675 lines
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675 lines
107 KiB
XML
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<mods:title id="71C2B15AA92FF26CD11310DCAF3FCE88">Ophioderma hendleri sp. nov. (Echinodermata: Ophiuroidea: Ophiodermatidae) and its congeners from the Eastern Pacific</mods:title>
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<mods:namePart id="52C9FAD497CE21541AEB5275D3C6DBF1">Granja-Fernández, Rebeca</mods:namePart>
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<mods:affiliation id="00D9927CA5BE59773CA4285239302C00">Laboratorio de Ecología Marina. Centro Universitario de la Costa, Universidad de Guadalajara, 48280 Puerto Vallarta, Mexico. Department of Biology, Division of Invertebrate Zoology, Florida Museum of Natural History, University of Florida, Gainesville, 32611 Florida, USA. Natural History Museum of Los Angeles County, 900 Exposition Blvd., 90007 Los Angeles, California, USA. Colección Nacional de Equinodermos “ Ma. E. Caso Muñoz ”, Laboratorio de Sistemática y Ecología de Equinodermos, Instituto de Ciencias del Mar y Limnología (ICML), Universidad Nacional Autónoma de Mexico (UNAM), 04510 Ciudad de Mexico, Mexico.</mods:affiliation>
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<taxonomicName id="4C4121995336FFF9FD9B9F6AFBCEC500" authority="Lutken, 1856" authorityName=", Lutken" authorityYear="1856" box="[564,1024,264,291]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="20" pageNumber="31" phylum="Echinodermata" rank="species" species="variegatum">
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<emphasis id="B93586085336FFF9FD9B9F6AFC92C501" bold="true" box="[564,860,264,291]" italics="true" pageId="20" pageNumber="31">Ophioderma variegatum</emphasis>
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<bibRefCitation id="EFD027EB5336FFF9FCCC9F6AFBCEC500" author="Lutken C. F." box="[867,1024,264,291]" pageId="20" pageNumber="31" pagination="20 - 26" refId="ref16758" refString="Lutken C. F. 1856. III. Bidrag til Kundskab om Ophiurerne ved Central-Amerikas Vestkyst. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening 1856: 20 - 26." type="journal article" year="1856">Lütken, 1856</bibRefCitation>
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<figureCitation id="137A469F5336FFF9FD1B9F4FFCD3C564" box="[692,797,301,327]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="21.[189,232,1964,1990]" captionStartId-1="22.[189,232,1752,1778]" captionTargetBox-0="[189,1398,1121,1926]" captionTargetBox-1="[289,1303,265,1714]" captionTargetId-0="figure-301@21.[189,1398,1121,1926]" captionTargetId-1="figure-151@22.[289,1303,265,1714]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="Fig. 8. Ophioderma variegatum Lütken, 1856, lectotype (NHMD-107680, DD = 12.4 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Dorsal arm. E. Ventral arm. F. Dorsal view." captionText-1="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi-0="http://doi.org/10.5281/zenodo.4326631" figureDoi-1="http://doi.org/10.5281/zenodo.4326633" httpUri-0="https://zenodo.org/record/4326631/files/figure.png" httpUri-1="https://zenodo.org/record/4326633/files/figure.png" pageId="20" pageNumber="31">Figs 8–9</figureCitation>
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;
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<tableCitation id="C6C36FA15336FFF9FC869F4FFC4EC564" box="[809,896,301,327]" captionStart="Table 1" captionStartId="10.[189,258,265,291]" captionTargetPageId="10" captionText="Table 1. Comparison of morphological characters between some species of Ophioderma from the Eastern Pacific." httpUri="http://table.plazi.org/id/DF3E0A925328FFE7FF129F6BFEBAC565" pageId="20" pageNumber="31" tableUuid="DF3E0A925328FFE7FF129F6BFEBAC565">Table 1</tableCitation>
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<emphasis id="B93586085336FFF9FF129F0DFE07C5AA" box="[189,457,367,393]" italics="true" pageId="20" pageNumber="31">Ophioderma variegata</emphasis>
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<bibRefCitation id="EFD027EB5336FFF9FE7F9F0DFD52C5AA" author="Lutken C. F." box="[464,668,367,394]" pageId="20" pageNumber="31" pagination="20 - 26" refId="ref16758" refString="Lutken C. F. 1856. III. Bidrag til Kundskab om Ophiurerne ved Central-Amerikas Vestkyst. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening 1856: 20 - 26." type="journal article" year="1856">Lütken, 1856: 21</bibRefCitation>
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.
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<emphasis id="B93586085336FFF9FF129FD3FE07C5E8" box="[189,457,433,459]" italics="true" pageId="20" pageNumber="31">Ophioderma variegata</emphasis>
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–
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<treatmentCitation id="0AE07C0B5336FFF9FE489FD3FD75C5EF" author="Lutken C. F." box="[487,699,433,460]" page="199" pageId="20" pageNumber="31" year="1859">
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<bibRefCitation id="EFD027EB5336FFF9FE489FD3FD75C5EF" author="Lutken C. F." box="[487,699,433,460]" pageId="20" pageNumber="31" pagination="75 - 169" refId="ref16788" refString="Lutken C. F. 1859. Additamenta ad Historiam Ophiuridarum. Beskrivelser af de fra Vestindien og Central Amerikas Vestkyst hidtil beckjendte Slangestjerner. Kongelige Danske Videnskabernes Selskab Biologiske Skrifter 2: 75 - 169." type="journal article" year="1859">Lütken 1859: 199</bibRefCitation>
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. —
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<treatmentCitation id="0AE07C0B5336FFF9FD439FD3FB53C5EF" author="Lutken C. F. & Mortensen T." box="[748,1181,433,460]" page="100" pageId="20" pageNumber="31" year="1899">
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<bibRefCitation id="EFD027EB5336FFF9FD439FD3FB53C5EF" author="Lutken C. F. & Mortensen T." box="[748,1181,433,460]" pageId="20" pageNumber="31" pagination="93 - 208" refId="ref16825" refString="Lutken C. F. & Mortensen T. 1899. Reports on an exploration off the west coast of Mexico, central and South America, and off the Galapagos Islands, in charge of Alexander Agassiz, by the U. S. Fish Commission Steamer " Albatross ", during 1891, Lieut. Commander Z. L. Tanner, U. S. N., commanding. XXV. The Ophiuridae. Memoirs of the Museum of Comparative Zoology at Harvard College 23 (2): 93 - 208. https: // doi. org / 10.5962 / bhl. title. 65637" type="journal article" year="1899">Lütken & Mortensen 1899: 100–101</bibRefCitation>
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, pl. 1, figs 1–3. —
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<treatmentCitation id="0AE07C0B5336FFF9FF459FB4FE00C5D3" author="Koehler R." box="[234,462,470,496]" page="282" pageId="20" pageNumber="31" year="1907">
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<bibRefCitation id="EFD027EB5336FFF9FF459FB4FE00C5D3" author="Koehler R." box="[234,462,470,496]" pageId="20" pageNumber="31" pagination="279 - 370" refId="ref16635" refString="Koehler R. 1907. Revision de la collection des ophiures du Museum D'Histoire Naturelle de Paris. Bulletin Scientifique de la France et de la Belgique 61: 279 - 370." type="journal article" year="1907">Koehler 1907: 282</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AE07C0B5336FFF9FE749FB4FDF9C5D3" author="Koehler R." box="[475,567,470,496]" page="7" pageId="20" pageNumber="31" year="1914">
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<bibRefCitation id="EFD027EB5336FFF9FE749FB4FDF9C5D3" author="Koehler R." box="[475,567,470,496]" pageId="20" pageNumber="31" pagination="1 - 173" refId="ref16668" refString="Koehler R. 1914. A contribution to the study of ophiurans of the United States National Museum. Bulletin of the United States National Museum 84: 1 - 173. https: // doi. org / 10.5962 / bhl. title. 27414" type="journal article" year="1914">1914: 7</bibRefCitation>
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</treatmentCitation>
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. — H.L.
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<treatmentCitation id="0AE07C0B5336FFF9FD059FB4FCBEC5D3" author="Clark H. L." box="[682,880,469,496]" page="205" pageId="20" pageNumber="31" year="1913">
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<bibRefCitation id="EFD027EB5336FFF9FD059FB4FCBEC5D3" author="Clark H. L." box="[682,880,469,496]" pageId="20" pageNumber="31" pagination="185 - 236" refId="ref15486" refString="Clark H. L. 1913. Echinoderms from Lower California, with descriptions of new species. Bulletin of the American Museum of Natural History 32: 185 - 236. https: // doi. org / 10.5962 / bhl. title. 1734" type="journal article" year="1913">Clark 1913: 205</bibRefCitation>
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</treatmentCitation>
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. —
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<treatmentCitation id="0AE07C0B5336FFF9FC0C9FB4FB75C5D3" author="Ziesenhenne F. C." box="[931,1211,469,496]" page="198" pageId="20" pageNumber="31" year="1955">
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<bibRefCitation id="EFD027EB5336FFF9FC0C9FB4FB75C5D3" author="Ziesenhenne F. C." box="[931,1211,469,496]" pageId="20" pageNumber="31" pagination="185 - 201" refId="ref18074" refString="Ziesenhenne F. C. 1955. A review of the genus Ophioderma Muller and Troschel. In: Allan Hancock Foundation for Scientific Research (eds) Essays in the Natural Sciences in Honor of Captain Allan Hancock: 185 - 201. University of Southern California Press, Los Angeles." type="book chapter" year="1955">Ziesenhenne 1955: 198</bibRefCitation>
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</treatmentCitation>
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. —
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<treatmentCitation id="0AE07C0B5336FFF9FB419FB4FEC5C637" author="Luke S. R." page="33" pageId="20" pageNumber="31" year="1982">
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<bibRefCitation id="EFD027EB5336FFF9FB419FB4FEC5C637" author="Luke S. R." pageId="20" pageNumber="31" refId="ref16714" refString="Luke S. R. 1982. Catalog of the Benthic Invertebrate Collections of the SCRIPPS Institution of Oceanography, Echinodermata. SCRIPPS Institution of Oceanography Reference Series, no. 82 - 5, University of California, San Diego. CA." type="book" year="1982">Luke 1982: 33</bibRefCitation>
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</treatmentCitation>
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. —
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||
<bibRefCitation id="EFD027EB5336FFF9FE949F98FD01C637" author="Granja-Fernandez R. & Herrero-Perezrul M. D. & Lopez-Perez A. & Hernandez-Morales A. & Rangel-Solis P. D." box="[315,719,506,533]" pageId="20" pageNumber="31" pagination="37 - 47" refId="ref16005" refString="Granja-Fernandez R., Herrero-Perezrul M. D., Lopez-Perez A., Hernandez-Morales A. & Rangel-Solis P. D. 2015 a. A literature review of the Ophiuroidea (Echinodermata) from the Pacific coast of Mexico. Revista de Biologia Tropical 63 (2): 37 - 47. https: // doi. org / 10.15517 / RBT. V 63 I 2.23127" type="journal article" year="2015">
|
||
Granja-Fernández
|
||
<emphasis id="B93586085336FFF9FDB99F99FD84C637" box="[534,586,506,532]" italics="true" pageId="20" pageNumber="31">et al</emphasis>
|
||
. 2015a: 41
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AE07C0B5336FFF9FD739F98FC98C636" author="Granja-Fernandez R. & Solis-Marin F. A. & Benitez-Villalobos F. & Herrero-Perezrul M. D. & Lopez-Perez A." box="[732,854,506,533]" page="95" pageId="20" pageNumber="31" year="2015">
|
||
<bibRefCitation id="EFD027EB5336FFF9FD739F98FC98C636" author="Granja-Fernandez R. & Solis-Marin F. A. & Benitez-Villalobos F. & Herrero-Perezrul M. D. & Lopez-Perez A." box="[732,854,506,533]" pageId="20" pageNumber="31" pagination="87 - 114" refId="ref16075" refString="Granja-Fernandez R., Solis-Marin F. A., Benitez-Villalobos F., Herrero-Perezrul M. D. & Lopez-Perez A. 2015 b. Checklist of echinoderms (Echinodermata) from the Southern Mexican Pacific: a historical review. Revista de Biologia Tropical 63 (2): 87 - 114. https: // doi. org / 10.15517 / rbt. v 63 i 2.23131" type="journal article" year="2015">2015b: 95</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5336FFF9FF129C7CFC2FC61B" blockId="20.[189,1399,433,713]" box="[189,993,542,569]" pageId="20" pageNumber="31">
|
||
<treatmentCitationGroup id="AB517D345336FFF9FF129C7CFC2FC61B" box="[189,993,542,569]" pageId="20" pageNumber="31">
|
||
<taxonomicName id="4C4121995336FFF9FF129C7CFE56C61B" baseAuthorityName="Lutken" baseAuthorityYear="1856" box="[189,408,542,568]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="20" pageNumber="31" phylum="Echinodermata" rank="species" species="variegata">
|
||
<emphasis id="B93586085336FFF9FF129C7CFE56C61B" box="[189,408,542,568]" italics="true" pageId="20" pageNumber="31">Ophiura variegata</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AE07C0B5336FFF9FE1A9C7CFD4FC61A" author="Verrill A. E." box="[437,641,542,569]" page="254" pageId="20" pageNumber="31" year="1867">
|
||
<bibRefCitation id="EFD027EB5336FFF9FE1A9C7CFD4FC61A" author="Verrill A. E." box="[437,641,542,569]" pageId="20" pageNumber="31" pagination="251 - 322" refId="ref17963" refString="Verrill A. E. 1867. V. Notes on the Radiata in the Museum of Yale College, with descriptions of new genera and species. No. 2. Notes on the echinoderms of Panama and West Coast of America, with descriptions of new genera and species. Transactions of the Connecticut Academy of Arts and Sciences 1 (2): 251 - 322." type="journal article" year="1867">Verrill 1867: 254</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AE07C0B5336FFF9FD1E9C7DFCABC61B" author="Lyman T." box="[689,869,542,569]" page="3" pageId="20" pageNumber="31" year="1875">
|
||
<bibRefCitation id="EFD027EB5336FFF9FD1E9C7DFCABC61B" author="Lyman T." box="[689,869,542,569]" pageId="20" pageNumber="31" pagination="1 - 34" refId="ref17019" refString="Lyman T. 1875. II. Ophiuridae and Astrophytidae. Zoological results of the " Hassler " Expedition. Illustrated Catalogue of the Museum of Comparative Zoology at Harvard College 8: 1 - 34." type="journal article" year="1875">Lyman 1875: 3</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AE07C0B5336FFF9FCDD9C7CFC13C61B" author="Lyman T." box="[882,989,542,568]" page="10" pageId="20" pageNumber="31" year="1882">
|
||
<bibRefCitation id="EFD027EB5336FFF9FCDD9C7CFC13C61B" author="Lyman T." box="[882,989,542,568]" pageId="20" pageNumber="31" pagination="1 - 386" refId="ref17056" refString="Lyman T. 1882. Report on the Ophiuroidea dredged by H. M. S. " Challenger " during the years 1873 - 1876. Report of the Scientific Results of the Voyage of H. M. S. " Challenger " 1873 - 1876 5 (14): 1 - 386." type="journal article" year="1882">1882: 10</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5336FFF9FF129C20FE97C6EA" blockId="20.[189,1399,433,713]" pageId="20" pageNumber="31">
|
||
<treatmentCitationGroup id="AB517D345336FFF9FF129C20FE97C6EA" pageId="20" pageNumber="31">
|
||
<taxonomicName id="4C4121995336FFF9FF129C20FE2EC67F" authorityName=", Lutken" authorityYear="1856" box="[189,480,578,604]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="20" pageNumber="31" phylum="Echinodermata" rank="species" species="variegatum">
|
||
<emphasis id="B93586085336FFF9FF129C20FE2EC67F" box="[189,480,578,604]" italics="true" pageId="20" pageNumber="31">Ophioderma variegatum</emphasis>
|
||
</taxonomicName>
|
||
– H.L.
|
||
<treatmentCitation id="0AE07C0B5336FFF9FD909C20FCCBC67F" author="Clark H. L." box="[575,773,578,605]" page="302" pageId="20" pageNumber="31" year="1915">
|
||
<bibRefCitation id="EFD027EB5336FFF9FD909C20FCCBC67F" author="Clark H. L." box="[575,773,578,605]" pageId="20" pageNumber="31" pagination="164 - 376" refId="ref15532" refString="Clark H. L. 1915. Catalogue of recent ophiurans: based on the collection of the Museum of Comparative Zoology. Memoirs of the Museum of Comparative Zoology 25 (4): 164 - 376. https: // doi. org / 10.5962 / bhl. title. 48598" type="journal article" year="1915">Clark 1915: 302</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 18, figs. 7–8; 1940: 343. —
|
||
<treatmentCitation id="0AE07C0B5336FFF9FB229C20FED7C6A3" author="Nielsen E." page="330" pageId="20" pageNumber="31" year="1932">
|
||
<bibRefCitation id="EFD027EB5336FFF9FB229C20FED7C6A3" author="Nielsen E." pageId="20" pageNumber="31" pagination="241 - 346" refId="ref17315" refString="Nielsen E. 1932. Ophiurans from the Gulf of Panama, California and the Strait of Georgia. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening 91: 241 - 346." type="journal article" year="1932">Nielsen 1932: 330– 332</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 36. —
|
||
<treatmentCitation id="0AE07C0B5336FFF9FE089C05FD60C6A2" author="Boone L." box="[423,686,614,641]" page="114" pageId="20" pageNumber="31" year="1933">
|
||
<bibRefCitation id="EFD027EB5336FFF9FE089C05FD60C6A2" author="Boone L." box="[423,686,614,641]" pageId="20" pageNumber="31" pagination="1 - 217" refId="ref15113" refString="Boone L. 1933. Scientific results of cruises of the yachts " Eagle " and " Ara ", 1921 - 1928, William K. Vanderbilt, commanding. Coelenterata, Echinodermata and Mollusca. Bulletin of the Vanderbilt Marine Museum 4: 1 - 217." type="journal article" year="1933">Boone 1933: 114–115</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 67. —
|
||
<treatmentCitation id="0AE07C0B5336FFF9FC9C9C04FB84C6A3" author="Ziesenhenne F. C." box="[819,1098,614,640]" page="227" pageId="20" pageNumber="31" year="1937">
|
||
<bibRefCitation id="EFD027EB5336FFF9FC9C9C04FB84C6A3" author="Ziesenhenne F. C." box="[819,1098,614,640]" pageId="20" pageNumber="31" pagination="209 - 239" refId="ref18033" refString="Ziesenhenne F. C. 1937. The Templeton Crocker Expedition. X. Echinoderms from the west coast of Lower California, the Gulf of California and Clarion Island. Zoologica 22 (15): 209 - 239." type="journal article" year="1937">Ziesenhenne 1937: 227</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AE07C0B5336FFF9FBD49C04FAF8C6A3" author="Maluf L. Y." box="[1147,1334,614,640]" page="84" pageId="20" pageNumber="31" year="1988">
|
||
<bibRefCitation id="EFD027EB5336FFF9FBD49C04FAF8C6A3" author="Maluf L. Y." box="[1147,1334,614,640]" pageId="20" pageNumber="31" pagination="1 - 242" refId="ref17113" refString="Maluf L. Y. 1988. Composition and distribution of the central Eastern Pacific echinoderms. Natural History Museum of Los Angeles County, Technical Report 2: 1 - 242." type="book chapter" year="1988">Maluf 1988: 84</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 203. —
|
||
<treatmentCitation id="0AE07C0B5336FFF9FEA09CE8FDAEC686" author="Solis-Marin F. A. & Laguarda-Figueras A. & Duran-Gonzalez A. & Ahearn C. G. & Torres-Vega J." box="[271,608,650,677]" page="128" pageId="20" pageNumber="31" year="2005">
|
||
<bibRefCitation id="EFD027EB5336FFF9FEA09CE8FDAEC686" author="Solis-Marin F. A. & Laguarda-Figueras A. & Duran-Gonzalez A. & Ahearn C. G. & Torres-Vega J." box="[271,608,650,677]" pageId="20" pageNumber="31" pagination="123 - 137" refId="ref17468" refString="Solis-Marin F. A., Laguarda-Figueras A., Duran-Gonzalez A., Ahearn C. G. & Torres-Vega J. 2005. Equinodermos (Echinodermata) del Golfo de California, Mexico. Revista de Biologia Tropical 53 (3): 123 - 137. https: // doi. org / 10.15517 / RBT. V 53 I 3.26770" type="journal article" year="2005">
|
||
Solís-Marín
|
||
<emphasis id="B93586085336FFF9FE0C9CE9FE19C687" box="[419,471,650,676]" italics="true" pageId="20" pageNumber="31">et al</emphasis>
|
||
. 2005: 128
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
; 2013: 570. —
|
||
<treatmentCitation id="0AE07C0B5336FFF9FCB89CE8FB56C687" author="Honey-Escandon M. & Solis-Marin F. A. & Laguarda-Figueras A." box="[791,1176,650,677]" page="63" pageId="20" pageNumber="31" year="2008">
|
||
<bibRefCitation id="EFD027EB5336FFF9FCB89CE8FB56C687" author="Honey-Escandon M. & Solis-Marin F. A. & Laguarda-Figueras A." box="[791,1176,650,677]" pageId="20" pageNumber="31" pagination="57 - 73" refId="ref16471" refString="Honey-Escandon M., Solis-Marin F. A. & Laguarda-Figueras A. 2008. Equinodermos (Echinodermata) del Pacifico Mexicano. Revista de Biologia Tropical 56 (3): 57 - 73. https: // doi. org / 10.15517 / RBT. V 56 I 3.27079" type="journal article" year="2008">
|
||
Honey-Escandón
|
||
<emphasis id="B93586085336FFF9FC469CE9FBD3C687" box="[1001,1053,650,676]" italics="true" pageId="20" pageNumber="31">et al</emphasis>
|
||
. 2008: 63
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AE07C0B5336FFF9FB699CE8FE98C6EA" author="Alvarado J. J. & Solis-Marin F. A. & Ahearn C. G." page="48" pageId="20" pageNumber="31" year="2010">
|
||
<bibRefCitation id="EFD027EB5336FFF9FB699CE8FE98C6EA" author="Alvarado J. J. & Solis-Marin F. A. & Ahearn C. G." pageId="20" pageNumber="31" pagination="45 - 56" refId="ref14865" refString="Alvarado J. J., Solis-Marin F. A. & Ahearn C. G. 2010. Echinoderm (Echinodermata) diversity in the Pacific coast of Central America. Marine Biodiversity 40: 45 - 56. https: // doi. org / 10.1007 / s 12526 - 009 - 0032 - 5" type="journal article" year="2010">
|
||
Alvarado
|
||
<emphasis id="B93586085336FFF9FA949CE9FAA1C687" box="[1339,1391,650,676]" italics="true" pageId="20" pageNumber="31">et al</emphasis>
|
||
. 2010: 48
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35B09915336FFF9FF129C93FE2FC7CC" pageId="20" pageNumber="31" type="diagnosis">
|
||
<paragraph id="8BFE5A1A5336FFF9FF129C93FEF8C728" blockId="20.[189,310,753,779]" box="[189,310,753,779]" pageId="20" pageNumber="31">
|
||
<heading id="D0B6ED765336FFF9FF129C93FEF8C728" bold="true" box="[189,310,753,779]" fontSize="11" level="3" pageId="20" pageNumber="31" reason="3">
|
||
<emphasis id="B93586085336FFF9FF129C93FEF8C728" bold="true" box="[189,310,753,779]" pageId="20" pageNumber="31">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5336FFF9FF129D42FE2FC7CC" blockId="20.[189,1399,800,1007]" pageId="20" pageNumber="31">Radial shields covered by granules. Base of the arms and distal genital slits with granulation. First ventral arm plate large, broader than long. Oral shields oval, larger than broad, proximal margin rounded with an obtuse angle, distal edge convex, lateral margins straight. Naked adoral shields, longer than broad. Up to 9 arm spines, very large and slender; dorsalmost and ventralmost a little bit shorter than the rest. In vivo coloration: dorsal disc olive-green with dark pink or red; arms with transverse olive-green and dark pink-red bands.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35B09915336FFF9FF129A7AFDD6C2C5" pageId="20" pageNumber="31" type="materials_examined">
|
||
<paragraph id="8BFE5A1A5336FFF9FF129A7AFE63C011" blockId="20.[189,429,1048,1074]" box="[189,429,1048,1074]" pageId="20" pageNumber="31">
|
||
<heading id="D0B6ED765336FFF9FF129A7AFE63C011" bold="true" box="[189,429,1048,1074]" fontSize="11" level="3" pageId="20" pageNumber="31" reason="3">
|
||
<emphasis id="B93586085336FFF9FF129A7AFE63C011" bold="true" box="[189,429,1048,1074]" pageId="20" pageNumber="31">Examined material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5336FFF9FF459A25FDF7C041" blockId="20.[189,1011,1095,1158]" box="[234,569,1095,1122]" pageId="20" pageNumber="31">
|
||
<emphasis id="B93586085336FFF9FF459A25FEA8C042" bold="true" box="[234,358,1095,1121]" pageId="20" pageNumber="31">
|
||
<typeStatus id="54FAE4B85336FFF9FF459A25FEA8C042" box="[234,358,1095,1121]" pageId="20" pageNumber="31" type="lectotype">Lectotype</typeStatus>
|
||
</emphasis>
|
||
(here designated)
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5336FFF9FF129A0EFC3DC0A6" blockId="20.[189,1011,1095,1158]" box="[189,1011,1131,1158]" pageId="20" pageNumber="31">
|
||
<materialsCitation id="3B2950475336FFF9FF129A0EFC3DC0A6" ID-GBIF-Occurrence="2995182313" box="[189,1011,1131,1158]" collectionCode="NHMD" country="Nicaragua" elevation="9" location="Realejo" pageId="20" pageNumber="31" specimenCode="NHMD-107680" specimenCount="1" typeStatus="lectotype">
|
||
<collectingCountry id="F3561A8A5336FFF9FF129A0EFEA5C0A5" box="[189,363,1132,1158]" name="Nicaragua" pageId="20" pageNumber="31">NICARAGUA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D4791935336FFF9FE2B9A0EFE17C0A5" box="[388,473,1132,1158]" pageId="20" pageNumber="31" type="generic">1 spec.</specimenCount>
|
||
;
|
||
<location id="8E9E0CC15336FFF9FE4A9A09FD8CC0A6" LSID="urn:lsid:plazi:treatment:03E8EB0C5336FFF5FD9B9F6AFD38C7B0:8E9E0CC15336FFF9FE4A9A09FD8CC0A6" box="[485,578,1131,1157]" country="Nicaragua" name="Realejo" pageId="20" pageNumber="31">Realejo</location>
|
||
;
|
||
<quantity id="4CB9F7FF5336FFF9FDE19A0EFD5FC0A5" box="[590,657,1132,1158]" metricMagnitude="0" metricUnit="m" metricValue="9.1" pageId="20" pageNumber="31" unit="m" value="9.1">
|
||
<elevation id="006CBD295336FFF9FDE19A0EFD5FC0A5" box="[590,657,1132,1158]" metricMagnitude="0" metricUnit="m" metricValue="9.1" pageId="20" pageNumber="31" unit="m" value="9.1">9.1 m</elevation>
|
||
</quantity>
|
||
(5 fathoms);
|
||
<specimenCode id="DBE7F2615336FFF9FC9F9A09FC3EC0A6" box="[816,1008,1131,1157]" pageId="20" pageNumber="31">NHMD-107680</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5336FFF9FF459ACCFDBBC0EA" blockId="20.[189,1128,1198,1261]" box="[234,629,1198,1225]" pageId="20" pageNumber="31">
|
||
<emphasis id="B93586085336FFF9FF459ACCFE6CC0EB" bold="true" box="[234,418,1198,1224]" pageId="20" pageNumber="31">
|
||
<typeStatus id="54FAE4B85336FFF9FF459ACCFE6CC0EB" box="[234,418,1198,1224]" pageId="20" pageNumber="31" type="paralectotype">Paralectotypes</typeStatus>
|
||
</emphasis>
|
||
(here designated)
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5336FFF9FF129AB0FBA6C0CF" blockId="20.[189,1128,1198,1261]" box="[189,1128,1234,1261]" pageId="20" pageNumber="31">
|
||
<materialsCitation id="3B2950475336FFF9FF129AB0FBA6C0CF" ID-GBIF-Occurrence="2995182319" box="[189,1128,1234,1261]" collectionCode="NHMD" country="Nicaragua" location="NICARAGUA" pageId="20" pageNumber="31" specimenCode="NHMD-619214" specimenCount="2">
|
||
<collectingCountry id="F3561A8A5336FFF9FF129AB0FEA5C0CF" box="[189,363,1234,1260]" name="Nicaragua" pageId="20" pageNumber="31">NICARAGUA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D4791935336FFF9FE2B9AB0FE13C0CE" box="[388,477,1234,1261]" pageId="20" pageNumber="31" type="generic">2 specs</specimenCount>
|
||
; same collection data as for lectotype;
|
||
<specimenCode id="DBE7F2615336FFF9FC0A9AB0FBAAC0CF" box="[933,1124,1234,1260]" pageId="20" pageNumber="31">NHMD-619214</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5336FFF9FF129B76FD1CC10C" blockId="20.[189,722,1300,1327]" box="[189,722,1300,1327]" pageId="20" pageNumber="31">
|
||
<heading id="D0B6ED765336FFF9FF129B76FD1CC10C" bold="true" box="[189,722,1300,1327]" fontSize="11" level="3" pageId="20" pageNumber="31" reason="3">
|
||
<emphasis id="B93586085336FFF9FF129B76FD1CC10C" bold="true" box="[189,722,1300,1327]" pageId="20" pageNumber="31">
|
||
Designation of
|
||
<typeStatus id="54FAE4B85336FFF9FED79B77FE26C10C" box="[376,488,1301,1327]" pageId="20" pageNumber="31" type="lectotype">lectotype</typeStatus>
|
||
and
|
||
<typeStatus id="54FAE4B85336FFF9FD889B77FD1CC10C" box="[551,722,1301,1327]" pageId="20" pageNumber="31" type="paralectotype">paralectotype</typeStatus>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5336FFF9FF129B26FAF9C2A3" blockId="20.[189,1399,1348,1664]" pageId="20" pageNumber="31">
|
||
In the original description of
|
||
<taxonomicName id="4C4121995336FFF9FDB69B26FC3DC17C" authority=", Lutken (1856)" authorityName=", Lutken" authorityYear="1856" box="[537,1011,1348,1375]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="20" pageNumber="31" phylum="Echinodermata" rank="species" species="variegatum">
|
||
<emphasis id="B93586085336FFF9FDB69B26FCF5C17D" box="[537,827,1348,1374]" italics="true" pageId="20" pageNumber="31">Ophioderma variegatum</emphasis>
|
||
,
|
||
<bibRefCitation id="EFD027EB5336FFF9FCE59B26FC3DC17C" author="Lutken C. F." box="[842,1011,1348,1375]" pageId="20" pageNumber="31" pagination="20 - 26" refId="ref16758" refString="Lutken C. F. 1856. III. Bidrag til Kundskab om Ophiurerne ved Central-Amerikas Vestkyst. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening 1856: 20 - 26." type="journal article" year="1856">Lütken (1856)</bibRefCitation>
|
||
</taxonomicName>
|
||
only mentioned the existence of one uncatalogued specimen (DD =
|
||
<quantity id="4CB9F7FF5336FFF9FDCF9B0AFD78C1A0" box="[608,694,1384,1411]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="20" pageNumber="31" unit="mm" value="14.0">14 mm</quantity>
|
||
) as part of the type material. However, three specimens are present in this lot (pers. com. Schiøtte, 2019) all of similar size (DD = ~
|
||
<quantity id="4CB9F7FF5336FFF9FBB59BEEFB47C184" box="[1050,1161,1420,1447]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="20" pageNumber="31" unit="mm" value="12.5">12.5 mm</quantity>
|
||
). The similarities of the morphological characters and sizes make it impossible to determine which of them corresponds to the specimen described by
|
||
<bibRefCitation id="EFD027EB5336FFF9FDA79BB7FD79C1CC" author="Lutken C. F." box="[520,695,1492,1519]" pageId="20" pageNumber="31" pagination="20 - 26" refId="ref16758" refString="Lutken C. F. 1856. III. Bidrag til Kundskab om Ophiurerne ved Central-Amerikas Vestkyst. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening 1856: 20 - 26." type="journal article" year="1856">Lütken (1856)</bibRefCitation>
|
||
. There are some discrepancies between the measurements from
|
||
<bibRefCitation id="EFD027EB5336FFF9FEAF9B9BFE65C237" author="Lutken C. F." box="[256,427,1529,1556]" pageId="20" pageNumber="31" pagination="20 - 26" refId="ref16758" refString="Lutken C. F. 1856. III. Bidrag til Kundskab om Ophiurerne ved Central-Amerikas Vestkyst. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening 1856: 20 - 26." type="journal article" year="1856">Lütken (1856)</bibRefCitation>
|
||
and our measurements which could be due to the preservation of the specimens through time. Because of this, and according to Recommendation 73F of the International Code of Zoological Nomenclature (
|
||
<bibRefCitation id="EFD027EB5336FFF9FE579823FD4DC278" author="ICZN & International Commission on Zoological Nomenclature" box="[504,643,1601,1627]" pageId="20" pageNumber="31" refId="ref16522" refString="ICZN, International Commission on Zoological Nomenclature. 1999. International Code of Zoological Nomenclature. Available from https: // www. iczn. org / the-code / the-international-code-of-zoological-nomenclature / the-code-online / [accessed 1 Feb. 2020]." type="url" year="1999">ICZN 1999</bibRefCitation>
|
||
), we propose the designation of the specimen NHMD-107680 as the
|
||
<typeStatus id="54FAE4B85336FFF9FF479807FE9AC25C" box="[232,340,1637,1663]" pageId="20" pageNumber="31" type="lectotype">lectotype</typeStatus>
|
||
and NHMD-619214 contains the
|
||
<typeStatus id="54FAE4B85336FFF9FD4B9807FC43C25C" box="[740,909,1637,1663]" pageId="20" pageNumber="31" type="paralectotype">paralectotypes</typeStatus>
|
||
(two specimens) for
|
||
<taxonomicName id="4C4121995336FFF9FB299804FAFEC25C" authorityName=", Lutken" authorityYear="1856" box="[1158,1328,1638,1664]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="20" pageNumber="31" phylum="Echinodermata" rank="species" species="variegatum">
|
||
<emphasis id="B93586085336FFF9FB299804FAFEC25C" box="[1158,1328,1638,1664]" italics="true" pageId="20" pageNumber="31">O. variegatum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5336FFF9FF4598C5FE68C2E2" blockId="20.[189,536,1703,1766]" box="[234,422,1703,1729]" pageId="20" pageNumber="31">
|
||
<emphasis id="B93586085336FFF9FF4598C5FE68C2E2" bold="true" box="[234,422,1703,1729]" pageId="20" pageNumber="31">Other material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5336FFF9FF1298AEFDD6C2C5" blockId="20.[189,536,1703,1766]" box="[189,536,1740,1766]" pageId="20" pageNumber="31">See: Supplementary material.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35B09915336FFFAFF12996CFCFEC06E" lastPageId="23" lastPageNumber="34" pageId="20" pageNumber="31" type="description">
|
||
<paragraph id="8BFE5A1A5336FFF9FF12996CFE10C30B" blockId="20.[189,478,1806,1832]" box="[189,478,1806,1832]" pageId="20" pageNumber="31">
|
||
<emphasis id="B93586085336FFF9FF12996CFE80C30B" bold="true" box="[189,334,1806,1832]" pageId="20" pageNumber="31">Description</emphasis>
|
||
(
|
||
<typeStatus id="54FAE4B85336FFF9FEF2996CFE18C30B" box="[349,470,1806,1832]" pageId="20" pageNumber="31" type="lectotype">Lectotype</typeStatus>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5336FFF9FF12995CFAB9C3CB" blockId="20.[189,1399,1854,2025]" pageId="20" pageNumber="31">
|
||
DD =
|
||
<quantity id="4CB9F7FF5336FFF9FEA6995CFEB8C37B" box="[265,374,1854,1880]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="20" pageNumber="31" unit="mm" value="12.4">12.4 mm</quantity>
|
||
; 5 arms, AL =
|
||
<quantity id="4CB9F7FF5336FFF9FD86995CFDB1C37B" box="[553,639,1854,1880]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="20" pageNumber="31" unit="mm" value="62.0">62 mm</quantity>
|
||
. Disc flat and rounded. Dorsal disc densely covered by rounded and very small granules, granule density 222
|
||
<superScript id="7C34F7525336FFF9FD499900FCEFC34C" attach="left" box="[742,801,1889,1916]" fontSize="6" pageId="20" pageNumber="31">mm-2</superScript>
|
||
; granules extend to base of arms. Some granules rubbed off in some areas of disc. Radial shields covered by granules (rubbed off in some parts due to preservation) (
|
||
<figureCitation id="137A469F5336FFF9FE2599C8FE24C3E7" box="[394,490,1962,1988]" captionStart="Fig" captionStartId="21.[189,232,1964,1990]" captionTargetBox="[189,1398,1121,1926]" captionTargetId="figure-301@21.[189,1398,1121,1926]" captionTargetPageId="21" captionText="Fig. 8. Ophioderma variegatum Lütken, 1856, lectotype (NHMD-107680, DD = 12.4 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Dorsal arm. E. Ventral arm. F. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326631" httpUri="https://zenodo.org/record/4326631/files/figure.png" pageId="20" pageNumber="31">Fig. 8A</figureCitation>
|
||
). Interradii covered with granules, similar to those on dorsal disc (
|
||
<figureCitation id="137A469F5336FFF9FAA599C8FAA6C3E7" box="[1290,1384,1962,1988]" captionStart="Fig" captionStartId="21.[189,232,1964,1990]" captionTargetBox="[189,1398,1121,1926]" captionTargetId="figure-301@21.[189,1398,1121,1926]" captionTargetPageId="21" captionText="Fig. 8. Ophioderma variegatum Lütken, 1856, lectotype (NHMD-107680, DD = 12.4 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Dorsal arm. E. Ventral arm. F. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326631" httpUri="https://zenodo.org/record/4326631/files/figure.png" pageId="20" pageNumber="31">Fig. 8B</figureCitation>
|
||
). Four genital slits on each interradius; proximal genital slits elongated and shorter than distal slits, in
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5337FFF8FF129F6BFE2FC565" blockId="21.[189,1398,265,326]" pageId="21" pageNumber="32">
|
||
contact with distal part of oral shield and with first LAP; distal genital slits elongated and surrounded by granulation (
|
||
<figureCitation id="137A469F5337FFF8FED89F4EFE1DC565" box="[375,467,300,326]" captionStart="Fig" captionStartId="21.[189,232,1964,1990]" captionTargetBox="[189,1398,1121,1926]" captionTargetId="figure-301@21.[189,1398,1121,1926]" captionTargetPageId="21" captionText="Fig. 8. Ophioderma variegatum Lütken, 1856, lectotype (NHMD-107680, DD = 12.4 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Dorsal arm. E. Ventral arm. F. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326631" httpUri="https://zenodo.org/record/4326631/files/figure.png" pageId="21" pageNumber="32">Fig. 8B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5337FFF8FF129F0CFD30C65D" blockId="21.[189,1399,366,639]" pageId="21" pageNumber="32">
|
||
Oral shields longer than broad, oval, proximal margin convex obtuse angled, distal edge convex, lateral margins straight. Madreporite with a small, circular and shallow depression located in center of oral shield. Adoral shields naked, triangular, longer than broad, separated. Jaws bear 8–9 papillae on each side: LyOs very small, as long as broad, angled upwards; AdShSp largest, rounded; 2°AdShSp smaller but similar in shape to AdShSp; LOPa 4–5 conical, pointed and separated, 1
|
||
<superScript id="7C34F7525337FFF8FB919F98FB87C62B" attach="left" box="[1086,1097,506,520]" fontSize="6" pageId="21" pageNumber="32">st</superScript>
|
||
LOPa similar in shape to and smaller than 2°AdShSp; IPa sometimes large, elongated, pointed and separated; TPa 2 at jaw tips, larger than LOPa, elongated, robust and pointed, separated. Granules covering oral plates slightly longer than those on dorsal disc and interradii (
|
||
<figureCitation id="137A469F5337FFF8FD3B9C06FD3EC65D" box="[660,752,612,638]" captionStart="Fig" captionStartId="21.[189,232,1964,1990]" captionTargetBox="[189,1398,1121,1926]" captionTargetId="figure-301@21.[189,1398,1121,1926]" captionTargetPageId="21" captionText="Fig. 8. Ophioderma variegatum Lütken, 1856, lectotype (NHMD-107680, DD = 12.4 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Dorsal arm. E. Ventral arm. F. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326631" httpUri="https://zenodo.org/record/4326631/files/figure.png" pageId="21" pageNumber="32">Fig. 8C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5337FFF8FF129CC7FDE1C795" blockId="21.[189,1399,677,950]" pageId="21" pageNumber="32">
|
||
Dorsal base of arms with granules. Dorsal arm plates, 2× broader than long, trapezoidal, overlapping (
|
||
<figureCitation id="137A469F5337FFF8FF699CABFEEDC6C0" box="[198,291,713,739]" captionStart="Fig" captionStartId="21.[189,232,1964,1990]" captionTargetBox="[189,1398,1121,1926]" captionTargetId="figure-301@21.[189,1398,1121,1926]" captionTargetPageId="21" captionText="Fig. 8. Ophioderma variegatum Lütken, 1856, lectotype (NHMD-107680, DD = 12.4 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Dorsal arm. E. Ventral arm. F. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326631" httpUri="https://zenodo.org/record/4326631/files/figure.png" pageId="21" pageNumber="32">Fig. 8D</figureCitation>
|
||
). First ventral arm plate large, broader than long, oval (
|
||
<figureCitation id="137A469F5337FFF8FC089CABFBCDC6C0" box="[935,1027,713,739]" captionStart="Fig" captionStartId="21.[189,232,1964,1990]" captionTargetBox="[189,1398,1121,1926]" captionTargetId="figure-301@21.[189,1398,1121,1926]" captionTargetPageId="21" captionText="Fig. 8. Ophioderma variegatum Lütken, 1856, lectotype (NHMD-107680, DD = 12.4 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Dorsal arm. E. Ventral arm. F. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326631" httpUri="https://zenodo.org/record/4326631/files/figure.png" pageId="21" pageNumber="32">Fig. 8C</figureCitation>
|
||
). Subsequent ventral arm plates quadrangular, slightly broader than long; distal edge slightly concave (
|
||
<figureCitation id="137A469F5337FFF8FBA49C8EFBA7C725" box="[1035,1129,748,774]" captionStart="Fig" captionStartId="21.[189,232,1964,1990]" captionTargetBox="[189,1398,1121,1926]" captionTargetId="figure-301@21.[189,1398,1121,1926]" captionTargetPageId="21" captionText="Fig. 8. Ophioderma variegatum Lütken, 1856, lectotype (NHMD-107680, DD = 12.4 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Dorsal arm. E. Ventral arm. F. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326631" httpUri="https://zenodo.org/record/4326631/files/figure.png" pageId="21" pageNumber="32">Fig. 8E</figureCitation>
|
||
). Paired, rounded, and very small holes between 1
|
||
<superScript id="7C34F7525337FFF8FE549D6DFDC8C73E" attach="left" box="[507,518,783,797]" fontSize="6" pageId="21" pageNumber="32">st</superScript>
|
||
and 2
|
||
<superScript id="7C34F7525337FFF8FDE19D6CFD91C73F" attach="none" box="[590,607,782,796]" fontSize="6" pageId="21" pageNumber="32">nd</superScript>
|
||
, and 2
|
||
<superScript id="7C34F7525337FFF8FD029D6CFD70C73F" attach="left" box="[685,702,782,796]" fontSize="6" pageId="21" pageNumber="32">nd</superScript>
|
||
and 3
|
||
<superScript id="7C34F7525337FFF8FCA99D6CFCDAC73F" attach="left" box="[774,788,782,796]" fontSize="6" pageId="21" pageNumber="32">rd</superScript>
|
||
ventral arm plates (
|
||
<figureCitation id="137A469F5337FFF8FC539D6DFB8EC70A" box="[1020,1088,783,809]" captionStart="Fig" captionStartId="21.[189,232,1964,1990]" captionTargetBox="[189,1398,1121,1926]" captionTargetId="figure-301@21.[189,1398,1121,1926]" captionTargetPageId="21" captionText="Fig. 8. Ophioderma variegatum Lütken, 1856, lectotype (NHMD-107680, DD = 12.4 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Dorsal arm. E. Ventral arm. F. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326631" httpUri="https://zenodo.org/record/4326631/files/figure.png" pageId="21" pageNumber="32">Fig. 8</figureCitation>
|
||
B–C). LAP with up to 7–8 arm spines.Arm spines with a blunt tip, slender, and very large. All arm spines of almost same size except dorsalmost and ventralmost, which are just a little shortest than rest (
|
||
<figureCitation id="137A469F5337FFF8FC7A9D37FBE1C753" box="[981,1071,853,880]" captionStart="Fig" captionStartId="21.[189,232,1964,1990]" captionTargetBox="[189,1398,1121,1926]" captionTargetId="figure-301@21.[189,1398,1121,1926]" captionTargetPageId="21" captionText="Fig. 8. Ophioderma variegatum Lütken, 1856, lectotype (NHMD-107680, DD = 12.4 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Dorsal arm. E. Ventral arm. F. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326631" httpUri="https://zenodo.org/record/4326631/files/figure.png" pageId="21" pageNumber="32">Fig. 8E</figureCitation>
|
||
); length approximately ¾ of size of LAP. Tentacle scales 2; adradial tentacle scale ovoid, elongated; abradial tentacle scale slightly shorter, subtriangular (
|
||
<figureCitation id="137A469F5337FFF8FE689DFEFDECC795" box="[455,546,924,950]" captionStart="Fig" captionStartId="21.[189,232,1964,1990]" captionTargetBox="[189,1398,1121,1926]" captionTargetId="figure-301@21.[189,1398,1121,1926]" captionTargetPageId="21" captionText="Fig. 8. Ophioderma variegatum Lütken, 1856, lectotype (NHMD-107680, DD = 12.4 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Dorsal arm. E. Ventral arm. F. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326631" httpUri="https://zenodo.org/record/4326631/files/figure.png" pageId="21" pageNumber="32">Fig. 8E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5337FFF8FF129DBFFD4AC038" blockId="21.[189,1398,989,1051]" pageId="21" pageNumber="32">
|
||
Specimen (ethanol preservation) discolored but with a disc with different shades of brown and arms with transverse dark brown bands (
|
||
<figureCitation id="137A469F5337FFF8FDB29A62FDB9C038" box="[541,631,1024,1051]" captionStart="Fig" captionStartId="21.[189,232,1964,1990]" captionTargetBox="[189,1398,1121,1926]" captionTargetId="figure-301@21.[189,1398,1121,1926]" captionTargetPageId="21" captionText="Fig. 8. Ophioderma variegatum Lütken, 1856, lectotype (NHMD-107680, DD = 12.4 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Dorsal arm. E. Ventral arm. F. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326631" httpUri="https://zenodo.org/record/4326631/files/figure.png" pageId="21" pageNumber="32">Fig. 8F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF3E0A925337FFF8FF1299CEFB92C3C9" ID-DOI="http://doi.org/10.5281/zenodo.4326631" ID-Zenodo-Dep="4326631" httpUri="https://zenodo.org/record/4326631/files/figure.png" pageId="21" pageNumber="32" startId="21.[189,232,1964,1990]" targetBox="[189,1398,1121,1926]" targetPageId="21">
|
||
<paragraph id="8BFE5A1A5337FFF8FF1299CEFB92C3C9" blockId="21.[189,1398,1964,2026]" pageId="21" pageNumber="32">
|
||
<emphasis id="B93586085337FFF8FF1299CEFEC5C3E5" bold="true" box="[189,267,1964,1990]" pageId="21" pageNumber="32">Fig. 8.</emphasis>
|
||
<taxonomicName id="4C4121995337FFF8FEBB99CFFD10C3E4" authority="Lutken, 1856" authorityName=", Lutken" authorityYear="1856" box="[276,734,1964,1991]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="21" pageNumber="32" phylum="Echinodermata" rank="species" species="variegatum">
|
||
<emphasis id="B93586085337FFF8FEBB99CFFDF9C3E5" box="[276,567,1965,1991]" italics="true" pageId="21" pageNumber="32">Ophioderma variegatum</emphasis>
|
||
<bibRefCitation id="EFD027EB5337FFF8FD9099CFFD10C3E4" author="Lutken C. F." box="[575,734,1964,1991]" pageId="21" pageNumber="32" pagination="20 - 26" refId="ref16758" refString="Lutken C. F. 1856. III. Bidrag til Kundskab om Ophiurerne ved Central-Amerikas Vestkyst. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening 1856: 20 - 26." type="journal article" year="1856">Lütken, 1856</bibRefCitation>
|
||
</taxonomicName>
|
||
, lectotype (NHMD-107680, DD = 12.4 mm).
|
||
<emphasis id="B93586085337FFF8FAAC99CEFAD6C3E5" bold="true" box="[1283,1304,1964,1990]" pageId="21" pageNumber="32">A</emphasis>
|
||
. Dorsal disc.
|
||
<emphasis id="B93586085337FFF8FF5499B2FEC1C3C9" bold="true" box="[251,271,2000,2026]" pageId="21" pageNumber="32">B</emphasis>
|
||
. Ventral disc.
|
||
<emphasis id="B93586085337FFF8FE1899B2FE02C3C9" bold="true" box="[439,460,2000,2026]" pageId="21" pageNumber="32">C</emphasis>
|
||
. Mouth.
|
||
<emphasis id="B93586085337FFF8FD9899B2FD82C3C9" bold="true" box="[567,588,2000,2026]" pageId="21" pageNumber="32">D</emphasis>
|
||
. Dorsal arm.
|
||
<emphasis id="B93586085337FFF8FD4299B2FCCFC3C9" bold="true" box="[749,769,2000,2026]" pageId="21" pageNumber="32">E</emphasis>
|
||
. Ventral arm.
|
||
<emphasis id="B93586085337FFF8FC0899B2FC77C3C9" bold="true" box="[935,953,2000,2026]" pageId="21" pageNumber="32">F</emphasis>
|
||
. Dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3E0A925334FFFBFF1298BAFEB1C3CB" ID-DOI="http://doi.org/10.5281/zenodo.4326633" ID-Zenodo-Dep="4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="22" pageNumber="33" startId="22.[189,232,1752,1778]" targetBox="[289,1303,265,1714]" targetPageId="22">
|
||
<paragraph id="8BFE5A1A5334FFFBFF1298BAFEB1C3CB" blockId="22.[189,1399,1752,2024]" pageId="22" pageNumber="33">
|
||
<emphasis id="B93586085334FFFBFF1298BAFEC0C2D1" bold="true" box="[189,270,1752,1778]" pageId="22" pageNumber="33">Fig. 9.</emphasis>
|
||
SEM images of skeletal elements of
|
||
<taxonomicName id="4C4121995334FFFBFD4E98BAFBC9C2D1" authorityName=", Lutken" authorityYear="1856" box="[737,1031,1752,1778]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="22" pageNumber="33" phylum="Echinodermata" rank="species" species="variegatum">
|
||
<emphasis id="B93586085334FFFBFD4E98BAFBC9C2D1" box="[737,1031,1752,1778]" italics="true" pageId="22" pageNumber="33">Ophioderma variegatum</emphasis>
|
||
</taxonomicName>
|
||
(ICML-UNAM 3.20.4, DD = 13.6 mm).
|
||
<emphasis id="B93586085334FFFBFEEC9899FE96C336" bold="true" box="[323,344,1787,1813]" pageId="22" pageNumber="33">A</emphasis>
|
||
. Radial shield (external view), deepest pores marked with arrows.
|
||
<emphasis id="B93586085334FFFBFB2D9899FB58C336" bold="true" box="[1154,1174,1787,1813]" pageId="22" pageNumber="33">B</emphasis>
|
||
. Dorsal arm plate, with five spurs.
|
||
<emphasis id="B93586085334FFFBFED5997CFE41C31B" bold="true" box="[378,399,1822,1848]" pageId="22" pageNumber="33">C</emphasis>
|
||
. Ventral arm plate, with one spur.
|
||
<emphasis id="B93586085334FFFBFC89997CFCF5C31B" bold="true" box="[806,827,1822,1848]" pageId="22" pageNumber="33">D</emphasis>
|
||
. Lateral arm plate, external face, with two spurs and one condyle.
|
||
<emphasis id="B93586085334FFFBFE269920FE53C37F" bold="true" box="[393,413,1858,1884]" pageId="22" pageNumber="33">E</emphasis>
|
||
. Lateral arm plate, lateral face, with spine articulations.
|
||
<emphasis id="B93586085334FFFBFB869920FBF5C37F" bold="true" box="[1065,1083,1858,1884]" pageId="22" pageNumber="33">F</emphasis>
|
||
. Lateral arm plate, internal face.
|
||
<emphasis id="B93586085334FFFBFF519906FEDBC35D" bold="true" box="[254,277,1892,1918]" pageId="22" pageNumber="33">G</emphasis>
|
||
. Outer surface of the LAP (D) with fine mesh and rounded knobs.
|
||
<emphasis id="B93586085334FFFBFB999907FB83C35C" bold="true" box="[1078,1101,1893,1919]" pageId="22" pageNumber="33">H</emphasis>
|
||
. Spine articulations with a weak sigmoidal fold and tilted lobes.
|
||
<emphasis id="B93586085334FFFBFD3099EAFD64C381" bold="true" box="[671,682,1928,1954]" pageId="22" pageNumber="33">I</emphasis>
|
||
. Arm vertebra, distal face.
|
||
<emphasis id="B93586085334FFFBFC5699EAFBC6C381" bold="true" box="[1017,1032,1928,1954]" pageId="22" pageNumber="33">J</emphasis>
|
||
. Arm vertebra, proximal face.
|
||
<emphasis id="B93586085334FFFBFF1299C9FF1AC3E6" bold="true" box="[189,212,1963,1989]" pageId="22" pageNumber="33">K</emphasis>
|
||
. Arm spine.
|
||
<emphasis id="B93586085334FFFBFEC599C9FEB0C3E6" bold="true" box="[362,382,1963,1989]" pageId="22" pageNumber="33">L</emphasis>
|
||
. Dental plate fragments.
|
||
<emphasis id="B93586085334FFFBFD0A99C9FD0FC3E6" bold="true" box="[677,705,1963,1989]" pageId="22" pageNumber="33">M</emphasis>
|
||
. Oral plate, adradial face, with arrows pointing to four oral papillae sockets.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFE5A1A5335FFFAFF459F6AFE30C501" blockId="23.[189,1399,264,1101]" box="[234,510,264,290]" pageId="23" pageNumber="34">
|
||
<emphasis id="B93586085335FFFAFF459F6AFE30C501" bold="true" box="[234,510,264,290]" pageId="23" pageNumber="34">Disarticulated ossicles</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5335FFFAFF129F49FCFEC06E" blockId="23.[189,1399,264,1101]" pageId="23" pageNumber="34">
|
||
Specimen analyzed:
|
||
<specimenCount id="9D4791935335FFFAFE189F49FDC1C565" box="[439,527,299,326]" pageId="23" pageNumber="34" type="generic">1 spec.</specimenCount>
|
||
, ICML-UNAM 3.20.4 (DD =
|
||
<quantity id="4CB9F7FF5335FFFAFC299F49FC38C565" box="[902,1014,299,326]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="23" pageNumber="34" unit="mm" value="13.6">13.6 mm</quantity>
|
||
, AL =
|
||
<quantity id="4CB9F7FF5335FFFAFBFF9F49FB0CC565" box="[1104,1218,299,326]" metricMagnitude="-2" metricUnit="m" metricValue="6.65" pageId="23" pageNumber="34" unit="mm" value="66.5">66.5 mm</quantity>
|
||
). Radial shield (external view) irregularly rounded, with an incised proximal margin, a slightly indented abradial edge with a tiny projection, a convex distal edge, and convex adradial margin, bordered by 9 pores along median to proximal margin (which are covered/overlapped by disc scales and granules in intact animal) (
|
||
<figureCitation id="137A469F5335FFFAFE839FDAFE5FC5F1" box="[300,401,440,466]" captionStart="Fig" captionStartId="22.[189,232,1752,1778]" captionTargetBox="[289,1303,265,1714]" captionTargetId="figure-151@22.[289,1303,265,1714]" captionTargetPageId="22" captionText="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi="http://doi.org/10.5281/zenodo.4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="23" pageNumber="34">Fig. 9A</figureCitation>
|
||
). Dorsal arm plate arched and somewhat trapezoid-shaped, 2 × as wide as long; proximal edge convex with 5 spurs on proximal portion of external surface, distally straight and incised (
|
||
<figureCitation id="137A469F5335FFFAFF699F9DFEEAC63A" box="[198,292,511,537]" captionStart="Fig" captionStartId="22.[189,232,1752,1778]" captionTargetBox="[289,1303,265,1714]" captionTargetId="figure-151@22.[289,1303,265,1714]" captionTargetPageId="22" captionText="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi="http://doi.org/10.5281/zenodo.4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="23" pageNumber="34">Fig. 9B</figureCitation>
|
||
). Ventral arm plate 2× as wide as long, rectangular-shaped with proximal side truncated and with a pointed-shape spur, lateral sides concave forming border of tentacle pore, distal margin convex (
|
||
<figureCitation id="137A469F5335FFFAFF699C27FEEBC67C" box="[198,293,581,607]" captionStart="Fig" captionStartId="22.[189,232,1752,1778]" captionTargetBox="[289,1303,265,1714]" captionTargetId="figure-151@22.[289,1303,265,1714]" captionTargetPageId="22" captionText="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi="http://doi.org/10.5281/zenodo.4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="23" pageNumber="34">Fig. 9C</figureCitation>
|
||
). LAP D-shaped, 2× as high as wide, with 9 spine articulations sunken in notches of distal edge (
|
||
<figureCitation id="137A469F5335FFFAFEA99C0AFE83C6A0" box="[262,333,616,643]" captionStart="Fig" captionStartId="22.[189,232,1752,1778]" captionTargetBox="[289,1303,265,1714]" captionTargetId="figure-151@22.[289,1303,265,1714]" captionTargetPageId="22" captionText="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi="http://doi.org/10.5281/zenodo.4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="23" pageNumber="34">Fig. 9</figureCitation>
|
||
D–E). Ventral portion of LAP projecting ventro-proximalwards (
|
||
<figureCitation id="137A469F5335FFFAFBFC9C0AFB7DC6A0" box="[1107,1203,616,643]" captionStart="Fig" captionStartId="22.[189,232,1752,1778]" captionTargetBox="[289,1303,265,1714]" captionTargetId="figure-151@22.[289,1303,265,1714]" captionTargetPageId="22" captionText="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi="http://doi.org/10.5281/zenodo.4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="23" pageNumber="34">Fig. 9D</figureCitation>
|
||
). Proximal edge of LAP with 2 prominent and elongated spurs, which are protruding and modify central-proximal edge (
|
||
<figureCitation id="137A469F5335FFFAFF699CCDFEEAC6EA" box="[198,292,687,713]" captionStart="Fig" captionStartId="22.[189,232,1752,1778]" captionTargetBox="[289,1303,265,1714]" captionTargetId="figure-151@22.[289,1303,265,1714]" captionTargetPageId="22" captionText="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi="http://doi.org/10.5281/zenodo.4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="23" pageNumber="34">Fig. 9D</figureCitation>
|
||
), between spurs and across remaining proximal margin a discernible band of different stereom structure is present (
|
||
<figureCitation id="137A469F5335FFFAFE069CB0FDC9C6CF" box="[425,519,722,748]" captionStart="Fig" captionStartId="22.[189,232,1752,1778]" captionTargetBox="[289,1303,265,1714]" captionTargetId="figure-151@22.[289,1303,265,1714]" captionTargetPageId="22" captionText="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi="http://doi.org/10.5281/zenodo.4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="23" pageNumber="34">Fig. 9D</figureCitation>
|
||
). Outer surface finely meshed with rounded stereom knobs (
|
||
<figureCitation id="137A469F5335FFFAFB699CB0FAEAC6CF" box="[1222,1316,722,748]" captionStart="Fig" captionStartId="22.[189,232,1752,1778]" captionTargetBox="[289,1303,265,1714]" captionTargetId="figure-151@22.[289,1303,265,1714]" captionTargetPageId="22" captionText="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi="http://doi.org/10.5281/zenodo.4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="23" pageNumber="34">Fig. 9G</figureCitation>
|
||
). Inner surface of LAP with a continuous prominent ridge on proximal edge (see arched view) and on ventrodistal margin 2 spurs matching those on external surface, with 1 perforation on vertical furrow (
|
||
<figureCitation id="137A469F5335FFFAFABE9D7AFAA7C710" box="[1297,1385,792,819]" captionStart="Fig" captionStartId="22.[189,232,1752,1778]" captionTargetBox="[289,1303,265,1714]" captionTargetId="figure-151@22.[289,1303,265,1714]" captionTargetPageId="22" captionText="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi="http://doi.org/10.5281/zenodo.4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="23" pageNumber="34">Fig. 9F</figureCitation>
|
||
). Spine articulations ventralwards increasing in size (
|
||
<figureCitation id="137A469F5335FFFAFCB09D5EFCB5C775" box="[799,891,828,854]" captionStart="Fig" captionStartId="22.[189,232,1752,1778]" captionTargetBox="[289,1303,265,1714]" captionTargetId="figure-151@22.[289,1303,265,1714]" captionTargetPageId="22" captionText="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi="http://doi.org/10.5281/zenodo.4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="23" pageNumber="34">Fig. 9E</figureCitation>
|
||
). Lobes with a weak sigmoidal fold, tilted, and curved (
|
||
<figureCitation id="137A469F5335FFFAFEE19D3DFE62C75A" box="[334,428,863,889]" captionStart="Fig" captionStartId="22.[189,232,1752,1778]" captionTargetBox="[289,1303,265,1714]" captionTargetId="figure-151@22.[289,1303,265,1714]" captionTargetPageId="22" captionText="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi="http://doi.org/10.5281/zenodo.4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="23" pageNumber="34">Fig. 9H</figureCitation>
|
||
). Ventral lobe less conspicuous than dorsal lobe (
|
||
<figureCitation id="137A469F5335FFFAFC459D3DFB89C75A" box="[1002,1095,863,889]" captionStart="Fig" captionStartId="22.[189,232,1752,1778]" captionTargetBox="[289,1303,265,1714]" captionTargetId="figure-151@22.[289,1303,265,1714]" captionTargetPageId="22" captionText="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi="http://doi.org/10.5281/zenodo.4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="23" pageNumber="34">Fig. 9H</figureCitation>
|
||
). Proximal vertebrae with a wing-like muscle flange (
|
||
<figureCitation id="137A469F5335FFFAFDAE9DE0FD98C7BF" box="[513,598,898,924]" captionStart="Fig" captionStartId="22.[189,232,1752,1778]" captionTargetBox="[289,1303,265,1714]" captionTargetId="figure-151@22.[289,1303,265,1714]" captionTargetPageId="22" captionText="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi="http://doi.org/10.5281/zenodo.4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="23" pageNumber="34">Fig. 9J</figureCitation>
|
||
). Distal face of vertebrae almost round, as wide as long, with large and irregular (sinuous) aboral muscle flange, with typical zygospondylous articulation (
|
||
<figureCitation id="137A469F5335FFFAFB6F9DC7FADDC7E3" box="[1216,1299,933,960]" captionStart="Fig" captionStartId="22.[189,232,1752,1778]" captionTargetBox="[289,1303,265,1714]" captionTargetId="figure-151@22.[289,1303,265,1714]" captionTargetPageId="22" captionText="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi="http://doi.org/10.5281/zenodo.4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="23" pageNumber="34">Fig. 9I</figureCitation>
|
||
). Spines with a scale-like surface, a blunt tip and laterally flattened (
|
||
<figureCitation id="137A469F5335FFFAFC259DABFC25C7C0" box="[906,1003,969,995]" captionStart="Fig" captionStartId="22.[189,232,1752,1778]" captionTargetBox="[289,1303,265,1714]" captionTargetId="figure-151@22.[289,1303,265,1714]" captionTargetPageId="22" captionText="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi="http://doi.org/10.5281/zenodo.4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="23" pageNumber="34">Fig. 9K</figureCitation>
|
||
). Dental plate consists of several pieces, 1
|
||
<superScript id="7C34F7525335FFFAFE8D9D8EFEE3C7D9" attach="left" box="[290,301,1004,1018]" fontSize="6" pageId="23" pageNumber="34">st</superScript>
|
||
piece has 2 TPa and 1 wide tooth, while rest of pieces with 1 tooth socket per piece (
|
||
<quantity id="4CB9F7FF5335FFFAFB509D8EFAE5C025" box="[1279,1323,1004,1031]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="23" pageNumber="34" unit="in" value="5.0">5 in</quantity>
|
||
total), none of sockets penetrate plate (
|
||
<figureCitation id="137A469F5335FFFAFD969A6DFD5BC00A" box="[569,661,1039,1065]" captionStart="Fig" captionStartId="22.[189,232,1752,1778]" captionTargetBox="[289,1303,265,1714]" captionTargetId="figure-151@22.[289,1303,265,1714]" captionTargetPageId="22" captionText="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi="http://doi.org/10.5281/zenodo.4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="23" pageNumber="34">Fig. 9L</figureCitation>
|
||
). Oral plate as long as high, adradial plate with a less indented middle section, and 4 oral papillae sockets (
|
||
<figureCitation id="137A469F5335FFFAFD109A50FCEDC06E" box="[703,803,1074,1101]" captionStart="Fig" captionStartId="22.[189,232,1752,1778]" captionTargetBox="[289,1303,265,1714]" captionTargetId="figure-151@22.[289,1303,265,1714]" captionTargetPageId="22" captionText="Fig. 9. SEM images of skeletal elements of Ophioderma variegatum (ICML-UNAM 3.20.4, DD = 13.6 mm). A. Radial shield (external view), deepest pores marked with arrows. B. Dorsal arm plate, with five spurs. C. Ventral arm plate, with one spur. D. Lateral arm plate, external face, with two spurs and one condyle. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, distal face. J. Arm vertebra, proximal face. K. Arm spine. L. Dental plate fragments. M. Oral plate, adradial face, with arrows pointing to four oral papillae sockets." figureDoi="http://doi.org/10.5281/zenodo.4326633" httpUri="https://zenodo.org/record/4326633/files/figure.png" pageId="23" pageNumber="34">Fig. 9M</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35B09915335FFFAFF129A16FD71C169" pageId="23" pageNumber="34" type="distribution">
|
||
<paragraph id="8BFE5A1A5335FFFAFF129A16FE21C0AD" blockId="23.[189,495,1140,1166]" box="[189,495,1140,1166]" pageId="23" pageNumber="34">
|
||
<heading id="D0B6ED765335FFFAFF129A16FE21C0AD" bold="true" box="[189,495,1140,1166]" fontSize="11" level="3" pageId="23" pageNumber="34" reason="3">
|
||
<emphasis id="B93586085335FFFAFF129A16FE21C0AD" bold="true" box="[189,495,1140,1166]" pageId="23" pageNumber="34">Habitat and distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5335FFFAFF129AC1FD71C169" blockId="23.[189,1398,1186,1354]" pageId="23" pageNumber="34">
|
||
<collectingCountry id="F3561A8A5335FFFAFF129AC1FF38C09E" box="[189,246,1187,1213]" name="United States of America" pageId="23" pageNumber="34">USA</collectingCountry>
|
||
(
|
||
<collectingRegion id="498594F85335FFFAFEA09AC0FE42C09E" box="[271,396,1186,1213]" country="United States of America" name="California" pageId="23" pageNumber="34">California</collectingRegion>
|
||
),
|
||
<collectingCountry id="F3561A8A5335FFFAFE049AC0FDCBC09F" box="[427,517,1186,1212]" name="Mexico" pageId="23" pageNumber="34">Mexico</collectingCountry>
|
||
(
|
||
<collectingRegion id="498594F85335FFFAFDB29AC0FD2EC09F" box="[541,736,1186,1212]" country="Mexico" name="Baja California" pageId="23" pageNumber="34">Baja California</collectingRegion>
|
||
, Gulf of California,
|
||
<collectingRegion id="498594F85335FFFAFC589AC0FB84C09F" box="[1015,1098,1186,1212]" country="Mexico" name="Jalisco" pageId="23" pageNumber="34">Jalisco</collectingRegion>
|
||
,
|
||
<collectingRegion id="498594F85335FFFAFBF09AC1FB74C09E" box="[1119,1210,1187,1213]" country="Mexico" name="Oaxaca" pageId="23" pageNumber="34">Oaxaca</collectingRegion>
|
||
, Isabel Island, Revillagigedo Islands, Marías Islands),
|
||
<collectingCountry id="F3561A8A5335FFFAFD3A9AA4FCDFC0C3" box="[661,785,1222,1248]" name="Nicaragua" pageId="23" pageNumber="34">Nicaragua</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3561A8A5335FFFAFCB29AA4FC51C0C3" box="[797,927,1222,1248]" name="Costa Rica" pageId="23" pageNumber="34">Costa Rica</collectingCountry>
|
||
(Cocos Island),
|
||
<collectingCountry id="F3561A8A5335FFFAFBCA9AA4FB0CC0C3" box="[1125,1218,1222,1248]" name="Panama" pageId="23" pageNumber="34">Panama</collectingCountry>
|
||
(Pearl Islands) and Galapagos Islands. From intertidal zone to
|
||
<quantity id="4CB9F7FF5335FFFAFCAE9A8BFC80C127" box="[769,846,1257,1284]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="23" pageNumber="34" unit="m" value="110.0">110 m</quantity>
|
||
depth. Collected in sand, rock, rubble, coral, algae, shell, oysters, and mangrove leaves (
|
||
<bibRefCitation id="EFD027EB5335FFFAFD689B6EFC68C105" author="Ziesenhenne F. C." box="[711,934,1292,1318]" pageId="23" pageNumber="34" pagination="209 - 239" refId="ref18033" refString="Ziesenhenne F. C. 1937. The Templeton Crocker Expedition. X. Echinoderms from the west coast of Lower California, the Gulf of California and Clarion Island. Zoologica 22 (15): 209 - 239." type="journal article" year="1937">Ziesenhenne 1937</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD027EB5335FFFAFC1B9B6EFBF3C105" author="Clark H. L." box="[948,1085,1292,1318]" pageId="23" pageNumber="34" pagination="331 - 352" refId="ref15672" refString="Clark H. L. 1940. XXI. Notes on Echinoderms from the west coast of Central America. Eastern Pacific Expeditions of the New York Zoological Society. Zoologica 25 (22): 331 - 352." type="journal article" year="1940">Clark 1940</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD027EB5335FFFAFBE39B6EFB12C105" author="Maluf L. Y." box="[1100,1244,1292,1318]" pageId="23" pageNumber="34" pagination="1 - 242" refId="ref17113" refString="Maluf L. Y. 1988. Composition and distribution of the central Eastern Pacific echinoderms. Natural History Museum of Los Angeles County, Technical Report 2: 1 - 242." type="book chapter" year="1988">Maluf 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD027EB5335FFFAFB459B6EFEF3C169" author="Solis-Marin F. A. & Alvarado J. J. & Abreu-Perez M. & Aguilera O. & Alio J. & Bacallado-Aranega J. J. & Williams S. M." pageId="23" pageNumber="34" pagination="543 - 658" refId="ref17532" refString="Solis-Marin F. A., Alvarado J. J., Abreu-Perez M., Aguilera O., Alio J., Bacallado-Aranega J. J. & Williams S. M. 2013. Appendix. In: Alvarado-Barrientos J. J. & Solis-Marin F. A. (eds) Echinoderm Research and Diversity in Latin America: 543 - 658. Springer-Verlag, Berlin. https: // doi. org / 10.1007 / 978 - 3 - 642 - 20051 - 9" type="book chapter" year="2013">
|
||
Solís-Marín
|
||
<emphasis id="B93586085335FFFAFF129B52FF3EC16A" box="[189,240,1327,1353]" italics="true" pageId="23" pageNumber="34">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD027EB5335FFFAFEE69B4DFD7CC169" author="Granja-Fernandez R. & Herrero-Perezrul M. D. & Lopez-Perez A. & Hernandez-Morales A. & Rangel-Solis P. D." box="[329,690,1327,1354]" pageId="23" pageNumber="34" pagination="37 - 47" refId="ref16005" refString="Granja-Fernandez R., Herrero-Perezrul M. D., Lopez-Perez A., Hernandez-Morales A. & Rangel-Solis P. D. 2015 a. A literature review of the Ophiuroidea (Echinodermata) from the Pacific coast of Mexico. Revista de Biologia Tropical 63 (2): 37 - 47. https: // doi. org / 10.15517 / RBT. V 63 I 2.23127" type="journal article" year="2015">
|
||
Granja-Fernández
|
||
<emphasis id="B93586085335FFFAFD8B9B52FD96C16A" box="[548,600,1327,1353]" italics="true" pageId="23" pageNumber="34">et al</emphasis>
|
||
. 2015a
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35B09915335FFF5FF129B13FD38C7B0" lastPageId="24" lastPageNumber="35" pageId="23" pageNumber="34" type="discussion">
|
||
<paragraph id="8BFE5A1A5335FFFAFF129B13FEE1C1A8" blockId="23.[189,303,1393,1419]" box="[189,303,1393,1419]" pageId="23" pageNumber="34">
|
||
<heading id="D0B6ED765335FFFAFF129B13FEE1C1A8" bold="true" box="[189,303,1393,1419]" fontSize="11" level="3" pageId="23" pageNumber="34" reason="3">
|
||
<emphasis id="B93586085335FFFAFF129B13FEE1C1A8" bold="true" box="[189,303,1393,1419]" pageId="23" pageNumber="34">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5335FFFAFF129BC2FCC7C292" blockId="23.[189,1399,1439,1713]" pageId="23" pageNumber="34">
|
||
In the original description of
|
||
<taxonomicName id="4C4121995335FFFAFDB49BC2FCB1C199" authority=", Lutken (1856)" authorityName=", Lutken" authorityYear="1856" box="[539,895,1439,1466]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="23" pageNumber="34" phylum="Echinodermata" rank="species" species="variegatum">
|
||
<emphasis id="B93586085335FFFAFDB49BC2FD08C19A" box="[539,710,1440,1466]" italics="true" pageId="23" pageNumber="34">O. variegatum</emphasis>
|
||
,
|
||
<bibRefCitation id="EFD027EB5335FFFAFD7A9BFDFCB1C199" author="Lutken C. F." box="[725,895,1439,1466]" pageId="23" pageNumber="34" pagination="20 - 26" refId="ref16758" refString="Lutken C. F. 1856. III. Bidrag til Kundskab om Ophiurerne ved Central-Amerikas Vestkyst. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening 1856: 20 - 26." type="journal article" year="1856">Lütken (1856)</bibRefCitation>
|
||
</taxonomicName>
|
||
provided just a brief description, but later he extended it (
|
||
<bibRefCitation id="EFD027EB5335FFFAFE2F9BA1FDE9C1FD" author="Lutken C. F." box="[384,551,1475,1502]" pageId="23" pageNumber="34" pagination="75 - 169" refId="ref16788" refString="Lutken C. F. 1859. Additamenta ad Historiam Ophiuridarum. Beskrivelser af de fra Vestindien og Central Amerikas Vestkyst hidtil beckjendte Slangestjerner. Kongelige Danske Videnskabernes Selskab Biologiske Skrifter 2: 75 - 169." type="journal article" year="1859">Lütken, 1859</bibRefCitation>
|
||
). However, the description provided in the present work is the most complete to date. All the characters mentioned by
|
||
<bibRefCitation id="EFD027EB5335FFFAFCB89B84FC74C222" author="Lutken C. F." box="[791,954,1510,1537]" pageId="23" pageNumber="34" pagination="20 - 26" refId="ref16758" refString="Lutken C. F. 1856. III. Bidrag til Kundskab om Ophiurerne ved Central-Amerikas Vestkyst. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening 1856: 20 - 26." type="journal article" year="1856">Lütken (1856</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD027EB5335FFFAFC679B84FBCAC222" author="Lutken C. F." box="[968,1028,1510,1537]" pageId="23" pageNumber="34" pagination="75 - 169" refId="ref16788" refString="Lutken C. F. 1859. Additamenta ad Historiam Ophiuridarum. Beskrivelser af de fra Vestindien og Central Amerikas Vestkyst hidtil beckjendte Slangestjerner. Kongelige Danske Videnskabernes Selskab Biologiske Skrifter 2: 75 - 169." type="journal article" year="1859">1859</bibRefCitation>
|
||
) correspond with the
|
||
<typeStatus id="54FAE4B85335FFFAFAA49B84FAB9C223" box="[1291,1399,1510,1536]" pageId="23" pageNumber="34" type="lectotype">lectotype</typeStatus>
|
||
and
|
||
<typeStatus id="54FAE4B85335FFFAFF5F986BFE40C200" box="[240,398,1545,1571]" pageId="23" pageNumber="34" type="paralectotype">paralectotype</typeStatus>
|
||
designated in this work. Although the type specimens are discolored, many authors (
|
||
<bibRefCitation id="EFD027EB5335FFFAFF6A984EFEAEC264" author="Lutken C. F." box="[197,352,1580,1607]" pageId="23" pageNumber="34" pagination="75 - 169" refId="ref16788" refString="Lutken C. F. 1859. Additamenta ad Historiam Ophiuridarum. Beskrivelser af de fra Vestindien og Central Amerikas Vestkyst hidtil beckjendte Slangestjerner. Kongelige Danske Videnskabernes Selskab Biologiske Skrifter 2: 75 - 169." type="journal article" year="1859">Lütken 1859</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD027EB5335FFFAFEC2984EFE31C265" author="Verrill A. E." box="[365,511,1580,1606]" pageId="23" pageNumber="34" pagination="251 - 322" refId="ref17963" refString="Verrill A. E. 1867. V. Notes on the Radiata in the Museum of Yale College, with descriptions of new genera and species. No. 2. Notes on the echinoderms of Panama and West Coast of America, with descriptions of new genera and species. Transactions of the Connecticut Academy of Arts and Sciences 1 (2): 251 - 322." type="journal article" year="1867">Verrill 1867</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD027EB5335FFFAFDA3984EFC82C264" author="Lutken C. F. & Mortensen T." box="[524,844,1580,1607]" pageId="23" pageNumber="34" pagination="93 - 208" refId="ref16825" refString="Lutken C. F. & Mortensen T. 1899. Reports on an exploration off the west coast of Mexico, central and South America, and off the Galapagos Islands, in charge of Alexander Agassiz, by the U. S. Fish Commission Steamer " Albatross ", during 1891, Lieut. Commander Z. L. Tanner, U. S. N., commanding. XXV. The Ophiuridae. Memoirs of the Museum of Comparative Zoology at Harvard College 23 (2): 93 - 208. https: // doi. org / 10.5962 / bhl. title. 65637" type="journal article" year="1899">Lütken & Mortensen 1899</bibRefCitation>
|
||
; H.L.
|
||
<bibRefCitation id="EFD027EB5335FFFAFC38984EFBD0C264" author="Clark H. L." box="[919,1054,1580,1607]" pageId="23" pageNumber="34" pagination="185 - 236" refId="ref15486" refString="Clark H. L. 1913. Echinoderms from Lower California, with descriptions of new species. Bulletin of the American Museum of Natural History 32: 185 - 236. https: // doi. org / 10.5962 / bhl. title. 1734" type="journal article" year="1913">Clark 1913</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD027EB5335FFFAFB83984FFBA4C264" author="Clark H. L." box="[1068,1130,1581,1607]" pageId="23" pageNumber="34" pagination="331 - 352" refId="ref15672" refString="Clark H. L. 1940. XXI. Notes on Echinoderms from the west coast of Central America. Eastern Pacific Expeditions of the New York Zoological Society. Zoologica 25 (22): 331 - 352." type="journal article" year="1940">1940</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD027EB5335FFFAFBD8984EFAD2C264" author="Koehler R." box="[1143,1308,1580,1607]" pageId="23" pageNumber="34" pagination="1 - 173" refId="ref16668" refString="Koehler R. 1914. A contribution to the study of ophiurans of the United States National Museum. Bulletin of the United States National Museum 84: 1 - 173. https: // doi. org / 10.5962 / bhl. title. 27414" type="journal article" year="1914">Koehler 1914</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD027EB5335FFFAFA85984FFF37C249" author="Boone L." pageId="23" pageNumber="34" pagination="1 - 217" refId="ref15113" refString="Boone L. 1933. Scientific results of cruises of the yachts " Eagle " and " Ara ", 1921 - 1928, William K. Vanderbilt, commanding. Coelenterata, Echinodermata and Mollusca. Bulletin of the Vanderbilt Marine Museum 4: 1 - 217." type="journal article" year="1933">Boone 1933</bibRefCitation>
|
||
) and our present work coincide that the coloration of this species consists of an olive-green disc with irregular blotches of bright rose-red and the arms banded with lighter and darker grayish green, brown or white every two to five dorsal segments.
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5335FFF5FF1298BAFE78C565" blockId="23.[189,1399,1752,2025]" lastBlockId="24.[189,1399,265,327]" lastPageId="24" lastPageNumber="35" pageId="23" pageNumber="34">
|
||
Although
|
||
<taxonomicName id="4C4121995335FFFAFE9A98BAFE2EC2D2" authorityName=", Lutken" authorityYear="1856" box="[309,480,1752,1778]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="23" pageNumber="34" phylum="Echinodermata" rank="species" species="variegatum">
|
||
<emphasis id="B93586085335FFFAFE9A98BAFE2EC2D2" box="[309,480,1752,1778]" italics="true" pageId="23" pageNumber="34">O. variegatum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4121995335FFFAFDB498BAFD0AC2D2" authorityName="Lutken" authorityYear="1859" box="[539,708,1752,1778]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="23" pageNumber="34" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B93586085335FFFAFDB498BAFD0AC2D2" box="[539,708,1752,1778]" italics="true" pageId="23" pageNumber="34">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
are completely different in overall morphology (
|
||
<tableCitation id="C6C36FA15335FFFAFAA498BAFAAAC2D1" box="[1291,1380,1752,1778]" captionStart="Table 1" captionStartId="10.[189,258,265,291]" captionTargetPageId="10" captionText="Table 1. Comparison of morphological characters between some species of Ophioderma from the Eastern Pacific." httpUri="http://table.plazi.org/id/DF3E0A925328FFE7FF129F6BFEBAC565" pageId="23" pageNumber="34" tableUuid="DF3E0A925328FFE7FF129F6BFEBAC565">Table 1</tableCitation>
|
||
), during our revisions we observed that it is very common to find specimens of
|
||
<taxonomicName id="4C4121995335FFFAFB949899FB2AC336" authorityName=", Lutken" authorityYear="1856" box="[1083,1252,1787,1813]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="23" pageNumber="34" phylum="Echinodermata" rank="species" species="variegatum">
|
||
<emphasis id="B93586085335FFFAFB949899FB2AC336" box="[1083,1252,1787,1813]" italics="true" pageId="23" pageNumber="34">O. variegatum</emphasis>
|
||
</taxonomicName>
|
||
identified as
|
||
<taxonomicName id="4C4121995335FFFAFF12997CFEABC31B" authorityName="Lutken" authorityYear="1859" box="[189,357,1822,1848]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="23" pageNumber="34" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B93586085335FFFAFF12997CFEABC31B" box="[189,357,1822,1848]" italics="true" pageId="23" pageNumber="34">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
, and vice versa. This tendency is very common in some published works (see references below); for example, we detected that in the description (i.e., naked radial shields and covered adoral shields) as well as the images of the species by
|
||
<bibRefCitation id="EFD027EB5335FFFAFD4B9907FCA3C35C" author="Caso M. E." box="[740,877,1892,1919]" pageId="23" pageNumber="34" pagination="219 - 312" refId="ref15224" refString="Caso M. E. 1951. Contribucion al conocimiento de los ofiuroideos de Mexico. 1. Algunas especies de ofiuroideos litorales. Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico 22 (1): 219 - 312." type="journal article" year="1951">Caso (1951</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD027EB5335FFFAFCD79906FC7BC35D" author="Caso M. E." box="[888,949,1892,1918]" pageId="23" pageNumber="34" pagination="293 - 330" refId="ref15268" refString="Caso M. E. 1962. Estudios sobre equinodermos de Mexico. Contribucion al conocimiento de los equinodermos de la Islas Revillagigedo. Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico 33 (1 - 2): 293 - 330." type="journal article" year="1962">1962</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD027EB5335FFFAFC6E9907FC30C35C" author="Caso M. E." box="[961,1022,1893,1919]" pageId="23" pageNumber="34" pagination="197 - 368" refId="ref15314" refString="Caso M. E. 1979. Los equinodermos de la Bahia de Mazatlan, Sinaloa. Anales del Centro de Ciencias del Mar y Limnologia, Universidad Nacional Autonoma de Mexico 6 (1): 197 - 368." type="journal article" year="1979">1979</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD027EB5335FFFAFBA59906FB86C35C" author="Caso M. E." box="[1034,1096,1892,1919]" pageId="23" pageNumber="34" pagination="223 - 248" refId="ref15355" refString="Caso M. E. 1986. Descripcion de una nueva especie de ofiuroideo de la Bahia de Mazatlan, Sin. Ophioderma sodipallaresi sp. nov. y comparacion con Ophioderma variegatum Lutken. Anales del Centro de Ciencias del Mar y Limnologia, Universidad Nacional Autonoma de Mexico 13 (2): 223 - 248." type="journal article" year="1986">1986</bibRefCitation>
|
||
), the specimens identified as
|
||
<taxonomicName id="4C4121995335FFFAFF7199EAFE46C381" authorityName=", Lutken" authorityYear="1856" box="[222,392,1928,1954]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="23" pageNumber="34" phylum="Echinodermata" rank="species" species="variegatum">
|
||
<emphasis id="B93586085335FFFAFF7199EAFE46C381" box="[222,392,1928,1954]" italics="true" pageId="23" pageNumber="34">O. variegatum</emphasis>
|
||
</taxonomicName>
|
||
were misidentified and corresponded to
|
||
<taxonomicName id="4C4121995335FFFAFCC599EAFBDDC381" authorityName="Lutken" authorityYear="1859" box="[874,1043,1928,1954]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="23" pageNumber="34" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B93586085335FFFAFCC599EAFBDDC381" box="[874,1043,1928,1954]" italics="true" pageId="23" pageNumber="34">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
. All the specimens studied by Caso are deposited at ICML-UNAM, and this identity was corroborated (see: Supplementary material). It is very feasible that the material mentioned in
|
||
<bibRefCitation id="EFD027EB5335FFFAFD5199ACFC1CC3CA" author="Caso M. E. & Laguarda-Figueras A. & Solis-Marin F. A. & Ortega-Salas A. & Duran-Gonzalez A. L." box="[766,978,1998,2025]" pageId="23" pageNumber="34" pagination="101 - 120" refId="ref15414" refString="Caso M. E., Laguarda-Figueras A., Solis-Marin F. A., Ortega-Salas A. & Duran-Gonzalez A. L. 1996. Contribucion al conocimiento de la ecologia de las comunidades de equinodermos de la Bahia de Mazatlan, Sinaloa, Mexico. Anales del Instituto de Ciencias del Mar y Limnologia, Universidad Nacional Autonoma de Mexico 22 (1): 101 - 120." type="journal article" year="1996">
|
||
Caso
|
||
<emphasis id="B93586085335FFFAFCEF99ADFCBBC3CB" box="[832,885,1998,2024]" italics="true" pageId="23" pageNumber="34">et al</emphasis>
|
||
. (1996)
|
||
</bibRefCitation>
|
||
also corresponds to
|
||
<emphasis id="B93586085335FFFAFB6999ADFAB8C3CB" box="[1222,1398,1999,2025]" italics="true" pageId="23" pageNumber="34">
|
||
<taxonomicName id="4C4121995335FFFAFB6999ADFABCC3CB" authorityName="Lutken" authorityYear="1859" box="[1222,1394,1999,2025]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="23" pageNumber="34" phylum="Echinodermata" rank="species" species="panamense">O. panamense</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but neither a description nor images of the studied specimens are provided in this work and therefore we cannot confirm it.
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A533AFFF5FF129F0CFB01C619" blockId="24.[189,1399,366,571]" pageId="24" pageNumber="35">
|
||
The jaw of the
|
||
<typeStatus id="54FAE4B8533AFFF5FED99F0DFE67C5AA" box="[374,425,367,393]" pageId="24" pageNumber="35">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="4C412199533AFFF5FD929F0DFD27C5AB" authorityName=", Lutken" authorityYear="1856" box="[573,745,367,393]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="24" pageNumber="35" phylum="Echinodermata" rank="species" species="variegatum">
|
||
<emphasis id="B9358608533AFFF5FD929F0DFD9CC5AA" box="[573,594,367,393]" italics="true" pageId="24" pageNumber="35">O</emphasis>
|
||
.
|
||
<emphasis id="B9358608533AFFF5FDCC9F0DFD27C5AB" box="[611,745,367,392]" italics="true" pageId="24" pageNumber="35">variegatum</emphasis>
|
||
</taxonomicName>
|
||
was closed, but like the other species of
|
||
<taxonomicName id="4C412199533AFFF5FB749F0CFAA1C5AB" authorityName="Muller & Troschel" authorityYear="1840" box="[1243,1391,366,392]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="24" pageNumber="35" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B9358608533AFFF5FB749F0CFAA1C5AB" box="[1243,1391,366,392]" italics="true" pageId="24" pageNumber="35">Ophioderma</emphasis>
|
||
</taxonomicName>
|
||
, voucher specimens with open jaws have one large and pointed OPRSp on each side, visible within the buccal cavity. It is important to recall that OPRSp were previously mentioned as spiniform structures or as a conspicuous pointed papilla (e.g., H.L.
|
||
<bibRefCitation id="EFD027EB533AFFF5FD169FBBFCF1C5D0" author="Clark H. L." box="[697,831,473,499]" pageId="24" pageNumber="35" pagination="429 - 453" refId="ref15585" refString="Clark H. L. 1917. Reports on the scientific results of the expedition to the Eastern Tropical Pacific in charge of Alexander Agassiz, by the U. S. Fish Commission Steamer Albatross from October, 1904, to March, 1905, Lieut. Commander L. M. Garrett, U. S. N., Commanding. XXX. Ophiuroidea. Bulletin of the Museum of Comparative Zoology at Harvard College 61 (12): 429 - 453." type="journal article" year="1917">Clark 1917</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFD027EB533AFFF5FCFC9FBBFBCBC5D0" author="Hendler G." box="[851,1029,473,499]" pageId="24" pageNumber="35" pagination="189 - 311" refId="ref16304" refString="Hendler G. 2018. Armed to the teeth: a new paradigm for the buccal skeleton of brittle stars (Echinodermata: Ophiuroidea). Contributions in Science 526: 189 - 311." type="journal article" year="2018">Hendler (2018)</bibRefCitation>
|
||
coined the term suggesting that OPRSp in
|
||
<taxonomicName id="4C412199533AFFF5FE969F9FFE03C634" authorityName="Muller & Troschel" authorityYear="1840" box="[313,461,509,535]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="24" pageNumber="35" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B9358608533AFFF5FE969F9FFE03C634" box="[313,461,509,535]" italics="true" pageId="24" pageNumber="35">Ophioderma</emphasis>
|
||
</taxonomicName>
|
||
(along with OPa and teeth) are used to grip food between the closed jaws and to retain items in the buccal cavity, facilitating a feeding habit of predation and scavenging.
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A533AFFF5FF129C00FD7AC6CB" blockId="24.[189,1399,610,744]" pageId="24" pageNumber="35">
|
||
According to our study,
|
||
<taxonomicName id="4C412199533AFFF5FE7B9C01FDB2C65F" authorityName=", Lutken" authorityYear="1856" box="[468,636,611,637]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="24" pageNumber="35" phylum="Echinodermata" rank="species" species="variegatum">
|
||
<emphasis id="B9358608533AFFF5FE7B9C01FDB2C65F" box="[468,636,611,637]" italics="true" pageId="24" pageNumber="35">O. variegatum</emphasis>
|
||
</taxonomicName>
|
||
is less abundant and less conspicuous in rock and coral reef zones of the Eastern Pacific compared to
|
||
<taxonomicName id="4C412199533AFFF5FDFD9CE4FD36C6BC" authorityName="Lutken" authorityYear="1859" box="[594,760,646,672]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="24" pageNumber="35" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9358608533AFFF5FDFD9CE4FD36C6BC" box="[594,760,646,672]" italics="true" pageId="24" pageNumber="35">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C412199533AFFF5FCA99CE4FBE9C6BC" authorityName=", Lutken" authorityYear="1856" box="[774,1063,646,672]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="24" pageNumber="35" phylum="Echinodermata" rank="species" species="variegatum">
|
||
<emphasis id="B9358608533AFFF5FCA99CE4FBE9C6BC" box="[774,1063,646,672]" italics="true" pageId="24" pageNumber="35">Ophioderma variegatum</emphasis>
|
||
</taxonomicName>
|
||
tends to inhabit deeper areas (up to
|
||
<quantity id="4CB9F7FF533AFFF5FEA39CCBFE99C6E7" box="[268,343,681,708]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="24" pageNumber="35" unit="m" value="110.0">110 m</quantity>
|
||
) and it is commonly collected by dredging, whereas
|
||
<taxonomicName id="4C412199533AFFF5FC629CC8FBB8C6E0" authorityName="Lutken" authorityYear="1859" box="[973,1142,682,708]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="24" pageNumber="35" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9358608533AFFF5FC629CC8FBB8C6E0" box="[973,1142,682,708]" italics="true" pageId="24" pageNumber="35">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
is easily collected by hand in shallow waters (up to
|
||
<quantity id="4CB9F7FF533AFFF5FDB09CAFFD94C6C4" box="[543,602,717,743]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="35" unit="m" value="20.0">20 m</quantity>
|
||
depth).
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A533AFFF5FF129D6DFD38C7B0" blockId="24.[189,1399,782,916]" pageId="24" pageNumber="35">
|
||
<bibRefCitation id="EFD027EB533AFFF5FF129D6DFE2BC70A" author="Maluf L. Y. & Brusca R. C." box="[189,485,782,809]" pageId="24" pageNumber="35" pagination="327 - 343" refId="ref17214" refString="Maluf L. Y. & Brusca R. C. 2005. Chapter 18. Echinodermata. In: Hendrickx M. E., Brusca R. C. & Findley L. T. (eds) A Distributional Checklist of the Macrofauna of the Gulf of California, Mexico. Part I. Invertebrates: 327 - 343. Arizona-Sonora Desert Museum, Tucson." type="book chapter" year="2005">Maluf & Brusca (2005)</bibRefCitation>
|
||
reported
|
||
<taxonomicName id="4C412199533AFFF5FDCC9D6DFCDAC70B" authorityName=", Lutken" authorityYear="1856" box="[611,788,783,809]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="24" pageNumber="35" phylum="Echinodermata" rank="species" species="variegatum">
|
||
<emphasis id="B9358608533AFFF5FDCC9D6DFDB6C70A" box="[611,632,783,809]" italics="true" pageId="24" pageNumber="35">O</emphasis>
|
||
.
|
||
<emphasis id="B9358608533AFFF5FD219D6DFCDAC70B" box="[654,788,783,808]" italics="true" pageId="24" pageNumber="35">variegatum</emphasis>
|
||
</taxonomicName>
|
||
for Gorgona Island (
|
||
<collectingCountry id="F3561A8A533AFFF5FB8C9D6DFB54C709" box="[1059,1178,783,810]" name="Colombia" pageId="24" pageNumber="35">Colombia</collectingCountry>
|
||
) but without any supporting data; therefore, we suspect that this record is invalid. Nevertheless, due to the wide geographic distribution of the species, it is very plausible to find it in
|
||
<collectingCountry id="F3561A8A533AFFF5FC449D34FB91C753" box="[1003,1119,854,880]" name="Colombia" pageId="24" pageNumber="35">Colombia</collectingCountry>
|
||
(or even further south), but specimens are needed to confirm this record.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |