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<document id="EE0ECA78EA99A0B65F957A77B50A6429" ID-CLB-Dataset="30629" ID-DOI="10.11646/zootaxa.4387.3.11" ID-GBIF-Dataset="c4a29c39-169e-4f88-8303-38ec5207aff6" ID-ISSN="1175-5326" ID-Zenodo-Dep="1187629" ID-ZooBank="F712D710-6557-441F-9F20-D17EAD850A96" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520059213900" checkinUser="plazi" docAuthor="Guilbert, Eric, Pham, Hong Thai &amp; Soulier-Perkins, Adeline" docDate="2018" docId="1D63878B9E0FFFC499ECFD3EFE9D0614" docLanguage="en" docName="zootaxa.4387.3.11.pdf" docOrigin="Zootaxa 4387 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cetiothucha constanti Guilbert &amp; Pham &amp; Soulier-Perkins 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="595" masterDocId="E15AFFF39E0CFFC0997BFFB9FFB30303" masterDocTitle="New species and new records of Tingidae (Insecta: Heteroptera) from Vietnam" masterLastPageNumber="600" masterPageNumber="591" pageNumber="594" updateTime="1698525736698" updateUser="ExternalLinkService">
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<mods:title id="178C34D24E7EC027E2F497F1671B62B9">New species and new records of Tingidae (Insecta: Heteroptera) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="8DB68F456C7B26AA17B0F70FA4C7A691" type="personal">
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<mods:namePart id="448FECC2C4FE1ADF4C7BEEC1216388BC">Guilbert, Eric</mods:namePart>
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<mods:namePart id="E531F2BAC27A65C396ECC36506310CBA">Pham, Hong Thai</mods:namePart>
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<mods:namePart id="AA018B7EC80161C1F263B26372915692">Soulier-Perkins, Adeline</mods:namePart>
</mods:name>
<mods:typeOfResource id="52B99552DC3C6FBB85D4DAB29100C8D9">text</mods:typeOfResource>
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<mods:title id="F45836454F37CDA1D45153C6F01BDC9C">Zootaxa</mods:title>
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<mods:part id="3DC49F4FB31E0843B5946E8418627E29">
<mods:date id="2FF0DC6FF9756DAF84414EFC75D3B4E5">2018</mods:date>
<mods:detail id="579B55C356ED5586B81BEEB5E36EBCC5" type="pubDate">
<mods:number id="CA45C35FF94F02700002D9E91992D537">2018-02-28</mods:number>
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<mods:number id="F9FB13FCC4C2C2D801CF0F1839F1402B">4387</mods:number>
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<mods:number id="077763CFC306B0F89C2B3932D477195C">3</mods:number>
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<mods:classification id="4B2531F4BD99402DD56F58D95FABB832">journal article</mods:classification>
<mods:identifier id="863E290CA797FAFF43513F762CABA941" type="CLB-Dataset">30629</mods:identifier>
<mods:identifier id="7E64A9F2B90CDA25412FA88DEDADA4BA" type="DOI">10.11646/zootaxa.4387.3.11</mods:identifier>
<mods:identifier id="99502F55AB4C86E2CC64310DB4201A79" type="GBIF-Dataset">c4a29c39-169e-4f88-8303-38ec5207aff6</mods:identifier>
<mods:identifier id="22FDB328CCC54CA28DF0CCA6B703BB9E" type="ISSN">1175-5326</mods:identifier>
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<treatment id="1D63878B9E0FFFC499ECFD3EFE9D0614" ID-DOI="http://doi.org/10.5281/zenodo.5970278" ID-GBIF-Taxon="141263953" ID-Zenodo-Dep="5970278" LSID="urn:lsid:plazi:treatment:1D63878B9E0FFFC499ECFD3EFE9D0614" httpUri="http://treatment.plazi.org/id/1D63878B9E0FFFC499ECFD3EFE9D0614" lastPageId="4" lastPageNumber="595" pageId="3" pageNumber="594">
<subSubSection id="DDD065169E0FFFC399ECFD3EFEA201C0" pageId="3" pageNumber="594" type="nomenclature">
<paragraph id="9575369D9E0FFFC399ECFD3EFDB601A1" blockId="3.[151,517,647,707]" box="[151,517,647,674]" pageId="3" pageNumber="594">
<heading id="CE3D81F19E0FFFC399ECFD3EFDB601A1" bold="true" box="[151,517,647,674]" fontSize="11" level="1" pageId="3" pageNumber="594" reason="1">
<emphasis id="A7BEEA8F9E0FFFC399ECFD3EFDB601A1" bold="true" box="[151,517,647,674]" pageId="3" pageNumber="594">
<taxonomicName id="52CA4D1E9E0FFFC399ECFD3EFE1301A2" authority="Guilbert &amp; Pham &amp; Soulier-Perkins, 2018" authorityName="Guilbert &amp; Pham &amp; Soulier-Perkins" authorityYear="2018" box="[151,416,647,673]" class="Insecta" family="Tingidae" genus="Cetiothucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="constanti" status="sp. nov.">
<emphasis id="A7BEEA8F9E0FFFC399ECFD3EFE1301A2" bold="true" box="[151,416,647,673]" italics="true" pageId="3" pageNumber="594">Cetiothucha constanti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BC8D57F49E0FFFC398D3FD31FDB601A1" box="[424,517,648,674]" pageId="3" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9575369D9E0FFFC399ECFD13FEA201C0" blockId="3.[151,517,647,707]" box="[151,273,682,707]" pageId="3" pageNumber="594">
(
<figureCitation id="0DF12A189E0FFFC399E4FD13FF5E01C0" box="[159,237,682,707]" captionStart="FIGURE 3" captionStartId="3.[151,250,1865,1887]" captionTargetBox="[179,1397,1066,1833]" captionTargetId="figure@3.[162,1411,1047,1846]" captionTargetPageId="3" captionText="FIGURE 3. Dorsal habitus of Cetiothucha constanti (scale bar is 1 mm)." httpUri="https://zenodo.org/record/1187635/files/figure.png" pageId="3" pageNumber="594">Figs. 3</figureCitation>
,
<figureCitation id="0DF12A189E0FFFC39981FD12FEBA01C0" box="[250,265,683,707]" captionStart="FIGURE 4" captionStartId="4.[151,250,599,621]" captionTargetBox="[170,1420,210,560]" captionTargetId="figure@4.[151,1437,193,578]" captionTargetPageId="4" captionText="FIGURE 4. Lateral habitus of Cetiothucha constanti (scale bar is 1 mm)." httpUri="https://zenodo.org/record/1187637/files/figure.png" pageId="3" pageNumber="594">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DDD065169E0FFFC399ECFD4BFD96002C" pageId="3" pageNumber="594" type="materials_examined">
<paragraph id="9575369D9E0FFFC399ECFD4BFD96002C" blockId="3.[151,1437,754,995]" pageId="3" pageNumber="594">
<emphasis id="A7BEEA8F9E0FFFC399ECFD4BFEFB0008" bold="true" box="[151,328,754,779]" pageId="3" pageNumber="594">Type material.</emphasis>
Holotype, 1 M, Cat Tien NP, 11°26’N, 107°26’E, 6/
<date id="E174105D9E0FFFC39ACAFD4BFB860008" box="[945,1077,754,779]" pageId="3" pageNumber="594" value="2012-04-16">16-IV-2012</date>
,
<collectingMethod id="4C8B4E8A9E0FFFC39D38FD4BFB670009" box="[1091,1236,754,779]" pageId="3" pageNumber="594">Malaise trap</collectingMethod>
, Leg Constant &amp; Bresseel, I.G. 32.161, IRScNB coll.
</paragraph>
</subSubSection>
<subSubSection id="DDD065169E0FFFC499BCFC83FBBE00F5" lastPageId="4" lastPageNumber="595" pageId="3" pageNumber="594" type="description">
<paragraph id="9575369D9E0FFFC399BCFC83FBD00050" blockId="3.[151,1437,754,995]" box="[199,1123,826,851]" pageId="3" pageNumber="594">
<emphasis id="A7BEEA8F9E0FFFC399BCFC83FEEA0050" bold="true" box="[199,345,826,851]" pageId="3" pageNumber="594">Description.</emphasis>
Color uniformly yellowish, Body glabrous, length, 2.92; width, 1.88.
</paragraph>
<paragraph id="9575369D9E0FFFC399BCFCE7FBEA00E0" blockId="3.[151,1437,754,995]" pageId="3" pageNumber="594">Head small, short, armed with three spines, an occipital pair and a median spine, spines long, the occipital spine extending to the anterior margin of eyes, slender, lying on head and directed forward; bucculae short, not meeting in front; labium long, extending to the first visible abdominal segment; antennae long and slender, with short setae all along, antennal segment measurements I, 0.16; II, 0.0 8, III, 0.8; IV, 0.8.</paragraph>
<caption id="C1B566159E0FFFC399ECF8F0FCCF045C" box="[151,892,1865,1887]" httpUri="https://zenodo.org/record/1187635/files/figure.png" pageId="3" pageNumber="594" startId="3.[151,250,1865,1887]" targetBox="[179,1397,1066,1833]" targetPageId="3">
<paragraph id="9575369D9E0FFFC399ECF8F0FCCF045C" blockId="3.[151,892,1865,1887]" box="[151,892,1865,1887]" pageId="3" pageNumber="594">
<emphasis id="A7BEEA8F9E0FFFC399ECF8F0FEA1045D" bold="true" box="[151,274,1865,1887]" pageId="3" pageNumber="594">FIGURE 3.</emphasis>
Dorsal habitus of
<taxonomicName id="52CA4D1E9E0FFFC398B6F8F0FD1E045C" authorityName="Guilbert &amp; Pham &amp; Soulier-Perkins" authorityYear="2018" box="[461,685,1865,1887]" class="Insecta" family="Tingidae" genus="Cetiothucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="constanti">
<emphasis id="A7BEEA8F9E0FFFC398B6F8F0FD1E045C" box="[461,685,1865,1887]" italics="true" pageId="3" pageNumber="594">Cetiothucha constanti</emphasis>
</taxonomicName>
(scale bar is 1 mm).
</paragraph>
</caption>
<caption id="C1B566159E08FFC499ECFDEEFC33016E" box="[151,896,599,622]" httpUri="https://zenodo.org/record/1187637/files/figure.png" pageId="4" pageNumber="595" startId="4.[151,250,599,621]" targetBox="[170,1420,210,560]" targetPageId="4">
<paragraph id="9575369D9E08FFC499ECFDEEFC33016E" blockId="4.[151,896,599,622]" box="[151,896,599,622]" pageId="4" pageNumber="595">
<emphasis id="A7BEEA8F9E08FFC499ECFDEEFEA1016F" bold="true" box="[151,274,599,621]" pageId="4" pageNumber="595">FIGURE 4.</emphasis>
Lateral habitus of
<taxonomicName id="52CA4D1E9E08FFC498A9FDEEFD02016D" authorityName="Guilbert &amp; Pham &amp; Soulier-Perkins" authorityYear="2018" box="[466,689,599,622]" class="Insecta" family="Tingidae" genus="Cetiothucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="constanti">
<emphasis id="A7BEEA8F9E08FFC498A9FDEEFD02016D" box="[466,689,599,622]" italics="true" pageId="4" pageNumber="595">Cetiothucha constanti</emphasis>
</taxonomicName>
(scale bar is 1 mm).
</paragraph>
</caption>
<paragraph id="9575369D9E08FFC499BCFD22FDFC0040" blockId="4.[151,1436,666,1303]" pageId="4" pageNumber="595">Pronotum concealed by hood and median carina developed to form a cyst covering all the pronotum; hood longitudinally oval, covering head except the eyes, with large, polygonal areolae, length, 1.38; width, 0.84; height, 0.80; paranota wide, raised and strongly curved upward to conceal the hood, with two rows of areolae, areolae smaller than on hood; mesosternal and metasternal laminae narrow, with one row of small areolae; rostral sulcus widened posteriorly and closed behind.</paragraph>
<paragraph id="9575369D9E08FFC499BCFCF7FBBE00F5" blockId="4.[151,1436,666,1303]" pageId="4" pageNumber="595">Hemelytra much larger than body beneath, progressively widening posteriorly, apex divergent; costal area triseriate at the middle, biseriate in front and posteriorly, areolae polygonal, as large as on hood; subcostal area narrow and uniseriate, areolae small opposite to discoidal area, larger posterior to discoidal area; discoidal area shorter than half the length of hemelytra, five areolae wide at widest part, areolae smaller than on costal area; sutural area three areolae wide at widest part, areolae as large as on costal area.</paragraph>
</subSubSection>
<subSubSection id="DDD065169E08FFC499BCFBBBFE0D073D" pageId="4" pageNumber="595" type="diagnosis">
<paragraph id="9575369D9E08FFC499BCFBBBFE0D073D" blockId="4.[151,1436,666,1303]" pageId="4" pageNumber="595">
<emphasis id="A7BEEA8F9E08FFC499BCFBBBFEF20718" bold="true" box="[199,321,1026,1051]" pageId="4" pageNumber="595">Diagnosis.</emphasis>
This species is characterized by the two occipital spines and a median frontal spines on the head and the biseriate paranota.
</paragraph>
</subSubSection>
<subSubSection id="DDD065169E08FFC499BCFBF3FD510784" pageId="4" pageNumber="595" type="etymology">
<paragraph id="9575369D9E08FFC499BCFBF3FD510784" blockId="4.[151,1436,666,1303]" pageId="4" pageNumber="595">
<emphasis id="A7BEEA8F9E08FFC499BCFBF3FEFD0760" bold="true" box="[199,334,1098,1123]" pageId="4" pageNumber="595">Etymology.</emphasis>
This new species is named in honor of Jérôme Constant (IRScNB,
<collectingCountry id="EDDD760D9E08FFC49D3FFBF3FB1A0760" box="[1092,1193,1098,1123]" name="Belgium" pageId="4" pageNumber="595">Belgium</collectingCountry>
), who provided many of the Vietnamese specimens studied by the authors.
</paragraph>
</subSubSection>
<subSubSection id="DDD065169E08FFC499BCFB2BFE9D0614" pageId="4" pageNumber="595" type="discussion">
<paragraph id="9575369D9E08FFC499BCFB2BFE9D0614" blockId="4.[151,1436,666,1303]" pageId="4" pageNumber="595">
<emphasis id="A7BEEA8F9E08FFC499BCFB2BFEFC07A8" bold="true" box="[199,335,1170,1195]" pageId="4" pageNumber="595">Comments.</emphasis>
The habitus of this new species is similar to the type species
<taxonomicName id="52CA4D1E9E08FFC49D5BFB2AFAD807A8" authority="Drake &amp; Ruhoff" authorityName="Drake &amp; Ruhoff" authorityYear="1965" box="[1056,1387,1170,1195]" class="Insecta" family="Tingidae" genus="Cetiothucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="physalia">
<emphasis id="A7BEEA8F9E08FFC49D5BFB2AFB1107A8" box="[1056,1186,1170,1195]" italics="true" pageId="4" pageNumber="595">C. physalia</emphasis>
Drake &amp; Ruhoff
</taxonomicName>
, but differs by the presence of two occipital spines and a median frontal spine on head, the paranota are biseriate with an outer row of large quadrate areolae and an inner row of smaller areolae.
<taxonomicName id="52CA4D1E9E08FFC49AC1FB63FB1B07F0" authorityName="Drake &amp; Ruhoff" authorityYear="1965" box="[954,1192,1242,1267]" class="Insecta" family="Tingidae" genus="Cetiothucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="physalia">
<emphasis id="A7BEEA8F9E08FFC49AC1FB63FB1B07F0" box="[954,1192,1242,1267]" italics="true" pageId="4" pageNumber="595">Cetiothucha physalia</emphasis>
</taxonomicName>
is known from Sabah and Sarawak.
</paragraph>
</subSubSection>
</treatment>
</document>
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