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<document id="7A2F972DAA34A2487A8F7AE4855BAFE2" ID-CLB-Dataset="35993" ID-DOI="http://dx.doi.org/10.3897/zookeys.339.6033" ID-GBIF-Dataset="80562a28-2e9e-47c5-9bf7-dc54f9ccecb1" ID-PMC="PMC3800829" ID-Pensoft-Pub="1313-2970-339-67" ID-PubMed="24146587" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-339-67" ModsDocOrigin="ZooKeys 339" ModsDocTitle="Revision of the Australian Ceratocanthinae (Coleoptera, Scarabaeoidea, Hybosoridae)" checkinTime="1451246837575" checkinUser="pensoft" docAuthor="Ballerio, Alberto" docDate="2013" docId="FE83EC4C0349A97E5E21606814DD61AF" docLanguage="en" docName="ZooKeys 339: 67-91" docOrigin="ZooKeys 339" docSource="http://dx.doi.org/10.3897/zookeys.339.6033" docTitle="Pterorthochaetes cribricollis Gestro 1899" docType="treatment" docVersion="5" lastPageNumber="80" masterDocId="FF955E61B74EFF9F585E376FFF8DB824" masterDocTitle="Revision of the Australian Ceratocanthinae (Coleoptera, Scarabaeoidea, Hybosoridae)" masterLastPageNumber="91" masterPageNumber="67" pageNumber="78" updateTime="1732752689784" updateUser="ExternalLinkService">
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<mods:title id="4B3CED42A244D4A6B29EB005230A2EE4">Revision of the Australian Ceratocanthinae (Coleoptera, Scarabaeoidea, Hybosoridae)</mods:title>
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<mods:namePart id="46FB6562B83E5EB99DA64A355313ED34">Ballerio, Alberto</mods:namePart>
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<mods:classification id="FEE642917A893A0747BE9C9DB5FFE6E0">journal article</mods:classification>
<mods:identifier id="890422DB76FE785669F6CEFD663F336A" type="DOI">http://dx.doi.org/10.3897/zookeys.339.6033</mods:identifier>
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<treatment id="FE83EC4C0349A97E5E21606814DD61AF" ID-GBIF-Taxon="152048600" LSID="urn:lsid:plazi:treatment:FE83EC4C0349A97E5E21606814DD61AF" httpUri="http://treatment.plazi.org/id/FE83EC4C0349A97E5E21606814DD61AF" lastPageId="13" lastPageNumber="80" pageId="11" pageNumber="78" scope_family="Hybosoridae" scope_order="Coleoptera" scope_subFamily="Ceratocanthinae" scope_superFamily="Scarabaeoidea">
<subSubSection id="B7340E3B10FAC9D174CBD0E9ADAA323C" pageId="11" pageNumber="78" type="nomenclature">
<paragraph id="361034E29B9245B74EF06BE61451D9FB" pageId="11" pageNumber="78">
<taxonomicName id="30CD6A8C8F25E6388FB7194C2D326302" ID-CoL="4Q35L" LSID="http://species-id.net/wiki/Pterorthochaetes_cribricollis" authority="Gestro, 1899" authorityName="Gestro" authorityYear="1899" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterorthochaetes cribricollis" order="Coleoptera" pageId="11" pageNumber="78" phylum="Arthropoda" rank="species" species="cribricollis">Pterorthochaetes cribricollis Gestro, 1899</taxonomicName>
Figs 6, 7
<normalizedToken id="0484CE3966B01F36535F8C30EEA012B6" originalValue="AD">A-D</normalizedToken>
, 11D, 12A, 13
<normalizedToken id="F62B3CA4316727E61B77B1E22A8D8E79" originalValue="AC">A-C</normalizedToken>
, 14C
</paragraph>
</subSubSection>
<subSubSection id="78580AF054315D848AD5ED3C57F79139" pageId="11" pageNumber="78" type="reference_group">
<paragraph id="1729C6AB81DAE940C559EDF07C0E842E" pageId="11" pageNumber="78">
<taxonomicName id="5D4075574AAFE2ECCA149AF7ADB26C14" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterorthochaetes cribricollis" order="Coleoptera" pageId="11" pageNumber="78" phylum="Arthropoda" rank="species" species="cribricollis">Pterorthochaetes cribricollis</taxonomicName>
Gestro, 1899: 37 (description, distribution, key);
<bibRefCitation id="E1901141E63CAE2D4D0EB04798455CC1" author="Paulian, R" journalOrPublisher="Annales de la Societe Entomologique de France (N. S.)" pageId="23" pageNumber="90" pagination="479 - 514" title="Revision des Ceratocanthidae [Col. Scarabaeoidea] II - Les especes orientales et australiennes." volume="14" year="1978">Paulian 1978</bibRefCitation>
(key, distribution);
<bibRefCitation id="B225114FF4BEC98A9F886F9AF390C830" author="Cassis, G" editor="Houston, WWK" journalOrPublisher="Canberra, AGPS" pageId="23" pageNumber="90" pagination="65 - 66" title="Ceratocanthidae." volume="9" volumeTitle="Zoological catalogue of Australia. Coleoptera: Scarabaeoidea." year="1992">Cassis and Weir 1992</bibRefCitation>
(catalogue);
<bibRefCitation id="D237EA6D282F80551E4DEDB2A6A6AEDB" author="Ocampo, FC" journalOrPublisher="Bulletin of the University of Nebraska State Museum" pageId="23" pageNumber="90" pagination="178 - 209" title="Phylogenetic analysis of the scarab family Hybosoridae and monographic revision of the New World subfamily Anaidinae (Coleoptera: Scarabaeoidea). 4. Catalog of the subfamilies Anaidinae, Ceratocanthinae, Hybosorinae, Liparochrinae, and Pachyplectrinae (Scarabaeoidea: Hybosoridae)." volume="19" year="2006">Ocampo and Ballerio 2006</bibRefCitation>
(checklist)
</paragraph>
</subSubSection>
<subSubSection id="A5F117DF823F3CB510346CE5AA3DDEA8" pageId="11" pageNumber="78" type="material examined">
<paragraph id="20EF07554C49A6C069D8C2076F7B0D74" pageId="11" pageNumber="78">Material examined.</paragraph>
<paragraph id="CC6F85E32E36FDA92B7D00A6D030DC23" pageId="11" pageNumber="78">
10 specimens [two males and two females dissected]: 4 males and 2 females, Iron Range, Cape York Pen., N. Qld. 28 Apr.-5 May 1968. G. Monteith (QM); 1 female, Iron Range, Cape York Pen., N. Qld. 11-17 May 1968. G. Monteith (QM); 1 female, Iron Range, Cape York Pen., N. Qld. 26
<normalizedToken id="8C913FB7147174C8BAE99B963EF4D367" originalValue="May">May-</normalizedToken>
2 June 1971 B. K. Cantrell (QM); 1 female, QLD:
<geoCoordinate id="7EEE6A9874F95542A71BAE813AD29F24" direction="south" orientation="latitude" precision="55" value="-12.71">12.710°S</geoCoordinate>
,
<geoCoordinate id="AB711BA01FE05D637115F9364155D263" direction="east" orientation="longitude" precision="55" value="143.291">143.291°E</geoCoordinate>
, Cooks Hut, Iron Range, 5 m, 15 Dec 2010, Monteith, Escalona &amp; Will, hand and at HV light 34817 (QM).
</paragraph>
</subSubSection>
<subSubSection id="B91B7CE27C3BB807752DE6D9A2538A92" lastPageId="12" lastPageNumber="79" pageId="11" pageNumber="78" type="description">
<paragraph id="6070506C345A2120D83767CCED57DB40" pageId="11" pageNumber="78">Description.</paragraph>
<paragraph id="C6E518EB72E01B481C8220BDF9AB5FE9" pageId="11" pageNumber="78">Size: HL = 0.90 mm; HW = 1.30 mm; PL = 1.32 mm; PW = 2.20 mm; EL = 2.25 mm; EW = 2.15 mm. Overall morphology as in generic description. Dark brown, shiny, setation yellowish, sternum, tarsi and antennae reddish-brown.</paragraph>
<paragraph id="7A4EED6D694F51A9108964C369165948" lastPageId="12" lastPageNumber="79" pageId="11" pageNumber="78">
Head: completely and uniformly covered by impressed comma-shaped punctures with posterior openings, spaced out by a distance of about half their diameter. Anterior
<pageBreakToken id="8D023B6B531A393F803480F2C60871D7" pageId="12" pageNumber="79" start="start">portion</pageBreakToken>
of clypeus with one or two irregular transverse anastomosing lines. Interocular distance about 9 times the maximum width of dorsal ocular area.
</paragraph>
<paragraph id="CDFDE5847139EC94963307E3C20D88EE" pageId="12" pageNumber="79">Pronotum: margins completely bordered, lateral margins with a row of erect thick yellowish slightly clavate setae, about as long as the distance between them. Pronotal setation made of thick medium sized clavate yellowish setae, punctation as follows: disc covered by impressed short transverse comma-shaped punctures, with posterior openings and containing a small fine setigerous pore, sides with a few larger more curved comma-shaped punctures opening backwards.</paragraph>
<paragraph id="80AECB0E4CAEA98B7F824C8F25A8CE81" pageId="12" pageNumber="79">Scutellum: basally with two longitudinal irregular rows of horseshoe-shaped punctures, uniting towards apex.</paragraph>
<paragraph id="E97D957AE4FD9E47462D897BB6A57C67" pageId="12" pageNumber="79">Elytra: humeral callus poorly pronounced, sutural stria occupying medial and distal third. Elytral punctation as follows: uniformly covered by irregular longitudinal rows of mixed simple impressed small punctures and medium-sized horseshoe-shaped punctures opening backwards, larger on sides and apical third. Interpunctural distance subequal to puncturelength.</paragraph>
<paragraph id="43FFE890AF9BF218183C713F99E5BA3D" pageId="12" pageNumber="79">
Aedeagus: basal piece about three times as long as parameres. Parameres slightly asymmetrical, internal sac distally with some irregular weak sclerotisations (Fig. 12A, Fig. 13
<normalizedToken id="0745797C529DE3E6922769BBCF679917" originalValue="AC">A-C</normalizedToken>
).
</paragraph>
<paragraph id="F7E852788D4C378F6FE0255AF59678D8" pageId="12" pageNumber="79">Male genital segment: as in Fig. 11D.</paragraph>
<paragraph id="21A8989D1A7FB493D1C9D05CD8E0524D" pageId="12" pageNumber="79">Bursal sclerites: slightly asymmetrical, as in Fig. 14C.</paragraph>
<caption id="C3BE3E63FF902D9A16B76067AE10BB8D" pageId="12" pageNumber="79">
<paragraph id="35C562AC260920EA75381F03BC261495" pageId="12" pageNumber="79">
Figure 7.
<taxonomicName id="A62C5138F7D49EF2C0A323407CE96431" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterorthochaetes cribricollis" order="Coleoptera" pageId="12" pageNumber="79" phylum="Arthropoda" rank="species" species="cribricollis">Pterorthochaetes cribricollis</taxonomicName>
Gestro, 1899 A Extended specimen, dorsal view B enrolled specimen, ventral view C enrolled specimen, dorsal view D enrolled specimen, lateral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C3A764D40D5C2CBF9242DB36CC60CC57" lastPageId="13" lastPageNumber="80" pageId="12" pageNumber="79" type="diagnosis">
<paragraph id="E3571760EFAD5D7451AB29CF36C8040E" pageId="12" pageNumber="79">Diagnosis.</paragraph>
<paragraph id="389C5CA8D37C0FE9515AD5F0E9F963F0" lastPageId="13" lastPageNumber="80" pageId="12" pageNumber="79">
Easily distinguished from all other Australian
<taxonomicName id="F3A7A510722A7D4D747047E70F47523F" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterorthochaetes" order="Coleoptera" pageId="13" pageNumber="80" phylum="Arthropoda" rank="genus">
<pageBreakToken id="5BB9251465B985E912B5D24192C91B8C" pageId="13" pageNumber="80" start="start">Pterorthochaetes</pageBreakToken>
</taxonomicName>
because of the combination of smaller size, the pattern of punctation of pronotum and elytra, which on pronotum is only made of short impressed transverse comma-shaped punctures, whereas all other Australian species have, at least partly, horse-shoe shaped punctures, often with a very small posterior openings, while on elytra is made of a much smaller punctation compared to
<taxonomicName id="3A123E9FACB6E8F10FBD02DD9D2584F3" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterorthochaetes danielsi" order="Coleoptera" pageId="13" pageNumber="80" phylum="Arthropoda" rank="species" species="danielsi">Pterorthochaetes danielsi</taxonomicName>
sp. n. and much denser compared to
<taxonomicName id="2AA429333B1654685A53B96844C0A4E7" genus="Pterorthochaeres" lsidName="Pterorthochaeres storeyi" pageId="13" pageNumber="80" rank="species" species="storeyi">Pterorthochaeres storeyi</taxonomicName>
sp. n. and
<taxonomicName id="A76335E4BD077A1A490B00213B6ECF9E" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterorthochaetes simplex" order="Coleoptera" pageId="13" pageNumber="80" phylum="Arthropoda" rank="species" species="simplex">Pterorthochaetes simplex</taxonomicName>
Gestro, 1899.
</paragraph>
</subSubSection>
<subSubSection id="E3604C2BF0B696AC890CE3F07D8FA400" pageId="13" pageNumber="80" type="etymology">
<paragraph id="CA857C8A3C0717BB940FC8DA47A2863A" pageId="13" pageNumber="80">Etymology.</paragraph>
<paragraph id="BB14C60663444E16E295F7C2D75114D8" pageId="13" pageNumber="80">From Latin cribratus (profusely perforated) and collis (pronotum), due to the dense and impressed punctation.</paragraph>
</subSubSection>
<subSubSection id="CDCE6B7285BD59024F456280BCE4657E" pageId="13" pageNumber="80" type="distribution">
<paragraph id="C1D7758EC55EB79C608FDB49C237162A" pageId="13" pageNumber="80">Distribution and habitat.</paragraph>
<paragraph id="E1483F043E372D4B6D743C73AC1D8017" pageId="13" pageNumber="80">In Australia known from the lowland rainforests of Iron Range (Northern Queensland). This species occurs also in New Guinea (type locality: Papua New Guinea, Central Province, lower Kemp Welch River, Ighibirei).</paragraph>
</subSubSection>
<subSubSection id="737A272532FEE641D28E3D845B339926" pageId="13" pageNumber="80" type="remaks">
<paragraph id="91FEC8909378091CAC40F299112E50E6" pageId="13" pageNumber="80">Remaks.</paragraph>
<paragraph id="7551509E41E6228FCB253324DF9D7629" pageId="13" pageNumber="80">identification was made by comparison with the holotype from New Guinea in Museo Civico di Storia Naturale &quot;G. Doria&quot;, Genova.</paragraph>
</subSubSection>
</treatment>
</document>