285 lines
56 KiB
XML
285 lines
56 KiB
XML
<document id="519C2EBF8C91DB40CE5A386EC98DEA4A" ID-DOI="10.5281/zenodo.173868" ID-GBIF-Dataset="3057cbcd-031a-4a23-b2df-c3f768da9262" ID-ISSN="11755326" ID-Zenodo-Dep="173868" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459785867585" checkinUser="plazi" docAuthor="González, Guillermo & Vandenberg, Natalia J." docDate="2006" docId="527187C41009BC640650F9FF825FFD38" docLanguage="en" docName="zt01311p050.pdf.imd" docOrigin="Zootaxa 1311" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Cycloneda germainii González & Vandenberg 2006" docType="key" docVersion="11" lastPageNumber="20" masterDocId="AE48FFBC100CBC630758FFE38774FFD6" masterDocTitle="Review of lady beetles in the Cycloneda germainii species complex (Coleoptera; Coccinellidae: Coccinellinae: Coccinellini) with descriptions of new and unusual species from Chile and surrounding countries" masterLastPageNumber="50" masterPageNumber="13" pageNumber="18" updateTime="1698218499776" updateUser="plazi">
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<mods:titleInfo id="EE35B0EB41DF5FEEC28B90423C84F560">
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<mods:title id="67CCF7909FFE15CBDE9694AD136622F7">Review of lady beetles in the Cycloneda germainii species complex (Coleoptera; Coccinellidae: Coccinellinae: Coccinellini) with descriptions of new and unusual species from Chile and surrounding countries</mods:title>
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<mods:namePart id="82E6478C5E31C1005971849C148D9CD8">González, Guillermo</mods:namePart>
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<mods:roleTerm id="667AA6E0FA0E5F905B43C5D6CB6BF3C2">Author</mods:roleTerm>
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<mods:namePart id="DF58B6C9510663974EA039A646A9C68D">Vandenberg, Natalia J.</mods:namePart>
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<mods:typeOfResource id="E47C27A655B3ACBEBA15AFFEF193284F">text</mods:typeOfResource>
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<mods:title id="D2DDBDA7F88BE92542B6739A4682DF08">Zootaxa</mods:title>
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<mods:part id="F4ACE7CC228AA12465D3A78ECAD2CC2D">
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<mods:date id="F23A1C647985E061F7BCF6A0A5AD89F1">2006</mods:date>
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<mods:number id="C1A0E912B662933975436676DCDC2011">2006-09-11</mods:number>
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<mods:detail id="303BFF321B7D9A68F3CD5AAB70BA65C5" type="volume">
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<mods:number id="BE4BA38A89555046AA7C54B40F75A6CB">1311</mods:number>
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<mods:classification id="BB204E5654D240DA05EBDAAED525BBBE">journal article</mods:classification>
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<mods:identifier id="620CA70119313B2FA15FE913E1E3D8D2" type="DOI">10.5281/zenodo.173868</mods:identifier>
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<treatment id="527187C41009BC640650F9FF825FFD38" ID-GBIF-Taxon="119349075" LSID="urn:lsid:plazi:treatment:527187C41009BC640650F9FF825FFD38" httpUri="http://treatment.plazi.org/id/527187C41009BC640650F9FF825FFD38" lastPageId="7" lastPageNumber="20" pageId="5" pageNumber="18">
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<subSubSection id="92C265591009BC660650F9FF84A2F9E0" box="[264,982,1564,1590]" pageId="5" pageNumber="18" type="nomenclature">
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<paragraph id="DA6736D21009BC660650F9FF84A2F9E0" blockId="5.[264,982,1564,1590]" box="[264,982,1564,1590]" pageId="5" pageNumber="18">
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<heading id="812F81BE1009BC660650F9FF84A2F9E0" bold="true" box="[264,982,1564,1590]" fontSize="11" level="1" pageId="5" pageNumber="18" reason="1">
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<emphasis id="E8ACEAC01009BC660650F9FF84A2F9E0" bold="true" box="[264,982,1564,1590]" pageId="5" pageNumber="18">
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Key to species in the
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<taxonomicName id="1DD84D511009BC660555F9FF847CF9E0" authorityName="González & Vandenberg" authorityYear="2006" baseAuthorityName="González & Vandenberg" baseAuthorityYear="2006" box="[525,776,1564,1590]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="germainii">
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<emphasis id="E8ACEAC01009BC660555F9FF847CF9E0" bold="true" box="[525,776,1564,1590]" italics="true" pageId="5" pageNumber="18">Cycloneda germainii</emphasis>
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</taxonomicName>
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species complex
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="92C265591009BC640650F984825FFD38" lastPageId="7" lastPageNumber="20" pageId="5" pageNumber="18" type="key">
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<keyStep id="612C2E771009BC650650F9848259FE28" lastPageId="6" lastPageNumber="19" pageId="5" pageNumber="18">
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<paragraph id="DA6736D21009BC660650F984825FF8A7" blockId="5.[264,1323,1639,1905]" pageId="5" pageNumber="18">
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<keyLead id="612995E71009BC660650F984825FF8A7" pageId="5" pageNumber="18">
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1. Form somewhat depressed, elongate elliptical; elytral epipleuron flat, ascending externally, visible in lateral view (
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<figureCitation id="42E32A571009BC6605B2F96C8433F97F" box="[746,839,1679,1705]" captionStart="FIGURE 3" captionStartId="15.[264,367,1565,1589]" captionTargetBox="[272,1316,323,1530]" captionTargetId="figure@15.[256,1331,310,1551]" captionTargetPageId="15" captionText="FIGURE 3. Morphology of Cycloneda species. Lateral views of bodies (elytral length standardized to facilitate comparison), A–G: A, C. lacrimosa González & Vandenberg, new species, paratype; B, C. disconsolata Vandenberg & González, new species, holotype; C, C. patagonica González & Vandenberg, new species, paratype; D, Cycloneda germainii (Crotch), representative specimen; E, C. eryngii (Mulsant), representative specimen; F, C. sicardi (Brèthes), specimen from Salta; G, C. boliviana (Mulsant), holotype. Pronotal dorsofrontal views, H–L: H, C. lacrimosa González & Vandenberg, new species, paratype; I, C. boliviana (Mulsant), holotype; J, C. sicardi (Brèthes), specimen from El Rincón, Catamarca; K, C. sicardi (Brèthes), specimen from Tacuil, Salta; L, C. sicardi (Brèthes), common aberration in much of range." httpUri="https://zenodo.org/record/173871/files/figure.png" pageId="5" pageNumber="18">Fig. 3C</figureCitation>
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); elytron narrow, parallelsided for much of length (
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<figureCitation id="42E32A571009BC660682F954851EF907" box="[474,618,1719,1745]" captionStart="FIGURE 5" captionStartId="17.[264,368,1530,1554]" captionTargetBox="[355,1242,322,1499]" captionTargetId="figure@17.[264,1323,310,1523]" captionTargetPageId="17" captionText="FIGURE 5. Elytral color patterns of Cycloneda species, left elytron: A–D, C. patagonica González & Vandenberg, new species, spectrum of observed variation from lightest to darkest; E –H, C. sicardi (Brèthes), spectrum of observed variation from lightest to darkest; I–K, C. boliviana (Mulsant), spectrum of observed variation from lightest to darkest." httpUri="https://zenodo.org/record/173873/files/figure.png" pageId="5" pageNumber="18">Figs. 5A–D</figureCitation>
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); punctation on pronotum and elytron dense (separated by 1.0–2.5X a diameter), sharply defined; antenna short, composed of 10 antennomeres, strongly, abruptly clubbed (
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<figureCitation id="42E32A571009BC6605A9F8E4843BF8F7" box="[753,847,1799,1825]" captionStart="FIGURE 9" captionStartId="26.[264,368,885,909]" captionTargetBox="[276,1316,321,884]" captionTargetId="figure@26.[264,1323,305,886]" captionTargetPageId="26" captionText="FIGURE 9. Left antenna of Cycloneda species, dorsal view: AC, C. lacrimosa Gonzlez & Vandenberg, new species (showing variation from 9 to 11 antennomeres); D, C. patagonica Gonzlez & Vandenberg, new species; E, Cycloneda germainii (Crotch); F, C. eryngii (Mulsant); G, C. sicardi (Brèthes); H, C. boliviana (Mulsant)." figureDoi="http://doi.org/10.5281/zenodo.3924340" httpUri="https://zenodo.org/record/3924340/files/figure.png" pageId="5" pageNumber="18">Fig. 9D</figureCitation>
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). Recorded from Magellanes,
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<collectingCountry id="A2CF76421009BC6603E0F8E48383F8F7" box="[1208,1271,1799,1825]" name="Chile" pageId="5" pageNumber="18">Chile</collectingCountry>
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and
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<collectingRegion id="181CF8301009BC660603F8CC8554F89F" box="[347,544,1839,1865]" country="Argentina" name="Tierra del Fuego - Antartida e Islas del Atlantico Sur" pageId="5" pageNumber="18">Tierra del Fuego</collectingRegion>
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,
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<collectingCountry id="A2CF76421009BC660574F8CC85D4F89F" box="[556,672,1839,1865]" name="Argentina" pageId="5" pageNumber="18">Argentina</collectingCountry>
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(
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<figureCitation id="42E32A571009BC6605E8F8CC8472F89F" box="[688,774,1839,1865]" captionStart="FIGURE 10" captionStartId="27.[264,368,1868,1892]" captionTargetBox="[287,1302,680,1870]" captionTargetId="figure@27.[287,1312,671,1870]" captionTargetPageId="27" captionText="FIGURE 10. Distributions of Cycloneda species." figureDoi="http://doi.org/10.5281/zenodo.3924342" httpUri="https://zenodo.org/record/3924342/files/figure.png" pageId="5" pageNumber="18">Fig. 10</figureCitation>
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, triangles).........................................................
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<emphasis id="E8ACEAC01009BC660607F8B48484F8A6" box="[351,1008,1879,1904]" italics="true" pageId="5" pageNumber="18">
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...................................................................
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<taxonomicName id="1DD84D511009BC660408F8B48484F8A6" authority="Gonzalez and Vandenberg" authorityName="Gonzalez & Vandenberg" authorityYear="2006" box="[848,1008,1879,1904]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="patagonica">C. patagonica</taxonomicName>
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</emphasis>
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González and Vandenberg
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</keyLead>
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</paragraph>
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<paragraph id="DA6736D2100ABC650650FEFF8259FE28" blockId="6.[264,1326,284,1870]" pageId="6" pageNumber="19">
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<keyLead id="612995E7100ABC650650FEFF8259FE28" pageId="6" pageNumber="19">
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1’. Form moderately to strongly convex, ovoid to oblong; elytral epipleuron flat to concave, approximately horizontal, not visible in lateral view or only very narrowly so in anterior half (
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<figureCitation id="42E32A57100ABC650567FE8F846CFE50" box="[575,792,364,390]" captionStart="FIGURE 3" captionStartId="15.[264,367,1565,1589]" captionTargetBox="[272,1316,323,1530]" captionTargetId="figure@15.[256,1331,310,1551]" captionTargetPageId="15" captionText="FIGURE 3. Morphology of Cycloneda species. Lateral views of bodies (elytral length standardized to facilitate comparison), A–G: A, C. lacrimosa González & Vandenberg, new species, paratype; B, C. disconsolata Vandenberg & González, new species, holotype; C, C. patagonica González & Vandenberg, new species, paratype; D, Cycloneda germainii (Crotch), representative specimen; E, C. eryngii (Mulsant), representative specimen; F, C. sicardi (Brèthes), specimen from Salta; G, C. boliviana (Mulsant), holotype. Pronotal dorsofrontal views, H–L: H, C. lacrimosa González & Vandenberg, new species, paratype; I, C. boliviana (Mulsant), holotype; J, C. sicardi (Brèthes), specimen from El Rincón, Catamarca; K, C. sicardi (Brèthes), specimen from Tacuil, Salta; L, C. sicardi (Brèthes), common aberration in much of range." httpUri="https://zenodo.org/record/173871/files/figure.png" pageId="6" pageNumber="19">Figs. 3A, B, D–G</figureCitation>
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); elytron broader, with external (lateral) bor der more arcuate (
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<figureCitation id="42E32A57100ABC650560FE7785CBFE78" box="[568,703,404,430]" captionStart="FIGURE 4" captionStartId="16.[264,367,1625,1649]" captionTargetBox="[395,1202,315,1596]" captionTargetId="figure@16.[316,1279,307,1620]" captionTargetPageId="16" captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)." httpUri="https://zenodo.org/record/173872/files/figure.png" pageId="6" pageNumber="19">Figs. 4A–P</figureCitation>
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,
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<figureCitation id="42E32A57100ABC650596FE778463FE78" box="[718,791,404,430]" captionStart="FIGURE 5" captionStartId="17.[264,368,1530,1554]" captionTargetBox="[355,1242,322,1499]" captionTargetId="figure@17.[264,1323,310,1523]" captionTargetPageId="17" captionText="FIGURE 5. Elytral color patterns of Cycloneda species, left elytron: A–D, C. patagonica González & Vandenberg, new species, spectrum of observed variation from lightest to darkest; E –H, C. sicardi (Brèthes), spectrum of observed variation from lightest to darkest; I–K, C. boliviana (Mulsant), spectrum of observed variation from lightest to darkest." httpUri="https://zenodo.org/record/173873/files/figure.png" pageId="6" pageNumber="19">5E–K</figureCitation>
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); punctation less dense, variable, sometimes indistinct; antenna longer (
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<figureCitation id="42E32A57100ABC6505CDFE5F8412FE00" box="[661,870,444,470]" captionStart="FIGURE 9" captionStartId="26.[264,368,885,909]" captionTargetBox="[276,1316,321,884]" captionTargetId="figure@26.[264,1323,305,886]" captionTargetPageId="26" captionText="FIGURE 9. Left antenna of Cycloneda species, dorsal view: AC, C. lacrimosa Gonzlez & Vandenberg, new species (showing variation from 9 to 11 antennomeres); D, C. patagonica Gonzlez & Vandenberg, new species; E, Cycloneda germainii (Crotch); F, C. eryngii (Mulsant); G, C. sicardi (Brèthes); H, C. boliviana (Mulsant)." figureDoi="http://doi.org/10.5281/zenodo.3924340" httpUri="https://zenodo.org/record/3924340/files/figure.png" pageId="6" pageNumber="19">Figs. 9A–C, E–H</figureCitation>
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), of 9–11 antennomeres, club variable.
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<collectingCountry id="A2CF7642100ABC650603FE0786C0FE28" box="[347,436,484,510]" name="Bolivia" pageId="6" pageNumber="19">Bolivia</collectingCountry>
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,
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<collectingCountry id="A2CF7642100ABC650698FE078577FE28" box="[448,515,484,510]" name="Chile" pageId="6" pageNumber="19">Chile</collectingCountry>
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,
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<collectingCountry id="A2CF7642100ABC650557FE0785F3FE28" box="[527,647,484,510]" name="Argentina" pageId="6" pageNumber="19">Argentina</collectingCountry>
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; not known from southeast of the straits of Magellan..... 2
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="612C2E77100ABC650650FDEF8259FD38" pageId="6" pageNumber="19">
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<paragraph id="DA6736D2100ABC650650FDEF8259FD48" blockId="6.[264,1326,284,1870]" pageId="6" pageNumber="19">
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<keyLead id="612995E7100ABC650650FDEF8259FD48" pageId="6" pageNumber="19">
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2 (1’). Pronotum with white to creamcolored anterior and lateral borders uneven (
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<figureCitation id="42E32A57100ABC6503A8FDEF86B7FD98" captionStart="FIGURE 3" captionStartId="15.[264,367,1565,1589]" captionTargetBox="[272,1316,323,1530]" captionTargetId="figure@15.[256,1331,310,1551]" captionTargetPageId="15" captionText="FIGURE 3. Morphology of Cycloneda species. Lateral views of bodies (elytral length standardized to facilitate comparison), A–G: A, C. lacrimosa González & Vandenberg, new species, paratype; B, C. disconsolata Vandenberg & González, new species, holotype; C, C. patagonica González & Vandenberg, new species, paratype; D, Cycloneda germainii (Crotch), representative specimen; E, C. eryngii (Mulsant), representative specimen; F, C. sicardi (Brèthes), specimen from Salta; G, C. boliviana (Mulsant), holotype. Pronotal dorsofrontal views, H–L: H, C. lacrimosa González & Vandenberg, new species, paratype; I, C. boliviana (Mulsant), holotype; J, C. sicardi (Brèthes), specimen from El Rincón, Catamarca; K, C. sicardi (Brèthes), specimen from Tacuil, Salta; L, C. sicardi (Brèthes), common aberration in much of range." httpUri="https://zenodo.org/record/173871/files/figure.png" pageId="6" pageNumber="19">Figs. 3H, J–K</figureCitation>
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): lateral border distinctly wider than anterior border; anterior border sometimes broken or obsolete, in some specimens with spurlike markings projecting posteriorly onto disk (
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<figureCitation id="42E32A57100ABC650506FD6785C6FD48" box="[606,690,644,670]" captionStart="FIGURE 3" captionStartId="15.[264,367,1565,1589]" captionTargetBox="[272,1316,323,1530]" captionTargetId="figure@15.[256,1331,310,1551]" captionTargetPageId="15" captionText="FIGURE 3. Morphology of Cycloneda species. Lateral views of bodies (elytral length standardized to facilitate comparison), A–G: A, C. lacrimosa González & Vandenberg, new species, paratype; B, C. disconsolata Vandenberg & González, new species, holotype; C, C. patagonica González & Vandenberg, new species, paratype; D, Cycloneda germainii (Crotch), representative specimen; E, C. eryngii (Mulsant), representative specimen; F, C. sicardi (Brèthes), specimen from Salta; G, C. boliviana (Mulsant), holotype. Pronotal dorsofrontal views, H–L: H, C. lacrimosa González & Vandenberg, new species, paratype; I, C. boliviana (Mulsant), holotype; J, C. sicardi (Brèthes), specimen from El Rincón, Catamarca; K, C. sicardi (Brèthes), specimen from Tacuil, Salta; L, C. sicardi (Brèthes), common aberration in much of range." httpUri="https://zenodo.org/record/173871/files/figure.png" pageId="6" pageNumber="19">Fig.3L</figureCitation>
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). ................................................................................. 3
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</keyLead>
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</paragraph>
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<paragraph id="DA6736D2100ABC650650FD4F8259FD38" blockId="6.[264,1326,284,1870]" pageId="6" pageNumber="19">
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<keyLead id="612995E7100ABC650650FD4F8259FD38" pageId="6" pageNumber="19">
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2’. Pronotum with white to creamcolored border of uniform width (
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<figureCitation id="42E32A57100ABC650308FD4F83D5FD10" box="[1104,1185,684,710]" captionStart="FIGURE 3" captionStartId="15.[264,367,1565,1589]" captionTargetBox="[272,1316,323,1530]" captionTargetId="figure@15.[256,1331,310,1551]" captionTargetPageId="15" captionText="FIGURE 3. Morphology of Cycloneda species. Lateral views of bodies (elytral length standardized to facilitate comparison), A–G: A, C. lacrimosa González & Vandenberg, new species, paratype; B, C. disconsolata Vandenberg & González, new species, holotype; C, C. patagonica González & Vandenberg, new species, paratype; D, Cycloneda germainii (Crotch), representative specimen; E, C. eryngii (Mulsant), representative specimen; F, C. sicardi (Brèthes), specimen from Salta; G, C. boliviana (Mulsant), holotype. Pronotal dorsofrontal views, H–L: H, C. lacrimosa González & Vandenberg, new species, paratype; I, C. boliviana (Mulsant), holotype; J, C. sicardi (Brèthes), specimen from El Rincón, Catamarca; K, C. sicardi (Brèthes), specimen from Tacuil, Salta; L, C. sicardi (Brèthes), common aberration in much of range." httpUri="https://zenodo.org/record/173871/files/figure.png" pageId="6" pageNumber="19">Fig. 3I</figureCitation>
|
||
), or at most only slightly narrower anteriorly than laterally, without spurlike markings ............ 5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="612C2E77100ABC650650FD1F825EFC08" pageId="6" pageNumber="19">
|
||
<paragraph id="DA6736D2100ABC650650FD1F825AFCB0" blockId="6.[264,1326,284,1870]" pageId="6" pageNumber="19">
|
||
<keyLead id="612995E7100ABC650650FD1F825AFCB0" pageId="6" pageNumber="19">
|
||
3 (2). Elytra with dark maculae framing a common pale teardropshaped mark or pair of such marks near suture at midlength (
|
||
<figureCitation id="42E32A57100ABC650479FCC784C4FCE8" box="[801,944,804,830]" captionStart="FIGURE 1" captionStartId="4.[264,367,1784,1808]" captionTargetBox="[344,1263,723,1752]" captionTargetId="figure@4.[279,1332,713,1777]" captionTargetPageId="4" captionText="FIGURE 1. Habitus views of Cycloneda species: A, C. lacrimosa González & Vandenberg, new species, male, paratype (3.0 mm); B, C. disconsolata Vandenberg & González, new species, female, holotype (3.3 mm); C, C. patagonica González & Vandenberg, new species, paratype (3.0 mm); D, C. germainii (Crotch), female, common form (3.5 mm)." httpUri="https://zenodo.org/record/173869/files/figure.png" pageId="6" pageNumber="19">Figs. 1A, B</figureCitation>
|
||
); body form shortened, oblong, strongly convex (
|
||
<figureCitation id="42E32A57100ABC65057FFCAF85DFFCB0" box="[551,683,844,870]" captionStart="FIGURE 3" captionStartId="15.[264,367,1565,1589]" captionTargetBox="[272,1316,323,1530]" captionTargetId="figure@15.[256,1331,310,1551]" captionTargetPageId="15" captionText="FIGURE 3. Morphology of Cycloneda species. Lateral views of bodies (elytral length standardized to facilitate comparison), A–G: A, C. lacrimosa González & Vandenberg, new species, paratype; B, C. disconsolata Vandenberg & González, new species, holotype; C, C. patagonica González & Vandenberg, new species, paratype; D, Cycloneda germainii (Crotch), representative specimen; E, C. eryngii (Mulsant), representative specimen; F, C. sicardi (Brèthes), specimen from Salta; G, C. boliviana (Mulsant), holotype. Pronotal dorsofrontal views, H–L: H, C. lacrimosa González & Vandenberg, new species, paratype; I, C. boliviana (Mulsant), holotype; J, C. sicardi (Brèthes), specimen from El Rincón, Catamarca; K, C. sicardi (Brèthes), specimen from Tacuil, Salta; L, C. sicardi (Brèthes), common aberration in much of range." httpUri="https://zenodo.org/record/173871/files/figure.png" pageId="6" pageNumber="19">Figs 3A, B</figureCitation>
|
||
); antenna with 9, 10, or 11 antennomeres .................. 4
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="DA6736D2100ABC650650FC97825EFC08" blockId="6.[264,1326,284,1870]" pageId="6" pageNumber="19">
|
||
<keyLead id="612995E7100ABC650650FC97825EFC08" pageId="6" pageNumber="19">
|
||
3’. Elytra with pair of discrete pale circular marks near suture at midlength (
|
||
<figureCitation id="42E32A57100ABC650390FC978268FC58" box="[1224,1308,884,910]" captionStart="FIGURE 2" captionStartId="5.[264,368,1403,1427]" captionTargetBox="[302,1293,302,1387]" captionTargetId="figure@5.[264,1323,280,1401]" captionTargetPageId="5" captionText="FIGURE 2. Habitus views of Cycloneda species: A, C. eryngii (Mulsant), female, common form (3.8 mm); B, C. sicardi (Brèthes), female from Salta, Argentina (3.8 mm); C, C. boliviana (Mulsant), female, holotype (4.6 mm)." httpUri="https://zenodo.org/record/173870/files/figure.png" pageId="6" pageNumber="19">Fig.2B</figureCitation>
|
||
); body form slightly elongate oval, moderately convex (
|
||
<figureCitation id="42E32A57100ABC6504BBFC7F8338FC60" box="[995,1100,924,950]" captionStart="FIGURE 2" captionStartId="5.[264,368,1403,1427]" captionTargetBox="[302,1293,302,1387]" captionTargetId="figure@5.[264,1323,280,1401]" captionTargetPageId="5" captionText="FIGURE 2. Habitus views of Cycloneda species: A, C. eryngii (Mulsant), female, common form (3.8 mm); B, C. sicardi (Brèthes), female from Salta, Argentina (3.8 mm); C, C. boliviana (Mulsant), female, holotype (4.6 mm)." httpUri="https://zenodo.org/record/173870/files/figure.png" pageId="6" pageNumber="19">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="42E32A57100ABC650301FC7F8308FC60" box="[1113,1148,924,950]" captionStart="FIGURE 3" captionStartId="15.[264,367,1565,1589]" captionTargetBox="[272,1316,323,1530]" captionTargetId="figure@15.[256,1331,310,1551]" captionTargetPageId="15" captionText="FIGURE 3. Morphology of Cycloneda species. Lateral views of bodies (elytral length standardized to facilitate comparison), A–G: A, C. lacrimosa González & Vandenberg, new species, paratype; B, C. disconsolata Vandenberg & González, new species, holotype; C, C. patagonica González & Vandenberg, new species, paratype; D, Cycloneda germainii (Crotch), representative specimen; E, C. eryngii (Mulsant), representative specimen; F, C. sicardi (Brèthes), specimen from Salta; G, C. boliviana (Mulsant), holotype. Pronotal dorsofrontal views, H–L: H, C. lacrimosa González & Vandenberg, new species, paratype; I, C. boliviana (Mulsant), holotype; J, C. sicardi (Brèthes), specimen from El Rincón, Catamarca; K, C. sicardi (Brèthes), specimen from Tacuil, Salta; L, C. sicardi (Brèthes), common aberration in much of range." httpUri="https://zenodo.org/record/173871/files/figure.png" pageId="6" pageNumber="19">3F</figureCitation>
|
||
); antenna with 11 antennomeres (
|
||
<figureCitation id="42E32A57100ABC65056AFC2785FEFC08" box="[562,650,964,990]" captionStart="FIGURE 9" captionStartId="26.[264,368,885,909]" captionTargetBox="[276,1316,321,884]" captionTargetId="figure@26.[264,1323,305,886]" captionTargetPageId="26" captionText="FIGURE 9. Left antenna of Cycloneda species, dorsal view: AC, C. lacrimosa Gonzlez & Vandenberg, new species (showing variation from 9 to 11 antennomeres); D, C. patagonica Gonzlez & Vandenberg, new species; E, Cycloneda germainii (Crotch); F, C. eryngii (Mulsant); G, C. sicardi (Brèthes); H, C. boliviana (Mulsant)." figureDoi="http://doi.org/10.5281/zenodo.3924340" httpUri="https://zenodo.org/record/3924340/files/figure.png" pageId="6" pageNumber="19">Fig.9G</figureCitation>
|
||
).
|
||
<collectingCountry id="A2CF7642100ABC6505FDFC27858AFC08" box="[677,766,964,990]" name="Bolivia" pageId="6" pageNumber="19">Bolivia</collectingCountry>
|
||
,
|
||
<collectingCountry id="A2CF7642100ABC650452FC27840BFC08" box="[778,895,964,990]" name="Argentina" pageId="6" pageNumber="19">Argentina</collectingCountry>
|
||
(
|
||
<figureCitation id="42E32A57100ABC6504D7FC2784A9FC08" box="[911,989,964,990]" captionStart="FIGURE 11" captionStartId="32.[264,367,1618,1642]" captionTargetBox="[275,1312,375,1593]" captionTargetId="figure@32.[264,1323,366,1607]" captionTargetPageId="32" captionText="FIGURE 11. Distributions of Cycloneda species." httpUri="https://zenodo.org/record/173877/files/figure.png" pageId="6" pageNumber="19">Fig 11</figureCitation>
|
||
, stars)
|
||
<emphasis id="E8ACEAC0100ABC650375FC278337FC0B" box="[1069,1091,964,989]" italics="true" pageId="6" pageNumber="19">...</emphasis>
|
||
<taxonomicName id="1DD84D51100ABC65031BFC27825EFC08" authority="(Brethes)" authorityName="Brethes" baseAuthorityName="Brethes" box="[1091,1322,964,990]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="sicardi">
|
||
<emphasis id="E8ACEAC0100ABC65031BFC2783C2FC0B" box="[1091,1206,964,989]" italics="true" pageId="6" pageNumber="19">C. sicardi</emphasis>
|
||
(Brèthes)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="612C2E77100ABC650650FC0F825EFAC8" pageId="6" pageNumber="19">
|
||
<paragraph id="DA6736D2100ABC650650FC0F825FFBA8" blockId="6.[264,1326,284,1870]" pageId="6" pageNumber="19">
|
||
<keyLead id="612995E7100ABC650650FC0F825FFBA8" pageId="6" pageNumber="19">
|
||
4 (3). Elytra bicolored, or at least with orange to paler cream areas strongly suffused: common teardropshaped mark not divided at suture; dark maculae as shown (
|
||
<figureCitation id="42E32A57100ABC6503A8FBF786F5FB80" captionStart="FIGURE 1" captionStartId="4.[264,367,1784,1808]" captionTargetBox="[344,1263,723,1752]" captionTargetId="figure@4.[279,1332,713,1777]" captionTargetPageId="4" captionText="FIGURE 1. Habitus views of Cycloneda species: A, C. lacrimosa González & Vandenberg, new species, male, paratype (3.0 mm); B, C. disconsolata Vandenberg & González, new species, female, holotype (3.3 mm); C, C. patagonica González & Vandenberg, new species, paratype (3.0 mm); D, C. germainii (Crotch), female, common form (3.5 mm)." httpUri="https://zenodo.org/record/173869/files/figure.png" pageId="6" pageNumber="19">Figs. 1A</figureCitation>
|
||
,
|
||
<figureCitation id="42E32A57100ABC6506D6FBDF8694FB80" box="[398,480,1084,1110]" captionStart="FIGURE 4" captionStartId="16.[264,367,1625,1649]" captionTargetBox="[395,1202,315,1596]" captionTargetId="figure@16.[316,1279,307,1620]" captionTargetPageId="16" captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)." httpUri="https://zenodo.org/record/173872/files/figure.png" pageId="6" pageNumber="19">4M–N</figureCitation>
|
||
). Northern
|
||
<collectingCountry id="A2CF7642100ABC65053CFBDF85D3FB80" box="[612,679,1084,1110]" name="Chile" pageId="6" pageNumber="19">Chile</collectingCountry>
|
||
, northern
|
||
<collectingCountry id="A2CF7642100ABC650445FBDF84E1FB80" box="[797,917,1084,1110]" name="Argentina" pageId="6" pageNumber="19">Argentina</collectingCountry>
|
||
,
|
||
<collectingCountry id="A2CF7642100ABC6504F9FBDF8483FB80" box="[929,1015,1084,1110]" name="Bolivia" pageId="6" pageNumber="19">Bolivia</collectingCountry>
|
||
(
|
||
<figureCitation id="42E32A57100ABC65035FFBDF833BFB80" box="[1031,1103,1084,1110]" captionStart="FIGURE 10" captionStartId="27.[264,368,1868,1892]" captionTargetBox="[287,1302,680,1870]" captionTargetId="figure@27.[287,1312,671,1870]" captionTargetPageId="27" captionText="FIGURE 10. Distributions of Cycloneda species." figureDoi="http://doi.org/10.5281/zenodo.3924342" httpUri="https://zenodo.org/record/3924342/files/figure.png" pageId="6" pageNumber="19">Fig10</figureCitation>
|
||
, squares) ..............
|
||
<emphasis id="E8ACEAC0100ABC650607FB878484FBAB" box="[351,1008,1124,1149]" italics="true" pageId="6" pageNumber="19">
|
||
.....................................................................
|
||
<taxonomicName id="1DD84D51100ABC650406FB878484FBAB" authority="Gonzalez and Vandenberg" authorityName="Gonzalez & Vandenberg" authorityYear="2006" box="[862,1008,1124,1149]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="lacrimosa">C. lacrimosa</taxonomicName>
|
||
</emphasis>
|
||
González and Vandenberg
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="DA6736D2100ABC650650FB6F825EFAC8" blockId="6.[264,1326,284,1870]" pageId="6" pageNumber="19">
|
||
<keyLead id="612995E7100ABC650650FB6F825EFAC8" pageId="6" pageNumber="19">
|
||
4’. Elytra distinctly tricolored: pale central elytral figure divided by ferrugineous suture to form pair of creamcolored teardropshaped marks, one on each elytron; dark maculae as shown (
|
||
<figureCitation id="42E32A57100ABC650525FB3F8597FB20" box="[637,739,1244,1270]" captionStart="FIGURE 1" captionStartId="4.[264,367,1784,1808]" captionTargetBox="[344,1263,723,1752]" captionTargetId="figure@4.[279,1332,713,1777]" captionTargetPageId="4" captionText="FIGURE 1. Habitus views of Cycloneda species: A, C. lacrimosa González & Vandenberg, new species, male, paratype (3.0 mm); B, C. disconsolata Vandenberg & González, new species, female, holotype (3.3 mm); C, C. patagonica González & Vandenberg, new species, paratype (3.0 mm); D, C. germainii (Crotch), female, common form (3.5 mm)." httpUri="https://zenodo.org/record/173869/files/figure.png" pageId="6" pageNumber="19">Figs. 1B</figureCitation>
|
||
,
|
||
<figureCitation id="42E32A57100ABC6505A8FB3F844CFB20" box="[752,824,1244,1270]" captionStart="FIGURE 4" captionStartId="16.[264,367,1625,1649]" captionTargetBox="[395,1202,315,1596]" captionTargetId="figure@16.[316,1279,307,1620]" captionTargetPageId="16" captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)." httpUri="https://zenodo.org/record/173872/files/figure.png" pageId="6" pageNumber="19">4O–P</figureCitation>
|
||
). Known from disjunct localities in
|
||
<collectingCountry id="A2CF7642100ABC6503BEFB3F825DFB20" box="[1254,1321,1244,1270]" name="Chile" pageId="6" pageNumber="19">Chile</collectingCountry>
|
||
,
|
||
<collectingCountry id="A2CF7642100ABC650603FAE786A4FAC8" box="[347,464,1284,1310]" name="Argentina" pageId="6" pageNumber="19">Argentina</collectingCountry>
|
||
(
|
||
<figureCitation id="42E32A57100ABC6506B8FAE78543FAC8" box="[480,567,1284,1310]" captionStart="FIGURE 10" captionStartId="27.[264,368,1868,1892]" captionTargetBox="[287,1302,680,1870]" captionTargetId="figure@27.[287,1312,671,1870]" captionTargetPageId="27" captionText="FIGURE 10. Distributions of Cycloneda species." figureDoi="http://doi.org/10.5281/zenodo.3924342" httpUri="https://zenodo.org/record/3924342/files/figure.png" pageId="6" pageNumber="19">Fig. 10</figureCitation>
|
||
, circles with cross)
|
||
<emphasis id="E8ACEAC0100ABC650446FAE78484FACB" box="[798,1008,1284,1309]" italics="true" pageId="6" pageNumber="19">
|
||
...
|
||
<taxonomicName id="1DD84D51100ABC650462FAE78484FACB" authority="Vandenberg and Gonzalez" authorityName="Vandenberg & Gonzalez" authorityYear="2006" box="[826,1008,1284,1309]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="disconsolata">C. disconsolata</taxonomicName>
|
||
</emphasis>
|
||
Vandenberg and González
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="612C2E77100ABC650650FACF825EF978" pageId="6" pageNumber="19">
|
||
<paragraph id="DA6736D2100ABC650650FACF825FFA30" blockId="6.[264,1326,284,1870]" pageId="6" pageNumber="19">
|
||
<keyLead id="612995E7100ABC650650FACF825FFA30" pageId="6" pageNumber="19">
|
||
5 (2’). Posterior third of each elytron with pale spot positioned near apex (
|
||
<figureCitation id="42E32A57100ABC65032BFACF8385FA90" box="[1139,1265,1324,1350]" captionStart="FIGURE 4" captionStartId="16.[264,367,1625,1649]" captionTargetBox="[395,1202,315,1596]" captionTargetId="figure@16.[316,1279,307,1620]" captionTargetPageId="16" captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)." httpUri="https://zenodo.org/record/173872/files/figure.png" pageId="6" pageNumber="19">Figs. 4I–L</figureCitation>
|
||
) and often obliquely flattened against posterolateral margin (pale spot poorly distinguished from orangy ground color in most
|
||
<taxonomicName id="1DD84D51100ABC650431FA9F8495FA43" authorityName="González & Vandenberg" authorityYear="2006" baseAuthorityName="González & Vandenberg" baseAuthorityYear="2006" box="[873,993,1404,1429]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="eryngii">
|
||
<emphasis id="E8ACEAC0100ABC650431FA9F8495FA43" box="[873,993,1404,1429]" italics="true" pageId="6" pageNumber="19">C. eryngii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="42E32A57100ABC6504ABFA9F83F7FA40" box="[1011,1155,1404,1430]" captionStart="FIGURE 4" captionStartId="16.[264,367,1625,1649]" captionTargetBox="[395,1202,315,1596]" captionTargetId="figure@16.[316,1279,307,1620]" captionTargetPageId="16" captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)." httpUri="https://zenodo.org/record/173872/files/figure.png" pageId="6" pageNumber="19">Figs. 4A–H</figureCitation>
|
||
). Dorsal surfaces shiny; punctation sharply defined; surface between punctures with less apparent reticulation. Widespread in
|
||
<collectingCountry id="A2CF7642100ABC65044AFA2F8425FA30" box="[786,849,1484,1510]" name="Chile" pageId="6" pageNumber="19">Chile</collectingCountry>
|
||
and
|
||
<collectingCountry id="A2CF7642100ABC6504D3FA2F848BFA30" box="[907,1023,1484,1510]" name="Argentina" pageId="6" pageNumber="19">Argentina</collectingCountry>
|
||
..................................... 6
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="DA6736D2100ABC650650FA17825EF978" blockId="6.[264,1326,284,1870]" pageId="6" pageNumber="19">
|
||
<keyLead id="612995E7100ABC650650FA17825EF978" pageId="6" pageNumber="19">
|
||
5’. Posterior third of each elytron with circular to irregular pale spot centered some distance from apex (
|
||
<figureCitation id="42E32A57100ABC65050AF9FF85A3F9E0" box="[594,727,1564,1590]" captionStart="FIGURE 5" captionStartId="17.[264,368,1530,1554]" captionTargetBox="[355,1242,322,1499]" captionTargetId="figure@17.[264,1323,310,1523]" captionTargetPageId="17" captionText="FIGURE 5. Elytral color patterns of Cycloneda species, left elytron: A–D, C. patagonica González & Vandenberg, new species, spectrum of observed variation from lightest to darkest; E –H, C. sicardi (Brèthes), spectrum of observed variation from lightest to darkest; I–K, C. boliviana (Mulsant), spectrum of observed variation from lightest to darkest." httpUri="https://zenodo.org/record/173873/files/figure.png" pageId="6" pageNumber="19">Figs. 5I–K</figureCitation>
|
||
), but with outer edge approaching apex if spot is large (
|
||
<figureCitation id="42E32A57100ABC6506F1F9A7868BF988" box="[425,511,1604,1630]" captionStart="FIGURE 5" captionStartId="17.[264,368,1530,1554]" captionTargetBox="[355,1242,322,1499]" captionTargetId="figure@17.[264,1323,310,1523]" captionTargetPageId="17" captionText="FIGURE 5. Elytral color patterns of Cycloneda species, left elytron: A–D, C. patagonica González & Vandenberg, new species, spectrum of observed variation from lightest to darkest; E –H, C. sicardi (Brèthes), spectrum of observed variation from lightest to darkest; I–K, C. boliviana (Mulsant), spectrum of observed variation from lightest to darkest." httpUri="https://zenodo.org/record/173873/files/figure.png" pageId="6" pageNumber="19">Fig. 5I</figureCitation>
|
||
). Dorsal surfaces matt; punctation shallow and indistinct; surface between punctures with pronounced reticulation (alutaceous). Known from
|
||
<collectingCountry id="A2CF7642100ABC65038DF98F8258F950" box="[1237,1324,1644,1670]" name="Bolivia" pageId="6" pageNumber="19">Bolivia</collectingCountry>
|
||
and northern
|
||
<collectingCountry id="A2CF7642100ABC6506AFF9778518F978" box="[503,620,1684,1710]" name="Argentina" pageId="6" pageNumber="19">Argentina</collectingCountry>
|
||
(
|
||
<figureCitation id="42E32A57100ABC650524F97785A5F978" box="[636,721,1684,1710]" captionStart="FIGURE 11" captionStartId="32.[264,367,1618,1642]" captionTargetBox="[275,1312,375,1593]" captionTargetId="figure@32.[264,1323,366,1607]" captionTargetPageId="32" captionText="FIGURE 11. Distributions of Cycloneda species." httpUri="https://zenodo.org/record/173877/files/figure.png" pageId="6" pageNumber="19">Fig. 11</figureCitation>
|
||
, circles with dot)
|
||
<emphasis id="E8ACEAC0100ABC6504FAF9778357F97B" box="[930,1059,1684,1709]" italics="true" pageId="6" pageNumber="19">.................</emphasis>
|
||
<taxonomicName id="1DD84D51100ABC65037BF977825EF978" authority="(Mulsant)" authorityName="Mulsant" baseAuthorityName="Mulsant" box="[1059,1322,1684,1710]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="boliviana">
|
||
<emphasis id="E8ACEAC0100ABC65037BF97783C4F97B" box="[1059,1200,1684,1709]" italics="true" pageId="6" pageNumber="19">C. boliviana</emphasis>
|
||
(Mulsant)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="612C2E77100ABC640650F95F825FFD38" lastPageId="7" lastPageNumber="20" pageId="6" pageNumber="19">
|
||
<paragraph id="DA6736D2100ABC640650F95F825EFE50" blockId="6.[264,1326,284,1870]" lastBlockId="7.[264,1324,284,750]" lastPageId="7" lastPageNumber="20" pageId="6" pageNumber="19">
|
||
<keyLead id="612995E7100ABC640650F95F825EFE50" lastPageId="7" lastPageNumber="20" pageId="6" pageNumber="19">
|
||
6 (5). Antenna with 11 antennomeres (
|
||
<figureCitation id="42E32A57100ABC650585F95F8445F900" box="[733,817,1724,1750]" captionStart="FIGURE 9" captionStartId="26.[264,368,885,909]" captionTargetBox="[276,1316,321,884]" captionTargetId="figure@26.[264,1323,305,886]" captionTargetPageId="26" captionText="FIGURE 9. Left antenna of Cycloneda species, dorsal view: AC, C. lacrimosa Gonzlez & Vandenberg, new species (showing variation from 9 to 11 antennomeres); D, C. patagonica Gonzlez & Vandenberg, new species; E, Cycloneda germainii (Crotch); F, C. eryngii (Mulsant); G, C. sicardi (Brèthes); H, C. boliviana (Mulsant)." figureDoi="http://doi.org/10.5281/zenodo.3924340" httpUri="https://zenodo.org/record/3924340/files/figure.png" pageId="6" pageNumber="19">Fig 9F</figureCitation>
|
||
). Most specimens with dark elytral marks well separated at midline, less commonly with marks touching or longitudinally connected midway between sutural and median area of each elytron (
|
||
<figureCitation id="42E32A57100ABC6503D6F8EF826AF8F0" box="[1166,1310,1804,1830]" captionStart="FIGURE 4" captionStartId="16.[264,367,1625,1649]" captionTargetBox="[395,1202,315,1596]" captionTargetId="figure@16.[316,1279,307,1620]" captionTargetPageId="16" captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)." httpUri="https://zenodo.org/record/173872/files/figure.png" pageId="6" pageNumber="19">Figs. 4A–H</figureCitation>
|
||
); elytron in lateral view evenly convex and tapering toward apex, broader at anterior 1/4 than at posterior 1/4 (
|
||
<figureCitation id="42E32A57100BBC6405D9FEFF85ACFEE0" box="[641,728,284,310]" captionStart="FIGURE 3" captionStartId="15.[264,367,1565,1589]" captionTargetBox="[272,1316,323,1530]" captionTargetId="figure@15.[256,1331,310,1551]" captionTargetPageId="15" captionText="FIGURE 3. Morphology of Cycloneda species. Lateral views of bodies (elytral length standardized to facilitate comparison), A–G: A, C. lacrimosa González & Vandenberg, new species, paratype; B, C. disconsolata Vandenberg & González, new species, holotype; C, C. patagonica González & Vandenberg, new species, paratype; D, Cycloneda germainii (Crotch), representative specimen; E, C. eryngii (Mulsant), representative specimen; F, C. sicardi (Brèthes), specimen from Salta; G, C. boliviana (Mulsant), holotype. Pronotal dorsofrontal views, H–L: H, C. lacrimosa González & Vandenberg, new species, paratype; I, C. boliviana (Mulsant), holotype; J, C. sicardi (Brèthes), specimen from El Rincón, Catamarca; K, C. sicardi (Brèthes), specimen from Tacuil, Salta; L, C. sicardi (Brèthes), common aberration in much of range." httpUri="https://zenodo.org/record/173871/files/figure.png" pageId="7" pageNumber="20">Fig. 3E</figureCitation>
|
||
); in dorsal view, outer margin of elytron distinctly arcuate (
|
||
<figureCitation id="42E32A57100BBC640698FEA7855DFE88" box="[448,553,324,350]" captionStart="FIGURE 2" captionStartId="5.[264,368,1403,1427]" captionTargetBox="[302,1293,302,1387]" captionTargetId="figure@5.[264,1323,280,1401]" captionTargetPageId="5" captionText="FIGURE 2. Habitus views of Cycloneda species: A, C. eryngii (Mulsant), female, common form (3.8 mm); B, C. sicardi (Brèthes), female from Salta, Argentina (3.8 mm); C, C. boliviana (Mulsant), female, holotype (4.6 mm)." httpUri="https://zenodo.org/record/173870/files/figure.png" pageId="7" pageNumber="20">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="42E32A57100BBC64056DFEA785F5FE88" box="[565,641,324,350]" captionStart="FIGURE 4" captionStartId="16.[264,367,1625,1649]" captionTargetBox="[395,1202,315,1596]" captionTargetId="figure@16.[316,1279,307,1620]" captionTargetPageId="16" captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)." httpUri="https://zenodo.org/record/173872/files/figure.png" pageId="7" pageNumber="20">4A–H</figureCitation>
|
||
). Male genitalia with basal lobe slender, broadest at base
|
||
<emphasis id="E8ACEAC0100BBC640607FE8F8341FE53" box="[351,1077,364,389]" italics="true" pageId="7" pageNumber="20">...................................................................................................</emphasis>
|
||
<taxonomicName id="1DD84D51100BBC640363FE8F825EFE50" authority="(Mulsant)" authorityName="Mulsant" baseAuthorityName="Mulsant" box="[1083,1322,364,390]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="20" phylum="Arthropoda" rank="species" species="eryngii">
|
||
<emphasis id="E8ACEAC0100BBC640363FE8F83DBFE53" box="[1083,1199,364,389]" italics="true" pageId="7" pageNumber="20">C. eryngii</emphasis>
|
||
(Mulsant)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="DA6736D2100BBC640650FE77825FFD38" blockId="7.[264,1324,284,750]" pageId="7" pageNumber="20">
|
||
<keyLead id="612995E7100BBC640650FE77825FFD38" pageId="7" pageNumber="20">
|
||
6’. Antenna with 10 antennomeres (
|
||
<figureCitation id="42E32A57100BBC640585FE778447FE78" box="[733,819,404,430]" captionStart="FIGURE 9" captionStartId="26.[264,368,885,909]" captionTargetBox="[276,1316,321,884]" captionTargetId="figure@26.[264,1323,305,886]" captionTargetPageId="26" captionText="FIGURE 9. Left antenna of Cycloneda species, dorsal view: AC, C. lacrimosa Gonzlez & Vandenberg, new species (showing variation from 9 to 11 antennomeres); D, C. patagonica Gonzlez & Vandenberg, new species; E, Cycloneda germainii (Crotch); F, C. eryngii (Mulsant); G, C. sicardi (Brèthes); H, C. boliviana (Mulsant)." figureDoi="http://doi.org/10.5281/zenodo.3924340" httpUri="https://zenodo.org/record/3924340/files/figure.png" pageId="7" pageNumber="20">Fig 9E</figureCitation>
|
||
). Most specimens with dark elytral marks connected by median longitudinal stripe (
|
||
<figureCitation id="42E32A57100BBC64041BFE5F84BCFE00" box="[835,968,444,470]" captionStart="FIGURE 4" captionStartId="16.[264,367,1625,1649]" captionTargetBox="[395,1202,315,1596]" captionTargetId="figure@16.[316,1279,307,1620]" captionTargetPageId="16" captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)." httpUri="https://zenodo.org/record/173872/files/figure.png" pageId="7" pageNumber="20">Figs. 4J–K</figureCitation>
|
||
), some also with narrow parasutural and/or paralateral longitudinal connections, less commonly with median connection obsolete (
|
||
<figureCitation id="42E32A57100BBC64050DFDEF85B2FDF0" box="[597,710,524,550]" captionStart="FIGURE 4" captionStartId="16.[264,367,1625,1649]" captionTargetBox="[395,1202,315,1596]" captionTargetId="figure@16.[316,1279,307,1620]" captionTargetPageId="16" captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)." httpUri="https://zenodo.org/record/173872/files/figure.png" pageId="7" pageNumber="20">Fig. 4I, L</figureCitation>
|
||
) or indicated only in ground color; elytron in lateral view less convex in anterior half, less strongly tapered, equally broad at anterior and posterior 1/4 (
|
||
<figureCitation id="42E32A57100BBC640569FDBF85FBFDA0" box="[561,655,604,630]" captionStart="FIGURE 3" captionStartId="15.[264,367,1565,1589]" captionTargetBox="[272,1316,323,1530]" captionTargetId="figure@15.[256,1331,310,1551]" captionTargetPageId="15" captionText="FIGURE 3. Morphology of Cycloneda species. Lateral views of bodies (elytral length standardized to facilitate comparison), A–G: A, C. lacrimosa González & Vandenberg, new species, paratype; B, C. disconsolata Vandenberg & González, new species, holotype; C, C. patagonica González & Vandenberg, new species, paratype; D, Cycloneda germainii (Crotch), representative specimen; E, C. eryngii (Mulsant), representative specimen; F, C. sicardi (Brèthes), specimen from Salta; G, C. boliviana (Mulsant), holotype. Pronotal dorsofrontal views, H–L: H, C. lacrimosa González & Vandenberg, new species, paratype; I, C. boliviana (Mulsant), holotype; J, C. sicardi (Brèthes), specimen from El Rincón, Catamarca; K, C. sicardi (Brèthes), specimen from Tacuil, Salta; L, C. sicardi (Brèthes), common aberration in much of range." httpUri="https://zenodo.org/record/173871/files/figure.png" pageId="7" pageNumber="20">Fig. 3D</figureCitation>
|
||
); in dorsal view, outer margin of elytron linear to weakly arcuate for much of length (
|
||
<figureCitation id="42E32A57100BBC6405F2FD678460FD48" box="[682,788,644,670]" captionStart="FIGURE 1" captionStartId="4.[264,367,1784,1808]" captionTargetBox="[344,1263,723,1752]" captionTargetId="figure@4.[279,1332,713,1777]" captionTargetPageId="4" captionText="FIGURE 1. Habitus views of Cycloneda species: A, C. lacrimosa González & Vandenberg, new species, male, paratype (3.0 mm); B, C. disconsolata Vandenberg & González, new species, female, holotype (3.3 mm); C, C. patagonica González & Vandenberg, new species, paratype (3.0 mm); D, C. germainii (Crotch), female, common form (3.5 mm)." httpUri="https://zenodo.org/record/173869/files/figure.png" pageId="7" pageNumber="20">Figs. 1D</figureCitation>
|
||
,
|
||
<figureCitation id="42E32A57100BBC64047AFD678414FD48" box="[802,864,644,670]" captionStart="FIGURE 4" captionStartId="16.[264,367,1625,1649]" captionTargetBox="[395,1202,315,1596]" captionTargetId="figure@16.[316,1279,307,1620]" captionTargetPageId="16" captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)." httpUri="https://zenodo.org/record/173872/files/figure.png" pageId="7" pageNumber="20">4I–L</figureCitation>
|
||
). Male genitalia with basal lobe more robust than in preceding species, broadest at swollen region in apical ½..................
|
||
<emphasis id="E8ACEAC0100BBC640636FD378353FD3B" box="[366,1063,724,749]" italics="true" pageId="7" pageNumber="20">...............................................................................................</emphasis>
|
||
<taxonomicName id="1DD84D51100BBC640375FD37825FFD38" authority="(Crotch)" authorityName="Crotch" baseAuthorityName="Crotch" box="[1069,1323,724,750]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="20" phylum="Arthropoda" rank="species" species="germainii">
|
||
<emphasis id="E8ACEAC0100BBC640375FD3783B4FD3B" box="[1069,1216,724,749]" italics="true" pageId="7" pageNumber="20">C. germainii</emphasis>
|
||
(Crotch)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |