480 lines
67 KiB
XML
480 lines
67 KiB
XML
<document id="CDE591A16AE8E69F3872CE1C407C8530" ID="10.11646/zootaxa.4178.3.4" ID-DOI="10.11646/zootaxa.4178.3.4" ID-GBIF-Dataset="148afb57-ea4e-4600-a16e-8085326ed3a4" ID-ISSN="1175-5326" ID-Zenodo-Dep="163357" ID-ZooBank="CBAD9B97-AD9A-41F2-A341-64F899861247" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1477468339595" checkinUser="plazi" docAuthor="Shear, William A., Ferreira, Rodrigo Lopes, Iniesta, Luiz Felipe Moretti & Marek, Paul" docDate="2016" docId="946F87AAFF8EFFCEFF7C21ABFAE4BA81" docLanguage="en" docName="zootaxa.4178.3.4.pdf" docOrigin="Zootaxa 4178 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dobrodesmus mirabilis Shear, Ferreira & Iniesta, n. sp." docType="treatment" docVersion="9" lastPageNumber="377" masterDocId="6856FFD2FF8CFFC8FFEB201CFFBDBC12" masterDocTitle="A millipede missing link: Dobrodesmidae, a remarkable new polydesmidan millipede family from Brazil with supernumerary rings (Diplopoda, Polydesmida), and the establishment of a new suborder Dobrodesmidea" masterLastPageNumber="390" masterPageNumber="371" pageNumber="373" updateTime="1698390828981" updateUser="plazi">
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<mods:titleInfo id="C8837D0256944DEC5EAEC196AC06B8EE">
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<mods:title id="A88C539AE009D070D27D95A6F4BDA728">A millipede missing link: Dobrodesmidae, a remarkable new polydesmidan millipede family from Brazil with supernumerary rings (Diplopoda, Polydesmida), and the establishment of a new suborder Dobrodesmidea</mods:title>
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<mods:name id="9F1F2389EBDA991441BBEB215EADFC2C" type="personal">
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<mods:namePart id="19AF7F6969FC8F99C16141F6760A257D">Shear, William A.</mods:namePart>
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<mods:namePart id="5BE5E2978E4F4E8D19199390A036366B">Ferreira, Rodrigo Lopes</mods:namePart>
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<mods:roleTerm id="A252BE9822A5D330AE9D30334CA1A860">Author</mods:roleTerm>
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<mods:namePart id="202B27899C46AA189BAA8B8A87DA7D60">Iniesta, Luiz Felipe Moretti</mods:namePart>
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<mods:namePart id="7AD4AA2D5730C88EA0C04894856B6FBE">Marek, Paul</mods:namePart>
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<mods:typeOfResource id="638A85D49A6857272401F0EFF26D1AE9">text</mods:typeOfResource>
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<mods:title id="BF3C222459B31C47B304E250432C7D83">Zootaxa</mods:title>
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<mods:date id="806DBDE35EF98B608DDB84A41B4F544C">2016</mods:date>
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<mods:number id="C0854DFDE6168F25E27555A3081132E7">4178</mods:number>
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<treatment id="946F87AAFF8EFFCEFF7C21ABFAE4BA81" ID-DOI="http://doi.org/10.5281/zenodo.6073631" ID-GBIF-Taxon="124558136" ID-Zenodo-Dep="6073631" LSID="urn:lsid:plazi:treatment:946F87AAFF8EFFCEFF7C21ABFAE4BA81" httpUri="http://treatment.plazi.org/id/946F87AAFF8EFFCEFF7C21ABFAE4BA81" lastPageId="6" lastPageNumber="377" pageId="2" pageNumber="373">
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<subSubSection id="54DC6537FF8EFFCAFF7C21ABFE38BDE6" pageId="2" pageNumber="373" type="nomenclature">
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<paragraph id="1C7936BCFF8EFFCAFF7C21ABFC9EBDC2" blockId="2.[151,803,439,501]" box="[151,803,439,464]" pageId="2" pageNumber="373">
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<heading id="473181D0FF8EFFCAFF7C21ABFC9EBDC2" bold="true" box="[151,803,439,464]" fontSize="10" level="2" pageId="2" pageNumber="373" reason="2">
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<emphasis id="2EB2EAAEFF8EFFCAFF7C21ABFC9EBDC2" bold="true" box="[151,803,439,464]" pageId="2" pageNumber="373">
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<taxonomicName id="DBC64D3FFF8EFFCAFF7C21ABFD6ABDC2" authority="Shear, Ferreira & Iniesta" authorityName="Shear, Ferreira & Iniesta" box="[151,727,439,464]" class="Diplopoda" family="Rhachodesmidae" genus="Dobrodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="mirabilis" status="sp. nov.">
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<emphasis id="2EB2EAAEFF8EFFCAFF7C21ABFE19BDC2" bold="true" box="[151,420,439,464]" italics="true" pageId="2" pageNumber="373">Dobrodesmus mirabilis</emphasis>
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Shear, Ferreira & Iniesta
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</taxonomicName>
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,
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<taxonomicNameLabel id="358157D5FF8EFFCAFD0F21ABFC9EBDC2" box="[740,803,439,464]" pageId="2" pageNumber="373" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="1C7936BCFF8EFFCAFF7C21C0FE38BDE6" blockId="2.[151,803,439,501]" box="[151,389,476,501]" pageId="2" pageNumber="373">
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<figureCitation id="84FD2A39FF8EFFCAFF7C21C0FE8FBDE7" box="[151,306,476,501]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 7" captionStart-2="FIGURES 8 – 10" captionStart-3="FIGURE 11" captionStart-4="FIGURES 12, 13" captionStart-5="FIGURES 14 – 16" captionStartId-0="3.[199,298,819,841]" captionStartId-1="3.[151,264,1949,1971]" captionStartId-2="4.[151,264,1298,1320]" captionStartId-3="5.[151,250,1139,1161]" captionStartId-4="6.[151,264,1202,1224]" captionStartId-5="7.[151,264,1268,1290]" captionTargetBox-1="[340,1246,891,1927]" captionTargetBox-2="[351,1228,183,1269]" captionTargetBox-3="[340,1246,193,1117]" captionTargetBox-4="[340,1246,337,1180]" captionTargetBox-5="[340,1246,193,1245]" captionTargetId-1="figure@3.[340,1247,891,1928]" captionTargetId-2="figure@4.[351,1229,183,1270]" captionTargetId-3="figure@5.[340,1247,193,1118]" captionTargetId-4="figure@6.[340,1247,337,1181]" captionTargetId-5="figure@7.[340,1247,193,1246]" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionText-0="FIGURE 1. Dobrodesmus mirabilis, living male in habitat." captionText-1="FIGURES 2 – 7. Dobrodesmus mirabilis. Fig 2, collum. Fig 3, sensilla from sixth antennal article. Fig 4, midbody segment, dorsal view. Fig 5, midbody segment, ventral view. Fig 6, anterior right corner of midbody segment, dorsal view. Fig 7, posterior right corner of nonporiferous midbody segment, dorsal view." captionText-2="FIGURES 8 – 10. Dobrodesmus mirabilis. Fig 8, Ozopore, lateral view. Fig 9, intercalary microscutes of metazonital dorsum, dorsal view. Fig 10, same." captionText-3="FIGURE 11. Dobrodesmus mirabilis, posterior end, dorsolateral view. Abbreviations: s, spinneret; dp, dorsal division of paraproct; vp, ventral division of paraproct.; t, epiproct." captionText-4="FIGURES 12, 13. Dobrodesmus mirabilis. Fig 12, ventral view of right lateral part of midbody segment, showing positions of spiracles. Abbreviations: as, anterior spiracle; ps, posterior spiracle. Fig 13. Posterior spiracle." captionText-5="FIGURES 14 – 16. Fig 14, spinnerets of Dobrodesmus mirabilis. Fig 15, spinnerets of Tetracion jonesi Hoffman 1956 (Callipodida, Abacionidae). Abbreviations: sl, extended aperture of spinneret; dp, dorsal division of paraproct; vp, ventral division of paraproct. Fig 16, gonopods of Dobrodesmus mirabilis, ventral view. Abbreviations: cx, gonopod coxa; f, flagellum; p 1, sheathing process of coxa; p 2, triangular process of telopodite; tel, telopodite of gonopo; a?, possible acropodite." httpUri-1="https://zenodo.org/record/257272/files/figure.png" httpUri-2="https://zenodo.org/record/257273/files/figure.png" httpUri-3="https://zenodo.org/record/257274/files/figure.png" httpUri-4="https://zenodo.org/record/257275/files/figure.png" httpUri-5="https://zenodo.org/record/257276/files/figure.png" pageId="2" pageNumber="373">Figs 1–14, 16</figureCitation>
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,
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<figureCitation id="84FD2A39FF8EFFCAFED521C0FE38BDE6" box="[318,389,476,500]" captionStart-0="FIGURES 17, 18" captionStart-1="FIGURES 19 – 22" captionStartId-0="7.[151,264,1936,1958]" captionStartId-1="8.[151,264,1950,1972]" captionTargetBox-0="[340,1246,1462,1914]" captionTargetBox-1="[218,1385,193,1908]" captionTargetId-0="figure@7.[340,1247,1462,1914]" captionTargetId-1="figure@8.[203,1385,193,1923]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 17, 18. Fig 17, gonopods of Sinocallipus simplipodicus (Callipodida, Sinocallipodidae). Fig 18, left gonopod of Dobrodesmus mirabilis, ventral view. Abbreviations as for Fig 16." captionText-1="FIGURES 19 – 22. Fig 19, gonopods of Dobrodesmus mirabilis, posterior view. Fig 20, gonopods of Dobrodesmus mirabilis, anterior view. Fig 21, male, dorsal view, diplosegments numbered; t, telson. Fig 22, gonopods of Eostemmiulus caecus, (Stemmiulida, Stemmiulidae), posterior view. Abbreviations as for Fig 16; Fig 22 redrawn from Mauriès et al. (2010)." httpUri-0="https://zenodo.org/record/257277/files/figure.png" httpUri-1="https://zenodo.org/record/163364/files/figure.png" pageId="2" pageNumber="373">18–21</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="54DC6537FF8EFFCAFF7C2238FEE8BEDB" pageId="2" pageNumber="373" type="description">
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<paragraph id="1C7936BCFF8EFFCAFF7C2238FEE8BEDB" blockId="2.[151,1437,548,2016]" pageId="2" pageNumber="373">
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<materialsCitation id="ACAE3CE1FF8EFFCAFF7C2238FDBFBE4D" ID-GBIF-Occurrence="1324200587" collectorName="Gruta de Mangabeira & Ituacu Municipality" country="Iceland" location="Bahia" pageId="2" pageNumber="373" specimenCount="4" stateProvince="Bahia" typeStatus="holotype">
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||
<emphasis id="2EB2EAAEFF8EFFCAFF7C2238FF5EBE2F" bold="true" box="[151,227,548,573]" pageId="2" pageNumber="373">
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||
<typeStatus id="C37D881EFF8EFFCAFF7C2238FF62BE2F" box="[151,223,548,573]" pageId="2" pageNumber="373">Types</typeStatus>
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.
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</emphasis>
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Male
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<typeStatus id="C37D881EFF8EFFCAFEC42238FE2CBE2F" box="[303,401,548,573]" pageId="2" pageNumber="373" type="holotype">holotype</typeStatus>
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(
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<collectingCountry id="64D1762CFF8EFFCAFE4A2238FE5CBE2E" box="[417,481,548,572]" name="Iceland" pageId="2" pageNumber="373">ISLA</collectingCountry>
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3637) male
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<typeStatus id="C37D881EFF8EFFCAFD9F2239FD6ABE2F" box="[628,727,549,573]" pageId="2" pageNumber="373" type="paratype">paratype</typeStatus>
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, and numerous juvenile
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<typeStatus id="C37D881EFF8EFFCAFC1B2239FBE6BE2F" box="[1008,1115,549,573]" pageId="2" pageNumber="373" type="paratype">paratypes</typeStatus>
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from
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<collectorName id="B133536AFF8EFFCAFB482238FA25BE2F" box="[1187,1432,548,573]" pageId="2" pageNumber="373">Gruta de Mangabeira</collectorName>
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||
,
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||
<collectorName id="B133536AFF8EFFCAFF7C2254FEC8BE72" box="[151,373,583,608]" pageId="2" pageNumber="373">Ituaçu Municipality</collectorName>
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,
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<collectingRegion id="DE02F85EFF8EFFCAFE6B225BFE7CBE72" box="[384,449,583,608]" country="Brazil" name="Bahia" pageId="2" pageNumber="373">Bahia</collectingRegion>
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State
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</materialsCitation>
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,
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<materialsCitation id="ACAE3CE1FF8EFFCAFDE6225BFDDCBEB4" ID-GBIF-Occurrence="1324200588" collectingDate="2006-12-30" collectorName="R. Ferreira & Zoology Collection & Secao de Invertebrados Subterraneos & Universidade Federal de Lavras & Campus Universitario de Lavras" country="Brazil" location="Minas Gerais" pageId="2" pageNumber="373" specimenCount="1" stateProvince="Minas Gerais" typeStatus="holotype">
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<collectingCountry id="64D1762CFF8EFFCAFDE6225BFDECBE72" box="[525,593,583,608]" name="Brazil" pageId="2" pageNumber="373">Brazil</collectingCountry>
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||
(
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<figureCitation id="84FD2A39FF8EFFCAFD8B225BFD5FBE4D" box="[608,738,583,608]" captionStart-0="FIGURES 23 – 27" captionStart-1="FIGURE 28" captionStartId-0="9.[151,264,1836,1858]" captionStartId-1="11.[151,250,1362,1384]" captionTargetBox-0="[247,1335,193,1804]" captionTargetBox-1="[207,1340,596,1322]" captionTargetId-0="figure@9.[247,1335,193,1812]" captionTargetId-1="figure@11.[207,1341,589,1326]" captionTargetPageId-0="9" captionTargetPageId-1="11" captionText-0="FIGURES 23 – 27. Habitat of Dobrodesmus mirabilis in Mangebeira Cave. Fig 23, course of Mangebeira Cave (red) superimposed on aerial view, with part of municipality of Ituaçu. Fig 24, sinkhole entrance to the cave showing modifications for religious purposes. Fig 25, chapel area inside cave. Fig 26, pilgrims visiting the cave. Fig 27, deeper interior of the cave, with speleothems, close to collection locality of D. mirabilis holotype." captionText-1="FIGURE 28. Map showing location of type locality of Dobrodesmus mirabilis. Expansion is of Bahia State; red is known extent of formations of the Una Group, black is Ituaçu Municipality. Arrow indicates location of Mangebeira Cave." httpUri-0="https://zenodo.org/record/163365/files/figure.png" httpUri-1="https://zenodo.org/record/163366/files/figure.png" pageId="2" pageNumber="373">Figs 23–28</figureCitation>
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), collected by
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<collectorName id="B133536AFF8EFFCAFC682254FC42BE72" box="[899,1023,583,608]" pageId="2" pageNumber="373">R. Ferreira</collectorName>
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,
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<collectingDate id="783CE994FF8EFFCAFBE1225BFB5CBE72" box="[1034,1249,583,608]" pageId="2" pageNumber="373" value="2006-12-30">
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||
<date id="6878107CFF8EFFCAFBE1225BFB5CBE72" box="[1034,1249,583,608]" pageId="2" pageNumber="373" value="2006-12-30">30 December 2006</date>
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</collectingDate>
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, deposited in the
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<collectorName id="B133536AFF8EFFCAFF7C2276FEC8BE91" box="[151,373,618,643]" pageId="2" pageNumber="373">Zoology Collection</collectorName>
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||
,
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||
<collectorName id="B133536AFF8EFFCAFE6A2277FC9FBE91" box="[385,802,618,643]" pageId="2" pageNumber="373">Seção de Invertebrados Subterrâneos</collectorName>
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(ISLA) at the
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<collectorName id="B133536AFF8EFFCAFC212276FA89BE90" box="[970,1332,618,643]" pageId="2" pageNumber="373">Universidade Federal de Lavras</collectorName>
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,
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<collectorName id="B133536AFF8EFFCAFAAB2277FE12BEB7" pageId="2" pageNumber="373">Campus Universitário de Lavras</collectorName>
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||
,
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<collectingRegion id="DE02F85EFF8EFFCAFE2A2291FDDCBEB4" box="[449,609,653,678]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="373">Minas Gerais</collectingRegion>
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||
</materialsCitation>
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||
,
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||
<materialsCitation id="ACAE3CE1FF8EFFCAFD982291FEE8BEDB" ID-GBIF-Occurrence="1324200586" country="Brazil" location="Brazil" pageId="2" pageNumber="373" specimenCount="1" typeStatus="paratype">
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||
<collectingCountry id="64D1762CFF8EFFCAFD982291FD06BEB4" box="[627,699,653,678]" name="Brazil" pageId="2" pageNumber="373">Brazil</collectingCountry>
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. The male
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||
<typeStatus id="C37D881EFF8EFFCAFCA22292FC14BEB4" box="[841,937,654,678]" pageId="2" pageNumber="373" type="paratype">paratype</typeStatus>
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remains mounted on a scanning electron microscope stub.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="54DC6537FF8EFFCAFF2C22C8FE88BF02" pageId="2" pageNumber="373" type="etymology">
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<paragraph id="1C7936BCFF8EFFCAFF2C22C8FE88BF02" blockId="2.[151,1437,548,2016]" pageId="2" pageNumber="373">
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||
<emphasis id="2EB2EAAEFF8EFFCAFF2C22C8FEF3BEFF" bold="true" box="[199,334,724,749]" pageId="2" pageNumber="373">Etymology.</emphasis>
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The species epithet “
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<taxonomicName id="DBC64D3FFF8EFFCAFDD422C8FD1ABEFF" box="[575,679,724,749]" class="Diplopoda" family="Rhachodesmidae" genus="Dobrodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="mirabilis">mirabilis</taxonomicName>
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” means amazing in Latin and refers to the characteristics observed in the species.
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</paragraph>
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</subSubSection>
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<subSubSection id="54DC6537FF8EFFCAFF2C2306FD39BF21" box="[199,644,794,819]" pageId="2" pageNumber="373" type="diagnosis">
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||
<paragraph id="1C7936BCFF8EFFCAFF2C2306FD39BF21" blockId="2.[151,1437,548,2016]" box="[199,644,794,819]" pageId="2" pageNumber="373">
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||
<emphasis id="2EB2EAAEFF8EFFCAFF2C2306FEFCBF21" bold="true" box="[199,321,794,819]" pageId="2" pageNumber="373">Diagnosis.</emphasis>
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As for the genus, see above.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="54DC6537FF8EFFCEFF2C2321FAE4BA81" lastPageId="6" lastPageNumber="377" pageId="2" pageNumber="373" type="description">
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<paragraph id="1C7936BCFF8EFFCAFF2C2321FBDFBF44" blockId="2.[151,1437,548,2016]" box="[199,1122,829,854]" pageId="2" pageNumber="373">
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<emphasis id="2EB2EAAEFF8EFFCAFF2C2321FEE4BF44" bold="true" box="[199,345,829,854]" pageId="2" pageNumber="373">Description.</emphasis>
|
||
Male (
|
||
<figureCitation id="84FD2A39FF8EFFCAFE4C2321FE59BF44" box="[423,484,829,854]" captionStart="FIGURE 1" captionStartId="3.[199,298,819,841]" captionText="FIGURE 1. Dobrodesmus mirabilis, living male in habitat." pageId="2" pageNumber="373">Fig 1</figureCitation>
|
||
; now fragmentary) about
|
||
<quantity id="DB3E9B59FF8EFFCAFCE32322FC3CBF47" box="[776,897,830,854]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="2" pageNumber="373" unit="mm" value="21.5" valueMax="22.0" valueMin="21.0">21–22 mm</quantity>
|
||
long, 1.0 mm wide.
|
||
</paragraph>
|
||
<paragraph id="1C7936BCFF8EFFCAFF2C237CFB53BF8E" blockId="2.[151,1437,548,2016]" pageId="2" pageNumber="373">
|
||
Head subglobular, setose. Antennae
|
||
<quantity id="DB3E9B59FF8EFFCAFD8C237CFD4DBF6A" box="[615,752,864,889]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="2" pageNumber="373" unit="mm" value="1.65" valueMax="1.7" valueMin="1.6">1.6–1.7 mm</quantity>
|
||
long, if extended posteriad, reaching to posterior margin of fifth ring. Penultimate antennal segment with cluster of club-shaped setae lining outer apex (
|
||
<figureCitation id="84FD2A39FF8EFFCAFB482398FB5CBF8E" box="[1187,1249,900,925]" captionStart="FIGURES 2 – 7" captionStartId="3.[151,264,1949,1971]" captionTargetBox="[340,1246,891,1927]" captionTargetId="figure@3.[340,1247,891,1928]" captionTargetPageId="3" captionText="FIGURES 2 – 7. Dobrodesmus mirabilis. Fig 2, collum. Fig 3, sensilla from sixth antennal article. Fig 4, midbody segment, dorsal view. Fig 5, midbody segment, ventral view. Fig 6, anterior right corner of midbody segment, dorsal view. Fig 7, posterior right corner of nonporiferous midbody segment, dorsal view." httpUri="https://zenodo.org/record/257272/files/figure.png" pageId="2" pageNumber="373">Fig 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1C7936BCFF8EFFCAFF2C23BBFA83BFD2" blockId="2.[151,1437,548,2016]" box="[199,1342,935,960]" pageId="2" pageNumber="373">With troglobiotic character states; colorless, cuticle thin, poorly sclerotized, brittle, legs long and thin.</paragraph>
|
||
<paragraph id="1C7936BCFF8EFFCAFF2C23D6FE46B814" blockId="2.[151,1437,548,2016]" pageId="2" pageNumber="373">
|
||
Collum (
|
||
<figureCitation id="84FD2A39FF8EFFCAFED323D6FE3EBFF1" box="[312,387,970,995]" captionStart="FIGURES 2 – 7" captionStartId="3.[151,264,1949,1971]" captionTargetBox="[340,1246,891,1927]" captionTargetId="figure@3.[340,1247,891,1928]" captionTargetPageId="3" captionText="FIGURES 2 – 7. Dobrodesmus mirabilis. Fig 2, collum. Fig 3, sensilla from sixth antennal article. Fig 4, midbody segment, dorsal view. Fig 5, midbody segment, ventral view. Fig 6, anterior right corner of midbody segment, dorsal view. Fig 7, posterior right corner of nonporiferous midbody segment, dorsal view." httpUri="https://zenodo.org/record/257272/files/figure.png" pageId="2" pageNumber="373">Fig 2</figureCitation>
|
||
) densely setose, subelliptical, anteriorly rounded, posterior margin nearly straight, posteriolateral corners rounded.
|
||
</paragraph>
|
||
<paragraph id="1C7936BCFF8EFFCAFF2C240CFAF6B881" blockId="2.[151,1437,548,2016]" pageId="2" pageNumber="373">
|
||
Typical ring (
|
||
<figureCitation id="84FD2A39FF8EFFCAFEB6240CFE02B83B" box="[349,447,1040,1065]" captionStart="FIGURES 2 – 7" captionStartId="3.[151,264,1949,1971]" captionTargetBox="[340,1246,891,1927]" captionTargetId="figure@3.[340,1247,891,1928]" captionTargetPageId="3" captionText="FIGURES 2 – 7. Dobrodesmus mirabilis. Fig 2, collum. Fig 3, sensilla from sixth antennal article. Fig 4, midbody segment, dorsal view. Fig 5, midbody segment, ventral view. Fig 6, anterior right corner of midbody segment, dorsal view. Fig 7, posterior right corner of nonporiferous midbody segment, dorsal view." httpUri="https://zenodo.org/record/257272/files/figure.png" pageId="2" pageNumber="373">Figs 4, 5</figureCitation>
|
||
) with dense coat of fine, short setae, 12 longer, stouter setae with prominent apertures in two transverse rows on each metazonite. Midbody rings 1.0 mm wide across paranota, paranota level, projecting directly laterad, squarish, anteriolateral corners (
|
||
<figureCitation id="84FD2A39FF8EFFCAFD20244BFCB1B862" box="[715,780,1111,1136]" captionStart="FIGURES 2 – 7" captionStartId="3.[151,264,1949,1971]" captionTargetBox="[340,1246,891,1927]" captionTargetId="figure@3.[340,1247,891,1928]" captionTargetPageId="3" captionText="FIGURES 2 – 7. Dobrodesmus mirabilis. Fig 2, collum. Fig 3, sensilla from sixth antennal article. Fig 4, midbody segment, dorsal view. Fig 5, midbody segment, ventral view. Fig 6, anterior right corner of midbody segment, dorsal view. Fig 7, posterior right corner of nonporiferous midbody segment, dorsal view." httpUri="https://zenodo.org/record/257272/files/figure.png" pageId="2" pageNumber="373">Fig 6</figureCitation>
|
||
) strongly angular, with numerous stout triangular teeth, posteriorlateral corners (
|
||
<figureCitation id="84FD2A39FF8EFFCAFE432466FE58B881" box="[424,485,1146,1171]" captionStart="FIGURES 2 – 7" captionStartId="3.[151,264,1949,1971]" captionTargetBox="[340,1246,891,1927]" captionTargetId="figure@3.[340,1247,891,1928]" captionTargetPageId="3" captionText="FIGURES 2 – 7. Dobrodesmus mirabilis. Fig 2, collum. Fig 3, sensilla from sixth antennal article. Fig 4, midbody segment, dorsal view. Fig 5, midbody segment, ventral view. Fig 6, anterior right corner of midbody segment, dorsal view. Fig 7, posterior right corner of nonporiferous midbody segment, dorsal view." httpUri="https://zenodo.org/record/257272/files/figure.png" pageId="2" pageNumber="373">Fig 7</figureCitation>
|
||
) acutely produced, lateral edges finely and irregularly serrate in anterior half.
|
||
</paragraph>
|
||
<paragraph id="1C7936BCFF8EFFCAFF2C2481FC86B8CB" blockId="2.[151,1437,548,2016]" pageId="2" pageNumber="373">
|
||
Metazonite cuticle with numerous intercalary microscutes, (
|
||
<figureCitation id="84FD2A39FF8EFFCAFC7E2481FBABB8A4" box="[917,1046,1181,1206]" captionStart="FIGURES 8 – 10" captionStartId="4.[151,264,1298,1320]" captionTargetBox="[351,1228,183,1269]" captionTargetId="figure@4.[351,1229,183,1270]" captionTargetPageId="4" captionText="FIGURES 8 – 10. Dobrodesmus mirabilis. Fig 8, Ozopore, lateral view. Fig 9, intercalary microscutes of metazonital dorsum, dorsal view. Fig 10, same." httpUri="https://zenodo.org/record/257273/files/figure.png" pageId="2" pageNumber="373">Figs 9, 10</figureCitation>
|
||
) limbus unmodified, completely smooth. Prozonite sculpture regular rows of polygonal cells.
|
||
</paragraph>
|
||
<paragraph id="1C7936BCFF8EFFCAFF2C24F8FA2EB8EE" blockId="2.[151,1437,548,2016]" box="[199,1427,1252,1277]" pageId="2" pageNumber="373">
|
||
Ozopores (
|
||
<figureCitation id="84FD2A39FF8EFFCAFEAA24F8FEC0B8EE" box="[321,381,1252,1277]" captionStart="FIGURES 8 – 10" captionStartId="4.[151,264,1298,1320]" captionTargetBox="[351,1228,183,1269]" captionTargetId="figure@4.[351,1229,183,1270]" captionTargetPageId="4" captionText="FIGURES 8 – 10. Dobrodesmus mirabilis. Fig 8, Ozopore, lateral view. Fig 9, intercalary microscutes of metazonital dorsum, dorsal view. Fig 10, same." httpUri="https://zenodo.org/record/257273/files/figure.png" pageId="2" pageNumber="373">Fig 8</figureCitation>
|
||
) on rings 5, 7, 9, 10, 12, 13, 15–38, located on produced posteriolateral corner of paranotum.
|
||
</paragraph>
|
||
<paragraph id="1C7936BCFF8EFFCAFF2C251BFC70B9BF" blockId="2.[151,1437,548,2016]" pageId="2" pageNumber="373">
|
||
Epiproct (
|
||
<figureCitation id="84FD2A39FF8EFFCAFED2251BFE39B90D" box="[313,388,1287,1312]" captionStart="FIGURE 11" captionStartId="5.[151,250,1139,1161]" captionTargetBox="[340,1246,193,1117]" captionTargetId="figure@5.[340,1247,193,1118]" captionTargetPageId="5" captionText="FIGURE 11. Dobrodesmus mirabilis, posterior end, dorsolateral view. Abbreviations: s, spinneret; dp, dorsal division of paraproct; vp, ventral division of paraproct.; t, epiproct." httpUri="https://zenodo.org/record/257274/files/figure.png" pageId="2" pageNumber="373">Fig 11</figureCitation>
|
||
) twice as long as wide, densely setose, terminal process appears set off by groove, deeply divided into two subtriangular processes, each bearing spinneret for complement of two spinnerets. Paraprocts strongly margined, with oblique transverse sulcus dividing paraprocts into smaller dorsal and larger ventral portions (
|
||
<figureCitation id="84FD2A39FF8EFFCAFEE8256CFEF2B99B" box="[259,335,1392,1417]" captionStart="FIGURE 11" captionStartId="5.[151,250,1139,1161]" captionTargetBox="[340,1246,193,1117]" captionTargetId="figure@5.[340,1247,193,1118]" captionTargetPageId="5" captionText="FIGURE 11. Dobrodesmus mirabilis, posterior end, dorsolateral view. Abbreviations: s, spinneret; dp, dorsal division of paraproct; vp, ventral division of paraproct.; t, epiproct." httpUri="https://zenodo.org/record/257274/files/figure.png" pageId="2" pageNumber="373">Fig 11</figureCitation>
|
||
). Paraprocts densely setose, without evident marginal or submarginal macrosetae, but with large, distinct depression or pore at distal angle. Hypoproct entire, semicircular.
|
||
</paragraph>
|
||
<paragraph id="1C7936BCFF8EFFCAFF2C25ABFB05B9C2" blockId="2.[151,1437,548,2016]" box="[199,1208,1463,1488]" pageId="2" pageNumber="373">
|
||
Ventral surfaces of rings with transverse depression between insertions of legpairs (
|
||
<figureCitation id="84FD2A39FF8EFFCAFB8625ABFB16B9C2" box="[1133,1195,1463,1488]" captionStart="FIGURES 2 – 7" captionStartId="3.[151,264,1949,1971]" captionTargetBox="[340,1246,891,1927]" captionTargetId="figure@3.[340,1247,891,1928]" captionTargetPageId="3" captionText="FIGURES 2 – 7. Dobrodesmus mirabilis. Fig 2, collum. Fig 3, sensilla from sixth antennal article. Fig 4, midbody segment, dorsal view. Fig 5, midbody segment, ventral view. Fig 6, anterior right corner of midbody segment, dorsal view. Fig 7, posterior right corner of nonporiferous midbody segment, dorsal view." httpUri="https://zenodo.org/record/257272/files/figure.png" pageId="2" pageNumber="373">Fig 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1C7936BCFF8EFFCAFF2C25C6FB4BB9E1" blockId="2.[151,1437,548,2016]" box="[199,1270,1498,1523]" pageId="2" pageNumber="373">
|
||
Spiracles small (
|
||
<figureCitation id="84FD2A39FF8EFFCAFE6B25C6FE71B9E1" box="[384,460,1498,1523]" captionStart="FIGURES 12, 13" captionStartId="6.[151,264,1202,1224]" captionTargetBox="[340,1246,337,1180]" captionTargetId="figure@6.[340,1247,337,1181]" captionTargetPageId="6" captionText="FIGURES 12, 13. Dobrodesmus mirabilis. Fig 12, ventral view of right lateral part of midbody segment, showing positions of spiracles. Abbreviations: as, anterior spiracle; ps, posterior spiracle. Fig 13. Posterior spiracle." httpUri="https://zenodo.org/record/257275/files/figure.png" pageId="2" pageNumber="373">Fig 13</figureCitation>
|
||
), strongly reduced, advanced directly anterior of each leg coxa (
|
||
<figureCitation id="84FD2A39FF8EFFCAFB7025C6FB55B9E1" box="[1179,1256,1498,1523]" captionStart="FIGURES 12, 13" captionStartId="6.[151,264,1202,1224]" captionTargetBox="[340,1246,337,1180]" captionTargetId="figure@6.[340,1247,337,1181]" captionTargetPageId="6" captionText="FIGURES 12, 13. Dobrodesmus mirabilis. Fig 12, ventral view of right lateral part of midbody segment, showing positions of spiracles. Abbreviations: as, anterior spiracle; ps, posterior spiracle. Fig 13. Posterior spiracle." httpUri="https://zenodo.org/record/257275/files/figure.png" pageId="2" pageNumber="373">Fig 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1C7936BCFF8EFFCAFF2C25E2FD45BA04" blockId="2.[151,1437,548,2016]" box="[199,760,1533,1558]" pageId="2" pageNumber="373">
|
||
Legs long, thin; midbody legs about
|
||
<quantity id="DB3E9B59FF8EFFCAFD8925E2FD05BA07" box="[610,696,1534,1558]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="373" unit="mm" value="1.2">1.2 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
<paragraph id="1C7936BCFF8EFFCAFF2C263DFC0FBA4F" blockId="2.[151,1437,548,2016]" pageId="2" pageNumber="373">Sternum of second legpair not incorporated in second ring, freely articulating; second leg coxae quadrate, with opening of vas deferens mesal on each. No anterior legpairs incrassate.</paragraph>
|
||
<paragraph id="1C7936BCFF8EFFCAFF2C267BFEDBBBAF" blockId="2.[151,1437,548,2016]" pageId="2" pageNumber="373">
|
||
Gonopods (
|
||
<figureCitation id="84FD2A39FF8EFFCAFEA7267BFE18BA92" box="[332,421,1639,1664]" captionStart="FIGURES 14 – 16" captionStartId="7.[151,264,1268,1290]" captionTargetBox="[340,1246,193,1245]" captionTargetId="figure@7.[340,1247,193,1246]" captionTargetPageId="7" captionText="FIGURES 14 – 16. Fig 14, spinnerets of Dobrodesmus mirabilis. Fig 15, spinnerets of Tetracion jonesi Hoffman 1956 (Callipodida, Abacionidae). Abbreviations: sl, extended aperture of spinneret; dp, dorsal division of paraproct; vp, ventral division of paraproct. Fig 16, gonopods of Dobrodesmus mirabilis, ventral view. Abbreviations: cx, gonopod coxa; f, flagellum; p 1, sheathing process of coxa; p 2, triangular process of telopodite; tel, telopodite of gonopo; a?, possible acropodite." httpUri="https://zenodo.org/record/257276/files/figure.png" pageId="2" pageNumber="373">Figs 16</figureCitation>
|
||
,
|
||
<figureCitation id="84FD2A39FF8EFFCAFE5F267BFE6FBA6D" box="[436,466,1639,1663]" captionStart="FIGURES 17, 18" captionStartId="7.[151,264,1936,1958]" captionTargetBox="[340,1246,1462,1914]" captionTargetId="figure@7.[340,1247,1462,1914]" captionTargetPageId="7" captionText="FIGURES 17, 18. Fig 17, gonopods of Sinocallipus simplipodicus (Callipodida, Sinocallipodidae). Fig 18, left gonopod of Dobrodesmus mirabilis, ventral view. Abbreviations as for Fig 16." httpUri="https://zenodo.org/record/257277/files/figure.png" pageId="2" pageNumber="373">18</figureCitation>
|
||
,
|
||
<figureCitation id="84FD2A39FF8EFFCAFE0A267BFD96BA6D" box="[481,555,1639,1663]" captionStart="FIGURES 19 – 22" captionStartId="8.[151,264,1950,1972]" captionTargetBox="[218,1385,193,1908]" captionTargetId="figure@8.[203,1385,193,1923]" captionTargetPageId="8" captionText="FIGURES 19 – 22. Fig 19, gonopods of Dobrodesmus mirabilis, posterior view. Fig 20, gonopods of Dobrodesmus mirabilis, anterior view. Fig 21, male, dorsal view, diplosegments numbered; t, telson. Fig 22, gonopods of Eostemmiulus caecus, (Stemmiulida, Stemmiulidae), posterior view. Abbreviations as for Fig 16; Fig 22 redrawn from Mauriès et al. (2010)." httpUri="https://zenodo.org/record/163364/files/figure.png" pageId="2" pageNumber="373">19, 20</figureCitation>
|
||
) in tightly constricting oval aperture, coxae entirely exposed, constricted by margin of aperture. Sternum complete, articulated with, rather than partly or entirely fused to, coxae. Coxae large, egg-shaped, closely appressed, perhaps fused in midline, with clusters of 5–7 strong setae on lateral side, mesally with distinct angular flange; mesal coxal surface behind flange either fimbricate or with dense, fine setae. Coxae evidently immovable with respect to one another. Cannula not detected, but long, distally multihastate flagellum articulated with posterior apical coxal lobe sheathed in broad, distally acuminate coxal process. Telopodite singlearticled, subglobular with thin, triangular process guiding coxal sheathing process; small, distal, acute process (possibly representing acropodite) subtended by two setae, one of which has many small setulae, mesal group of three stout setae. No seminal (prostatic) groove or channel detected. Telopodite not divided into prefemoral and acropodal regions.
|
||
</paragraph>
|
||
<paragraph id="1C7936BCFF8EFFCAFF2C27DBFE7EBBF2" blockId="2.[151,1437,548,2016]" box="[199,451,1991,2016]" pageId="2" pageNumber="373">Females not collected.</paragraph>
|
||
<caption id="48B96634FF8FFFCBFF2C232FFC9FBF5B" box="[199,802,819,841]" pageId="3" pageNumber="374">
|
||
<paragraph id="1C7936BCFF8FFFCBFF2C232FFC9FBF5B" blockId="3.[199,802,819,841]" box="[199,802,819,841]" pageId="3" pageNumber="374">
|
||
<emphasis id="2EB2EAAEFF8FFFCBFF2C232FFE80BF5B" bold="true" box="[199,317,819,841]" pageId="3" pageNumber="374">FIGURE 1</emphasis>
|
||
.
|
||
<emphasis id="2EB2EAAEFF8FFFCBFEA2232FFD80BF5B" box="[329,573,819,841]" italics="true" pageId="3" pageNumber="374">
|
||
<taxonomicName id="DBC64D3FFF8FFFCBFEA2232FFD84BF5B" box="[329,569,819,841]" class="Diplopoda" family="Rhachodesmidae" genus="Dobrodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="mirabilis">Dobrodesmus mirabilis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
living male in habitat.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="48B96634FF8FFFCBFF7C2781FCDDBBE3" httpUri="https://zenodo.org/record/257272/files/figure.png" pageId="3" pageNumber="374" targetBox="[340,1246,891,1927]" targetPageId="3">
|
||
<paragraph id="1C7936BCFF8FFFCBFF7C2781FCDDBBE3" blockId="3.[151,1436,1949,2033]" pageId="3" pageNumber="374">
|
||
<emphasis id="2EB2EAAEFF8FFFCBFF7C2781FE88BBA1" bold="true" box="[151,309,1949,1971]" pageId="3" pageNumber="374">FIGURES 2–7</emphasis>
|
||
.
|
||
<emphasis id="2EB2EAAEFF8FFFCBFEAF2781FD86BBA1" box="[324,571,1949,1971]" italics="true" pageId="3" pageNumber="374">
|
||
<taxonomicName id="DBC64D3FFF8FFFCBFEAF2781FD8ABBA1" box="[324,567,1949,1971]" class="Diplopoda" family="Rhachodesmidae" genus="Dobrodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="mirabilis">Dobrodesmus mirabilis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="2EB2EAAEFF8FFFCBFDA92781FDC7BBA1" bold="true" box="[578,634,1949,1971]" pageId="3" pageNumber="374">Fig 2</emphasis>
|
||
, collum.
|
||
<emphasis id="2EB2EAAEFF8FFFCBFD352781FCABBBA1" bold="true" box="[734,790,1949,1971]" pageId="3" pageNumber="374">Fig 3</emphasis>
|
||
, sensilla from sixth antennal article.
|
||
<emphasis id="2EB2EAAEFF8FFFCBFB732781FB6CBBA1" bold="true" box="[1176,1233,1949,1971]" pageId="3" pageNumber="374">Fig 4</emphasis>
|
||
, midbody segment, dorsal view.
|
||
<emphasis id="2EB2EAAEFF8FFFCBFEF427A0FEE4BBC0" bold="true" box="[287,345,1980,2002]" pageId="3" pageNumber="374">Fig 5</emphasis>
|
||
, midbody segment, ventral view.
|
||
<emphasis id="2EB2EAAEFF8FFFCBFD2827A0FCBFBBC0" bold="true" box="[707,770,1980,2002]" pageId="3" pageNumber="374">Fig 6,</emphasis>
|
||
anterior right corner of midbody segment, dorsal view.
|
||
<emphasis id="2EB2EAAEFF8FFFCBFAB727A0FA28BBC0" bold="true" box="[1372,1429,1980,2002]" pageId="3" pageNumber="374">Fig 7</emphasis>
|
||
, posterior right corner of nonporiferous midbody segment, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="48B96634FF88FFCCFF7C250EFE1FB954" httpUri="https://zenodo.org/record/257273/files/figure.png" pageId="4" pageNumber="375" targetBox="[351,1228,183,1269]" targetPageId="4">
|
||
<paragraph id="1C7936BCFF88FFCCFF7C250EFE1FB954" blockId="4.[151,1436,1298,1350]" pageId="4" pageNumber="375">
|
||
<emphasis id="2EB2EAAEFF88FFCCFF7C250EFEFCB93A" bold="true" box="[151,321,1298,1320]" pageId="4" pageNumber="375">FIGURES 8–10</emphasis>
|
||
.
|
||
<emphasis id="2EB2EAAEFF88FFCCFEA4250EFDF8B93A" box="[335,581,1298,1320]" italics="true" pageId="4" pageNumber="375">
|
||
<taxonomicName id="DBC64D3FFF88FFCCFEA4250EFDFCB93A" box="[335,577,1298,1320]" class="Diplopoda" family="Rhachodesmidae" genus="Dobrodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="mirabilis">Dobrodesmus mirabilis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="2EB2EAAEFF88FFCCFDA0250EFD3CB93A" bold="true" box="[587,641,1298,1320]" pageId="4" pageNumber="375">Fig 8</emphasis>
|
||
, Ozopore, lateral view.
|
||
<emphasis id="2EB2EAAEFF88FFCCFC93250EFC12B93A" bold="true" box="[888,943,1298,1320]" pageId="4" pageNumber="375">Fig 9</emphasis>
|
||
, intercalary microscutes of metazonital dorsum, dorsal view.
|
||
<emphasis id="2EB2EAAEFF88FFCCFEFC252CFEE7B954" bold="true" box="[279,346,1328,1350]" pageId="4" pageNumber="375">Fig 10</emphasis>
|
||
, same.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1C7936BCFF88FFCCFF2C2571FA8CBB24" blockId="4.[151,1436,1389,2026]" pageId="4" pageNumber="375">
|
||
<emphasis id="2EB2EAAEFF88FFCCFF2C2571FEACB994" bold="true" box="[199,273,1389,1414]" pageId="4" pageNumber="375">Notes.</emphasis>
|
||
Observations using optical microscopy on the anatomy of the male
|
||
<typeStatus id="C37D881EFF88FFCCFBE72571FBD3B994" box="[1036,1134,1389,1414]" pageId="4" pageNumber="375" type="holotype">holotype</typeStatus>
|
||
were made difficult by the transparency (intra- and inter-ring muscles and digestive tract are clearly visible) and brittleness of the cuticle, so subsequent observations may alter some details. Certain features were clarified using the
|
||
<typeStatus id="C37D881EFF88FFCCFB2125ABFA97B9DD" box="[1226,1322,1463,1487]" pageId="4" pageNumber="375" type="paratype">paratype</typeStatus>
|
||
male for scanning electron microscopy. It was particularly difficult to detect the ozopores and while their distribution appeared to fit the “normal” polydesmidan pattern (except for their continuation beyond ring 19) we are not entirely sure of this; on one specimen of 24 rings, ozopores appeared not to begin until ring seven. It may be that the chemical defense system has been reduced due to relaxed predator pressure in the subterranean environment. The spiracles are so small as to be undetectable with light microscopy (
|
||
<figureCitation id="84FD2A39FF88FFCCFC5E2675FBBCBA90" box="[949,1025,1641,1666]" captionStart="FIGURES 12, 13" captionStartId="6.[151,264,1202,1224]" captionTargetBox="[340,1246,337,1180]" captionTargetId="figure@6.[340,1247,337,1181]" captionTargetPageId="6" captionText="FIGURES 12, 13. Dobrodesmus mirabilis. Fig 12, ventral view of right lateral part of midbody segment, showing positions of spiracles. Abbreviations: as, anterior spiracle; ps, posterior spiracle. Fig 13. Posterior spiracle." httpUri="https://zenodo.org/record/257275/files/figure.png" pageId="4" pageNumber="375">Fig 13</figureCitation>
|
||
); their position was only determined by careful searching using scanning electron microscopy. The positions of the spiracles on each ring are advanced anteriorly so that the posterior spiracles are between the legpairs of each side, and the anterior spiracles are near the posterior margin of the prozonite (
|
||
<figureCitation id="84FD2A39FF88FFCCFDD026C9FD32BAFC" box="[571,655,1749,1774]" captionStart="FIGURES 12, 13" captionStartId="6.[151,264,1202,1224]" captionTargetBox="[340,1246,337,1180]" captionTargetId="figure@6.[340,1247,337,1181]" captionTargetPageId="6" captionText="FIGURES 12, 13. Dobrodesmus mirabilis. Fig 12, ventral view of right lateral part of midbody segment, showing positions of spiracles. Abbreviations: as, anterior spiracle; ps, posterior spiracle. Fig 13. Posterior spiracle." httpUri="https://zenodo.org/record/257275/files/figure.png" pageId="4" pageNumber="375">Fig 12</figureCitation>
|
||
). Reduced or apparently absent spiracles occur elsewhere in the
|
||
<taxonomicName id="DBC64D3FFF88FFCCFF7C26E5FE96BB00" box="[151,299,1785,1810]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="375" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
|
||
.
|
||
<bibRefCitation id="78574B4DFF88FFCCFEDE26E5FE54BB00" author="Mesibov" box="[309,489,1785,1810]" pageId="4" pageNumber="375" refString="Mesibov, R. (2009) Revision of Agathodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae). ZooKeys, 12, 87 - 110. http: // dx. doi. org / 10.3897 / zookeys. 12.206" type="journal article" year="2009">Mesibov (2009)</bibRefCitation>
|
||
at first thought the Australian haplodesmid
|
||
<taxonomicName id="DBC64D3FFF88FFCCFC3E26E6FB61BB00" authority="Silvestri" authorityName="Silvestri" box="[981,1244,1785,1810]" class="Diplopoda" family="Haplodesmidae" genus="Agathodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2EB2EAAEFF88FFCCFC3E26E6FBC4BB00" box="[981,1145,1786,1810]" italics="true" pageId="4" pageNumber="375">Agathodesmus</emphasis>
|
||
Silvestri
|
||
</taxonomicName>
|
||
lacked spiracles, but later examination with scanning electron microscopy revealed them as very reduced (
|
||
<bibRefCitation id="78574B4DFF88FFCCFB962701FA98BB24" author="Mesibov" box="[1149,1317,1821,1846]" pageId="4" pageNumber="375" refString="Mesibov, R. (2013) New species of Agathodesmus Silvestri, 1910 from Australia (Diplopoda, Polydesmida, Haplodesmidae). ZooKeys, 325, 33 - 64. http: // dx. doi. org / 10.3897 / zookeys. 325.5932" type="journal article" year="2013">Mesibov 2013</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1C7936BCFF88FFCCFF7C275DFC64BBF8" blockId="4.[151,1436,1389,2026]" pageId="4" pageNumber="375">
|
||
Spiracles could not be detected in species of the haplodesmid genus
|
||
<taxonomicName id="DBC64D3FFF88FFCCFC7D275EFAD5BB48" authority="Silvestri (Mesibov 2012)" authorityName="Silvestri (Mesibov" authorityYear="2012" box="[918,1384,1857,1882]" class="Diplopoda" family="Haplodesmidae" genus="Prosopodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2EB2EAAEFF88FFCCFC7D275EFBFBBB48" box="[918,1094,1858,1882]" italics="true" pageId="4" pageNumber="375">Prosopodesmus</emphasis>
|
||
Silvestri (
|
||
<bibRefCitation id="78574B4DFF88FFCCFB52275DFADDBB48" author="Mesibov" box="[1209,1376,1857,1882]" pageId="4" pageNumber="375" refString="Mesibov, R. (2012) New species of Prosopodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae) from Queensland, Australia. ZooKeys, 190, 33 - 54. http: // dx. doi. org / 10.3897 / zookeys. 190.3276" type="journal article" year="2012">Mesibov 2012</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and Golovatch & VandenSpiegel (2014, 2015) failed to find spiracles in
|
||
<taxonomicName id="DBC64D3FFF88FFCCFC2A277BFA21BB6C" authority="Golovatch & VandenSpiegel" authorityName="Golovatch & VandenSpiegel" box="[961,1436,1893,1918]" class="Diplopoda" family="Haplodesmidae" genus="Koponenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2EB2EAAEFF88FFCCFC2A277BFBFABB6C" box="[961,1095,1895,1918]" italics="true" pageId="4" pageNumber="375">Koponenius</emphasis>
|
||
Golovatch & VandenSpiegel
|
||
</taxonomicName>
|
||
species, also haplodesmids. It seems doubtful that openings to the tracheal system could be entirely lacking in a millipede of any size, but their position may be anomalous or they may be so small and misshapen (as in the case of
|
||
<taxonomicName id="DBC64D3FFF88FFCCFF7C27CEFEA3BBF8" box="[151,286,2002,2026]" class="Diplopoda" family="Rhachodesmidae" genus="Dobrodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="mirabilis">
|
||
<emphasis id="2EB2EAAEFF88FFCCFF7C27CEFEA3BBF8" box="[151,286,2002,2026]" italics="true" pageId="4" pageNumber="375">D. mirabilis</emphasis>
|
||
</taxonomicName>
|
||
) as to be overlooked even with scanning electron microscopy.
|
||
</paragraph>
|
||
<caption id="48B96634FF89FFCDFF7C246FFD73B8BA" httpUri="https://zenodo.org/record/257274/files/figure.png" pageId="5" pageNumber="376" targetBox="[340,1246,193,1117]" targetPageId="5">
|
||
<paragraph id="1C7936BCFF89FFCDFF7C246FFD73B8BA" blockId="5.[151,1435,1139,1192]" pageId="5" pageNumber="376">
|
||
<emphasis id="2EB2EAAEFF89FFCDFF7C246FFEA1B89B" bold="true" box="[151,284,1139,1161]" pageId="5" pageNumber="376">FIGURE 11</emphasis>
|
||
.
|
||
<emphasis id="2EB2EAAEFF89FFCDFEC7246FFD98B89B" box="[300,549,1139,1161]" italics="true" pageId="5" pageNumber="376">
|
||
<taxonomicName id="DBC64D3FFF89FFCDFEC7246FFD9CB89B" box="[300,545,1139,1161]" class="Diplopoda" family="Rhachodesmidae" genus="Dobrodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="mirabilis">Dobrodesmus mirabilis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
posterior end, dorsolateral view. Abbreviations: s, spinneret;
|
||
<emphasis id="2EB2EAAEFF89FFCDFB5E246FFB6CB89B" bold="true" box="[1205,1233,1139,1161]" pageId="5" pageNumber="376">dp</emphasis>
|
||
, dorsal division of paraproct;
|
||
<emphasis id="2EB2EAAEFF89FFCDFEEF248EFEA2B8BA" bold="true" box="[260,287,1170,1192]" pageId="5" pageNumber="376">vp</emphasis>
|
||
, ventral division of paraproct.;
|
||
<emphasis id="2EB2EAAEFF89FFCDFDB5248EFDDBB8BA" bold="true" box="[606,614,1170,1192]" pageId="5" pageNumber="376">t</emphasis>
|
||
, epiproct.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1C7936BCFF89FFCDFF2C24C8FB55B9FB" blockId="5.[151,1437,1236,2017]" pageId="5" pageNumber="376">
|
||
Spinnerets in the
|
||
<taxonomicName id="DBC64D3FFF89FFCDFE6224C8FDA7B8FF" box="[393,538,1236,1261]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
|
||
are typically four in number, arranged in a quadrangle (
|
||
<bibRefCitation id="78574B4DFF89FFCDFB6C24C8FAB1B8FF" author="Shear" box="[1159,1292,1236,1261]" pageId="5" pageNumber="376" refString="Shear, W. A. (2008) Spinnerets in the milliped order Polydesmida, and the phylogenetic significance of spinnerets in millipeds (Diplopoda). International Journal of Myriapodology, 1, 123 - 146. http: // dx. doi. org / 10.1163 / 187525408 X 395904" type="journal article" year="2008">Shear 2008</bibRefCitation>
|
||
). In contrast, two spinnerets are typical of the orders
|
||
<taxonomicName id="DBC64D3FFF89FFCDFD9B24E4FD4EB903" box="[624,755,1272,1297]" class="Diplopoda" kingdom="Animalia" order="Callipodida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="order">Callipodida</taxonomicName>
|
||
and Chordeumatida, and at least one species in the order
|
||
<taxonomicName id="DBC64D3FFF89FFCDFF7C2500FE9BB927" box="[151,294,1308,1333]" class="Diplopoda" kingdom="Animalia" order="Stemmiulida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="order">Stemmiulida</taxonomicName>
|
||
(
|
||
<bibRefCitation id="78574B4DFF89FFCDFED62500FE76B927" author="Shear" box="[317,459,1308,1333]" pageId="5" pageNumber="376" refString="Shear, W. A. (2008) Spinnerets in the milliped order Polydesmida, and the phylogenetic significance of spinnerets in millipeds (Diplopoda). International Journal of Myriapodology, 1, 123 - 146. http: // dx. doi. org / 10.1163 / 187525408 X 395904" type="journal article" year="2008">Shear 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="78574B4DFF89FFCDFE342500FD64B927" author="Mauries" box="[479,729,1308,1333]" pageId="5" pageNumber="376" refString="Mauries, J. - P., Golovatch, S. & Geoffroy, J. - J. (2010) A new genus and species of the order Stemmiulida from Vietnam. Arthropoda Selecta, 19, 73 - 80." type="journal article" year="2010">
|
||
Mauriès
|
||
<emphasis id="2EB2EAAEFF89FFCDFDA02501FD31B927" box="[587,652,1309,1333]" italics="true" pageId="5" pageNumber="376">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
). The spinnerets of
|
||
<taxonomicName id="DBC64D3FFF89FFCDFC312501FBD7B927" box="[986,1130,1309,1333]" class="Diplopoda" family="Rhachodesmidae" genus="Dobrodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="mirabilis">
|
||
<emphasis id="2EB2EAAEFF89FFCDFC312501FBD7B927" box="[986,1130,1309,1333]" italics="true" pageId="5" pageNumber="376">D. mirabilis</emphasis>
|
||
</taxonomicName>
|
||
do not resemble typical polydesmidan spinnerets (
|
||
<figureCitation id="84FD2A39FF89FFCDFE57255CFDB4B94B" box="[444,521,1344,1369]" captionStart="FIGURES 14 – 16" captionStartId="7.[151,264,1268,1290]" captionTargetBox="[340,1246,193,1245]" captionTargetId="figure@7.[340,1247,193,1246]" captionTargetPageId="7" captionText="FIGURES 14 – 16. Fig 14, spinnerets of Dobrodesmus mirabilis. Fig 15, spinnerets of Tetracion jonesi Hoffman 1956 (Callipodida, Abacionidae). Abbreviations: sl, extended aperture of spinneret; dp, dorsal division of paraproct; vp, ventral division of paraproct. Fig 16, gonopods of Dobrodesmus mirabilis, ventral view. Abbreviations: cx, gonopod coxa; f, flagellum; p 1, sheathing process of coxa; p 2, triangular process of telopodite; tel, telopodite of gonopo; a?, possible acropodite." httpUri="https://zenodo.org/record/257276/files/figure.png" pageId="5" pageNumber="376">Fig 14</figureCitation>
|
||
), but their long basal sleeves and setal shafts are very similar to the spinnerets of callipodidans (
|
||
<figureCitation id="84FD2A39FF89FFCDFED72578FE36B96F" box="[316,395,1380,1405]" captionStart="FIGURES 14 – 16" captionStartId="7.[151,264,1268,1290]" captionTargetBox="[340,1246,193,1245]" captionTargetId="figure@7.[340,1247,193,1246]" captionTargetPageId="7" captionText="FIGURES 14 – 16. Fig 14, spinnerets of Dobrodesmus mirabilis. Fig 15, spinnerets of Tetracion jonesi Hoffman 1956 (Callipodida, Abacionidae). Abbreviations: sl, extended aperture of spinneret; dp, dorsal division of paraproct; vp, ventral division of paraproct. Fig 16, gonopods of Dobrodesmus mirabilis, ventral view. Abbreviations: cx, gonopod coxa; f, flagellum; p 1, sheathing process of coxa; p 2, triangular process of telopodite; tel, telopodite of gonopo; a?, possible acropodite." httpUri="https://zenodo.org/record/257276/files/figure.png" pageId="5" pageNumber="376">Fig 15</figureCitation>
|
||
; further illustrations in
|
||
<bibRefCitation id="78574B4DFF89FFCDFD7C2578FCA1B96F" author="Shear" box="[663,796,1380,1405]" pageId="5" pageNumber="376" refString="Shear, W. A. (2008) Spinnerets in the milliped order Polydesmida, and the phylogenetic significance of spinnerets in millipeds (Diplopoda). International Journal of Myriapodology, 1, 123 - 146. http: // dx. doi. org / 10.1163 / 187525408 X 395904" type="journal article" year="2008">Shear 2008</bibRefCitation>
|
||
).
|
||
<taxonomicName id="DBC64D3FFF89FFCDFCDB2579FA25B96F" authority="Mauries, Golovatch & Geoffroy" authorityName="Mauries, Golovatch & Geoffroy" box="[816,1432,1380,1405]" class="Diplopoda" family="Stemmiulidae" genus="Eostemmiulus" kingdom="Animalia" order="Stemmiulida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="caecus">
|
||
<emphasis id="2EB2EAAEFF89FFCDFCDB2579FB9FB96E" box="[816,1058,1381,1405]" italics="true" pageId="5" pageNumber="376">Eostemmiulus caecus</emphasis>
|
||
Mauriès, Golovatch & Geoffroy
|
||
</taxonomicName>
|
||
, arguably the most basal member of the
|
||
<taxonomicName id="DBC64D3FFF89FFCDFDB72594FD53B9B3" box="[604,750,1416,1441]" class="Diplopoda" kingdom="Animalia" order="Stemmiulida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="order">Stemmiulida</taxonomicName>
|
||
, also has two spinnerets, in contrast to other members of its order in which the number ranges from four to eight, and those spinnerets resemble polydesmidan spinnerets in having a short, basal sleeve instead of arising from unsocketed mounds as is usual in
|
||
<taxonomicName id="DBC64D3FFF89FFCDFBB925CCFB58B9FB" box="[1106,1253,1488,1513]" class="Diplopoda" kingdom="Animalia" order="Stemmiulida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="order">Stemmiulida</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1C7936BCFF89FFCDFF2C25E8FAECBAF7" blockId="5.[151,1437,1236,2017]" pageId="5" pageNumber="376">
|
||
The dividing ridge on the paraprocts (
|
||
<figureCitation id="84FD2A39FF89FFCDFD9B25E8FD04BA1F" box="[624,697,1524,1549]" captionStart="FIGURE 11" captionStartId="5.[151,250,1139,1161]" captionTargetBox="[340,1246,193,1117]" captionTargetId="figure@5.[340,1247,193,1118]" captionTargetPageId="5" captionText="FIGURE 11. Dobrodesmus mirabilis, posterior end, dorsolateral view. Abbreviations: s, spinneret; dp, dorsal division of paraproct; vp, ventral division of paraproct.; t, epiproct." httpUri="https://zenodo.org/record/257274/files/figure.png" pageId="5" pageNumber="376">Fig 11</figureCitation>
|
||
) seems to be unique in the
|
||
<taxonomicName id="DBC64D3FFF89FFCDFC0425E8FB3DBA1F" box="[1007,1152,1524,1549]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
|
||
and may represent either an existing division of the paraproct into two sclerites, or a recent fusion of an ancestral two sclerites. The dividing ridge can only be seen in specimens with distended anal regions; in other specimens the ridge is tightly appressed to the margin of the epiproct. In scanning electron micrographs, the ridge appears as a possible suture with flexible cuticle between the sclerites. Divided paraprocts are characteristic of at least some callipodidans. A pair of setae on each of the paraprocts appears to be part of the polydesmidan groundplan, but these setae are absent in
|
||
<emphasis id="2EB2EAAEFF89FFCDFAE526B5FA21BAD3" box="[1294,1436,1705,1729]" italics="true" pageId="5" pageNumber="376">
|
||
<taxonomicName id="DBC64D3FFF89FFCDFAE526B5FA25BAD3" box="[1294,1432,1705,1729]" class="Diplopoda" family="Rhachodesmidae" genus="Dobrodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="mirabilis">D. mirabilis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and instead, as in nematophorans, the paraprocts bear numerous fine setae without any regular arrangement.
|
||
</paragraph>
|
||
<paragraph id="1C7936BCFF89FFCEFF2C26ECFF5DBD0F" blockId="5.[151,1437,1236,2017]" lastBlockId="6.[151,1436,151,285]" lastPageId="6" lastPageNumber="377" pageId="5" pageNumber="376">
|
||
<bibRefCitation id="78574B4DFF89FFCDFF2C26ECFE5BBB1B" author="Akkari" box="[199,486,1776,1801]" pageId="5" pageNumber="376" refString="Akkari, N. & Enghoff, H. (2011) On some surface structures of potential taxonomic importance in families of the suborders Polydesmidea and Dalodesmidea (Polydesmida, Diplopoda). ZooKeys, 156, 1 - 24. http: // dx. doi. org / 10.3897 / zookeys. 156.2134" type="journal article" year="2011">Akkari & Enghoff (2011)</bibRefCitation>
|
||
illustrated intercalary microscutes for a number of polydesmidans. The microscutes of
|
||
<taxonomicName id="DBC64D3FFF89FFCDFF5F2709FE86BB3F" box="[180,315,1813,1837]" class="Diplopoda" family="Rhachodesmidae" genus="Dobrodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="mirabilis">
|
||
<emphasis id="2EB2EAAEFF89FFCDFF5F2709FE86BB3F" box="[180,315,1813,1837]" italics="true" pageId="5" pageNumber="376">D. mirabilis</emphasis>
|
||
</taxonomicName>
|
||
differ from those illustrated in their regularly wrinkled surface (
|
||
<figureCitation id="84FD2A39FF89FFCDFBEC2708FBCABB3F" box="[1031,1143,1812,1837]" captionStart="FIGURES 8 – 10" captionStartId="4.[151,264,1298,1320]" captionTargetBox="[351,1228,183,1269]" captionTargetId="figure@4.[351,1229,183,1270]" captionTargetPageId="4" captionText="FIGURES 8 – 10. Dobrodesmus mirabilis. Fig 8, Ozopore, lateral view. Fig 9, intercalary microscutes of metazonital dorsum, dorsal view. Fig 10, same." httpUri="https://zenodo.org/record/257273/files/figure.png" pageId="5" pageNumber="376">Figs 9, 10</figureCitation>
|
||
) rather than being smooth and flat; the wrinkling suggests a thin cuticle or membranous surface.
|
||
<bibRefCitation id="78574B4DFF89FFCDFC4C2724FB7EBB43" author="Akkari" box="[935,1219,1848,1873]" pageId="5" pageNumber="376" refString="Akkari, N. & Enghoff, H. (2011) On some surface structures of potential taxonomic importance in families of the suborders Polydesmidea and Dalodesmidea (Polydesmida, Diplopoda). ZooKeys, 156, 1 - 24. http: // dx. doi. org / 10.3897 / zookeys. 156.2134" type="journal article" year="2011">Akkari & Enghoff (2011)</bibRefCitation>
|
||
found the scutes in members of the families
|
||
<taxonomicName id="DBC64D3FFF89FFCDFE0B2740FD3DBB67" box="[480,640,1884,1909]" class="Diplopoda" family="Polydesmidae" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="family">Polydesmidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="DBC64D3FFF89FFCDFD732740FC2FBB67" box="[664,914,1884,1909]" class="Diplopoda" family="Macrosternodesmidae" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="family">Macrosternodesmidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="DBC64D3FFF89FFCDFC412740FB2FBB67" box="[938,1170,1884,1909]" class="Diplopoda" family="Trichopolydesmidae" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="family">Trichopolydesmidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="DBC64D3FFF89FFCDFB412740FA25BB67" box="[1194,1432,1884,1909]" class="Diplopoda" family="Fuhrmannodesmidae" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="family">Fuhrmannodesmidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="DBC64D3FFF89FFCDFF7C279CFE8CBB8B" box="[151,305,1920,1945]" class="Diplopoda" family="Opisotretidae" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="family">Opisotretidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="DBC64D3FFF89FFCDFEA8279CFDBBBB8B" box="[323,518,1920,1945]" class="Diplopoda" family="Nearctodesmidae" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="family">Nearctodesmidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="DBC64D3FFF89FFCDFDA7279CFD52BB8B" box="[588,751,1920,1945]" class="Diplopoda" family="Dalodesmidae" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="family">Dalodesmidae</taxonomicName>
|
||
, but not in species of
|
||
<taxonomicName id="DBC64D3FFF89FFCDFBFA279CFB76BB8B" box="[1041,1227,1920,1945]" class="Diplopoda" family="Ammodesmidae" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="family">Ammodesmidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="DBC64D3FFF89FFCDFB35279CFA25BB8B" box="[1246,1432,1920,1945]" class="Diplopoda" family="Cryptodesmidae" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="family">Cryptodesmidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="DBC64D3FFF89FFCDFF7C27B9FEF9BBAC" box="[151,324,1957,1982]" class="Diplopoda" family="Cyrtodesmidae" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="family">Cyrtodesmidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="DBC64D3FFF89FFCDFEB927B9FDBEBBAC" box="[338,515,1957,1982]" class="Diplopoda" family="Haplodesmidae" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="family">Haplodesmidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="DBC64D3FFF89FFCDFDF927B9FD76BBAC" box="[530,715,1957,1982]" class="Diplopoda" family="Oniscodesmidae" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="family">Oniscodesmidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="DBC64D3FFF89FFCDFCE227B9FC04BBAC" box="[777,953,1957,1982]" class="Diplopoda" family="Pyrgodesmidae" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="family">Pyrgodesmidae</taxonomicName>
|
||
. Intercalary microscutes are of no known functional significance, though it is worth noting that many of the members of the families that lack them cover themselves with debris and soil in life. Reboleira & Enghoff (2015) illustrate, but do not discuss, intercalary microscutes in the pleurotergal cuticle of the dorypetalid callipodidan
|
||
<taxonomicName id="DBC64D3FFF8AFFCEFBE420A1FA28BCC7" authority="Verhoeff" authorityName="Verhoeff" box="[1039,1429,188,213]" class="Diplopoda" family="Dorypetalidae" genus="Lusitanipus" kingdom="Animalia" order="Callipodida" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="alternans">
|
||
<emphasis id="2EB2EAAEFF8AFFCEFBE420A1FAB3BCC6" box="[1039,1294,188,212]" italics="true" pageId="6" pageNumber="377">Lusitanipus alternans</emphasis>
|
||
(Verhoeff)
|
||
</taxonomicName>
|
||
. Microscutes have also been detected in two spirostreptidan families (Enghoff 2014, 2016;
|
||
<bibRefCitation id="78574B4DFF8AFFCEFB4920C3FF69BD0F" author="Enghoff" pageId="6" pageNumber="377" refString="Enghoff, H. & Fredriksen, S. B. (2015) A mountain of millipedes II. The genus Aquattuor Fredriksen, 2013 - five new species from the Udzingawa Mountains and one from Mt. Kilamanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy, 150, 1 - 25." type="journal article" year="2015">Enghoff & Fredriksen 2015</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="48B96634FF8AFFCEFF7C24AEFBE9B8F5" httpUri="https://zenodo.org/record/257275/files/figure.png" pageId="6" pageNumber="377" targetBox="[340,1246,337,1180]" targetPageId="6">
|
||
<paragraph id="1C7936BCFF8AFFCEFF7C24AEFBE9B8F5" blockId="6.[151,1436,1202,1255]" pageId="6" pageNumber="377">
|
||
<emphasis id="2EB2EAAEFF8AFFCEFF7C24AEFEF1B8DA" bold="true" box="[151,332,1202,1224]" pageId="6" pageNumber="377">FIGURES 12, 13</emphasis>
|
||
.
|
||
<emphasis id="2EB2EAAEFF8AFFCEFEB124AEFDF3B8DA" box="[346,590,1202,1224]" italics="true" pageId="6" pageNumber="377">
|
||
<taxonomicName id="DBC64D3FFF8AFFCEFEB124AEFDF6B8DA" box="[346,587,1202,1224]" class="Diplopoda" family="Rhachodesmidae" genus="Dobrodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="mirabilis">Dobrodesmus mirabilis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="2EB2EAAEFF8AFFCEFDBF24AEFD2BB8DA" bold="true" box="[596,662,1202,1224]" pageId="6" pageNumber="377">Fig 12</emphasis>
|
||
, ventral view of right lateral part of midbody segment, showing positions of spiracles. Abbreviations:
|
||
<emphasis id="2EB2EAAEFF8AFFCEFE7224CDFE0DB8F5" bold="true" box="[409,432,1233,1255]" pageId="6" pageNumber="377">as</emphasis>
|
||
, anterior spiracle;
|
||
<emphasis id="2EB2EAAEFF8AFFCEFD8624CDFD38B8F5" bold="true" box="[621,645,1233,1255]" pageId="6" pageNumber="377">ps</emphasis>
|
||
, posterior spiracle.
|
||
<emphasis id="2EB2EAAEFF8AFFCEFCA624CDFC2DB8F5" bold="true" box="[845,912,1233,1255]" pageId="6" pageNumber="377">Fig 13</emphasis>
|
||
. Posterior spiracle.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1C7936BCFF8AFFCEFF2C250FFA8EB966" blockId="6.[151,1437,1299,1684]" pageId="6" pageNumber="377">
|
||
<emphasis id="2EB2EAAEFF8AFFCEFF2C250FFD92B93E" bold="true" box="[199,559,1299,1324]" pageId="6" pageNumber="377">
|
||
Anamorphosis of
|
||
<taxonomicName id="DBC64D3FFF8AFFCEFE752508FD94B93E" box="[414,553,1299,1324]" class="Diplopoda" family="Rhachodesmidae" genus="Dobrodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="mirabilis">
|
||
<emphasis id="2EB2EAAEFF8AFFCEFE752508FD94B93E" bold="true" box="[414,553,1299,1324]" italics="true" pageId="6" pageNumber="377">D. mirabilis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Of the 24 specimens of
|
||
<taxonomicName id="DBC64D3FFF8AFFCEFCA02508FC69B939" box="[843,980,1299,1323]" class="Diplopoda" family="Rhachodesmidae" genus="Dobrodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="mirabilis">
|
||
<emphasis id="2EB2EAAEFF8AFFCEFCA02508FC69B939" box="[843,980,1299,1323]" italics="true" pageId="6" pageNumber="377">D. mirabilis</emphasis>
|
||
</taxonomicName>
|
||
so far collected, only two, the holotype and paratype, are mature, and one other (ISLA 3638) was a juvenile male, with gonopod primordia on ring seven. The other juvenile specimens are either females or are at a stadium too early to show gonopod primordia.
|
||
</paragraph>
|
||
<paragraph id="1C7936BCFF8AFFCEFF2C2563FCD8BA16" blockId="6.[151,1437,1299,1684]" pageId="6" pageNumber="377">
|
||
The weak sclerotization and transparency of the juvenile specimens made it difficult to count rings, and some of our data do not correlate with the anamorphosis tables for other
|
||
<taxonomicName id="DBC64D3FFF8AFFCEFC0F25BFFBC5B9AE" box="[996,1144,1443,1468]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="377" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
|
||
. Nevertheless, since the anamorphosis of
|
||
<taxonomicName id="DBC64D3FFF8AFFCEFE8A25D4FE50B9CD" box="[353,493,1479,1503]" class="Diplopoda" family="Rhachodesmidae" genus="Dobrodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="mirabilis">
|
||
<emphasis id="2EB2EAAEFF8AFFCEFE8A25D4FE50B9CD" box="[353,493,1479,1503]" italics="true" pageId="6" pageNumber="377">D. mirabilis</emphasis>
|
||
</taxonomicName>
|
||
is probably atypical in any case, we present our data in the table below as we obtained it. In this table, ring numbers do not include the telson.
|
||
</paragraph>
|
||
<paragraph id="1C7936BCFF8AFFCEFF2C2613FAE4BA81" blockId="6.[151,1437,1299,1684]" pageId="6" pageNumber="377">
|
||
If the anamorphosis of this species proceeds as for
|
||
<taxonomicName id="DBC64D3FFF8AFFCEFD16260CFB18BA3A" authority="(Gray) (Miley 1927)" authorityName="(Gray) (Miley" authorityYear="1927" box="[765,1189,1551,1576]" class="Diplopoda" family="Aphelidesmidae" genus="Euryurus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="leachii">
|
||
<emphasis id="2EB2EAAEFF8AFFCEFD16260CFC04BA35" box="[765,953,1551,1575]" italics="true" pageId="6" pageNumber="377">Euryurus leachii</emphasis>
|
||
(Gray) (
|
||
<bibRefCitation id="78574B4DFF8AFFCEFBFC2613FB20BA3A" author="Miley" box="[1047,1181,1551,1576]" pageId="6" pageNumber="377" refString="Miley, H. H. (1927) Development of the male gonopods and life history studies of a polydesmid millipede. Ohio Journal of Science, 27, 25 - 43." type="journal article" year="1927">Miley 1927</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, specimens with eight or nine rings represent stadium II. Specimens with 11 rings are likely to be in stadium III, those with 14–16 rings in either stadium IV or V, and those with 17 rings in stadium VI. Polydesmidans with 19+1 (20) rings mature in stadium VIII, but the presence of juveniles with 24–27 rings indicates that molting continues in
|
||
<emphasis id="2EB2EAAEFF8AFFCEFB222660FAE4BA81" box="[1225,1369,1659,1683]" italics="true" pageId="6" pageNumber="377">
|
||
<taxonomicName id="DBC64D3FFF8AFFCEFB222660FAE8BA81" box="[1225,1365,1659,1683]" class="Diplopoda" family="Rhachodesmidae" genus="Dobrodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="mirabilis">D. mirabilis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |