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122 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.619.9951" ID-GBIF-Dataset="dedaf101-3b90-4f1e-adef-5f7ddd95ad4e" ID-PMC="PMC5090162" ID-Pensoft-Pub="1313-2970-619-45" ID-PubMed="27829789" ID-ZBK="80E9A40681954C1DBB3C0AF732CD478C" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-619-45" ModsDocOrigin="ZooKeys 619" ModsDocTitle="Taxonomic revision of New Guinea diving beetles of the Exocelinadanae group, with the description of ten new species (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1475024545068" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sagata, Katayo & Balke, Michael" docDate="2016" docId="A5B314DC8D9C36516D2D295E5D7F8A68" docLanguage="en" docName="ZooKeys 619: 45-102" docOrigin="ZooKeys 619" docSource="http://dx.doi.org/10.3897/zookeys.619.9951" docTitle="Exocelina varirata Shaverdo & Balke, sp. n." docType="treatment" docUuid="610FF2E1-A763-4AF1-AABA-FEA533636427" docUuidSource="ZooBank" docVersion="5" lastPageNumber="72" masterDocId="FFE3FFD6FFD3FFBB044E305DFFEA8F13" masterDocTitle="Taxonomic revision of New Guinea diving beetles of the Exocelinadanae group, with the description of ten new species (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="102" masterPageNumber="45" pageNumber="72" updateTime="1668163550721" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of New Guinea diving beetles of the Exocelinadanae group, with the description of ten new species (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaverdo, Helena</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sagata, Katayo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balke, Michael</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="volume">
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<mods:number>619</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>45</mods:start>
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<mods:end>102</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.619.9951</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.619.9951</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-619-45</mods:identifier>
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<mods:identifier type="ZBK">80E9A40681954C1DBB3C0AF732CD478C</mods:identifier>
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<mods:identifier type="ZooBank">80E9A40681954C1DBB3C0AF732CD478C</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127890994" LSID="urn:lsid:zoobank.org:act:610FF2E1-A763-4AF1-AABA-FEA533636427" httpUri="http://treatment.plazi.org/id/A5B314DC8D9C36516D2D295E5D7F8A68" lastPageNumber="72" pageId="27" pageNumber="72">
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<subSubSection pageId="27" pageNumber="72" type="multiple">
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<paragraph pageId="27" pageNumber="72">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="72" type="nomenclature">
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<paragraph pageId="27" pageNumber="72">
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13.
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<taxonomicName LSID="http://zoobank.org/610FF2E1-A763-4AF1-AABA-FEA533636427" authority="Shaverdo & Balke" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina varirata" order="Coleoptera" pageId="27" pageNumber="72" phylum="Arthropoda" rank="species" species="varirata">Exocelina varirata Shaverdo & Balke</taxonomicName>
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<taxonomicNameLabel pageId="27" pageNumber="72">sp. n.</taxonomicNameLabel>
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Figs 18, 34
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="72" type="reference_group">
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<paragraph pageId="27" pageNumber="72">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="27" pageNumber="72" phylum="Arthropoda" rank="genus">Exocelina</taxonomicName>
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undescribed sp. MB3303:
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<bibRefCitation author="Toussaint, EFA" journalOrPublisher="Nature Communications" pageId="36" pageNumber="81" pagination="1 - 10" title="The towering orogeny of New Guinea as a trigger for arthropod megadiversity." url="10.1038/ncomms5001" volume="1" year="2014">Toussaint et al. 2014</bibRefCitation>
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: supplementary figs 1-4, tab. 2.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="72" type="type locality">
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<paragraph pageId="27" pageNumber="72">Type locality.</paragraph>
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<paragraph pageId="27" pageNumber="72">
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Papua New Guinea: National Capital District Province, Varirata National Park,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-9.4355">09°26.13'S</geoCoordinate>
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;
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="147.36816">147°22.09'E</geoCoordinate>
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, 600 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="72" type="type material">
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<paragraph pageId="27" pageNumber="72">Type material.</paragraph>
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<paragraph pageId="27" pageNumber="72">
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Holotype: male "Papua New Guinea: National Capital District, Varirata NP, 600m, 16.xii.2007, 09.26.13S 147.22.09E, Balke & Sagata (PNG 159)", "DNA M.Balke 3303" [green] (ZSM). Paratype: Central: 1 male "Papua New Guinea: Central, Myola, 1110m, i.2008,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-9.2105">09 12.630S</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="147.53133">147 31.880E</geoCoordinate>
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, Posman (PNG 177)", "DNA M.Balke 3407" [green] (ZSM).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="72" type="diagnosis">
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<paragraph pageId="27" pageNumber="72">Diagnosis.</paragraph>
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<paragraph pageId="27" pageNumber="72">
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Beetle medium-sized, dark brown, with reddish-brown pronotal sides; dorsal surface with strong punctation and microreticulation, matt; pronotum with distinct lateral bead; male antennomeres simple; male protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta; median lobe slender, with slightly curved, short, broad apex and compact area of fine distal setae in lateral view, with slightly concave apex in ventral view; paramere with very shallow notch on dorsal side. The species is very similar to
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina wareaga" order="Coleoptera" pageId="27" pageNumber="72" phylum="Arthropoda" rank="species" species="wareaga">Exocelina wareaga</taxonomicName>
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sp. n. but differs from it in shape of the median lobe: it is more slender, lateral margins apically and subapically not very thick and not bordered with a carina; also the fine distal setae on lateral sides of the median lobe are not situated linearly, but in compact areas.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="72" type="description">
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<paragraph pageId="27" pageNumber="72">Description.</paragraph>
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<paragraph pageId="27" pageNumber="72">Size and shape: Beetle medium-sized (TL-H 4.25-4.35 mm, TL 4.5-4.85 mm, MW 2.25-2.45 mm), with oblong-oval habitus, broadest at elytral middle. Coloration: Head reddish brown to dark brown, paler on clypeus; pronotum dark brown on disc and reddish-brown on sides; elytra uniformly dark brown; ventrally dark brown; head appendages reddish-brown, legs darker distally (Fig. 34).</paragraph>
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<paragraph pageId="27" pageNumber="72">
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Surface sculpture: as in
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina wareaga" order="Coleoptera" pageId="27" pageNumber="72" phylum="Arthropoda" rank="species" species="wareaga">Exocelina wareaga</taxonomicName>
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sp. n.
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</paragraph>
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<paragraph pageId="27" pageNumber="72">Structures: Pronotum with distinct lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, smooth anteriorly, without anterolateral extensions. Blade of prosternal process lanceolate, relatively narrow, convex, with distinct bead and few setae laterally; neck and blade of prosternal process evenly jointed. Abdominal ventrite 6 broadly rounded.</paragraph>
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<paragraph pageId="27" pageNumber="72">
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Male: Antenna simple. Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior band of more than 70 short setae and posterior row of 12 relatively long, thin setae (Fig. 18A). Abdominal ventrite 6 with 4-6 lateral striae on each side. Median lobe slender, with slightly curved, short, broad apex and compact area of fine distal setae in lateral view, with slightly concave apex in ventral view (Figs 18
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<normalizedToken originalValue="B–C">B-C</normalizedToken>
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). Paramere with very shallow notch on dorsal side, with thin, sparse, inconspicuous proximal setae and thicker, denser, and longer subdistal setae (Fig. 18D).
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</paragraph>
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<paragraph pageId="27" pageNumber="72">Holotype: TL-H 4.35 mm, TL 4.85 mm, MW 2.45 mm.</paragraph>
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<paragraph pageId="27" pageNumber="72">Female: Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="72" type="distribution">
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<paragraph pageId="27" pageNumber="72">Distribution.</paragraph>
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<paragraph pageId="27" pageNumber="72">Papua New Guinea: National Capital District and Central Provinces (Fig. 40).</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="72" type="etymology">
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<paragraph pageId="27" pageNumber="72">Etymology.</paragraph>
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<paragraph pageId="27" pageNumber="72">The species is named after Varirata National Park. The name is a noun in the nominative singular standing in apposition.</paragraph>
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</subSubSection>
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</treatment>
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</document> |