148 lines
16 KiB
XML
148 lines
16 KiB
XML
<document ID-CLB-Dataset="22951" ID-DOI="http://dx.doi.org/10.3897/zookeys.927.49584" ID-GBIF-Dataset="46c58c23-7c82-40ee-868d-77d3b5058aaf" ID-PMC="PMC7180163" ID-Pensoft-Pub="1313-2970-927-65" ID-Pensoft-UUID="DDBCBAC4F1D65BB280F9A6989755996C" ID-PubMed="32341675" ID-ZooBank="F27544E1800140F5A879403CBD5BE52F" ModsDocID="1313-2970-927-65" checkinTime="1587138236753" checkinUser="pensoft" docAuthor="Erwin, Terry L., Kavanaugh, David H. & Maddison, David R." docDate="2020" docId="1DCA7A60040950E7BC61B4FC693618AE" docLanguage="en" docName="ZooKeys 927: 65-74" docOrigin="ZooKeys 927" docSource="http://dx.doi.org/10.3897/zookeys.927.49584" docTitle="Nototylus fryi 1863" docType="treatment" docVersion="7" id="DDBCBAC4F1D65BB280F9A6989755996C" lastPageNumber="65" masterDocId="DDBCBAC4F1D65BB280F9A6989755996C" masterDocTitle="After 157 years, a second specimen and species of the phylogenetically enigmatic and previously monobasic genus Nototylus Gemminger & Harold, 1868 (Coleoptera, Carabidae, Nototylini)" masterLastPageNumber="74" masterPageNumber="65" pageNumber="65" updateTime="1732938983461" updateUser="ExternalLinkService">
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<mods:title id="CA9A9F68D5FADA28AA1B4EE9721C4E2B">After 157 years, a second specimen and species of the phylogenetically enigmatic and previously monobasic genus Nototylus Gemminger & Harold, 1868 (Coleoptera, Carabidae, Nototylini)</mods:title>
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<mods:namePart id="5AC8DACDDAFA096E782CB378C4E96D28">Erwin, Terry L.</mods:namePart>
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<mods:affiliation id="DCEF0E5F3638B3F3F98499150E963E37">Hyper-diversity Group, Department of Entomology, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA</mods:affiliation>
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<mods:nameIdentifier id="57A5BF160F8A2C7E8E3AF3A6BB069AE3" type="email">erwint@si.edu</mods:nameIdentifier>
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<mods:namePart id="66D4A578288066B3636946BD3FB63BEF">Kavanaugh, David H.</mods:namePart>
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<mods:affiliation id="F79E272D250F8946D5393D4DAC99F10D">Curator Emeritus, Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:namePart id="3B1258CB2CF089363B2D599E34852F3B">Maddison, David R.</mods:namePart>
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<mods:affiliation id="A6F4A5CF147F4FFABC0BB1A1B7F0546D">Department of Integrative Biology, 3029 Cordley Hall, Oregon State University, Corvallis, OR 97331, USA</mods:affiliation>
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<mods:nameIdentifier id="B77D07E8BCB872B633B6E95A3B84A522" type="orcid">https://orcid.org/0000-0002-7152-3824</mods:nameIdentifier>
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<mods:typeOfResource id="5888E694658AF334953852B700C84F08">text</mods:typeOfResource>
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<mods:title id="10677D1699F38D0553AB9A18D8278DB2">ZooKeys</mods:title>
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<mods:date id="D2464485B9B5C5E6880F491B3E4C170A">2020</mods:date>
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<mods:detail id="0DE8FD382E83EAEE1CEF1F0393FEA111" type="volume">
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<mods:number id="2CCD8A78758D1E108102D95510629305">927</mods:number>
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<mods:start id="B9FBEB42B5D8BC423BBE20445B46085A">65</mods:start>
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<mods:end id="65C46AEE8C43452E4FD895B195EEAB9A">74</mods:end>
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<mods:location id="590AEE78D10D25304CDD35373F3A73D5">
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<mods:url id="5C7928EE0D0358FF727BB0AC9F597618">http://dx.doi.org/10.3897/zookeys.927.49584</mods:url>
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<mods:classification id="3CCE6DD2345C42E0526FEA3E8FBFE2A7">journal article</mods:classification>
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<mods:identifier id="4284C64E4382955B79D6A20DD36F82A5" type="DOI">http://dx.doi.org/10.3897/zookeys.927.49584</mods:identifier>
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<mods:identifier id="92CDB8B0B62433425603A8066689043D" type="Pensoft-Pub">1313-2970-927-65</mods:identifier>
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<treatment id="1DCA7A60040950E7BC61B4FC693618AE" ID-GBIF-Taxon="163643494" LSID="urn:lsid:plazi:treatment:1DCA7A60040950E7BC61B4FC693618AE" httpUri="http://treatment.plazi.org/id/1DCA7A60040950E7BC61B4FC693618AE" lastPageNumber="65" pageId="0" pageNumber="65" scope_family="Carabidae" scope_order="Coleoptera">
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<subSubSection id="3A67110DCB6DE54011E52BE8E8EBA4F0" pageId="0" pageNumber="65" type="nomenclature">
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<paragraph id="29842DA0A48FE1C8672D64C1E95382B9" pageId="0" pageNumber="65">
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<taxonomicName id="5E54B84A6CA60F3D5E4326E2489EEEF4" ID-CoL="47ZBB" LSID="1DCA7A60-0409-50E7-BC61-B4FC693618AE" authority="(Schaum), 1863" authorityName="" authorityYear="1863" baseAuthorityName="Schaum" baseAuthorityYear="1863" class="Insecta" family="Carabidae" genus="Nototylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nototylus fryi" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="fryi">Nototylus fryi (Schaum), 1863</taxonomicName>
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<figureCitation id="C0F9F66F3633E832072894F572965BBB" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Nototylus 1 Nototylus fryi (Schaum), habitus, dorsal aspect, apparent body length (ABL) = 8.2 mm 2 - 6 Nototylus balli sp. nov.: 2 habitus, dorsal aspect, ABL = 9.1 mm 3 habitus, left lateral aspect 4 head, dorsal aspect 5 head, left lateral aspect; arrow indicates location of sulcus beneath eye 6 left foreleg, lateral aspect; femur with antennal cleaner present subapicoventrally 7 left mesotibia, ventral aspect 8 closeup of a middle leg spatulate seta. Scale bars: 0.5 mm (1 - 8)." figureDoi="10.3897/zookeys.927.49584.figures1-8" httpUri="https://binary.pensoft.net/fig/400931" pageId="0" pageNumber="65">
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<normalizedToken id="A9D477EB0B62606484BB0B7B85837AB8" originalValue="Fry’s">Fry's</normalizedToken>
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strange-combed beetle Figures 1
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</figureCitation>
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<figureCitation id="2F8214B2DE934658BC2990612C16A979" captionStart="Figure 9" captionStartId="F2" captionText="Figure 9. Map illustrating known distributions of species of Nototylus. Key: ■ = precise locality for N. balli sp. nov. (see text); ● = generalized locality for N. fryi (Schaum) in Brazil." figureDoi="10.3897/zookeys.927.49584.figure9" httpUri="https://binary.pensoft.net/fig/400932" pageId="0" pageNumber="65">, 9</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="BDD707919BC5C33E31F844F8BB8215DA" pageId="0" pageNumber="65" type="reference_group">
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<paragraph id="FA4236C0AA499A12D615669491FB50CB" pageId="0" pageNumber="65">
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<taxonomicName id="BA135D3886504BE7E4B7D3B645EFE7A6" authorityName="Schaum" authorityYear="1863" class="Insecta" family="Carabidae" genus="Tylonotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylonotus fryi" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="fryi">Tylonotus fryi</taxonomicName>
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Schaum, 1863: 75.
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</paragraph>
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<paragraph id="9D38DF928A216BC8F99117BB1F729B0D" pageId="0" pageNumber="65">
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<taxonomicName id="343E6A335263E1C3C07B92EFB6D57D00" authorityName="" authorityYear="1863" baseAuthorityName="Schaum" baseAuthorityYear="1863" class="Insecta" family="Carabidae" genus="Nototylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nototylus fryi" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="fryi">Nototylus fryi</taxonomicName>
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(Schaum): Gemminger & Harold, 1868: 161.
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</paragraph>
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</subSubSection>
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<subSubSection id="C8FDB87BDEEC8D133CE45A2D747C895D" pageId="0" pageNumber="65" type="type material">
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<paragraph id="B639083EE797700E93F9970F149B2CC4" pageId="0" pageNumber="65">Type material.</paragraph>
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<paragraph id="9BE1966234DCABBD97E5C507984F0581" pageId="0" pageNumber="65">
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<emphasis id="8E3A1FAB9A0B9F31E6416833AD389054" bold="true" italics="true" pageId="0" pageNumber="65">Holotype</emphasis>
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female deposited in NHMUK. Detailed description and illustrations in
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<bibRefCitation id="A9D49A02CBB246553B171E1EACC77C86" author="Deuve, T" journalOrPublisher="Annales de la Societe Entomologique de France (nouvelle serie)" pageId="0" pageNumber="65" pagination="133 - 144" refId="B4" refString="Deuve, T, 1994. Les Nototylidae, une famille enigmatique de coleopteres Adephaga. Annales de la Societe Entomologique de France (nouvelle serie) 30: 133 - 144" title="Les Nototylidae, une famille enigmatique de coleopteres Adephaga." volume="30" year="1994">Deuve (1994)</bibRefCitation>
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. See also
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<bibRefCitation id="4B34008B0E17C7863F58859AB3E6E878" author="Schaum, H" journalOrPublisher="The Journal of Entomology (London)" pageId="0" pageNumber="65" pagination="74 - 78" refId="B15" refString="Schaum, H, 1863. Descriptions of four new genera of Carabidae. The Journal of Entomology (London) 2: 74 - 78" title="Descriptions of four new genera of Carabidae." volume="2" year="1863">Schaum (1863)</bibRefCitation>
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,
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<bibRefCitation id="D710D5138E6000A3BF590B9AB8852F86" DOI="https://doi.org/10.1002/mmnd.48019270301" author="Baenninger, M" journalOrPublisher="Deutsche Entomologische Zeitschrift," pageId="0" pageNumber="65" pagination="177 - 216" refId="B1" refString="Baenninger, M, 1927. Die Ozaenini (Col. Carab.). 10. Beitrag zur Kenntnis der Carabinae. Deutsche Entomologische Zeitschrift, 1927: 177 - 216, DOI: https://doi.org/10.1002/mmnd.48019270301" title="Die Ozaenini (Col. Carab.). 10. Beitrag zur Kenntnis der Carabinae." url="https://doi.org/10.1002/mmnd.48019270301" volume="1927" year="1927">
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<normalizedToken id="A010701B1155274DC6FCC64B52A9DF40" originalValue="Bänninger">Baenninger</normalizedToken>
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(1927)
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</bibRefCitation>
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, and
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<bibRefCitation id="FD64FF26B6999BED7462D6ACCAF718B8" author="Erwin, TL" journalOrPublisher="Pensoft, Sofia-Moscow" pageId="0" pageNumber="65" refId="B6" refString="Erwin, TL, 2011. A Treatise on the Western Hemisphere Caraboidea (Coleoptera): Their classification, distributions, and ways of life Volume III (Carabidae - Loxomeriformes, Melaeniformes). Pensoft, Sofia-Moscow" title="A Treatise on the Western Hemisphere Caraboidea (Coleoptera): Their classification, distributions, and ways of life Volume III (Carabidae - Loxomeriformes, Melaeniformes)." year="2011">Erwin (2011)</bibRefCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="665071A9D34A9CB38298D91CD26A305A" pageId="0" pageNumber="65" type="distribution">
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<paragraph id="60E0863100A5DBCA91003CE3651ACDBF" pageId="0" pageNumber="65">Geographical distribution</paragraph>
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<paragraph id="3C86E6CFCBC02984A14936759C53188C" pageId="0" pageNumber="65">
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(Fig.
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<figureCitation id="0124A97923794C0D77789B4F4916A6FE" captionStart="Figure 9" captionStartId="F2" captionText="Figure 9. Map illustrating known distributions of species of Nototylus. Key: ■ = precise locality for N. balli sp. nov. (see text); ● = generalized locality for N. fryi (Schaum) in Brazil." figureDoi="10.3897/zookeys.927.49584.figure9" httpUri="https://binary.pensoft.net/fig/400932" pageId="0" pageNumber="65">9</figureCitation>
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). Known only from Brazil,
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<normalizedToken id="677946326D4592B1C8C20505CFB55461" originalValue="Espíritu">Espiritu</normalizedToken>
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Santo, without precise locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="13C2ADA038F7601C9FA19A56F66D51C1" pageId="0" pageNumber="65" type="dispersal potential">
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<paragraph id="385121943C0A1954E645DD66F6C6055C" pageId="0" pageNumber="65">Dispersal potential.</paragraph>
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<paragraph id="4B2F52BF73C9FEE066072CAB51E7AC45" pageId="0" pageNumber="65">Brachypterous (wing truncated without distal venation), probably not capable of flight.</paragraph>
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</subSubSection>
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<subSubSection id="FF3190F6102A07FE23C165A0C947C819" pageId="0" pageNumber="65" type="way of life">
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<paragraph id="E771FA4A3586EBA4903A38C550E6F391" pageId="0" pageNumber="65">Way of life.</paragraph>
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<paragraph id="162C7A1E2F360D6E2CFAAA734AEF522D" pageId="0" pageNumber="65">
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Unknown, except that they live in Southern Atlantic Forest (Mata
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<normalizedToken id="C71B99C5BE6A20FB8E64E69D90852F5D" originalValue="Atlântica">Atlantica</normalizedToken>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="A46323786AC13169000DC45D8CBCE5BA" pageId="0" pageNumber="65" type="notes">
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<paragraph id="881753CFD949078A78A03E9BB409DD14" pageId="0" pageNumber="65">Note.</paragraph>
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<paragraph id="393FB8BC99272ED0E4D028659B6832E0" pageId="0" pageNumber="65">
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This taxon is known from a single disarticulated specimen in the NHMUK, which one of us (TLE) has re-examined twice. Its habitat, somewhere in the state of
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<normalizedToken id="836A070F81C57A380056D02DB96448A9" originalValue="Espíritu">Espiritu</normalizedToken>
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Santo, Brazil, likely has suffered forest conversion to sugar cane, cacao plantations, or cattle ranches. The Bahia Coastal Forests ecoregion, which includes the state of
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<normalizedToken id="352EFC8350E5D230402C2A386822D5F4" originalValue="Espíritu">Espiritu</normalizedToken>
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Santo, has less than 5% of the original forest vegetation remaining. See the web site http://www.worldwildlife.org/ecoregions/nt0103 (last accessed on 19 December 2019) for a very good description of what the area where this unique species lived was like previously and is like now (not good). Perhaps
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<taxonomicName id="3023B97FA1528A44A85CC8B7AD536987" lsidName="N. fryi" pageId="0" pageNumber="65" rank="species" species="fryi">
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<emphasis id="C7EC686555F287659F21C1BBCD525E13" italics="true" pageId="0" pageNumber="65">N. fryi</emphasis>
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</taxonomicName>
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is still extant in remaining protected areas such as Sooretama Biological Reserve and/or Linhares Forest Reserve. A major effort needs to be made to seek more specimens, particularly males, which remain unknown.
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</paragraph>
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<caption id="E7FA39821F3264D1357ED1D68507924A" doi="10.3897/zookeys.927.49584.figure9" httpUri="https://binary.pensoft.net/fig/400932" pageId="0" pageNumber="65" start="Figure 9" startId="F2">
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<paragraph id="BC85FF20A7520B1B83AB79E916F98B1B" pageId="0" pageNumber="65">
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<emphasis id="B7E370EF49FDC201A012DCE74A3C0250" bold="true" pageId="0" pageNumber="65">Figure 9.</emphasis>
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Map illustrating known distributions of species of
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<taxonomicName id="16099926252B5089783C1641C2CEA1A7" authorityName="Gemminger & Harold" authorityYear="1868" class="Insecta" family="Carabidae" genus="Nototylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nototylus" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="genus">
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<emphasis id="D2F64224699460F36FB66151433AE20D" italics="true" pageId="0" pageNumber="65">Nototylus</emphasis>
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</taxonomicName>
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. Key: ■ = precise locality for
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<taxonomicName id="1605394CDC8163A9766EE4D216091D90" lsidName="N. balli" pageId="0" pageNumber="65" rank="species" species="balli">
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<emphasis id="83948B9FAF55C606209D90E21AE3DC20" italics="true" pageId="0" pageNumber="65">N. balli</emphasis>
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</taxonomicName>
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sp. nov. (see text); ● = generalized locality for
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<taxonomicName id="E7430B11FC298D0FF3F8519CAF853AAF" lsidName="N. fryi" pageId="0" pageNumber="65" rank="species" species="fryi">
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<emphasis id="58F5C86FED8758EA6B2FE214CBADF345" italics="true" pageId="0" pageNumber="65">N. fryi</emphasis>
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</taxonomicName>
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(Schaum) in Brazil.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |