42">
<subSubSection id="819A6575FFB63316FF3CFE95B8D82945" pageId="9" pageNumber="42" type="nomenclature">
<paragraph id="C93F36FEFFB63316FF3CFE95BE052AD0" blockId="9.[151,632,364,461]" box="[151,556,364,391]" pageId="9" pageNumber="42">
<heading id="92778192FFB63316FF3CFE95BE052AD0" bold="true" box="[151,556,364,391]" fontSize="11" level="1" pageId="9" pageNumber="42" reason="1">
<taxonomicName id="0E804D7DFFB63316FF3CFE95BE052AD0" ID-CoL="3GLD3" authority="Nunomura, 1992" authorityName="Nunomura" authorityYear="1992" box="[151,556,364,391]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="FBF4EAECFFB63316FF3CFE95BE052AD0" bold="true" box="[151,556,364,391]" pageId="9" pageNumber="42">
<emphasis id="FBF4EAECFFB63316FF3CFE95BD7B2AD1" bold="true" box="[151,338,364,390]" italics="true" pageId="9" pageNumber="42">Gnathia nasuta</emphasis>
Nunomura, 1992
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C93F36FEFFB63316FF3CFE69BD232A9A" blockId="9.[151,632,364,461]" pageId="9" pageNumber="42">
(Japanese name: Hanadaka-umi-kuwagata) (
<figureCitation id="51BB2A7BFFB63316FF34FE4DBD2A2A9A" box="[159,259,436,461]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="11.[151,250,1954,1976]" captionStartId-1="12.[151,250,1953,1975]" captionStartId-2="13.[151,250,1955,1977]" captionTargetBox-0="[207,1379,193,1932]" captionTargetBox-1="[183,1403,193,1931]" captionTargetBox-2="[329,1261,193,1933]" captionTargetId-0="figure@11.[207,1379,193,1933]" captionTargetId-1="figure@12.[183,1403,193,1932]" captionTargetId-2="figure@13.[324,1262,193,1933]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionText-0="FIGURE 5. Gnathia nasuta Nunomura, 1992, (A, D; TOYA Cr- 23570), (B, C, E, F; TOYA Cr- 11172): A, body, dorsal view; B, cephalosome, dorsal view; C, cephalosome, lateral view; D, pleotelson, dorsal view; E, right antennule; F, right antenna." captionText-1="FIGURE 6. Gnathia nasuta Nunomura, 1992, (A, B, E; TOYA Cr- 11172), (C, D; TOYA Cr- 23570: A, right maxilliped, ventral view; B right pylopod, ventral view; C, left pereopod II, lateral view; D, left pleopod II; E, penes." captionText-2="FIGURE 7. Cephalosomes (A D) and pleotelsons (E H) of four holotypes: A and E, Gnathia hirayamai (TOYA Cr- 11167); B and F, Gnathia nagasakiensis (TOYA Cr- 23352); C and G, Gnathia shijikiensis (TOYA Cr- 23353); D and H, Gnathia brevicephala (TOYA Cr- 23356). The cephalosome of G. nagasakiensis was deformed." httpUri-0="https://zenodo.org/record/285220/files/figure.png" httpUri-1="https://zenodo.org/record/285221/files/figure.png" httpUri-2="https://zenodo.org/record/285222/files/figure.png" pageId="9" pageNumber="42">Figs 57</figureCitation>
)
</paragraph>
<paragraph id="C93F36FEFFB63316FF3CFE05B8D82945" blockId="9.[151,1436,508,867]" box="[151,1265,508,530]" pageId="9" pageNumber="42">
<taxonomicName id="0E804D7DFFB63316FF3CFE05BE242945" authority="Nunomura, 1992: 69" authorityName="Nunomura" authorityPageNumber="69" authorityYear="1992" box="[151,525,508,530]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="FBF4EAECFFB63316FF3CFE05BD1A2945" box="[151,307,508,530]" italics="true" pageId="9" pageNumber="42">Gnathia nasuta</emphasis>
Nunomura, 1992: 69
</taxonomicName>
71, fig. 6.—Nunomura, 1995: 209, fig. 21-196G.—Saito
<emphasis id="FBF4EAECFFB63316FBF2FE04B8AF2945" box="[1113,1158,508,530]" italics="true" pageId="9" pageNumber="42">et al</emphasis>
. 2000: 17.
</paragraph>
</subSubSection>
<subSubSection id="819A6575FFB63316FF3CFDE3BEE72834" pageId="9" pageNumber="42" type="description">
<paragraph id="C93F36FEFFB63316FF3CFDE3BD392918" blockId="9.[151,1436,508,867]" pageId="9" pageNumber="42">
<taxonomicName id="0E804D7DFFB63316FF3CFDE3BD592967" box="[151,368,538,560]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="FBF4EAECFFB63316FF3CFDE3BD592967" box="[151,368,538,560]" italics="true" pageId="9" pageNumber="42">Caecognathia nasuta</emphasis>
</taxonomicName>
.—Cohen &amp; Poore 1994: 288.—Smit &amp; Davies 2004: 296.—Nunomura &amp; Shimomura 2012: 585, 586, fig. 99.
</paragraph>
<paragraph id="C93F36FEFFB63316FF3CFDA1BEE729FC" blockId="9.[151,1436,508,867]" pageId="9" pageNumber="42">
<taxonomicName id="0E804D7DFFB63316FF3CFDA1BE192939" authority="Nunomura, 1992: 65" authorityName="Nunomura" authorityPageNumber="65" authorityYear="1992" box="[151,560,600,622]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="hirayamai">
<emphasis id="FBF4EAECFFB63316FF3CFDA1BD712939" box="[151,344,600,622]" italics="true" pageId="9" pageNumber="42">Gnathia hirayamai</emphasis>
Nunomura, 1992: 65
</taxonomicName>
67, fig. 4.—Cohen &amp; Poore 1994: 289.—Nunomura, 1995: 209, fig. 21-
<specimenCount id="DF86FD77FFB63316FAB8FDA1B9112939" box="[1299,1336,600,622]" pageId="9" pageNumber="42" type="generic">196</specimenCount>
F.—Saito
<emphasis id="FBF4EAECFFB63316FF6CFD81BCDD29DB" box="[199,244,630,653]" italics="true" pageId="9" pageNumber="42">et al</emphasis>
. 2000: 17.—Tanaka 2004: 58.—Nunomura, 2004: 354.—Nunomura &amp; Shimomura 2013a: 142, 145.—Nunomura &amp; Shimomura 2013b: 292, fig. 132. [New synonymy]
</paragraph>
<paragraph id="C93F36FEFFB63316FF3CFD4DBEE729BF" blockId="9.[151,1436,508,867]" pageId="9" pageNumber="42">
<taxonomicName id="0E804D7DFFB63316FF3CFD4DBE53299D" authority="Nunomura, 2012: 62" authorityName="Nunomura" authorityPageNumber="62" authorityYear="2012" box="[151,634,692,714]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="nagasakiensis">
<emphasis id="FBF4EAECFFB63316FF3CFD4DBDAF299D" box="[151,390,692,714]" italics="true" pageId="9" pageNumber="42">Gnathia nagasakiensis</emphasis>
Nunomura, 2012: 62
</taxonomicName>
, 63, fig. 3.—Nunomura &amp; Shimomura 2013a: 142, 144.—Nunomura &amp; Shimomura 2013b: 293, fig. 134. [New synonymy]
</paragraph>
<paragraph id="C93F36FEFFB63316FF3CFD08BE7B2871" blockId="9.[151,1436,508,867]" pageId="9" pageNumber="42">
<taxonomicName id="0E804D7DFFB63316FF3CFD08BE6E2850" authority="Nunomura, 2012a: 64" authorityName="Nunomura" authorityPageNumber="64" authorityYear="2012" box="[151,583,753,775]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="shijikiensis">
<emphasis id="FBF4EAECFFB63316FF3CFD08BD482850" box="[151,353,753,775]" italics="true" pageId="9" pageNumber="42">Gnathia shijikiensis</emphasis>
Nunomura, 2012a: 64
</taxonomicName>
, 65, fig. 4.—Nunomura &amp; Shimomura 2013a: 142, 144.—Nunomura &amp; Shimomura 2013b: 294, fig. 135. [New synonymy]
</paragraph>
<paragraph id="C93F36FEFFB63316FF3CFCD7BEE72834" blockId="9.[151,1436,508,867]" pageId="9" pageNumber="42">
<taxonomicName id="0E804D7DFFB63316FF3CFCD7BE532813" authority="Nunomura, 2012a: 65" authorityName="Nunomura" authorityPageNumber="65" authorityYear="2012" box="[151,634,814,836]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="brevicephala">
<emphasis id="FBF4EAECFFB63316FF3CFCD7BD552813" box="[151,380,814,836]" italics="true" pageId="9" pageNumber="42">Gnathia brevicephala</emphasis>
Nunomura, 2012a: 65
</taxonomicName>
67, fig. 5.—Nunomura &amp; Shimomura 2013a: 142, 144.—Nunomura &amp; Shimomura 2013b: 293, fig. 133. [New synonymy]
</paragraph>
</subSubSection>
<subSubSection id="819A6575FFB63315FF3CFC76B9302A4A" lastPageId="10" lastPageNumber="43" pageId="9" pageNumber="42" type="materials_examined">
<paragraph id="C93F36FEFFB63316FF3CFC76B8172F42" blockId="9.[151,1437,911,2017]" pageId="9" pageNumber="42">
<emphasis id="FBF4EAECFFB63316FF3CFC76BE0A28FF" bold="true" box="[151,547,911,936]" pageId="9" pageNumber="42">
Material examined of
<typeStatus id="163B885CFFB63316FE0FFC76BDFE28FF" box="[420,471,911,936]" pageId="9" pageNumber="42">type</typeStatus>
series
</emphasis>
. Male adult,
<typeStatus id="163B885CFFB63316FD16FC76BF3628FF" box="[701,799,911,936]" pageId="9" pageNumber="42" type="holotype">holotype</typeStatus>
of
<taxonomicName id="0E804D7DFFB63316FCE3FC69B93E28FF" authority="Nunomura, 1992" authorityName="Nunomura" authorityYear="1992" box="[840,1303,912,936]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="FBF4EAECFFB63316FCE3FC69B81028FF" box="[840,1081,912,936]" italics="true" pageId="9" pageNumber="42">Caecognathia nasuta</emphasis>
(Nunomura, 1992)
</taxonomicName>
, TOYA Cr- 11172,
<quantity id="0E789B1BFFB63316FF40FC4DBD6D289B" box="[235,324,948,972]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="9" pageNumber="42" unit="mm" value="3.3">3.3 mm</quantity>
, from
<quantity id="0E789B1BFFB63316FE3AFC4DBDFA289B" box="[401,467,948,973]" metricMagnitude="0" metricUnit="m" metricValue="8.5" pageId="9" pageNumber="42" unit="m" value="8.5">8.5 m</quantity>
depth, off Tomioka, Reihoku-cho, Kumamoto Pref., Kyushu, western
<collectingCountry id="B197766EFFB63316FB52FC4CB910289B" box="[1273,1337,949,972]" name="Japan" pageId="9" pageNumber="42">Japan</collectingCountry>
(32°20 22N, 130°0103E),
<date id="BD3E103EFFB63316FE20FC21BE0028A7" box="[395,553,984,1008]" pageId="9" pageNumber="42" value="1980-05-22">22 May 1980</date>
, coll. Taiji Kikuchi. Adult male,
<typeStatus id="163B885CFFB63316FC06FC2EB82628A7" box="[941,1039,983,1008]" pageId="9" pageNumber="42" type="holotype">holotype</typeStatus>
of
<taxonomicName id="0E804D7DFFB63316FB91FC21BCFB2F43" authority="Nunomura, 1992" authorityName="Nunomura" authorityYear="1992" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="hirayamai">
<emphasis id="FBF4EAECFFB63316FB91FC21B93A28A7" box="[1082,1299,984,1008]" italics="true" pageId="9" pageNumber="42">Gnathia hirayamai</emphasis>
Nunomura, 1992
</taxonomicName>
, TOYA Cr-11167,
<quantity id="0E789B1BFFB63316FE0EFC05BDD42F43" box="[421,509,1020,1044]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="9" pageNumber="42" unit="mm" value="3.3">3.3 mm</quantity>
, from same locality and date of
<emphasis id="FBF4EAECFFB63316FCCFFC05BFE22F43" box="[868,971,1020,1044]" italics="true" pageId="9" pageNumber="42">G. na s u t a</emphasis>
<typeStatus id="163B885CFFB63316FC7EFC05B8122F42" box="[981,1083,1020,1045]" pageId="9" pageNumber="42" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="C93F36FEFFB63316FF6CFBE6BE512E62" blockId="9.[151,1437,911,2017]" pageId="9" pageNumber="42">
Male adult,
<typeStatus id="163B885CFFB63316FEE7FBE6BD872F6F" box="[332,430,1055,1080]" pageId="9" pageNumber="42" type="holotype">holotype</typeStatus>
of
<taxonomicName id="0E804D7DFFB63316FE78FBD9BFB02F6F" authority="Nunomura, 2012" authorityName="Nunomura" authorityYear="2012" box="[467,921,1056,1080]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="nagasakiensis">
<emphasis id="FBF4EAECFFB63316FE78FBD9BEFB2F6F" box="[467,722,1056,1080]" italics="true" pageId="9" pageNumber="42">Gnathia nagasakiensis</emphasis>
Nunomura, 2012
</taxonomicName>
a, TOYA Cr-23312 (in original description but actually Cr-23352, 5.0 mm in original description but actually
<quantity id="0E789B1BFFB63316FCF3FBBDBF9B2F0B" box="[856,946,1092,1116]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="9" pageNumber="42" unit="mm" value="3.9">3.9 mm</quantity>
), from
<quantity id="0E789B1BFFB63316FBA8FBBDB8142F0B" box="[1027,1085,1092,1116]" metricMagnitude="1" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="42" unit="m" value="23.0">23 m</quantity>
depth of Shijiki Bay, Nagasaki Pref., western
<collectingCountry id="B197766EFFB63316FEEAFB91BDA82F28" box="[321,385,1128,1151]" name="Japan" pageId="9" pageNumber="42">Japan</collectingCountry>
(33°1113N, 129°2421E),
<date id="BD3E103EFFB63316FD42FB91BFA02FD7" box="[745,905,1128,1152]" pageId="9" pageNumber="42" value="1984-05-11">11 May 1984</date>
, coll. Mikio Azuma. Male adult,
<typeStatus id="163B885CFFB63316FABCFB9EB9502FD7" box="[1303,1401,1127,1152]" pageId="9" pageNumber="42" type="holotype">holotype</typeStatus>
of
<taxonomicName id="0E804D7DFFB63316FF3CFB75BE142FF3" authority="Nunomura, 2012" authorityName="Nunomura" authorityYear="2012" box="[151,573,1164,1188]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="shijikiensis">
<emphasis id="FBF4EAECFFB63316FF3CFB75BD5F2FF3" box="[151,374,1164,1188]" italics="true" pageId="9" pageNumber="42">Gnathia shijikiensis</emphasis>
Nunomura, 2012
</taxonomicName>
a, TOYA Cr-23353,
<quantity id="0E789B1BFFB63316FCB5FB75BF5D2FF3" box="[798,884,1164,1189]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="9" pageNumber="42" unit="mm" value="5.6">5.6 mm</quantity>
in original description but actually
<quantity id="0E789B1BFFB63316FAA9FB75B9732FF3" box="[1282,1370,1164,1188]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="9" pageNumber="42" unit="mm" value="4.4">4.4 mm</quantity>
, from
<quantity id="0E789B1BFFB63316FF3CFB56BCFD2F90" box="[151,212,1199,1224]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="42" unit="m" value="35.0">35 m</quantity>
depth of Shijiki Bay,
<date id="BD3E103EFFB63316FE49FB49BEA82F9F" box="[482,641,1200,1224]" pageId="9" pageNumber="42" value="1984-05-11">11 May 1984</date>
, coll. Mikio Azuma. Male adult,
<typeStatus id="163B885CFFB63316FBB8FB56B85C2F9F" box="[1043,1141,1199,1224]" pageId="9" pageNumber="42" type="holotype">holotype</typeStatus>
of
<taxonomicName id="0E804D7DFFB63316FB08FB49BD7E2FBB" authority="Nunomura, 1992" authorityName="Nunomura" authorityYear="1992" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="brevicephala">
<emphasis id="FBF4EAECFFB63316FB08FB49B9B52F9F" box="[1187,1436,1200,1224]" italics="true" pageId="9" pageNumber="42">Gnathia brevicephala</emphasis>
Nunomura, 1992
</taxonomicName>
, TOYA Cr-23356,
<quantity id="0E789B1BFFB63316FD80FB2DBEA82FBB" box="[555,641,1236,1261]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="9" pageNumber="42" unit="mm" value="5.6">5.6 mm</quantity>
in original description but actually
<quantity id="0E789B1BFFB63316FBA4FB2DB84E2FBB" box="[1039,1127,1236,1260]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="9" pageNumber="42" unit="mm" value="4.3">4.3 mm</quantity>
, from
<quantity id="0E789B1BFFB63316FB04FB2DB8C12FBB" box="[1199,1256,1236,1261]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="42" unit="m" value="25.0">25 m</quantity>
depth of Shijiki Bay,
<date id="BD3E103EFFB63316FF7FFB01BD4E2E47" box="[212,359,1272,1296]" pageId="9" pageNumber="42" value="1984-05-02">2 May 1984</date>
, coll. Mikio Azuma. All materials from Shijiki Bay were collected by Nagasaki University Sledge (NUS) net or Smith-McIntyre grab.
</paragraph>
<paragraph id="C93F36FEFFB63315FF6CFAC6B9302A4A" blockId="9.[151,1437,911,2017]" lastBlockId="10.[151,1437,151,2013]" lastPageId="10" lastPageNumber="43" pageId="9" pageNumber="42">
<emphasis id="FBF4EAECFFB63316FF6CFAC6BD412E0F" bold="true" box="[199,360,1343,1368]" pageId="9" pageNumber="42">New material</emphasis>
. Two male adults, TOYA Cr-23570 and 23571, 3.1 and
<quantity id="0E789B1BFFB63316FC71FAB9B81B2E00" box="[986,1074,1344,1368]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="9" pageNumber="42" unit="mm" value="3.1">3.1 mm</quantity>
, from the same locality and date of
<taxonomicName id="0E804D7DFFB63316FF1DFA9DBD0A2E2B" box="[182,291,1380,1404]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="FBF4EAECFFB63316FF1DFA9DBD0A2E2B" box="[182,291,1380,1404]" italics="true" pageId="9" pageNumber="42">C. nasuta</emphasis>
</taxonomicName>
<typeStatus id="163B885CFFB63316FE87FA9DBDBB2E2A" box="[300,402,1380,1405]" pageId="9" pageNumber="42" type="holotype">holotype</typeStatus>
. One male adult, KMNH IvR-500,708,
<quantity id="0E789B1BFFB63316FCFEFA9DBF862E2B" box="[853,943,1380,1404]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="9" pageNumber="42" unit="mm" value="2.4">2.4 mm</quantity>
, from
<quantity id="0E789B1BFFB63316FC51FA9DB86A2E2B" box="[1018,1091,1380,1404]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="42" unit="m" value="180.0">180 m</quantity>
depth off east Amami-oshima Island, southern
<collectingCountry id="B197766EFFB63316FECBFA71BD892EC8" box="[352,416,1416,1439]" name="Japan" pageId="9" pageNumber="42">Japan</collectingCountry>
(
<geoCoordinate id="ACB45039FFB63316FE1CFA7EBE122EF7" box="[439,571,1415,1440]" direction="north" orientation="latitude" pageId="9" pageNumber="42" precision="9" value="28.25">28°15.00N</geoCoordinate>
,
<geoCoordinate id="ACB45039FFB63316FDE5FA7EBEF42EF7" box="[590,733,1415,1440]" direction="east" orientation="longitude" pageId="9" pageNumber="42" precision="9" value="129.58333">129°35.00E</geoCoordinate>
), sledge net,
<date id="BD3E103EFFB63316FC2CFA71B8182EF7" box="[903,1073,1416,1440]" pageId="9" pageNumber="42" value="1994-11-10">10 Nov. 1994</date>
, coll. Takeo Yamauchi,
<collectingCountry id="B197766EFFB63316FAF7FA71B9A92EF7" box="[1372,1408,1416,1440]" name="Chile" pageId="9" pageNumber="42">TR</collectingCountry>
/V Toyoshio-maru. Six male adults, KMNH IvR-500,709,
<quantity id="0E789B1BFFB63316FC84FA55BFE92E93" box="[815,960,1452,1476]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="3.2" metricValueMin="2.2" pageId="9" pageNumber="42" unit="mm" value="2.7" valueMax="3.2" valueMin="2.2">2.23.2 mm</quantity>
, from
<quantity id="0E789B1BFFB63316FBBCFA55B84F2E93" box="[1047,1126,1452,1477]" metricMagnitude="2" metricUnit="m" metricValue="1.26" pageId="9" pageNumber="42" unit="m" value="126.0">126 m</quantity>
depth, sandy and muddy sediment, off west Mi-shima Island, Sea of
<collectingCountry id="B197766EFFB63316FD3CFA29BEFE2EB0" box="[663,727,1488,1511]" name="Japan" pageId="9" pageNumber="42">Japan</collectingCountry>
(34°36.163138N, 130°32.750894E), sledge net,
<date id="BD3E103EFFB63316FAEBFA29BCFB2D5B" pageId="9" pageNumber="42" value="1999-11-18">18 Nov. 1999</date>
, coll. Michitaka Shimomura,
<collectingCountry id="B197766EFFB63316FD86FA0DBE782D5B" box="[557,593,1524,1548]" name="Chile" pageId="9" pageNumber="42">TR</collectingCountry>
/V Toyoshio-maru. One male adult, KMNH IvR-500,710,
<quantity id="0E789B1BFFB63316FB51FA0DB97F2D5B" box="[1274,1366,1524,1549]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="42" unit="mm" value="2.6">2.6 mm</quantity>
, from
<quantity id="0E789B1BFFB63316FF3CF9EEBD302D67" box="[151,281,1559,1584]" metricMagnitude="2" metricUnit="m" metricValue="1.6649999999999998" metricValueMax="1.7" metricValueMin="1.63" pageId="9" pageNumber="42" unit="m" value="166.5" valueMax="170.0" valueMin="163.0">163170 m</quantity>
depth, off north Tokashiki-jima Island, Kerama Islands, southern
<collectingCountry id="B197766EFFB63316FBB0F9E0B8722D67" box="[1051,1115,1561,1584]" name="Japan" pageId="9" pageNumber="42">Japan</collectingCountry>
(
<geoCoordinate id="ACB45039FFB63316FBC6F9EEB8D82D67" box="[1133,1265,1559,1584]" direction="north" orientation="latitude" pageId="9" pageNumber="42" precision="9" value="26.061167">26°03.67N</geoCoordinate>
,
<geoCoordinate id="ACB45039FFB63316FAABF9EEB9A62D67" box="[1280,1423,1559,1584]" direction="east" orientation="longitude" pageId="9" pageNumber="42" precision="9" value="127.35233">127°21.14E</geoCoordinate>
), beam trawl,
<date id="BD3E103EFFB63316FE80F9C5BDE52D02" box="[299,460,1596,1621]" pageId="9" pageNumber="42" value="2006-05-26">26 May 2006</date>
, coll. Michitaka Shimomura,
<collectingCountry id="B197766EFFB63316FC80F9C5BF662D03" box="[811,847,1596,1620]" name="Chile" pageId="9" pageNumber="42">TR</collectingCountry>
/V Toyoshio-maru. Two male adults, KMNH IvR- 500,711, 2.3 and
<quantity id="0E789B1BFFB63316FECCF999BDED2D2F" box="[359,452,1632,1656]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="42" unit="mm" value="2.3">2.3 mm</quantity>
, from
<quantity id="0E789B1BFFB63316FDBDF9A6BE572D2F" box="[534,638,1631,1656]" metricMagnitude="1" metricUnit="m" metricValue="7.3" metricValueMax="7.5" metricValueMin="7.1" pageId="9" pageNumber="42" unit="m" value="73.0" valueMax="75.0" valueMin="71.0">7175 m</quantity>
depth, off south Okinawa-jima Island, southern
<collectingCountry id="B197766EFFB63316FB14F998B8D62D2F" box="[1215,1279,1633,1656]" name="Japan" pageId="9" pageNumber="42">Japan</collectingCountry>
(
<geoCoordinate id="ACB45039FFB63316FAB8F9A6B9BE2D2F" box="[1299,1431,1631,1656]" direction="north" orientation="latitude" pageId="9" pageNumber="42" precision="9" value="26.075">26°04.50N</geoCoordinate>
,
<geoCoordinate id="ACB45039FFB63316FF3CF97DBD0F2DCA" box="[151,294,1668,1693]" direction="east" orientation="longitude" pageId="9" pageNumber="42" precision="9" value="127.537">127°32.22E</geoCoordinate>
), beam trawl,
<date id="BD3E103EFFB63316FE7FF97DBE5F2DCA" box="[468,630,1668,1693]" pageId="9" pageNumber="42" value="2006-05-26">26 May 2006</date>
, coll. Michitaka Shimomura,
<collectingCountry id="B197766EFFB63316FC7DF97DBFD32DCB" box="[982,1018,1668,1692]" name="Chile" pageId="9" pageNumber="42">TR</collectingCountry>
/V Toyoshio-maru. One male adult, KMNH IvR-500,712,
<quantity id="0E789B1BFFB63316FE36F951BDD22D97" box="[413,507,1704,1728]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="9" pageNumber="42" unit="mm" value="3.2">3.2 mm</quantity>
, from
<quantity id="0E789B1BFFB63316FDFAF951BEFF2D97" box="[593,726,1704,1728]" metricMagnitude="2" metricUnit="m" metricValue="4.075" metricValueMax="4.12" metricValueMin="4.029999999999999" pageId="9" pageNumber="42" unit="m" value="407.5" valueMax="412.0" valueMin="403.0">403412 m</quantity>
depth, off southeast Amami-oshima Island, southern
<collectingCountry id="B197766EFFB63316FAF7F950B9B52D97" box="[1372,1436,1705,1728]" name="Japan" pageId="9" pageNumber="42">Japan</collectingCountry>
(
<geoCoordinate id="ACB45039FFB63316FF34F935BD0A2DB2" box="[159,291,1740,1765]" direction="north" orientation="latitude" pageId="9" pageNumber="42" precision="9" value="27.967">27°58.02N</geoCoordinate>
,
<geoCoordinate id="ACB45039FFB63316FE99F935BDE82DB2" box="[306,449,1740,1765]" direction="east" orientation="longitude" pageId="9" pageNumber="42" precision="9" value="129.4005">129°24.03E</geoCoordinate>
), inner net of beam trawl,
<date id="BD3E103EFFB63316FD50F935BFB02DB3" box="[763,921,1740,1765]" pageId="9" pageNumber="42" value="2009-05-22">22 May 2009</date>
, coll Susumu Otsuka,
<collectingCountry id="B197766EFFB63316FB0BF935B8ED2DB3" box="[1184,1220,1740,1764]" name="Chile" pageId="9" pageNumber="42">TR</collectingCountry>
/V Toyoshio-maru. Two male adults, KMNH IvR-500,713, 2.5 and 3.0 mm, from
<quantity id="0E789B1BFFB63316FCCCF909BF992C5F" box="[871,944,1776,1800]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="42" unit="m" value="130.0">130 m</quantity>
depth, muddy sediment with foraminiferal shells, off west Goto Islands, western
<collectingCountry id="B197766EFFB63316FD94F8ECBE562C7B" box="[575,639,1813,1836]" name="Japan" pageId="9" pageNumber="42">Japan</collectingCountry>
(
<geoCoordinate id="ACB45039FFB63316FD26F8EDBF2D2C7A" box="[653,772,1812,1837]" direction="north" orientation="latitude" pageId="9" pageNumber="42" precision="92" value="33.08">33°04.8N</geoCoordinate>
,
<geoCoordinate id="ACB45039FFB63316FCA4F8EDBFA62C7A" box="[783,911,1812,1837]" direction="east" orientation="longitude" pageId="9" pageNumber="42" precision="92" value="128.33667">128°20.2E</geoCoordinate>
), benthos net,
<date id="BD3E103EFFB63316FB9AF8EDB8942C7B" box="[1073,1213,1812,1837]" pageId="9" pageNumber="42" value="2010-07-25">25 Jul. 2010</date>
, coll. Keiich Kakui, R/V Soyo-maru. Two male adults, KMNH IvR-500,714, 2.7 and
<quantity id="0E789B1BFFB63316FCC4F8C1BFEE2C07" box="[879,967,1848,1872]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="9" pageNumber="42" unit="mm" value="3.4">3.4 mm</quantity>
, from
<quantity id="0E789B1BFFB63316FBBBF8C1B87E2C07" box="[1040,1111,1848,1872]" metricMagnitude="2" metricUnit="m" metricValue="1.42" pageId="9" pageNumber="42" unit="m" value="142.0">142 m</quantity>
depth, muddy sediment with foraminiferal shells, off west Goto Islands, western
<collectingCountry id="B197766EFFB63316FD76F8A4BF342C23" box="[733,797,1885,1908]" name="Japan" pageId="9" pageNumber="42">Japan</collectingCountry>
(
<geoCoordinate id="ACB45039FFB63316FC87F8A5BF8A2C22" box="[812,931,1884,1909]" direction="north" orientation="latitude" pageId="9" pageNumber="42" precision="92" value="33.268333">33°16.1N</geoCoordinate>
,
<geoCoordinate id="ACB45039FFB63316FC05F8A5B8062C22" box="[942,1071,1884,1909]" direction="east" orientation="longitude" pageId="9" pageNumber="42" precision="92" value="128.33667">128°20.2E</geoCoordinate>
), benthos net,
<date id="BD3E103EFFB63316FB79F8A5B9762C23" box="[1234,1375,1884,1909]" pageId="9" pageNumber="42" value="2010-07-25">25 Jul. 2010</date>
, coll. Keiich Kakui, R/V Soyo-maru.
<specimenCount id="DF86FD77FFB63316FDA9F879BE752CCF" box="[514,604,1919,1944]" pageId="9" pageNumber="42" type="male">17 male</specimenCount>
adults, KMNH IvR-500,715,
<quantity id="0E789B1BFFB63316FC1FF879B8172CCF" box="[948,1086,1920,1944]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.8" metricValueMin="2.4" pageId="9" pageNumber="42" unit="mm" value="3.0999999999999996" valueMax="3.8" valueMin="2.4">2.43.8 mm</quantity>
, from
<quantity id="0E789B1BFFB63316FB20F879B8FA2CCF" box="[1163,1235,1920,1944]" metricMagnitude="2" metricUnit="m" metricValue="2.13" pageId="9" pageNumber="42" unit="m" value="213.0">213 m</quantity>
depth, sandy and muddy sediment, off east Amakusa Town, Kumamoto Pref., western
<collectingCountry id="B197766EFFB63316FC1EF85CBFDC2CEB" box="[949,1013,1957,1980]" name="Japan" pageId="9" pageNumber="42">Japan</collectingCountry>
(
<geoCoordinate id="ACB45039FFB63316FBACF85DB8542CEA" box="[1031,1149,1956,1981]" direction="north" orientation="latitude" pageId="9" pageNumber="42" precision="92" value="32.22333">32°13.4N</geoCoordinate>
,
<geoCoordinate id="ACB45039FFB63316FB20F85DB9242CEA" box="[1163,1293,1956,1981]" direction="east" orientation="longitude" pageId="9" pageNumber="42" precision="92" value="128.65167">128°39.1E</geoCoordinate>
), dredge,
<date id="BD3E103EFFB63316FA2BF85DBD282CB7" pageId="9" pageNumber="42" value="2010-07-26">26 Jul. 2010</date>
, coll. Keiichi Kakui, R/V Soyo-maru. Nine male adults, KMNH IvR-500,716, 1.93.0 mm, from
<quantity id="0E789B1BFFB63316FAFFF831B9B52CB7" box="[1364,1436,1992,2017]" metricMagnitude="2" metricUnit="m" metricValue="2.05" pageId="9" pageNumber="42" unit="m" value="205.0">205 m</quantity>
depth of Tosa Bay, Shikoku, southern
<collectingCountry id="B197766EFFB53315FDE1FF61BEA32BF8" box="[586,650,152,175]" name="Japan" pageId="10" pageNumber="43">Japan</collectingCountry>
(
<geoCoordinate id="ACB45039FFB53315FD31FF6EBF7C2BE7" box="[666,853,151,176]" direction="north" orientation="latitude" pageId="10" pageNumber="43" precision="1" value="33.245903">33°1445.249N</geoCoordinate>
,
<geoCoordinate id="ACB45039FFB53315FCC9FF6EB8012BE7" box="[866,1064,151,176]" direction="east" orientation="longitude" pageId="10" pageNumber="43" precision="1" value="133.63168">133°3754.048E</geoCoordinate>
), sledge net with
<quantity id="0E789B1BFFB53315FB5AFF61B91A2BF8" box="[1265,1331,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="43" unit="mm" value="1.0">1 mm</quantity>
cod end,
<date id="BD3E103EFFB53315FF3CFF45BD192B83" box="[151,304,188,212]" pageId="10" pageNumber="43" value="2012-10-24">24 Oct. 2012</date>
, coll. Tomomi Saito. Six and
<specimenCount id="DF86FD77FFB53315FD27FF45BEF32B82" box="[652,730,188,213]" pageId="10" pageNumber="43" type="male">1 male</specimenCount>
adults, KMNH IvR-500,717 and 500,718,
<quantity id="0E789B1BFFB53315FB67FF45B97E2B83" box="[1228,1367,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.6" metricValueMin="2.9" pageId="10" pageNumber="43" unit="mm" value="3.25" valueMax="3.6" valueMin="2.9">2.93.6 mm</quantity>
, from
<quantity id="0E789B1BFFB53315FF3CFF26BD0D2BA0" box="[151,292,223,248]" metricMagnitude="1" metricUnit="m" metricValue="6.95" metricValueMax="7.59" metricValueMin="6.31" pageId="10" pageNumber="43" unit="m" value="69.5" valueMax="75.9" valueMin="63.1">63.175.9 m</quantity>
and
<quantity id="0E789B1BFFB53315FEF7FF19BDC02BA0" box="[348,489,224,248]" metricMagnitude="1" metricUnit="m" metricValue="2.7" metricValueMax="2.92" metricValueMin="2.48" pageId="10" pageNumber="43" unit="m" value="27.0" valueMax="29.2" valueMin="24.8">24.829.2 m</quantity>
depth, muddy sediments, off Tateyama, Boso Peninsula, central
<collectingCountry id="B197766EFFB53315FB6FFF19B92D2BA0" box="[1220,1284,224,247]" name="Japan" pageId="10" pageNumber="43">Japan</collectingCountry>
(
<geoCoordinate id="ACB45039FFB53315FAB8FF26B9B12BAF" box="[1299,1432,223,248]" direction="north" orientation="latitude" pageId="10" pageNumber="43" precision="9" value="34.991165">34°59.47N</geoCoordinate>
,
<geoCoordinate id="ACB45039FFB53315FF3CFEFDBD0A2A4A" box="[151,291,260,285]" direction="east" orientation="longitude" pageId="10" pageNumber="43" precision="9" value="139.783">139°46.98E</geoCoordinate>
and
<geoCoordinate id="ACB45039FFB53315FEF2FEFDBDF52A4A" box="[345,476,260,285]" direction="north" orientation="latitude" pageId="10" pageNumber="43" precision="9" value="34.994335">34°59.66N</geoCoordinate>
,
<geoCoordinate id="ACB45039FFB53315FE4CFEFDBE5F2A4A" box="[487,630,260,285]" direction="east" orientation="longitude" pageId="10" pageNumber="43" precision="9" value="139.80833">139°48.50E</geoCoordinate>
), dredge,
<date id="BD3E103EFFB53315FD4FFEFDBF532A4B" box="[740,890,260,285]" pageId="10" pageNumber="43" value="2013-04-16">16 Apr. 2013</date>
, coll. Yuzo Ota,
<collectingCountry id="B197766EFFB53315FB9FFEFDB8712A4B" box="[1076,1112,260,284]" name="Chile" pageId="10" pageNumber="43">TR</collectingCountry>
/V Hosaka-maru.
</paragraph>
</subSubSection>
<subSubSection id="819A6575FFB53315FF6CFEDEBD4C299B" pageId="10" pageNumber="43" type="distribution">
<paragraph id="C93F36FEFFB53315FF6CFEDEB8A62A17" blockId="10.[151,1437,151,2013]" box="[199,1167,295,320]" pageId="10" pageNumber="43">
<emphasis id="FBF4EAECFFB53315FF6CFEDEBD702A17" bold="true" box="[199,345,295,320]" pageId="10" pageNumber="43">Distribution</emphasis>
(
<figureCitation id="51BB2A7BFFB53315FEC3FEDEBD822A17" box="[360,427,295,320]" captionStart="FIGURE 1" captionStartId="1.[151,250,1758,1780]" captionTargetBox="[155,1430,763,1707]" captionTargetId="figure@1.[151,1436,756,1716]" captionTargetPageId="1" captionText="FIGURE 1. Sampling sites of holotype materials and new materials: k, Elaphognathia kikuchii; n, Gnathia nasuta; s, Gnathia sanrikuensis; m, Gnathia mutsuensis; b, Gnathia bungoensis. The numbers correspond to the specimen numbers in Table 1." httpUri="https://zenodo.org/record/285216/files/figure.png" pageId="10" pageNumber="43">Fig. 1</figureCitation>
,
<tableCitation id="84020345FFB53315FE12FEDEBE232A17" box="[441,522,295,320]" captionStart="TABLE 1" captionStartId="3.[151,235,152,174]" captionText="TABLE 1. Sample data of five gnathiid species. Female and larval specimens were collected but they were not able to identified due to mixture of sediment samples." pageId="10" pageNumber="43">Table 1</tableCitation>
). From southwestern to central
<collectingCountry id="B197766EFFB53315FCDAFED1BF982A68" box="[881,945,296,319]" name="Japan" pageId="10" pageNumber="43">Japan</collectingCountry>
(
<quantity id="0E789B1BFFB53315FC6BFEDEB8102A68" box="[960,1081,295,320]" metricMagnitude="2" metricUnit="m" metricValue="2.1025" metricValueMax="4.12" metricValueMin="0.08499999999999999" pageId="10" pageNumber="43" unit="m" value="210.25" valueMax="412.0" valueMin="8.5">8.5412 m</quantity>
depth).
</paragraph>
<paragraph id="C93F36FEFFB53315FF6CFEB5BD4C299B" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="43">
<emphasis id="FBF4EAECFFB53315FF6CFEB5BD442A32" bold="true" box="[199,365,332,357]" pageId="10" pageNumber="43">Redescription</emphasis>
(
<figureCitation id="51BB2A7BFFB53315FE2BFEB5BDC22A32" box="[384,491,332,357]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="11.[151,250,1954,1976]" captionStartId-1="12.[151,250,1953,1975]" captionStartId-2="13.[151,250,1955,1977]" captionTargetBox-0="[207,1379,193,1932]" captionTargetBox-1="[183,1403,193,1931]" captionTargetBox-2="[329,1261,193,1933]" captionTargetId-0="figure@11.[207,1379,193,1933]" captionTargetId-1="figure@12.[183,1403,193,1932]" captionTargetId-2="figure@13.[324,1262,193,1933]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionText-0="FIGURE 5. Gnathia nasuta Nunomura, 1992, (A, D; TOYA Cr- 23570), (B, C, E, F; TOYA Cr- 11172): A, body, dorsal view; B, cephalosome, dorsal view; C, cephalosome, lateral view; D, pleotelson, dorsal view; E, right antennule; F, right antenna." captionText-1="FIGURE 6. Gnathia nasuta Nunomura, 1992, (A, B, E; TOYA Cr- 11172), (C, D; TOYA Cr- 23570: A, right maxilliped, ventral view; B right pylopod, ventral view; C, left pereopod II, lateral view; D, left pleopod II; E, penes." captionText-2="FIGURE 7. Cephalosomes (A D) and pleotelsons (E H) of four holotypes: A and E, Gnathia hirayamai (TOYA Cr- 11167); B and F, Gnathia nagasakiensis (TOYA Cr- 23352); C and G, Gnathia shijikiensis (TOYA Cr- 23353); D and H, Gnathia brevicephala (TOYA Cr- 23356). The cephalosome of G. nagasakiensis was deformed." httpUri-0="https://zenodo.org/record/285220/files/figure.png" httpUri-1="https://zenodo.org/record/285221/files/figure.png" httpUri-2="https://zenodo.org/record/285222/files/figure.png" pageId="10" pageNumber="43">Figs 57</figureCitation>
).
<emphasis id="FBF4EAECFFB53315FDA9FEB5BE152A33" box="[514,572,332,356]" italics="true" pageId="10" pageNumber="43">Body</emphasis>
<quantity id="0E789B1BFFB53315FDEDFEB5BEF82A33" box="[582,721,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="3.8" metricValueMin="1.9" pageId="10" pageNumber="43" unit="mm" value="2.8499999999999996" valueMax="3.8" valueMin="1.9">1.93.8 mm</quantity>
(mean ± SD; 2.8 ±
<quantity id="0E789B1BFFB53315FC6EFEB5B8082A33" box="[965,1057,332,356]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="43" unit="mm" value="0.4">0.4 mm</quantity>
, n = 51, new materials) without tubercles. Color of fixed specimens white or light brown; digestive organs brown due to congealed host blood.
<emphasis id="FBF4EAECFFB53315FF3CFE6DBD192AFB" box="[151,304,404,428]" italics="true" pageId="10" pageNumber="43">Cephalosome</emphasis>
(
<figureCitation id="51BB2A7BFFB53315FEEAFE6DBDA32AFA" box="[321,394,404,429]" captionStart="FIGURE 5" captionStartId="11.[151,250,1954,1976]" captionTargetBox="[207,1379,193,1932]" captionTargetId="figure@11.[207,1379,193,1933]" captionTargetPageId="11" captionText="FIGURE 5. Gnathia nasuta Nunomura, 1992, (A, D; TOYA Cr- 23570), (B, C, E, F; TOYA Cr- 11172): A, body, dorsal view; B, cephalosome, dorsal view; C, cephalosome, lateral view; D, pleotelson, dorsal view; E, right antennule; F, right antenna." httpUri="https://zenodo.org/record/285220/files/figure.png" pageId="10" pageNumber="43">Figs 5</figureCitation>
AC, 7AD) almost square with posterior margin slightly concave, sparsely covered with setae. Anterior part of cephalosome acutely lowered in thickness at near eyes in lateral view. Frontal border convex with many setae. Mediofrontal process and 2 frontolateral processes visible on central frontal border and apex smooth or slightly dentate. Mediofrontal process extends 2 frontolateral processes. Superior frontolateral processes with several setae. Tips of ventral frontal border often visible under superior frontolateral processes. Protrusions apparently visible inner sides of basis of antennae. Low and pentagonal rising often visible near frontal border. Dorsal sulcus narrow or U-shaped. Oval-shaped translucent region visible anteromedianly on dorsal sulcus. Eyes composed with 44 ocelli. Paraocular ornamentation prominent as one protrusion. Supraocular lobe not prominent with smooth apex.
</paragraph>
</subSubSection>
<subSubSection id="819A6575FFB53315FF6CFD21BF0C2C27" pageId="10" pageNumber="43" type="description">
<paragraph id="C93F36FEFFB53315FF6CFD21BF90280B" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="43">
<emphasis id="FBF4EAECFFB53315FF6CFD21BD6D29A7" box="[199,324,728,752]" italics="true" pageId="10" pageNumber="43">Pereonite I</emphasis>
(
<figureCitation id="51BB2A7BFFB53315FEFFFD2EBDB329A7" box="[340,410,727,752]" captionStart="FIGURE 5" captionStartId="11.[151,250,1954,1976]" captionTargetBox="[207,1379,193,1932]" captionTargetId="figure@11.[207,1379,193,1933]" captionTargetPageId="11" captionText="FIGURE 5. Gnathia nasuta Nunomura, 1992, (A, D; TOYA Cr- 23570), (B, C, E, F; TOYA Cr- 11172): A, body, dorsal view; B, cephalosome, dorsal view; C, cephalosome, lateral view; D, pleotelson, dorsal view; E, right antennule; F, right antenna." httpUri="https://zenodo.org/record/285220/files/figure.png" pageId="10" pageNumber="43">Figs 5</figureCitation>
AC, 7AD) short, not fused or medially fused with cephalosome, reaching lateral margin of cephalosome. Widths and lengths of pereonites II, III, and IV almost same. Pereonites II and III anteriorly concave. Pereonite IV without constriction and anterolateral lobes slightly visible.
<taxonomicName id="0E804D7DFFB53315FBEFFCD9B8AE286F" box="[1092,1159,800,824]" class="Liliopsida" family="Araceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Alismatales" pageId="10" pageNumber="43" phylum="Magnoliophyta" rank="tribe" tribe="Areae">Areae</taxonomicName>
laterales slightly visible on pereonites V. Pereonite VII short and narrow, overlapping pleonite I.
</paragraph>
<paragraph id="C93F36FEFFB53315FF6CFC91B85928F2" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="43">
<emphasis id="FBF4EAECFFB53315FF6CFC91BD4928D7" box="[199,352,872,896]" italics="true" pageId="10" pageNumber="43">Pleonite IIV</emphasis>
(
<figureCitation id="51BB2A7BFFB53315FEDAFC9EBD9C28D7" box="[369,437,871,896]" captionStart="FIGURE 5" captionStartId="11.[151,250,1954,1976]" captionTargetBox="[207,1379,193,1932]" captionTargetId="figure@11.[207,1379,193,1933]" captionTargetPageId="11" captionText="FIGURE 5. Gnathia nasuta Nunomura, 1992, (A, D; TOYA Cr- 23570), (B, C, E, F; TOYA Cr- 11172): A, body, dorsal view; B, cephalosome, dorsal view; C, cephalosome, lateral view; D, pleotelson, dorsal view; E, right antennule; F, right antenna." httpUri="https://zenodo.org/record/285220/files/figure.png" pageId="10" pageNumber="43">Fig. 5</figureCitation>
A) almost equal length and narrower posteriorly, with 1 seta on lateral margin of each segment. Pleonite V longer than each segment of Pleonite IIV. Epimera not prominent.
</paragraph>
<paragraph id="C93F36FEFFB53315FF6CFC49BEE128BB" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="43">
<emphasis id="FBF4EAECFFB53315FF6CFC49BD14289F" box="[199,317,944,968]" italics="true" pageId="10" pageNumber="43">Pleotelson</emphasis>
(
<figureCitation id="51BB2A7BFFB53315FEE0FC56BDB1289F" box="[331,408,943,968]" captionStart="FIGURE 5" captionStartId="11.[151,250,1954,1976]" captionTargetBox="[207,1379,193,1932]" captionTargetId="figure@11.[207,1379,193,1933]" captionTargetPageId="11" captionText="FIGURE 5. Gnathia nasuta Nunomura, 1992, (A, D; TOYA Cr- 23570), (B, C, E, F; TOYA Cr- 11172): A, body, dorsal view; B, cephalosome, dorsal view; C, cephalosome, lateral view; D, pleotelson, dorsal view; E, right antennule; F, right antenna." httpUri="https://zenodo.org/record/285220/files/figure.png" pageId="10" pageNumber="43">Figs. 5</figureCitation>
D, 7EH) almost triangular with acute apex and often covered with scales. Three pair of seta on dorsal surface and 1 pair of seta on distal apex.
</paragraph>
<paragraph id="C93F36FEFFB53315FF6CFC01B9282F0F" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="43">
<emphasis id="FBF4EAECFFB53315FF6CFC01BD1A2F47" box="[199,307,1016,1040]" italics="true" pageId="10" pageNumber="43">Mandible</emphasis>
(
<figureCitation id="51BB2A7BFFB53315FEEFFC0EBDBA2F47" box="[324,403,1015,1040]" captionStart="FIGURE 5" captionStartId="11.[151,250,1954,1976]" captionTargetBox="[207,1379,193,1932]" captionTargetId="figure@11.[207,1379,193,1933]" captionTargetPageId="11" captionText="FIGURE 5. Gnathia nasuta Nunomura, 1992, (A, D; TOYA Cr- 23570), (B, C, E, F; TOYA Cr- 11172): A, body, dorsal view; B, cephalosome, dorsal view; C, cephalosome, lateral view; D, pleotelson, dorsal view; E, right antennule; F, right antenna." httpUri="https://zenodo.org/record/285220/files/figure.png" pageId="10" pageNumber="43">Figs. 5</figureCitation>
AC, 7AD) nipper-like shape, seems as triangular in dorsal view, from half to two-third length of cephalosome. Apex often bends backward. Dentate blade occupies over half-length of mandible. One mandibular seta presents near armed carina on mid-dorsal surface. Basal neck and erisma prominent.
</paragraph>
<paragraph id="C93F36FEFFB53315FF6CFB9DB9222F93" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="43">
<emphasis id="FBF4EAECFFB53315FF6CFB9DBD102F2B" box="[199,313,1124,1148]" italics="true" pageId="10" pageNumber="43">Antennule</emphasis>
(
<figureCitation id="51BB2A7BFFB53315FEE2FB9DBDA52F2A" box="[329,396,1124,1149]" captionStart="FIGURE 5" captionStartId="11.[151,250,1954,1976]" captionTargetBox="[207,1379,193,1932]" captionTargetId="figure@11.[207,1379,193,1933]" captionTargetPageId="11" captionText="FIGURE 5. Gnathia nasuta Nunomura, 1992, (A, D; TOYA Cr- 23570), (B, C, E, F; TOYA Cr- 11172): A, body, dorsal view; B, cephalosome, dorsal view; C, cephalosome, lateral view; D, pleotelson, dorsal view; E, right antennule; F, right antenna." httpUri="https://zenodo.org/record/285220/files/figure.png" pageId="10" pageNumber="43">Fig. 5</figureCitation>
E) composed with 3 peduncle and 5 flagellar articles. Two, 3, and 2 feather-like bristles on distal margins of peduncle article I, II, and III, respectively. One feather-like bristle on flagellar article I. One aesthetasc on flagellar article III and IV, and V, respectively. Flagellar article V terminating in 3 setae.
</paragraph>
<paragraph id="C93F36FEFFB53315FF6CFB29BCFF2E78" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="43">
<emphasis id="FBF4EAECFFB53315FF6CFB29BD0C2FB0" box="[199,293,1232,1255]" italics="true" pageId="10" pageNumber="43">Antenna</emphasis>
(
<figureCitation id="51BB2A7BFFB53315FE92FB36BDA92FBF" box="[313,384,1231,1256]" captionStart="FIGURE 5" captionStartId="11.[151,250,1954,1976]" captionTargetBox="[207,1379,193,1932]" captionTargetId="figure@11.[207,1379,193,1933]" captionTargetPageId="11" captionText="FIGURE 5. Gnathia nasuta Nunomura, 1992, (A, D; TOYA Cr- 23570), (B, C, E, F; TOYA Cr- 11172): A, body, dorsal view; B, cephalosome, dorsal view; C, cephalosome, lateral view; D, pleotelson, dorsal view; E, right antennule; F, right antenna." httpUri="https://zenodo.org/record/285220/files/figure.png" pageId="10" pageNumber="43">Fig. 5</figureCitation>
F) composed with 4 peduncle and 7 flagellar articles. One and 3 feather-like bristles on peduncle III and IV, respectively. 02 setae on distal margins of flagellar article IVI. Article VII terminates in 4 setae.
</paragraph>
<paragraph id="C93F36FEFFB53315FF6CFAC5B8022E2F" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="43">
<emphasis id="FBF4EAECFFB53315FF6CFAC5BD692E03" box="[199,320,1340,1364]" italics="true" pageId="10" pageNumber="43">Maxilliped</emphasis>
(
<figureCitation id="51BB2A7BFFB53315FEFBFAC5BDBB2E02" box="[336,402,1340,1365]" captionStart="FIGURE 6" captionStartId="12.[151,250,1953,1975]" captionTargetBox="[183,1403,193,1931]" captionTargetId="figure@12.[183,1403,193,1932]" captionTargetPageId="12" captionText="FIGURE 6. Gnathia nasuta Nunomura, 1992, (A, B, E; TOYA Cr- 11172), (C, D; TOYA Cr- 23570: A, right maxilliped, ventral view; B right pylopod, ventral view; C, left pereopod II, lateral view; D, left pleopod II; E, penes." httpUri="https://zenodo.org/record/285221/files/figure.png" pageId="10" pageNumber="43">Fig. 6</figureCitation>
A) with endite reaching palp article II. Three, 7, 5, and 7 plumose setae on external margins of palp articles IIV, respectively. Four setae on internal margin of palp article IV.
</paragraph>
<paragraph id="C93F36FEFFB53315FF6CFA7DBEF42EB2" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="43">
<emphasis id="FBF4EAECFFB53315FF6CFA7DBD0D2ECB" box="[199,292,1412,1436]" italics="true" pageId="10" pageNumber="43">Pylopod</emphasis>
(
<figureCitation id="51BB2A7BFFB53315FE9EFA7DBD512ECA" box="[309,376,1412,1437]" captionStart="FIGURE 6" captionStartId="12.[151,250,1953,1975]" captionTargetBox="[183,1403,193,1931]" captionTargetId="figure@12.[183,1403,193,1932]" captionTargetPageId="12" captionText="FIGURE 6. Gnathia nasuta Nunomura, 1992, (A, B, E; TOYA Cr- 11172), (C, D; TOYA Cr- 23570: A, right maxilliped, ventral view; B right pylopod, ventral view; C, left pereopod II, lateral view; D, left pleopod II; E, penes." httpUri="https://zenodo.org/record/285221/files/figure.png" pageId="10" pageNumber="43">Fig. 6</figureCitation>
B) 3-articled. Article I semicircular with posterior corner; 3 areolae visible, 32 plumose setae on internal margin, 12 setae on external margin, 9 setae on distal margin, and fine setae cover ventral surface. Article II elliptical with 10 setae. Article III minute.
</paragraph>
<paragraph id="C93F36FEFFB53315FF6CFA09BFA32D22" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="43">
<emphasis id="FBF4EAECFFB53315FF6CFA09BD662D5F" box="[199,335,1520,1544]" italics="true" pageId="10" pageNumber="43">Pereopod II</emphasis>
(
<figureCitation id="51BB2A7BFFB53315FECAFA16BD8F2D5F" box="[353,422,1519,1544]" captionStart="FIGURE 6" captionStartId="12.[151,250,1953,1975]" captionTargetBox="[183,1403,193,1931]" captionTargetId="figure@12.[183,1403,193,1932]" captionTargetPageId="12" captionText="FIGURE 6. Gnathia nasuta Nunomura, 1992, (A, B, E; TOYA Cr- 11172), (C, D; TOYA Cr- 23570: A, right maxilliped, ventral view; B right pylopod, ventral view; C, left pereopod II, lateral view; D, left pleopod II; E, penes." httpUri="https://zenodo.org/record/285221/files/figure.png" pageId="10" pageNumber="43">Fig. 6</figureCitation>
C) sparsely covered with setae. Basis oblong with 1 feather-like bristle. Ischium shorter than basis, becoming larger distally. Merus about half-length of ischium. Carpus slightly larger than merus with 1 denticulate compound spine on inner margin. Propodus rectangular, bearing 2 denticulate compound spines on inner-mid and inner-distal margins. Dactylus terminating in unguis.
</paragraph>
<paragraph id="C93F36FEFFB53315FF6CF979BE952DB7" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="43">
<emphasis id="FBF4EAECFFB53315FF6CF979BD172DCF" box="[199,318,1664,1688]" italics="true" pageId="10" pageNumber="43">Pleopod II</emphasis>
(
<figureCitation id="51BB2A7BFFB53315FEE6F986BDB92DCF" box="[333,400,1663,1688]" captionStart="FIGURE 6" captionStartId="12.[151,250,1953,1975]" captionTargetBox="[183,1403,193,1931]" captionTargetId="figure@12.[183,1403,193,1932]" captionTargetPageId="12" captionText="FIGURE 6. Gnathia nasuta Nunomura, 1992, (A, B, E; TOYA Cr- 11172), (C, D; TOYA Cr- 23570: A, right maxilliped, ventral view; B right pylopod, ventral view; C, left pereopod II, lateral view; D, left pleopod II; E, penes." httpUri="https://zenodo.org/record/285221/files/figure.png" pageId="10" pageNumber="43">Fig. 6</figureCitation>
D) with 1 seta on outer distal corner and coupling hook on inner margin. Both rami oval and equal in length. Appendix masculina of pleopod II extending half-length of endopod. Eight or 9 plumose setae on exopods. Seven or 8 plumose setae on endopods.
</paragraph>
<paragraph id="C93F36FEFFB53315FF6CF915BFDD2C1A" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="43">
<emphasis id="FBF4EAECFFB53315FF6CF915BD342C53" box="[199,285,1772,1796]" italics="true" pageId="10" pageNumber="43">Uropod</emphasis>
(
<figureCitation id="51BB2A7BFFB53315FE85F915BD552C52" box="[302,380,1772,1797]" captionStart="FIGURE 5" captionStartId="11.[151,250,1954,1976]" captionTargetBox="[207,1379,193,1932]" captionTargetId="figure@11.[207,1379,193,1933]" captionTargetPageId="11" captionText="FIGURE 5. Gnathia nasuta Nunomura, 1992, (A, D; TOYA Cr- 23570), (B, C, E, F; TOYA Cr- 11172): A, body, dorsal view; B, cephalosome, dorsal view; C, cephalosome, lateral view; D, pleotelson, dorsal view; E, right antennule; F, right antenna." httpUri="https://zenodo.org/record/285220/files/figure.png" pageId="10" pageNumber="43">Figs. 5</figureCitation>
D, 7EH,) fringed with fine setae laterally. Both rami subequal in length. Exopod not extend beyond apex of pleotelson and endopod almost reach apex. Exopod with 6 simple and 4 plumose setae and endopod with 3 simple and 6 plumose setae, and several feather-like bristles.
</paragraph>
<paragraph id="C93F36FEFFB53315FF6CF8A0BF0C2C27" blockId="10.[151,1437,151,2013]" box="[199,805,1879,1904]" pageId="10" pageNumber="43">
<emphasis id="FBF4EAECFFB53315FF6CF8A0BD232C27" box="[199,266,1881,1904]" italics="true" pageId="10" pageNumber="43">Penes</emphasis>
(
<figureCitation id="51BB2A7BFFB53315FEB2F8AEBD722C27" box="[281,347,1879,1904]" captionStart="FIGURE 6" captionStartId="12.[151,250,1953,1975]" captionTargetBox="[183,1403,193,1931]" captionTargetId="figure@12.[183,1403,193,1932]" captionTargetPageId="12" captionText="FIGURE 6. Gnathia nasuta Nunomura, 1992, (A, B, E; TOYA Cr- 11172), (C, D; TOYA Cr- 23570: A, right maxilliped, ventral view; B right pylopod, ventral view; C, left pereopod II, lateral view; D, left pleopod II; E, penes." httpUri="https://zenodo.org/record/285221/files/figure.png" pageId="10" pageNumber="43">Fig. 6</figureCitation>
E) composed with 2 contiguous papillae.
</paragraph>
</subSubSection>
<subSubSection id="819A6575FFB53311FF6CF885BFB42E97" lastPageId="14" lastPageNumber="47" pageId="10" pageNumber="43" type="discussion">
<paragraph id="C93F36FEFFB53311FF6CF885BF572A4A" blockId="10.[151,1437,151,2013]" lastBlockId="14.[151,1437,151,1473]" lastPageId="14" lastPageNumber="47" pageId="10" pageNumber="43">
<emphasis id="FBF4EAECFFB53315FF6CF885BD122CC2" bold="true" box="[199,315,1916,1941]" pageId="10" pageNumber="43">Remarks.</emphasis>
<taxonomicName id="0E804D7DFFB53315FEE7F885BE6C2CC3" box="[332,581,1916,1940]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="FBF4EAECFFB53315FEE7F885BE6C2CC3" box="[332,581,1916,1940]" italics="true" pageId="10" pageNumber="43">Caecognathia nasuta</emphasis>
</taxonomicName>
and
<taxonomicName id="0E804D7DFFB53315FD24F885BF462CC3" box="[655,879,1916,1940]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="hirayamai">
<emphasis id="FBF4EAECFFB53315FD24F885BF462CC3" box="[655,879,1916,1940]" italics="true" pageId="10" pageNumber="43">Gnathia hirayamai</emphasis>
</taxonomicName>
were described by Nunomura 1992, and
<taxonomicName id="0E804D7DFFB53315FA2FF885BD1D2CEF" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="nagasakiensis">
<emphasis id="FBF4EAECFFB53315FA2FF885BD1D2CEF" italics="true" pageId="10" pageNumber="43">G. nagasakiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="0E804D7DFFB53315FEEFF858BDC82CEF" box="[324,481,1952,1976]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="shijikiensis">
<emphasis id="FBF4EAECFFB53315FEEFF858BDC82CEF" box="[324,481,1952,1976]" italics="true" pageId="10" pageNumber="43">G. shijikiensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="0E804D7DFFB53315FD89F858BEFA2CEF" box="[546,723,1952,1976]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="brevicephala">
<emphasis id="FBF4EAECFFB53315FD89F858BEFA2CEF" box="[546,723,1952,1976]" italics="true" pageId="10" pageNumber="43">G. brevicephala</emphasis>
</taxonomicName>
were described by Nunomura, 2012a. A redescription of their five
<typeStatus id="163B885CFFB53315FF7BF83DBD142C8A" box="[208,317,1988,2013]" pageId="10" pageNumber="43" type="holotype">holotypes</typeStatus>
revealed that they shared the following characteristics: nipper-shaped mandibles; dentate blade occupying more than half the length of the mandible (apparently triangular in shape in the dorsal view); acutely lowered anterior part of the cephalosome; convex frontal border with several setae; mediofrontal process extending into two frontolateral processes; superior frontolateral processes with several setae; prominent paraocular ornamentation as one protrusion, and acute apex of the pleotelson.
</paragraph>
<caption id="9DFF6676FFB43314FF3CF85BB95E2C80" httpUri="https://zenodo.org/record/285220/files/figure.png" pageId="11" pageNumber="44" targetBox="[207,1379,193,1932]" targetPageId="11">
<paragraph id="C93F36FEFFB43314FF3CF85BB95E2C80" blockId="11.[151,1436,1954,2007]" pageId="11" pageNumber="44">
<emphasis id="FBF4EAECFFB43314FF3CF85BBD3D2CEF" bold="true" box="[151,276,1954,1976]" pageId="11" pageNumber="44">FIGURE 5.</emphasis>
<taxonomicName id="0E804D7DFFB43314FEB0F85BBE472CEF" authority="Nunomura, 1992" authorityName="Nunomura" authorityYear="1992" box="[283,622,1954,1976]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="FBF4EAECFFB43314FEB0F85BBD912CEF" box="[283,440,1954,1976]" italics="true" pageId="11" pageNumber="44">Gnathia nasuta</emphasis>
Nunomura, 1992
</taxonomicName>
, (A, D; TOYA Cr-23570), (B, C, E, F; TOYA Cr-11172): A, body, dorsal view; B, cephalosome, dorsal view; C, cephalosome, lateral view; D, pleotelson, dorsal view; E, right antennule; F, right antenna.
</paragraph>
</caption>
<caption id="9DFF6676FFB33313FF3CF858B85A2C81" httpUri="https://zenodo.org/record/285221/files/figure.png" pageId="12" pageNumber="45" targetBox="[183,1403,193,1931]" targetPageId="12">
<paragraph id="C93F36FEFFB33313FF3CF858B85A2C81" blockId="12.[151,1435,1953,2006]" pageId="12" pageNumber="45">
<emphasis id="FBF4EAECFFB33313FF3CF858BD3B2CE0" bold="true" box="[151,274,1953,1975]" pageId="12" pageNumber="45">FIGURE 6.</emphasis>
<taxonomicName id="0E804D7DFFB33313FEB3F858BE412CE0" authority="Nunomura, 1992" authorityName="Nunomura" authorityYear="1992" box="[280,616,1953,1975]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="FBF4EAECFFB33313FEB3F858BD9C2CE0" box="[280,437,1953,1975]" italics="true" pageId="12" pageNumber="45">Gnathia nasuta</emphasis>
Nunomura, 1992
</taxonomicName>
, (A, B, E; TOYA Cr-11172), (C, D; TOYA Cr-23570: A, right maxilliped, ventral view; B right pylopod, ventral view; C, left pereopod II, lateral view; D, left pleopod II; E, penes.
</paragraph>
</caption>
<caption id="9DFF6676FFB23312FF3CF85AB8282CA1" httpUri="https://zenodo.org/record/285222/files/figure.png" pageId="13" pageNumber="46" targetBox="[329,1261,193,1933]" targetPageId="13">
<paragraph id="C93F36FEFFB23312FF3CF85AB8282CA1" blockId="13.[151,1436,1955,2038]" pageId="13" pageNumber="46">
<emphasis id="FBF4EAECFFB23312FF3CF85ABD3A2CEE" bold="true" box="[151,275,1955,1977]" pageId="13" pageNumber="46">FIGURE 7.</emphasis>
Cephalosomes (AD) and pleotelsons (EH) of four holotypes: A and E,
<taxonomicName id="0E804D7DFFB23312FC56F85AB8972CEE" box="[1021,1214,1955,1977]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="hirayamai">
<emphasis id="FBF4EAECFFB23312FC56F85AB8972CEE" box="[1021,1214,1955,1977]" italics="true" pageId="13" pageNumber="46">Gnathia hirayamai</emphasis>
</taxonomicName>
(TOYA Cr-11167); B and F,
<taxonomicName id="0E804D7DFFB23312FF4DF838BDFD2C80" box="[230,468,1985,2007]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="nagasakiensis">
<emphasis id="FBF4EAECFFB23312FF4DF838BDFD2C80" box="[230,468,1985,2007]" italics="true" pageId="13" pageNumber="46">Gnathia nagasakiensis</emphasis>
</taxonomicName>
(TOYA Cr-23352); C and G,
<taxonomicName id="0E804D7DFFB23312FC8FF838BFDD2C80" box="[804,1012,1985,2007]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="shijikiensis">
<emphasis id="FBF4EAECFFB23312FC8FF838BFDD2C80" box="[804,1012,1985,2007]" italics="true" pageId="13" pageNumber="46">Gnathia shijikiensis</emphasis>
</taxonomicName>
(TOYA Cr-23353); D and H,
<taxonomicName id="0E804D7DFFB23312FAE2F838BD332CA1" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="brevicephala">
<emphasis id="FBF4EAECFFB23312FAE2F838BD332CA1" italics="true" pageId="13" pageNumber="46">Gnathia brevicephala</emphasis>
</taxonomicName>
(TOYA Cr-23356). The cephalosome of
<taxonomicName id="0E804D7DFFB23312FD10F818BF4C2CA1" box="[699,869,2016,2038]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="nagasakiensis">
<emphasis id="FBF4EAECFFB23312FD10F818BF4C2CA1" box="[699,869,2016,2038]" italics="true" pageId="13" pageNumber="46">G. nagasakiensis</emphasis>
</taxonomicName>
was deformed.
</paragraph>
</caption>
<paragraph id="C93F36FEFFB13311FF6CFEDEBDA4296B" blockId="14.[151,1437,151,1473]" pageId="14" pageNumber="47">
The following are the differences between the drawings by Nunomura (1992) and the present study.
<taxonomicName id="0E804D7DFFB13311FF3CFEB5BD9C2A33" box="[151,437,332,356]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="hirayamai">
<emphasis id="FBF4EAECFFB13311FF3CFEB5BD9C2A33" box="[151,437,332,356]" italics="true" pageId="14" pageNumber="47">Caecognathia hirayamai</emphasis>
</taxonomicName>
: (1) “a conspicuous incisor” was mentioned in the original description but it was inconspicuous in the present study; (2) the frontal border was almost straight without processes but there was actually a convex frontal border with several setae and three frontal processes; (3) the number of flagellar articles of the antenna was mentioned as nine but it was actually seven; (4) article III of the pylopod was not drawn.
<taxonomicName id="0E804D7DFFB13311FF3CFE25BDA02AA3" box="[151,393,476,500]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="FBF4EAECFFB13311FF3CFE25BDA02AA3" box="[151,393,476,500]" italics="true" pageId="14" pageNumber="47">Caecognathia nasuta</emphasis>
</taxonomicName>
: (1) a “big pentagonal frontal border” was drawn but there was actually a convex frontal border. Furthermore, Nunomura (1992) did not describe pereonite I and the endite of the maxilliped, although they were actually present.
</paragraph>
<paragraph id="C93F36FEFFB13311FF6CFDBEB8CC2828" blockId="14.[151,1437,151,1473]" pageId="14" pageNumber="47">
The following are the differences between the drawings by Nunomura (2012a) and the present study.
<taxonomicName id="0E804D7DFFB13311FAEBFDB1BD1D29D3" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="nagasakiensis">
<emphasis id="FBF4EAECFFB13311FAEBFDB1BD1D29D3" italics="true" pageId="14" pageNumber="47">Gnathia nagasakiensis</emphasis>
</taxonomicName>
: (1) the frontal process and setae were not clearly shown; (2) “small rings” were described, which were actually absent.
<taxonomicName id="0E804D7DFFB13311FE21FD69BE4029FF" box="[394,617,656,680]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="shijikiensis">
<emphasis id="FBF4EAECFFB13311FE21FD69BE4029FF" box="[394,617,656,680]" italics="true" pageId="14" pageNumber="47">Gnathia shijikiensis</emphasis>
</taxonomicName>
: (1) pereonite I was not drawn.
<taxonomicName id="0E804D7DFFB13311FC78FD69B8EE29FF" box="[979,1223,656,680]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="brevicephala">
<emphasis id="FBF4EAECFFB13311FC78FD69B8EE29FF" box="[979,1223,656,680]" italics="true" pageId="14" pageNumber="47">Gnathia brevicephala</emphasis>
</taxonomicName>
: (1) the number of flagellar articles of the antennule and antenna were mentioned as five and six, but there were actually five and seven, respectively; (2) article I was described as oval-shaped but it was actually semicircular. Furthermore, Nunomura (2012a) did not describe the endite of the maxilliped, although it was actually present. Prominent paraocular ornamentation was not observed in the original descriptions of these five species. Considering the similarities between
<taxonomicName id="0E804D7DFFB13311FE3EFCBCBE31280B" box="[405,536,836,860]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="kikuchii">
<emphasis id="FBF4EAECFFB13311FE3EFCBCBE31280B" box="[405,536,836,860]" italics="true" pageId="14" pageNumber="47">E. kikuchii</emphasis>
</taxonomicName>
and its synonyms, the present study determined that
<taxonomicName id="0E804D7DFFB13311FB65FCBDB942280B" box="[1230,1387,836,860]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="hirayamai">
<emphasis id="FBF4EAECFFB13311FB65FCBDB942280B" box="[1230,1387,836,860]" italics="true" pageId="14" pageNumber="47">G. hirayamai</emphasis>
</taxonomicName>
,
<taxonomicName id="0E804D7DFFB13311FA2FFCBDBD1D28D7" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="nagasakiensis">
<emphasis id="FBF4EAECFFB13311FA2FFCBDBD1D28D7" italics="true" pageId="14" pageNumber="47">G. nagasakiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="0E804D7DFFB13311FEE9FC91BDF428D7" box="[322,477,872,896]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="shijikiensis">
<emphasis id="FBF4EAECFFB13311FEE9FC91BDF428D7" box="[322,477,872,896]" italics="true" pageId="14" pageNumber="47">G. shijikiensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="0E804D7DFFB13311FDB1FC91BEE328D7" box="[538,714,872,896]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="brevicephala">
<emphasis id="FBF4EAECFFB13311FDB1FC91BEE328D7" box="[538,714,872,896]" italics="true" pageId="14" pageNumber="47">G. brevicephala</emphasis>
</taxonomicName>
were junior synonyms of
<taxonomicName id="0E804D7DFFB13311FC44FC91B8F728D7" box="[1007,1246,872,896]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="FBF4EAECFFB13311FC44FC91B8F728D7" box="[1007,1246,872,896]" italics="true" pageId="14" pageNumber="47">Caecognathia nasuta</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C93F36FEFFB13311FF6CFC75B9252F2B" blockId="14.[151,1437,151,1473]" pageId="14" pageNumber="47">
The present drawing of the
<taxonomicName id="0E804D7DFFB13311FDAAFC75BE9728F3" box="[513,702,908,932]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="nagasakiensis">
<emphasis id="FBF4EAECFFB13311FDAAFC75BE9728F3" box="[513,702,908,932]" italics="true" pageId="14" pageNumber="47">G. nagasakiensis</emphasis>
</taxonomicName>
<typeStatus id="163B885CFFB13311FD6DFC75BF0128F2" box="[710,808,908,933]" pageId="14" pageNumber="47" type="holotype">holotype</typeStatus>
(
<figureCitation id="51BB2A7BFFB13311FC92FC75BF5528F3" box="[825,892,908,933]" captionStart="FIGURE 7" captionStartId="13.[151,250,1955,1977]" captionTargetBox="[329,1261,193,1933]" captionTargetId="figure@13.[324,1262,193,1933]" captionTargetPageId="13" captionText="FIGURE 7. Cephalosomes (A D) and pleotelsons (E H) of four holotypes: A and E, Gnathia hirayamai (TOYA Cr- 11167); B and F, Gnathia nagasakiensis (TOYA Cr- 23352); C and G, Gnathia shijikiensis (TOYA Cr- 23353); D and H, Gnathia brevicephala (TOYA Cr- 23356). The cephalosome of G. nagasakiensis was deformed." httpUri="https://zenodo.org/record/285222/files/figure.png" pageId="14" pageNumber="47">Fig. 7</figureCitation>
B) appears to be different from other specimens with respect to the mandible shapes and eyes, the basal neck and armed carina are not distinct, the tip of the mandible is not curved inward, and the eyes are presented laterally. This is because the mandibles are closed and the basal neck is hidden behind the frontal border, while the armed carina is visible laterally. The cephalosome of this specimen was deformed during the previous dissection and the location of the right eye moved laterally (the left eye was located dorsally). This specimen was preserved with other male materials (non
<typeStatus id="163B885CFFB13311FB46FBB9B9362F0F" box="[1261,1311,1088,1112]" pageId="14" pageNumber="47">type</typeStatus>
) and their morphologies were similar. Thus,
<taxonomicName id="0E804D7DFFB13311FDBEFB9DBEF82F2B" box="[533,721,1124,1148]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="nagasakiensis">
<emphasis id="FBF4EAECFFB13311FDBEFB9DBEF82F2B" box="[533,721,1124,1148]" italics="true" pageId="14" pageNumber="47">G. nagasakiensis</emphasis>
</taxonomicName>
is considered to be a junior synonym of
<taxonomicName id="0E804D7DFFB13311FB31FB9DB92C2F2B" box="[1178,1285,1124,1148]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="FBF4EAECFFB13311FB31FB9DB92C2F2B" box="[1178,1285,1124,1148]" italics="true" pageId="14" pageNumber="47">C. nasuta</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C93F36FEFFB13311FF6CFB7EB86B2FBF" blockId="14.[151,1437,151,1473]" pageId="14" pageNumber="47">
The genus of the present species was considered to be
<taxonomicName id="0E804D7DFFB13311FC9BFB71BFE42FF7" box="[816,973,1160,1184]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="FBF4EAECFFB13311FC9BFB71BFE42FF7" box="[816,973,1160,1184]" italics="true" pageId="14" pageNumber="47">Caecognathia</emphasis>
</taxonomicName>
by Cohen &amp; Poore (1994) but it belongs to the genus
<taxonomicName id="0E804D7DFFB13311FE9CFB55BDBA2F93" box="[311,403,1196,1220]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="FBF4EAECFFB13311FE9CFB55BDBA2F93" box="[311,403,1196,1220]" italics="true" pageId="14" pageNumber="47">Gnathia</emphasis>
</taxonomicName>
because of the following features: the frontal border is not excavated and has frontal processes; paraocular ornamentation; a dorsal sulcus is present on the cephalosome.
</paragraph>
<paragraph id="C93F36FEFFB13311FF6CFB0DBFB42E97" blockId="14.[151,1437,151,1473]" pageId="14" pageNumber="47">
<taxonomicName id="0E804D7DFFB13311FF6CFB0DBD5C2E5B" box="[199,373,1268,1292]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="FBF4EAECFFB13311FF6CFB0DBD5C2E5B" box="[199,373,1268,1292]" italics="true" pageId="14" pageNumber="47">Gnathia nasuta</emphasis>
</taxonomicName>
was closely related to
<taxonomicName id="0E804D7DFFB13311FDDDFB0DBFE72E5A" authority="Monod, 1926" authorityName="Monod" authorityYear="1926" box="[630,974,1268,1293]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="FBF4EAECFFB13311FDDDFB0DBF342E5B" box="[630,797,1268,1292]" italics="true" pageId="14" pageNumber="47">C. nipponensis</emphasis>
(Monod, 1926)
</taxonomicName>
in terms of the following characteristics: nipper-shaped mandible, prominent paraocular ornamentation as one protrusion, acute apex of the pleotelson, and appendix masculina reaching half the length of the endopod of pleopod II.
<taxonomicName id="0E804D7DFFB13311FBA2FAC5B91E2E03" box="[1033,1335,1340,1364]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="FBF4EAECFFB13311FBA2FAC5B91E2E03" box="[1033,1335,1340,1364]" italics="true" pageId="14" pageNumber="47">Caecognathia nipponensis</emphasis>
</taxonomicName>
was not regarded as
<taxonomicName id="0E804D7DFFB13311FE88FA99BD562E2F" box="[291,383,1376,1400]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="FBF4EAECFFB13311FE88FA99BD562E2F" box="[291,383,1376,1400]" italics="true" pageId="14" pageNumber="47">Gnathia</emphasis>
</taxonomicName>
by Cohen &amp; Poore (1994). However, the original description may have failed to reveal the frontal processes on the frontal border (Fig.
<quantity id="0E789B1BFFB13311FD28FA7DBEE32ECA" box="[643,714,1412,1437]" metricMagnitude="0" metricUnit="m" metricValue="3.7845999999999997" pageId="14" pageNumber="47" unit="in" value="149.0">149 in</quantity>
Monod 1926). More detailed observations of the
<taxonomicName id="0E804D7DFFB13311FB5DFA7DB9B52ECB" box="[1270,1436,1412,1436]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="47" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="FBF4EAECFFB13311FB5DFA7DB9B52ECB" box="[1270,1436,1412,1436]" italics="true" pageId="14" pageNumber="47">C. nipponensis</emphasis>
</taxonomicName>
<typeStatus id="163B885CFFB13311FF3CFA5EBCD02E97" box="[151,249,1447,1472]" pageId="14" pageNumber="47" type="holotype">holotype</typeStatus>
and morphological comparisons will be necessary in future.
</paragraph>
</subSubSection>
</treatment>
</document>