643 lines
102 KiB
XML
643 lines
102 KiB
XML
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<mods:title id="E6C3D26F56107614D8EA19E7195EEC18">Ophioderma hendleri sp. nov. (Echinodermata: Ophiuroidea: Ophiodermatidae) and its congeners from the Eastern Pacific</mods:title>
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<taxonomicName id="4C412199532FFFE0FD9B9F6AFBCEC500" authority="Lutken, 1859" authorityName="Lutken" authorityYear="1859" box="[564,1024,264,291]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="24" phylum="Echinodermata" rank="species" species="panamense">
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<emphasis id="B9358608532FFFE0FD9B9F6AFC92C500" bold="true" box="[564,860,264,291]" italics="true" pageId="13" pageNumber="24">Ophioderma panamense</emphasis>
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<bibRefCitation id="EFD027EB532FFFE0FCCC9F6AFBCEC500" author="Lutken C. F." box="[867,1024,264,291]" pageId="13" pageNumber="24" pagination="75 - 169" refId="ref16788" refString="Lutken C. F. 1859. Additamenta ad Historiam Ophiuridarum. Beskrivelser af de fra Vestindien og Central Amerikas Vestkyst hidtil beckjendte Slangestjerner. Kongelige Danske Videnskabernes Selskab Biologiske Skrifter 2: 75 - 169." type="journal article" year="1859">Lütken, 1859</bibRefCitation>
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E–J, 6;
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<emphasis id="B9358608532FFFE0FF129F0CFE2AC5AB" box="[189,484,366,392]" italics="true" pageId="13" pageNumber="24">Ophioderma panamensis</emphasis>
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<bibRefCitation id="EFD027EB532FFFE0FE459F0CFD0BC5AB" author="Lutken C. F." box="[490,709,366,393]" pageId="13" pageNumber="24" pagination="75 - 169" refId="ref16788" refString="Lutken C. F. 1859. Additamenta ad Historiam Ophiuridarum. Beskrivelser af de fra Vestindien og Central Amerikas Vestkyst hidtil beckjendte Slangestjerner. Kongelige Danske Videnskabernes Selskab Biologiske Skrifter 2: 75 - 169." type="journal article" year="1859">Lütken, 1859: 193</bibRefCitation>
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.
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<paragraph id="8BFE5A1A532FFFE0FF129FF3FCA6C588" blockId="13.[189,872,366,428]" box="[189,872,401,428]" pageId="13" pageNumber="24">
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<taxonomicName id="4C412199532FFFE0FF129FF3FCC7C58F" authority="Nielsen, 1932: 334 - 335" authorityName="Nielsen" authorityPageNumber="334 - 335" authorityYear="1932" box="[189,777,401,428]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophiocryptus" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="24" phylum="Echinodermata" rank="species" species="granulosus">
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<emphasis id="B9358608532FFFE0FF129FF3FE28C588" box="[189,486,401,427]" italics="true" pageId="13" pageNumber="24">Ophiocryptus granulosus</emphasis>
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<bibRefCitation id="EFD027EB532FFFE0FE419FF3FCC7C58F" author="Nielsen E." box="[494,777,401,428]" pageId="13" pageNumber="24" pagination="241 - 346" refId="ref17315" refString="Nielsen E. 1932. Ophiurans from the Gulf of Panama, California and the Strait of Georgia. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening 91: 241 - 346." type="journal article" year="1932">Nielsen, 1932: 334–335</bibRefCitation>
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, fig. 38.
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<paragraph id="8BFE5A1A532FFFE0FF129FB1FB24C5CD" blockId="13.[189,1399,467,849]" box="[189,1258,467,494]" pageId="13" pageNumber="24">
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<emphasis id="B9358608532FFFE0FF129FB1FE7DC5CE" box="[189,435,467,493]" italics="true" pageId="13" pageNumber="24">Ophiura panamensis</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0AE07C0B532FFFE0FE7F9FB6FD71C5CE" author="Lyman T." box="[464,703,467,494]" page="32" pageId="13" pageNumber="24" year="1865">
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<bibRefCitation id="EFD027EB532FFFE0FE7F9FB6FD71C5CE" author="Lyman T." box="[464,703,467,494]" pageId="13" pageNumber="24" pagination="1 - 200" refId="ref16993" refString="Lyman T. 1865. Ophiuridae and Astrophytidae. Illustrated Catalogue of the Museum of Comparative Zoology at Harvard College 1: 1 - 200." type="journal article" year="1865">Lyman 1865: 32–34</bibRefCitation>
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</treatmentCitation>
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. —
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<treatmentCitation id="0AE07C0B532FFFE0FD409FB1FC38C5CD" author="Verrill A. E." box="[751,1014,467,494]" page="253" pageId="13" pageNumber="24" year="1867">
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<bibRefCitation id="EFD027EB532FFFE0FD409FB1FC38C5CD" author="Verrill A. E." box="[751,1014,467,494]" pageId="13" pageNumber="24" pagination="251 - 322" refId="ref17963" refString="Verrill A. E. 1867. V. Notes on the Radiata in the Museum of Yale College, with descriptions of new genera and species. No. 2. Notes on the echinoderms of Panama and West Coast of America, with descriptions of new genera and species. Transactions of the Connecticut Academy of Arts and Sciences 1 (2): 251 - 322." type="journal article" year="1867">Verrill 1867: 253–254</bibRefCitation>
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</treatmentCitation>
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. —
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<treatmentCitation id="0AE07C0B532FFFE0FB8A9FB6FB2BC5CD" author="Ives J. E." box="[1061,1253,467,494]" page="175" pageId="13" pageNumber="24" year="1889">
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<bibRefCitation id="EFD027EB532FFFE0FB8A9FB6FB2BC5CD" author="Ives J. E." box="[1061,1253,467,494]" pageId="13" pageNumber="24" pagination="169 - 179" refId="ref16594" refString="Ives J. E. 1889 b. Catalogue of the Asteroidea and Ophiuroidea in the Collection of the Academy of Natural Sciences of Philadelphia. Proceedings of the Academy of Natural Sciences of Philadelphia 41: 169 - 179." type="journal article" year="1889">Ives 1889b: 175</bibRefCitation>
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</treatmentCitation>
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.
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</paragraph>
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<paragraph id="8BFE5A1A532FFFE0FF129F95FBD1C65F" blockId="13.[189,1399,467,849]" pageId="13" pageNumber="24">
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<taxonomicName id="4C412199532FFFE0FF129F95FE2BC632" authorityName="Lutken" authorityYear="1859" box="[189,485,503,529]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="24" phylum="Echinodermata" rank="species" species="panamensis">
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<emphasis id="B9358608532FFFE0FF129F95FE2BC632" box="[189,485,503,529]" italics="true" pageId="13" pageNumber="24">Ophioderma panamensis</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0AE07C0B532FFFE0FDAC9F95FD2CC632" author="Ives J. E." box="[515,738,503,529]" page="76" pageId="13" pageNumber="24" year="1889">
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<bibRefCitation id="EFD027EB532FFFE0FDAC9F95FD2CC632" author="Ives J. E." box="[515,738,503,529]" pageId="13" pageNumber="24" pagination="76 - 77" refId="ref16563" refString="Ives J. E. 1889 a. Variation in Ophiura panamensis and Ophiura teres. Proceedings of the Academy of Natural Sciences of Philadelphia 41: 76 - 77." type="journal article" year="1889">Ives 1889a: 76–77</bibRefCitation>
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</treatmentCitation>
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. —
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<treatmentCitation id="0AE07C0B532FFFE0FCBC9F95FC3BC632" author="Koehler R." box="[787,1013,503,529]" page="282" pageId="13" pageNumber="24" year="1907">
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<bibRefCitation id="EFD027EB532FFFE0FCBC9F95FC3BC632" author="Koehler R." box="[787,1013,503,529]" pageId="13" pageNumber="24" pagination="279 - 370" refId="ref16635" refString="Koehler R. 1907. Revision de la collection des ophiures du Museum D'Histoire Naturelle de Paris. Bulletin Scientifique de la France et de la Belgique 61: 279 - 370." type="journal article" year="1907">Koehler 1907: 282</bibRefCitation>
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</treatmentCitation>
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. —
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<treatmentCitation id="0AE07C0B532FFFE0FB889F95FAEBC631" author="McClendon J. F." box="[1063,1317,503,530]" page="35" pageId="13" pageNumber="24" year="1909">
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<bibRefCitation id="EFD027EB532FFFE0FB889F95FAEBC631" author="McClendon J. F." box="[1063,1317,503,530]" pageId="13" pageNumber="24" pagination="33 - 64" refId="ref17283" refString="McClendon J. F. 1909. XXV. The ophiurans of the San Diego region. University of California Publications in Zoology 6 (3): 33 - 64." type="journal article" year="1909">McClendon 1909: 35</bibRefCitation>
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</treatmentCitation>
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, fig. 1. — H.L.
|
||
<treatmentCitation id="0AE07C0B532FFFE0FEFD9C78FDD7C616" author="Clark H. L." box="[338,537,538,565]" page="205" pageId="13" pageNumber="24" year="1913">
|
||
<bibRefCitation id="EFD027EB532FFFE0FEFD9C78FDD7C616" author="Clark H. L." box="[338,537,538,565]" pageId="13" pageNumber="24" pagination="185 - 236" refId="ref15486" refString="Clark H. L. 1913. Echinoderms from Lower California, with descriptions of new species. Bulletin of the American Museum of Natural History 32: 185 - 236. https: // doi. org / 10.5962 / bhl. title. 1734" type="journal article" year="1913">Clark 1913: 205</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AE07C0B532FFFE0FD879C78FD2EC617" author="Clark H. L." box="[552,736,538,565]" page="300" pageId="13" pageNumber="24" year="1915">
|
||
<bibRefCitation id="EFD027EB532FFFE0FD879C78FD2EC617" author="Clark H. L." box="[552,736,538,565]" pageId="13" pageNumber="24" pagination="164 - 376" refId="ref15532" refString="Clark H. L. 1915. Catalogue of recent ophiurans: based on the collection of the Museum of Comparative Zoology. Memoirs of the Museum of Comparative Zoology 25 (4): 164 - 376. https: // doi. org / 10.5962 / bhl. title. 48598" type="journal article" year="1915">1915: 300–301</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AE07C0B532FFFE0FCBA9C78FC21C617" author="Campbell A. S." box="[789,1007,538,564]" page="2" pageId="13" pageNumber="24" year="1921">
|
||
<bibRefCitation id="EFD027EB532FFFE0FCBA9C78FC21C617" author="Campbell A. S." box="[789,1007,538,564]" pageId="13" pageNumber="24" pagination="2 - 4" refId="ref15200" refString="Campbell A. S. 1921. Littoral ophiurans at Laguna Beach. Journal of Entomology and Zoology 13: 2 - 4." type="journal article" year="1921">Campbell 1921: 2</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 1. —
|
||
<treatmentCitation id="0AE07C0B532FFFE0FBD89C78FABDC616" author="Caso M. E." box="[1143,1395,538,565]" page="265" pageId="13" pageNumber="24" year="1951">
|
||
<bibRefCitation id="EFD027EB532FFFE0FBD89C78FABDC616" author="Caso M. E." box="[1143,1395,538,565]" pageId="13" pageNumber="24" pagination="219 - 312" refId="ref15224" refString="Caso M. E. 1951. Contribucion al conocimiento de los ofiuroideos de Mexico. 1. Algunas especies de ofiuroideos litorales. Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico 22 (1): 219 - 312." type="journal article" year="1951">Caso 1951: 265–272</bibRefCitation>
|
||
</treatmentCitation>
|
||
, figs 25–28; 1962: 296, 302; 1979: 210, pl. 97–100; 1986: 225, 239, pl. 14–19. —
|
||
<treatmentCitation id="0AE07C0B532FFFE0FB0C9C5CFE11C65F" author="Granja-Fernandez R. & Herrero-Perezrul M. D. & Lopez-Perez R. A. & Hernandez L. & Rodriguez-Zaragoza F. A. & Jones R. W. & Pineda-Lopez R." httpUri="http://treatment.plazi.org/id/1F54ED41913165EB0CA285FD8E5E43DB" page="132" pageId="13" pageNumber="24" year="2014">
|
||
<bibRefCitation id="EFD027EB532FFFE0FB0C9C5CFE11C65F" author="Granja-Fernandez R. & Herrero-Perezrul M. D. & Lopez-Perez R. A. & Hernandez L. & Rodriguez-Zaragoza F. A. & Jones R. W. & Pineda-Lopez R." pageId="13" pageNumber="24" pagination="101 - 145" refId="ref15937" refString="Granja-Fernandez R., Herrero-Perezrul M. D., Lopez-Perez R. A., Hernandez L., Rodriguez-Zaragoza F. A., Jones R. W. & Pineda-Lopez R. 2014. Ophiuroidea (Echinodermata) from coral reefs in the Mexican Pacific. Zookeys 406: 101 - 145. https: // doi. org / 10.3897 / zookeys. 406.6306" type="journal article" year="2014">
|
||
Granja-Fernández
|
||
<emphasis id="B9358608532FFFE0FF459C00FED0C658" box="[234,286,609,635]" italics="true" pageId="13" pageNumber="24">et al</emphasis>
|
||
. 2014: 132–134
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 6a–f; 2015a: 41; 2015b: 95; 2017: 172, 174.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A532FFFE0FF129CE7FDFDC729" blockId="13.[189,1399,467,849]" pageId="13" pageNumber="24">
|
||
<treatmentCitationGroup id="AB517D34532FFFE0FF129CE7FDFDC729" pageId="13" pageNumber="24">
|
||
<taxonomicName id="4C412199532FFFE0FF129CE7FE2EC6BC" authorityName="Lutken" authorityYear="1859" box="[189,480,645,671]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="24" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9358608532FFFE0FF129CE7FE2EC6BC" box="[189,480,645,671]" italics="true" pageId="13" pageNumber="24">Ophioderma panamense</emphasis>
|
||
</taxonomicName>
|
||
– H.L.
|
||
<treatmentCitation id="0AE07C0B532FFFE0FDEB9CE7FC89C6BC" author="Clark H. L." box="[580,839,645,672]" page="301" pageId="13" pageNumber="24" year="1915">
|
||
<bibRefCitation id="EFD027EB532FFFE0FDEB9CE7FC89C6BC" author="Clark H. L." box="[580,839,645,672]" pageId="13" pageNumber="24" pagination="164 - 376" refId="ref15532" refString="Clark H. L. 1915. Catalogue of recent ophiurans: based on the collection of the Museum of Comparative Zoology. Memoirs of the Museum of Comparative Zoology 25 (4): 164 - 376. https: // doi. org / 10.5962 / bhl. title. 48598" type="journal article" year="1915">Clark 1915: 301–302</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AE07C0B532FFFE0FCF99CE7FC1CC6BC" author="Clark H. L." box="[854,978,645,671]" page="443" pageId="13" pageNumber="24" year="1917">
|
||
<bibRefCitation id="EFD027EB532FFFE0FCF99CE7FC1CC6BC" author="Clark H. L." box="[854,978,645,671]" pageId="13" pageNumber="24" pagination="429 - 453" refId="ref15585" refString="Clark H. L. 1917. Reports on the scientific results of the expedition to the Eastern Tropical Pacific in charge of Alexander Agassiz, by the U. S. Fish Commission Steamer Albatross from October, 1904, to March, 1905, Lieut. Commander L. M. Garrett, U. S. N., Commanding. XXX. Ophiuroidea. Bulletin of the Museum of Comparative Zoology at Harvard College 61 (12): 429 - 453." type="journal article" year="1917">1917: 443</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AE07C0B532FFFE0FC4D9CE7FB57C6BC" author="Clark H. L." box="[994,1177,645,671]" page="341" pageId="13" pageNumber="24" year="1940">
|
||
<bibRefCitation id="EFD027EB532FFFE0FC4D9CE7FB57C6BC" author="Clark H. L." box="[994,1177,645,671]" pageId="13" pageNumber="24" pagination="331 - 352" refId="ref15672" refString="Clark H. L. 1940. XXI. Notes on Echinoderms from the west coast of Central America. Eastern Pacific Expeditions of the New York Zoological Society. Zoologica 25 (22): 331 - 352." type="journal article" year="1940">1940: 341–342</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AE07C0B532FFFE0FB7F9CE7FE9BC6E0" author="Nielsen E." page="327" pageId="13" pageNumber="24" year="1932">
|
||
<bibRefCitation id="EFD027EB532FFFE0FB7F9CE7FE9BC6E0" author="Nielsen E." pageId="13" pageNumber="24" pagination="241 - 346" refId="ref17315" refString="Nielsen E. 1932. Ophiurans from the Gulf of Panama, California and the Strait of Georgia. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening 91: 241 - 346." type="journal article" year="1932">Nielsen 1932: 327–330</bibRefCitation>
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||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AE07C0B532FFFE0FE279CCAFD6FC6E0" author="Ziesenhenne F. C." box="[392,673,680,707]" page="227" pageId="13" pageNumber="24" year="1937">
|
||
<bibRefCitation id="EFD027EB532FFFE0FE279CCAFD6FC6E0" author="Ziesenhenne F. C." box="[392,673,680,707]" pageId="13" pageNumber="24" pagination="209 - 239" refId="ref18033" refString="Ziesenhenne F. C. 1937. The Templeton Crocker Expedition. X. Echinoderms from the west coast of Lower California, the Gulf of California and Clarion Island. Zoologica 22 (15): 209 - 239." type="journal article" year="1937">Ziesenhenne 1937: 227</bibRefCitation>
|
||
</treatmentCitation>
|
||
,
|
||
<treatmentCitation id="0AE07C0B532FFFE0FD019CCAFCAAC6E0" author="Ziesenhenne F. C." box="[686,868,680,707]" page="192" pageId="13" pageNumber="24" year="1955">
|
||
<bibRefCitation id="EFD027EB532FFFE0FD019CCAFCAAC6E0" author="Ziesenhenne F. C." box="[686,868,680,707]" pageId="13" pageNumber="24" pagination="185 - 201" refId="ref18074" refString="Ziesenhenne F. C. 1955. A review of the genus Ophioderma Muller and Troschel. In: Allan Hancock Foundation for Scientific Research (eds) Essays in the Natural Sciences in Honor of Captain Allan Hancock: 185 - 201. University of Southern California Press, Los Angeles." type="book chapter" year="1955">1955: 192–197</bibRefCitation>
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||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AE07C0B532FFFE0FC379CCAFADAC6E0" author="Boolotian R. A. & Leighton D." box="[920,1300,680,707]" page="10" pageId="13" pageNumber="24" year="1966">
|
||
<bibRefCitation id="EFD027EB532FFFE0FC379CCAFADAC6E0" author="Boolotian R. A. & Leighton D." box="[920,1300,680,707]" pageId="13" pageNumber="24" pagination="1 - 20" refId="ref15069" refString="Boolotian R. A. & Leighton D. 1966. A key to the species of Ophiuroidea (brittle stars) of the Santa Monica Bay and adjacent areas. Los Angeles County Museum Contributions in Science 93: 1 - 20." type="journal article" year="1966">Boolotian & Leighton 1966: 10</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 23. —
|
||
<treatmentCitation id="0AE07C0B532FFFE0FEA09CAEFE70C6C5" author="Luke S. R." box="[271,446,716,742]" page="33" pageId="13" pageNumber="24" year="1982">
|
||
<bibRefCitation id="EFD027EB532FFFE0FEA09CAEFE70C6C5" author="Luke S. R." box="[271,446,716,742]" pageId="13" pageNumber="24" refId="ref16714" refString="Luke S. R. 1982. Catalog of the Benthic Invertebrate Collections of the SCRIPPS Institution of Oceanography, Echinodermata. SCRIPPS Institution of Oceanography Reference Series, no. 82 - 5, University of California, San Diego. CA." type="book" year="1982">Luke 1982: 33</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AE07C0B532FFFE0FE419CAEFCF3C6C5" author="Solis-Marin F. A. & Laguarda-Figueras A. & Duran-Gonzalez A. & Ahearn C. G. & Torres-Vega J." box="[494,829,716,743]" page="128" pageId="13" pageNumber="24" year="2005">
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||
<bibRefCitation id="EFD027EB532FFFE0FE419CAEFCF3C6C5" author="Solis-Marin F. A. & Laguarda-Figueras A. & Duran-Gonzalez A. & Ahearn C. G. & Torres-Vega J." box="[494,829,716,743]" pageId="13" pageNumber="24" pagination="123 - 137" refId="ref17468" refString="Solis-Marin F. A., Laguarda-Figueras A., Duran-Gonzalez A., Ahearn C. G. & Torres-Vega J. 2005. Equinodermos (Echinodermata) del Golfo de California, Mexico. Revista de Biologia Tropical 53 (3): 123 - 137. https: // doi. org / 10.15517 / RBT. V 53 I 3.26770" type="journal article" year="2005">
|
||
Solís-Marín
|
||
<emphasis id="B9358608532FFFE0FD2D9CAFFD7BC6C5" box="[642,693,716,742]" italics="true" pageId="13" pageNumber="24">et al</emphasis>
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||
. 2005: 128
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||
</bibRefCitation>
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||
</treatmentCitation>
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||
; 2013: 570. —
|
||
<treatmentCitation id="0AE07C0B532FFFE0FC5B9CAEFABDC6C5" author="Honey-Escandon M. & Solis-Marin F. A. & Laguarda-Figueras A." box="[1012,1395,716,743]" page="63" pageId="13" pageNumber="24" year="2008">
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<bibRefCitation id="EFD027EB532FFFE0FC5B9CAEFABDC6C5" author="Honey-Escandon M. & Solis-Marin F. A. & Laguarda-Figueras A." box="[1012,1395,716,743]" pageId="13" pageNumber="24" pagination="57 - 73" refId="ref16471" refString="Honey-Escandon M., Solis-Marin F. A. & Laguarda-Figueras A. 2008. Equinodermos (Echinodermata) del Pacifico Mexicano. Revista de Biologia Tropical 56 (3): 57 - 73. https: // doi. org / 10.15517 / RBT. V 56 I 3.27079" type="journal article" year="2008">
|
||
Honey-Escandón
|
||
<emphasis id="B9358608532FFFE0FB6A9CAFFB37C6C5" box="[1221,1273,716,742]" italics="true" pageId="13" pageNumber="24">et al</emphasis>
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. 2008: 63
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</bibRefCitation>
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</treatmentCitation>
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. —
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||
<treatmentCitation id="0AE07C0B532FFFE0FEA29C92FDE1C729" author="Alvarado J. J. & Solis-Marin F. A. & Ahearn C. G." box="[269,559,752,778]" page="48" pageId="13" pageNumber="24" year="2010">
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||
<bibRefCitation id="EFD027EB532FFFE0FEA29C92FDE1C729" author="Alvarado J. J. & Solis-Marin F. A. & Ahearn C. G." box="[269,559,752,778]" pageId="13" pageNumber="24" pagination="45 - 56" refId="ref14865" refString="Alvarado J. J., Solis-Marin F. A. & Ahearn C. G. 2010. Echinoderm (Echinodermata) diversity in the Pacific coast of Central America. Marine Biodiversity 40: 45 - 56. https: // doi. org / 10.1007 / s 12526 - 009 - 0032 - 5" type="journal article" year="2010">
|
||
Alvarado
|
||
<emphasis id="B9358608532FFFE0FE2D9C92FE7BC729" box="[386,437,752,778]" italics="true" pageId="13" pageNumber="24">et al</emphasis>
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||
. 2010: 48
|
||
</bibRefCitation>
|
||
</treatmentCitation>
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||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A532FFFE0FF129D71FAE9C772" blockId="13.[189,1399,467,849]" pageId="13" pageNumber="24">
|
||
<treatmentCitationGroup id="AB517D34532FFFE0FF129D71FAE9C772" pageId="13" pageNumber="24">
|
||
<taxonomicName id="4C412199532FFFE0FF129D71FE2DC70E" authorityName="Nielsen" authorityYear="1932" box="[189,483,787,813]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophiocryptus" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="24" phylum="Echinodermata" rank="species" species="granulosus">
|
||
<emphasis id="B9358608532FFFE0FF129D71FE2DC70E" box="[189,483,787,813]" italics="true" pageId="13" pageNumber="24">Ophiocryptus granulosus</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AE07C0B532FFFE0FE549D71FCA3C70E" author="Honey-Escandon M. & Solis-Marin F. A. & Laguarda-Figueras A." box="[507,877,787,814]" page="63" pageId="13" pageNumber="24" year="2008">
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||
<bibRefCitation id="EFD027EB532FFFE0FE549D71FCA3C70E" author="Honey-Escandon M. & Solis-Marin F. A. & Laguarda-Figueras A." box="[507,877,787,814]" pageId="13" pageNumber="24" pagination="57 - 73" refId="ref16471" refString="Honey-Escandon M., Solis-Marin F. A. & Laguarda-Figueras A. 2008. Equinodermos (Echinodermata) del Pacifico Mexicano. Revista de Biologia Tropical 56 (3): 57 - 73. https: // doi. org / 10.15517 / RBT. V 56 I 3.27079" type="journal article" year="2008">
|
||
Honey-Escandón
|
||
<emphasis id="B9358608532FFFE0FD669D76FD37C70E" box="[713,761,787,813]" italics="true" pageId="13" pageNumber="24">et al</emphasis>
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||
. 2008: 63
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||
</bibRefCitation>
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||
</treatmentCitation>
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||
. —
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||
<bibRefCitation id="EFD027EB532FFFE0FC399D71FB93C70E" author="Cortes J." box="[918,1117,787,813]" pageId="13" pageNumber="24" pagination="131 - 185" refId="ref15712" refString="Cortes J. 2012. Marine biodiversity of an Eastern Tropical Pacific oceanic island, Isla del Coco, Costa Rica. Revista de Biologia Tropical 60 (3): 131 - 185. https: // doi. org / 10.15517 / RBT. V 60 I 3.28356" type="journal article" year="2012">Cortés 2012:173</bibRefCitation>
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. —
|
||
<treatmentCitation id="0AE07C0B532FFFE0FB279D71FDCEC772" author="Granja-Fernandez M. R. & Lopez-Perez R. A." page="363" pageId="13" pageNumber="24" year="2012">
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<bibRefCitation id="EFD027EB532FFFE0FB279D71FDCEC772" author="Granja-Fernandez M. R. & Lopez-Perez R. A." pageId="13" pageNumber="24" pagination="357 - 370" refId="ref15806" refString="Granja-Fernandez M. R. & Lopez-Perez R. A. 2012. Biodiversidad de ofiuroideos (Echinodermata: Ophiuroidea) de Oaxaca y Chiapas. In: Sanchez A. J., Chiappa-Carrara X. & Brito-Perez R. (eds) Recursos Costeros del Sureste: Tendencias actuales en Investigacion y Estado del Arte: 357 - 370. CONCITEY, FOMIX, SIIDETEY, UNAM, CONACYT, ECOSUR, PROMEP, RECORECOS, UNACAR, UJAT, Merida." type="book chapter" year="2012">Granja-Fernández & López-Pérez 2012: 363</bibRefCitation>
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. —
|
||
<treatmentCitation id="0AE07C0B532FFFE0FD809D55FC6AC772" author="Bastida-Zavala J. R. & Garcia-Madrigal M. S. & Rosas-Alquicira E. F. & Lopez-Perez R. A. & Benitez-Villalobos F. & Meraz-Hernando J. F. & Torres-Huerta A. M. & Montoya-Marquez A. & Barrientos-Lujan N. A." box="[559,932,823,849]" page="367" pageId="13" pageNumber="24" year="2013">
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<bibRefCitation id="EFD027EB532FFFE0FD809D55FC6AC772" author="Bastida-Zavala J. R. & Garcia-Madrigal M. S. & Rosas-Alquicira E. F. & Lopez-Perez R. A. & Benitez-Villalobos F. & Meraz-Hernando J. F. & Torres-Huerta A. M. & Montoya-Marquez A. & Barrientos-Lujan N. A." box="[559,932,823,849]" pageId="13" pageNumber="24" pagination="329 - 390" refId="ref14922" refString="Bastida-Zavala J. R., Garcia-Madrigal M. S., Rosas-Alquicira E. F., Lopez-Perez R. A., Benitez-Villalobos F., Meraz-Hernando J. F., Torres-Huerta A. M., Montoya-Marquez A. & Barrientos-Lujan N. A. 2013. Marine and coastal biodiversity of Oaxaca, Mexico. Check List 9 (2): 329 - 390. https: // doi. org / 10.15560 / 9.2.329" type="journal article" year="2013">
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Bastida-Zavala
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<emphasis id="B9358608532FFFE0FD479D55FCD2C772" box="[744,796,823,849]" italics="true" pageId="13" pageNumber="24">et al</emphasis>
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. 2013: 367
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</bibRefCitation>
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||
. —
|
||
<treatmentCitation id="0AE07C0B532FFFE0FC7B9D55FAEDC772" author="Solis-Marin F. A. & Alvarado J. J. & Abreu-Perez M. & Aguilera O. & Alio J. & Bacallado-Aranega J. J. & Williams S. M." box="[980,1315,822,849]" page="569" pageId="13" pageNumber="24" year="2013">
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<bibRefCitation id="EFD027EB532FFFE0FC7B9D55FAEDC772" author="Solis-Marin F. A. & Alvarado J. J. & Abreu-Perez M. & Aguilera O. & Alio J. & Bacallado-Aranega J. J. & Williams S. M." box="[980,1315,822,849]" pageId="13" pageNumber="24" pagination="543 - 658" refId="ref17532" refString="Solis-Marin F. A., Alvarado J. J., Abreu-Perez M., Aguilera O., Alio J., Bacallado-Aranega J. J. & Williams S. M. 2013. Appendix. In: Alvarado-Barrientos J. J. & Solis-Marin F. A. (eds) Echinoderm Research and Diversity in Latin America: 543 - 658. Springer-Verlag, Berlin. https: // doi. org / 10.1007 / 978 - 3 - 642 - 20051 - 9" type="book chapter" year="2013">
|
||
Solís-Marín
|
||
<emphasis id="B9358608532FFFE0FBC89D55FB55C772" box="[1127,1179,823,849]" italics="true" pageId="13" pageNumber="24">et al</emphasis>
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||
. 2013: 569
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35B0991532FFFE0FF129D1AFC79C02B" pageId="13" pageNumber="24" type="diagnosis">
|
||
<paragraph id="8BFE5A1A532FFFE0FF129D1AFEF8C7B1" blockId="13.[189,310,888,914]" box="[189,310,888,914]" pageId="13" pageNumber="24">
|
||
<heading id="D0B6ED76532FFFE0FF129D1AFEF8C7B1" bold="true" box="[189,310,888,914]" fontSize="11" level="3" pageId="13" pageNumber="24" reason="3">
|
||
<emphasis id="B9358608532FFFE0FF129D1AFEF8C7B1" bold="true" box="[189,310,888,914]" pageId="13" pageNumber="24">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A532FFFE0FF129DC5FC79C02B" blockId="13.[189,1398,935,1033]" pageId="13" pageNumber="24">Radial shields naked, oval, small, and widely separated. Adoral shields covered by granules. Up to 12 arm spines, short and with a blunt tip. In vivo coloration: disc green, brown or grey mottled with red, orange or yellow; arms with transverse bands, lighter and darker.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35B0991532FFFE0FF129A52FDD6C121" pageId="13" pageNumber="24" type="materials_examined">
|
||
<paragraph id="8BFE5A1A532FFFE0FF129A52FE64C069" blockId="13.[189,426,1072,1098]" box="[189,426,1072,1098]" pageId="13" pageNumber="24">
|
||
<heading id="D0B6ED76532FFFE0FF129A52FE64C069" bold="true" box="[189,426,1072,1098]" fontSize="11" level="3" pageId="13" pageNumber="24" reason="3">
|
||
<emphasis id="B9358608532FFFE0FF129A52FE64C069" bold="true" box="[189,426,1072,1098]" pageId="13" pageNumber="24">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A532FFFE0FF459A3DFE92C05A" blockId="13.[189,737,1119,1181]" box="[234,348,1119,1145]" pageId="13" pageNumber="24">
|
||
<emphasis id="B9358608532FFFE0FF459A3DFE92C05A" bold="true" box="[234,348,1119,1145]" pageId="13" pageNumber="24">
|
||
<typeStatus id="54FAE4B8532FFFE0FF459A3DFE92C05A" box="[234,348,1119,1145]" pageId="13" pageNumber="24" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A532FFFE0FF129AE1FD2FC0BE" blockId="13.[189,737,1119,1181]" box="[189,737,1155,1181]" pageId="13" pageNumber="24">
|
||
<materialsCitation id="3B295047532FFFE0FF129AE1FD2FC0BE" ID-GBIF-Occurrence="2995182322" box="[189,737,1155,1181]" collectionCode="NHMD" country="Panama" location="Panama" pageId="13" pageNumber="24" specimenCode="NHMD-107679" specimenCount="1" stateProvince="Panama" typeStatus="holotype">
|
||
<collectingCountry id="F3561A8A532FFFE0FF129AE1FEF6C0BE" box="[189,312,1155,1181]" name="Panama" pageId="13" pageNumber="24">PANAMA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D479193532FFFE0FEFE9AE1FE68C0BE" box="[337,422,1155,1181]" pageId="13" pageNumber="24" type="generic">1 spec.</specimenCount>
|
||
;
|
||
<collectingRegion id="498594F8532FFFE0FE1D9AE1FDDDC0BE" box="[434,531,1155,1181]" country="Panama" name="Panama" pageId="13" pageNumber="24">Panama</collectingRegion>
|
||
;
|
||
<specimenCode id="DBE7F261532FFFE0FDB19AE1FD13C0BE" box="[542,733,1155,1181]" pageId="13" pageNumber="24">NHMD-107679</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A532FFFE0FF459AA6FE68C0FD" blockId="13.[189,536,1220,1282]" box="[234,422,1220,1246]" pageId="13" pageNumber="24">
|
||
<emphasis id="B9358608532FFFE0FF459AA6FE68C0FD" bold="true" box="[234,422,1220,1246]" pageId="13" pageNumber="24">Other material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A532FFFE0FF129A8AFDD6C121" blockId="13.[189,536,1220,1282]" box="[189,536,1256,1282]" pageId="13" pageNumber="24">See: Supplementary material.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35B0991532FFFE2FF129B48FDBEC312" lastPageId="15" lastPageNumber="26" pageId="13" pageNumber="24" type="description">
|
||
<paragraph id="8BFE5A1A532FFFE0FF129B48FE12C167" blockId="13.[189,476,1322,1348]" box="[189,476,1322,1348]" pageId="13" pageNumber="24">
|
||
<heading id="D0B6ED76532FFFE0FF129B48FE12C167" bold="true" box="[189,476,1322,1348]" fontSize="11" level="3" pageId="13" pageNumber="24" reason="3">
|
||
<emphasis id="B9358608532FFFE0FF129B48FE12C167" bold="true" box="[189,476,1322,1348]" pageId="13" pageNumber="24">
|
||
<typeStatus id="54FAE4B8532FFFE0FF129B48FEE1C167" box="[189,303,1322,1348]" pageId="13" pageNumber="24" type="holotype">Holotype</typeStatus>
|
||
redescription
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A532FFFE0FF129B3BFA9DC206" blockId="13.[189,1399,1369,1573]" pageId="13" pageNumber="24">
|
||
DD =
|
||
<quantity id="4CB9F7FF532FFFE0FEA19B3BFEB0C150" box="[270,382,1369,1395]" metricMagnitude="-2" metricUnit="m" metricValue="2.19" pageId="13" pageNumber="24" unit="mm" value="21.9">21.9 mm</quantity>
|
||
; 5 arms, AL =
|
||
<quantity id="4CB9F7FF532FFFE0FD909B3BFD60C150" box="[575,686,1369,1395]" metricMagnitude="-2" metricUnit="m" metricValue="9.19" pageId="13" pageNumber="24" unit="mm" value="91.9">91.9 mm</quantity>
|
||
. Disc flat and pentagonal. Dorsal disc densely covered by rounded and small granules, granule density 61
|
||
<superScript id="7C34F752532FFFE0FD589B1FFCFDC1A9" attach="left" box="[759,819,1404,1431]" fontSize="6" pageId="13" pageNumber="24">mm-2</superScript>
|
||
; rubbed off in some areas. Radial shields naked, small, oval, and separated (
|
||
<figureCitation id="137A469F532FFFE0FE509BC2FD94C198" box="[511,602,1440,1467]" captionStart="Fig" captionStartId="9.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@9.[189,1398,291,1855]" captionTargetPageId="9" captionText="Fig. 5. A–D. Ophioderma hendleri sp. nov. A–B. In vivo coloration in natural habitat. C–D. In vivo coloration of collected specimens. – E–J. Ophioderma panamense Lütken, 1859, holotype (NHMD- 107679, DD = 21.9 mm). E. Dorsal disc. F. Ventral disc. G. Mouth. H. Dorsal arm. I. Ventral arm. J. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326625" httpUri="https://zenodo.org/record/4326625/files/figure.png" pageId="13" pageNumber="24">Fig. 5E</figureCitation>
|
||
). Ventral interradii covered with granules similar to those on dorsal side of disc (
|
||
<figureCitation id="137A469F532FFFE0FEF99BA1FE7FC1FD" box="[342,433,1475,1502]" captionStart="Fig" captionStartId="9.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@9.[189,1398,291,1855]" captionTargetPageId="9" captionText="Fig. 5. A–D. Ophioderma hendleri sp. nov. A–B. In vivo coloration in natural habitat. C–D. In vivo coloration of collected specimens. – E–J. Ophioderma panamense Lütken, 1859, holotype (NHMD- 107679, DD = 21.9 mm). E. Dorsal disc. F. Ventral disc. G. Mouth. H. Dorsal arm. I. Ventral arm. J. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326625" httpUri="https://zenodo.org/record/4326625/files/figure.png" pageId="13" pageNumber="24">Fig. 5F</figureCitation>
|
||
). Four genital slits on each interradius; proximal genital slits rounded and shorter than distal slits, in contact with distal part of oral shield and with 1
|
||
<superScript id="7C34F752532FFFE0FC7C9B85FC10C1D6" attach="left" box="[979,990,1511,1525]" fontSize="6" pageId="13" pageNumber="24">st</superScript>
|
||
LAP; distal genital slits elongated and surrounded by granules and numerous and imbricated scales next to lateral arm plates (
|
||
<figureCitation id="137A469F532FFFE0FB439869FA88C206" box="[1260,1350,1546,1573]" captionStart="Fig" captionStartId="9.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@9.[189,1398,291,1855]" captionTargetPageId="9" captionText="Fig. 5. A–D. Ophioderma hendleri sp. nov. A–B. In vivo coloration in natural habitat. C–D. In vivo coloration of collected specimens. – E–J. Ophioderma panamense Lütken, 1859, holotype (NHMD- 107679, DD = 21.9 mm). E. Dorsal disc. F. Ventral disc. G. Mouth. H. Dorsal arm. I. Ventral arm. J. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326625" httpUri="https://zenodo.org/record/4326625/files/figure.png" pageId="13" pageNumber="24">Fig. 5F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A532FFFE0FF12982EFC50C3A7" blockId="13.[189,1399,1612,1924]" pageId="13" pageNumber="24">
|
||
Oral shields slightly broader than long, rounded triangular, with convex obtuse proximal angle, straight distal edge, obtuse lateral angles. Madreporite with a small, circular, and shallow depression located in distal part and larger than other oral shields. Adoral shields covered by granules, which are larger than those on rest of body. Jaws bear 7–8 papillae at each side: LyOs small, 1.5× as long as broad, angled upwards; AdShSp largest, rounded; 2°AdShSp a little larger and similar in shape to LOPa; LOPa 3–4 conical and pointed, separated; IPa similar and separated from LOPa; TPa 1–2 at jaw tips, robust and pointed, larger and more robust than LOPa, widely separated. vT similar in shape but slightly larger than TPa. OPRSp 1, large and pointed at each side, visible within buccal cavity. Granules covering oral plates, larger than those on dorsal disc and interradii (
|
||
<figureCitation id="137A469F532FFFE0FC9D990BFC5FC3A7" box="[818,913,1897,1924]" captionStart="Fig" captionStartId="9.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@9.[189,1398,291,1855]" captionTargetPageId="9" captionText="Fig. 5. A–D. Ophioderma hendleri sp. nov. A–B. In vivo coloration in natural habitat. C–D. In vivo coloration of collected specimens. – E–J. Ophioderma panamense Lütken, 1859, holotype (NHMD- 107679, DD = 21.9 mm). E. Dorsal disc. F. Ventral disc. G. Mouth. H. Dorsal arm. I. Ventral arm. J. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326625" httpUri="https://zenodo.org/record/4326625/files/figure.png" pageId="13" pageNumber="24">Fig. 5G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A532FFFE2FF1299C9FDE9C61D" blockId="13.[189,1399,1962,2025]" lastBlockId="15.[189,1399,265,574]" lastPageId="15" lastPageNumber="26" pageId="13" pageNumber="24">
|
||
Dorsal base of arms covered by granules and with approximately 15 scales laterally at dorsal arm plates. Dorsal arm plates 3× broader than long, overlapping and with an irregular to straight proximal and distal margin, with rounded lateral edges; a few dorsal arm plates can be fragmented in 2 pieces (
|
||
<figureCitation id="137A469F532DFFE2FF699F4EFEEBC564" box="[198,293,300,327]" captionStart="Fig" captionStartId="9.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@9.[189,1398,291,1855]" captionTargetPageId="9" captionText="Fig. 5. A–D. Ophioderma hendleri sp. nov. A–B. In vivo coloration in natural habitat. C–D. In vivo coloration of collected specimens. – E–J. Ophioderma panamense Lütken, 1859, holotype (NHMD- 107679, DD = 21.9 mm). E. Dorsal disc. F. Ventral disc. G. Mouth. H. Dorsal arm. I. Ventral arm. J. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326625" httpUri="https://zenodo.org/record/4326625/files/figure.png" pageId="15" pageNumber="26">Fig. 5H</figureCitation>
|
||
). First ventral arm plate small, broader than long; with a small indentation distally (
|
||
<figureCitation id="137A469F532DFFE2FAA59F4EFAA7C564" box="[1290,1385,300,327]" captionStart="Fig" captionStartId="9.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@9.[189,1398,291,1855]" captionTargetPageId="9" captionText="Fig. 5. A–D. Ophioderma hendleri sp. nov. A–B. In vivo coloration in natural habitat. C–D. In vivo coloration of collected specimens. – E–J. Ophioderma panamense Lütken, 1859, holotype (NHMD- 107679, DD = 21.9 mm). E. Dorsal disc. F. Ventral disc. G. Mouth. H. Dorsal arm. I. Ventral arm. J. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326625" httpUri="https://zenodo.org/record/4326625/files/figure.png" pageId="15" pageNumber="26">Fig. 5G</figureCitation>
|
||
). Subsequent ventral arm plates quadrangular, slightly broader than long; distal edge slightly concave (
|
||
<figureCitation id="137A469F532DFFE2FF699F11FED7C5AD" box="[198,281,371,398]" captionStart="Fig" captionStartId="9.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@9.[189,1398,291,1855]" captionTargetPageId="9" captionText="Fig. 5. A–D. Ophioderma hendleri sp. nov. A–B. In vivo coloration in natural habitat. C–D. In vivo coloration of collected specimens. – E–J. Ophioderma panamense Lütken, 1859, holotype (NHMD- 107679, DD = 21.9 mm). E. Dorsal disc. F. Ventral disc. G. Mouth. H. Dorsal arm. I. Ventral arm. J. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326625" httpUri="https://zenodo.org/record/4326625/files/figure.png" pageId="15" pageNumber="26">Fig. 5I</figureCitation>
|
||
). Paired and rounded pores between 1
|
||
<superScript id="7C34F752532DFFE2FD719F11FD27C5A2" attach="left" box="[734,745,371,385]" fontSize="6" pageId="15" pageNumber="26">st</superScript>
|
||
and 2
|
||
<superScript id="7C34F752532DFFE2FC9C9F10FC8AC5A3" attach="none" box="[819,836,370,384]" fontSize="6" pageId="15" pageNumber="26">nd</superScript>
|
||
, and 2
|
||
<superScript id="7C34F752532DFFE2FC3A9F10FC68C5A3" attach="left" box="[917,934,370,384]" fontSize="6" pageId="15" pageNumber="26">nd</superScript>
|
||
and 3
|
||
<superScript id="7C34F752532DFFE2FC409F10FC33C5A3" attach="left" box="[1007,1021,370,384]" fontSize="6" pageId="15" pageNumber="26">rd</superScript>
|
||
ventral arm plates (
|
||
<figureCitation id="137A469F532DFFE2FB449F11FAE1C5AD" box="[1259,1327,371,398]" captionStart="Fig" captionStartId="9.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@9.[189,1398,291,1855]" captionTargetPageId="9" captionText="Fig. 5. A–D. Ophioderma hendleri sp. nov. A–B. In vivo coloration in natural habitat. C–D. In vivo coloration of collected specimens. – E–J. Ophioderma panamense Lütken, 1859, holotype (NHMD- 107679, DD = 21.9 mm). E. Dorsal disc. F. Ventral disc. G. Mouth. H. Dorsal arm. I. Ventral arm. J. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326625" httpUri="https://zenodo.org/record/4326625/files/figure.png" pageId="15" pageNumber="26">Fig. 5</figureCitation>
|
||
F–G). LAP with up to 8–9 arm spines. Arm spines with blunt tip and robust; length approximately ½ of size of LAP. Dorsalmost arm spine shortest and narrowest; ventralmost arm spine longest and slightly more robust, in contact with tentacle scale of succeeding joint (
|
||
<figureCitation id="137A469F532DFFE2FCF39FBFFC61C5DB" box="[860,943,477,504]" captionStart="Fig" captionStartId="9.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@9.[189,1398,291,1855]" captionTargetPageId="9" captionText="Fig. 5. A–D. Ophioderma hendleri sp. nov. A–B. In vivo coloration in natural habitat. C–D. In vivo coloration of collected specimens. – E–J. Ophioderma panamense Lütken, 1859, holotype (NHMD- 107679, DD = 21.9 mm). E. Dorsal disc. F. Ventral disc. G. Mouth. H. Dorsal arm. I. Ventral arm. J. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326625" httpUri="https://zenodo.org/record/4326625/files/figure.png" pageId="15" pageNumber="26">Fig. 5I</figureCitation>
|
||
). Two tentacle scales; adradial tentacle scale ovoid, elongated, approximately half length of ventral arm plate; abradial tentacle scale slightly shorter, subtriangular (
|
||
<figureCitation id="137A469F532DFFE2FE689C41FDD7C61D" box="[455,537,547,574]" captionStart="Fig" captionStartId="9.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@9.[189,1398,291,1855]" captionTargetPageId="9" captionText="Fig. 5. A–D. Ophioderma hendleri sp. nov. A–B. In vivo coloration in natural habitat. C–D. In vivo coloration of collected specimens. – E–J. Ophioderma panamense Lütken, 1859, holotype (NHMD- 107679, DD = 21.9 mm). E. Dorsal disc. F. Ventral disc. G. Mouth. H. Dorsal arm. I. Ventral arm. J. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326625" httpUri="https://zenodo.org/record/4326625/files/figure.png" pageId="15" pageNumber="26">Fig. 5I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF3E0A92532CFFE3FF12989EFD6BC3C9" ID-DOI="http://doi.org/10.5281/zenodo.4326627" ID-Zenodo-Dep="4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="14" pageNumber="25" startId="14.[189,232,1788,1814]" targetBox="[244,1344,265,1751]" targetPageId="14">
|
||
<paragraph id="8BFE5A1A532CFFE3FF12989EFD6BC3C9" blockId="14.[189,1399,1788,2027]" pageId="14" pageNumber="25">
|
||
<emphasis id="B9358608532CFFE3FF12989EFEC3C334" bold="true" box="[189,269,1788,1815]" pageId="14" pageNumber="25">Fig. 6.</emphasis>
|
||
SEM images of skeletal elements of
|
||
<taxonomicName id="4C412199532CFFE3FD76989FFC32C334" authorityName="Lutken" authorityYear="1859" box="[729,1020,1789,1815]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="25" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9358608532CFFE3FD76989FFC32C334" box="[729,1020,1789,1815]" italics="true" pageId="14" pageNumber="25">Ophioderma panamense</emphasis>
|
||
</taxonomicName>
|
||
(ICML-UNAM 3.18.60, DD = 11.1 mm).
|
||
<emphasis id="B9358608532CFFE3FEEE9942FE98C319" bold="true" box="[321,342,1824,1850]" pageId="14" pageNumber="25">A</emphasis>
|
||
. Radial shield (external view), the pores marked with arrows.
|
||
<emphasis id="B9358608532CFFE3FBE99942FB94C319" bold="true" box="[1094,1114,1824,1850]" pageId="14" pageNumber="25">B</emphasis>
|
||
. Dorsal arm plate, with seven spurs.
|
||
<emphasis id="B9358608532CFFE3FEF69921FEA0C37E" bold="true" box="[345,366,1859,1885]" pageId="14" pageNumber="25">C</emphasis>
|
||
. Ventral arm plate with one spur.
|
||
<emphasis id="B9358608532CFFE3FCA29921FCECC37E" bold="true" box="[781,802,1859,1885]" pageId="14" pageNumber="25">D</emphasis>
|
||
. Lateral arm plate, external face, with two spurs.
|
||
<emphasis id="B9358608532CFFE3FF129904FF1FC3A3" bold="true" box="[189,209,1894,1920]" pageId="14" pageNumber="25">E</emphasis>
|
||
. Lateral arm plate, lateral face, with spine articulations.
|
||
<emphasis id="B9358608532CFFE3FCC09904FC4FC3A3" bold="true" box="[879,897,1894,1920]" pageId="14" pageNumber="25">F</emphasis>
|
||
. Lateral arm plate, internal face.
|
||
<emphasis id="B9358608532CFFE3FAA29904FAEAC3A3" bold="true" box="[1293,1316,1894,1920]" pageId="14" pageNumber="25">G</emphasis>
|
||
. Outer surface of the LAP (D) with fine mesh and rounded knobs.
|
||
<emphasis id="B9358608532CFFE3FCC499E8FC4CC387" bold="true" box="[875,898,1930,1956]" pageId="14" pageNumber="25">H</emphasis>
|
||
. Spine articulations with a weak sigmoidal fold and tilted lobes.
|
||
<emphasis id="B9358608532CFFE3FE6E99CFFE02C3E4" bold="true" box="[449,460,1965,1991]" pageId="14" pageNumber="25">I</emphasis>
|
||
. Arm vertebra, proximal face.
|
||
<emphasis id="B9358608532CFFE3FCEB99CFFC9DC3E4" bold="true" box="[836,851,1965,1991]" pageId="14" pageNumber="25">J</emphasis>
|
||
. Arm vertebra, distal face.
|
||
<emphasis id="B9358608532CFFE3FB0F99CFFB79C3E4" bold="true" box="[1184,1207,1965,1991]" pageId="14" pageNumber="25">K</emphasis>
|
||
. Tentacle scale.
|
||
<emphasis id="B9358608532CFFE3FF1299B2FF1FC3C9" bold="true" box="[189,209,2000,2026]" pageId="14" pageNumber="25">L</emphasis>
|
||
. Arm spine.
|
||
<emphasis id="B9358608532CFFE3FEC899B2FE4DC3C9" bold="true" box="[359,387,2000,2026]" pageId="14" pageNumber="25">M</emphasis>
|
||
. Dental plate fragments.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFE5A1A532DFFE2FF129C07FCE9C6A3" blockId="15.[189,807,613,640]" box="[189,807,613,640]" pageId="15" pageNumber="26">
|
||
Color pattern beige in ethanol, likely faded (
|
||
<figureCitation id="137A469F532DFFE2FD6A9C07FCD4C6A3" box="[709,794,613,640]" captionStart="Fig" captionStartId="9.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@9.[189,1398,291,1855]" captionTargetPageId="9" captionText="Fig. 5. A–D. Ophioderma hendleri sp. nov. A–B. In vivo coloration in natural habitat. C–D. In vivo coloration of collected specimens. – E–J. Ophioderma panamense Lütken, 1859, holotype (NHMD- 107679, DD = 21.9 mm). E. Dorsal disc. F. Ventral disc. G. Mouth. H. Dorsal arm. I. Ventral arm. J. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326625" httpUri="https://zenodo.org/record/4326625/files/figure.png" pageId="15" pageNumber="26">Fig. 5J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A532DFFE2FF459CC4FE30C6E3" blockId="15.[189,1399,678,1553]" box="[234,510,678,704]" pageId="15" pageNumber="26">
|
||
<emphasis id="B9358608532DFFE2FF459CC4FE30C6E3" bold="true" box="[234,510,678,704]" pageId="15" pageNumber="26">Disarticulated ossicles</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A532DFFE2FF129CA8FD66C232" blockId="15.[189,1399,678,1553]" pageId="15" pageNumber="26">
|
||
Specimen analyzed:
|
||
<specimenCount id="9D479193532DFFE2FE199CA8FDC3C6C7" box="[438,525,714,740]" pageId="15" pageNumber="26" type="generic">1 spec.</specimenCount>
|
||
, ICML-UNAM 3.18.60 (DD =
|
||
<quantity id="4CB9F7FF532DFFE2FC219CA8FC35C6C7" box="[910,1019,714,740]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="15" pageNumber="26" unit="mm" value="11.1">11.1 mm</quantity>
|
||
, AL =
|
||
<quantity id="4CB9F7FF532DFFE2FBFC9CA8FB0AC6C7" box="[1107,1220,714,740]" metricMagnitude="-2" metricUnit="m" metricValue="4.4399999999999995" pageId="15" pageNumber="26" unit="mm" value="44.4">44.4 mm</quantity>
|
||
). Radial shield (external view) irregularly rounded, with an indented proximal margin, an incised abradial edge with a projection, an indented distal edge, and convex adradial margin, it is bordered by 7 pores on median to proximal margin (which are covered/overlapped by disc scales and granules in intact animal) (
|
||
<figureCitation id="137A469F532DFFE2FAA49D56FAA4C76C" box="[1291,1386,820,847]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6A</figureCitation>
|
||
). Only exposed surface is domed center portion of radial shield. Outer surface of stereom is an open mesh of pores and knobs (
|
||
<figureCitation id="137A469F532DFFE2FE1A9D18FDD8C7B6" box="[437,534,890,917]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6A</figureCitation>
|
||
). Dorsal arm plate somewhat rectangular, 3× as wide as long; proximal margin convex to straight, and distal margin straight, 7 spurs on proximal portion of external surface (
|
||
<figureCitation id="137A469F532DFFE2FF699DA3FEEBC7FF" box="[198,293,961,988]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6B</figureCitation>
|
||
). Ventral arm plate as long as wide, quadrangular with proximal side truncated and with a pointed-shape spur, lateral sides concave forming border of a small tentacle pore, and distally indented (
|
||
<figureCitation id="137A469F532DFFE2FF699A6AFEEFC000" box="[198,289,1032,1059]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6C</figureCitation>
|
||
). LAP D-shaped, 3× as high as wide, with 8 spine articulations sunken in notches of distal edge (
|
||
<figureCitation id="137A469F532DFFE2FF699A49FEC2C065" box="[198,268,1067,1094]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6</figureCitation>
|
||
D–E). Ventral portion of LAP projecting ventro-proximalwards and ventro-distal tip projecting ventralwards (
|
||
<figureCitation id="137A469F532DFFE2FEC49A2CFE01C04A" box="[363,463,1102,1129]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6D</figureCitation>
|
||
). Proximal edge of LAP with 2 prominent and elongated spurs, which are protruding and modify central-proximal edge (
|
||
<figureCitation id="137A469F532DFFE2FD4B9A10FC8DC0AE" box="[740,835,1138,1165]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6D</figureCitation>
|
||
); between spurs and across remaining proximal margin a discernible band of different stereom structure is present (
|
||
<figureCitation id="137A469F532DFFE2FC6E9AF7FBD0C093" box="[961,1054,1173,1200]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6D</figureCitation>
|
||
). Outer surface finely meshed with relatively large rounded stereom knobs (
|
||
<figureCitation id="137A469F532DFFE2FD7F9ADAFCE0C0F0" box="[720,814,1208,1235]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6G</figureCitation>
|
||
). Inner surface of LAP with a continuous ridge on proximal edge, and on ventro-distal margin with 2 spurs matching those on external surface (
|
||
<figureCitation id="137A469F532DFFE2FAA09ABEFAA7C0D4" box="[1295,1385,1244,1271]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6F</figureCitation>
|
||
), and 4 pores in center. Spine articulations ventralwards increasing in size (
|
||
<figureCitation id="137A469F532DFFE2FB8A9A9DFB4CC139" box="[1061,1154,1279,1306]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6E</figureCitation>
|
||
). Lobes with a weak sigmoidal fold, tilted, and curved (
|
||
<figureCitation id="137A469F532DFFE2FDF69B40FD77C11E" box="[601,697,1314,1341]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6H</figureCitation>
|
||
). Ventral lobe smaller than dorsal lobe (
|
||
<figureCitation id="137A469F532DFFE2FB3A9B40FB3BC11E" box="[1173,1269,1314,1341]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6H</figureCitation>
|
||
). Proximal vertebrae (
|
||
<figureCitation id="137A469F532DFFE2FE969B24FE47C142" box="[313,393,1350,1377]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6I</figureCitation>
|
||
) almost oval, as wide as long, with large aboral muscle flange and smaller oral muscle flange. Distal face of vertebrae with large muscle flanges, with typical zygospondylous articulation (
|
||
<figureCitation id="137A469F532DFFE2FF699BEEFED5C184" box="[198,283,1420,1447]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6J</figureCitation>
|
||
). Tentacle scale longer than wide, with a scale-like surface (
|
||
<figureCitation id="137A469F532DFFE2FC479BEEFB86C184" box="[1000,1096,1420,1447]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6K</figureCitation>
|
||
). Spines with a scale-like surface, a blunt tip, and laterally flattened (
|
||
<figureCitation id="137A469F532DFFE2FD6A9BD2FCEDC1E9" box="[709,803,1455,1482]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6L</figureCitation>
|
||
). Dental plate consists of several pieces, 1
|
||
<superScript id="7C34F752532DFFE2FA8C9BD2FAE0C19D" attach="left" box="[1315,1326,1456,1470]" fontSize="6" pageId="15" pageNumber="26">st</superScript>
|
||
piece bears 2 TPa and a single wide tooth, while rest of pieces with a single tooth socket per piece (
|
||
<quantity id="4CB9F7FF532DFFE2FB529BB1FAE5C1CE" box="[1277,1323,1491,1517]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="15" pageNumber="26" unit="in" value="4.0">4 in</quantity>
|
||
total), none of sockets penetrate plate (
|
||
<figureCitation id="137A469F532DFFE2FD989B94FD55C232" box="[567,667,1526,1553]" captionStart="Fig" captionStartId="14.[189,232,1788,1814]" captionTargetBox="[244,1344,265,1751]" captionTargetId="figure-138@14.[244,1344,265,1751]" captionTargetPageId="14" captionText="Fig. 6. SEM images of skeletal elements of Ophioderma panamense (ICML-UNAM 3.18.60, DD = 11.1 mm). A. Radial shield (external view), the pores marked with arrows. B. Dorsal arm plate, with seven spurs. C. Ventral arm plate with one spur. D. Lateral arm plate, external face, with two spurs. E. Lateral arm plate, lateral face, with spine articulations. F. Lateral arm plate, internal face. G. Outer surface of the LAP (D) with fine mesh and rounded knobs. H. Spine articulations with a weak sigmoidal fold and tilted lobes. I. Arm vertebra, proximal face. J. Arm vertebra, distal face. K. Tentacle scale. L. Arm spine. M. Dental plate fragments." figureDoi="http://doi.org/10.5281/zenodo.4326627" httpUri="https://zenodo.org/record/4326627/files/figure.png" pageId="15" pageNumber="26">Fig. 6M</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A532DFFE2FF12985AFE21C271" blockId="15.[189,495,1592,1618]" box="[189,495,1592,1618]" pageId="15" pageNumber="26">
|
||
<heading id="D0B6ED76532DFFE2FF12985AFE21C271" bold="true" box="[189,495,1592,1618]" fontSize="11" level="3" pageId="15" pageNumber="26" reason="3">
|
||
<emphasis id="B9358608532DFFE2FF12985AFE21C271" bold="true" box="[189,495,1592,1618]" pageId="15" pageNumber="26">Habitat and distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A532DFFE2FF129805FDBEC312" blockId="15.[189,1399,1639,1842]" pageId="15" pageNumber="26">
|
||
<collectingCountry id="F3561A8A532DFFE2FF129805FF38C2A2" box="[189,246,1639,1665]" name="United States of America" pageId="15" pageNumber="26">USA</collectingCountry>
|
||
(
|
||
<collectingRegion id="498594F8532DFFE2FEA79805FE4BC2A1" box="[264,389,1639,1666]" country="United States of America" name="California" pageId="15" pageNumber="26">California</collectingRegion>
|
||
),
|
||
<collectingCountry id="F3561A8A532DFFE2FE339805FE38C2A2" box="[412,502,1639,1665]" name="Mexico" pageId="15" pageNumber="26">Mexico</collectingCountry>
|
||
(
|
||
<collectingRegion id="498594F8532DFFE2FDA89805FD0DC2A2" box="[519,707,1639,1665]" country="Mexico" name="Baja California" pageId="15" pageNumber="26">Baja California</collectingRegion>
|
||
, Gulf of California,
|
||
<collectingRegion id="498594F8532DFFE2FC149805FBD8C2A2" box="[955,1046,1639,1665]" country="Mexico" name="Nayarit" pageId="15" pageNumber="26">Nayarit</collectingRegion>
|
||
,
|
||
<collectingRegion id="498594F8532DFFE2FB8B9805FBB9C2A2" box="[1060,1143,1639,1665]" country="Mexico" name="Jalisco" pageId="15" pageNumber="26">Jalisco</collectingRegion>
|
||
,
|
||
<collectingRegion id="498594F8532DFFE2FB2A9805FB10C2A2" box="[1157,1246,1639,1665]" country="Mexico" name="Colima" pageId="15" pageNumber="26">Colima</collectingRegion>
|
||
,
|
||
<collectingRegion id="498594F8532DFFE2FB429805FABCC2A2" box="[1261,1394,1639,1665]" country="Mexico" name="Michoacan de Ocampo" pageId="15" pageNumber="26">Michoacán</collectingRegion>
|
||
,
|
||
<collectingRegion id="498594F8532DFFE2FF1298E8FEE4C287" box="[189,298,1674,1700]" country="Mexico" name="Guerrero" pageId="15" pageNumber="26">Guerrero</collectingRegion>
|
||
,
|
||
<collectingRegion id="498594F8532DFFE2FE9698E8FE5AC287" box="[313,404,1674,1700]" country="Mexico" name="Oaxaca" pageId="15" pageNumber="26">Oaxaca</collectingRegion>
|
||
,
|
||
<collectingRegion id="498594F8532DFFE2FE0C98E8FD9BC287" box="[419,597,1674,1700]" inferredBySuffix="true" pageId="15" pageNumber="26">Marías Islands</collectingRegion>
|
||
,
|
||
<collectingRegion id="498594F8532DFFE2FDCB98E8FCE5C287" box="[612,811,1674,1700]" inferredBySuffix="true" pageId="15" pageNumber="26">Marietas Islands</collectingRegion>
|
||
,
|
||
<collectingRegion id="498594F8532DFFE2FC9498E8FB8FC286" box="[827,1089,1674,1701]" inferredBySuffix="true" pageId="15" pageNumber="26">Revillagigedo Islands</collectingRegion>
|
||
),
|
||
<collectingCountry id="F3561A8A532DFFE2FBF698E8FB29C287" box="[1113,1255,1674,1700]" name="El Salvador" pageId="15" pageNumber="26">El Salvador</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3561A8A532DFFE2FB5898E8FABDC287" box="[1271,1395,1674,1700]" name="Nicaragua" pageId="15" pageNumber="26">Nicaragua</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3561A8A532DFFE2FF1298CCFEF3C2E4" box="[189,317,1709,1736]" name="Costa Rica" pageId="15" pageNumber="26">Costa Rica</collectingCountry>
|
||
(Caño Island, Cocos Island,
|
||
<collectingRegion id="498594F8532DFFE2FD3C98CFFC4AC2EB" box="[659,900,1709,1736]" inferredBySuffix="true" pageId="15" pageNumber="26">Murciélagos Islands</collectingRegion>
|
||
),
|
||
<collectingCountry id="F3561A8A532DFFE2FC3698CCFC37C2EB" box="[921,1017,1710,1736]" name="Panama" pageId="15" pageNumber="26">Panama</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3561A8A532DFFE2FBAA98CFFBB2C2E4" box="[1029,1148,1709,1735]" name="Colombia" pageId="15" pageNumber="26">Colombia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3561A8A532DFFE2FB2898CFFB27C2E4" box="[1159,1257,1709,1735]" name="Ecuador" pageId="15" pageNumber="26">Ecuador</collectingCountry>
|
||
(
|
||
<collectingRegion id="498594F8532DFFE2FB5698CFFEDAC2CF" country="Ecuador" name="Galapagos" pageId="15" pageNumber="26">Galapagos Islands</collectingRegion>
|
||
), and
|
||
<collectingCountry id="F3561A8A532DFFE2FEF798B3FE40C2C8" box="[344,398,1745,1771]" name="Peru" pageId="15" pageNumber="26">Peru</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFD027EB532DFFE2FE3498B3FD63C2C8" author="Solis-Marin F. A. & Alvarado J. J. & Abreu-Perez M. & Aguilera O. & Alio J. & Bacallado-Aranega J. J. & Williams S. M." box="[411,685,1745,1771]" pageId="15" pageNumber="26" pagination="543 - 658" refId="ref17532" refString="Solis-Marin F. A., Alvarado J. J., Abreu-Perez M., Aguilera O., Alio J., Bacallado-Aranega J. J. & Williams S. M. 2013. Appendix. In: Alvarado-Barrientos J. J. & Solis-Marin F. A. (eds) Echinoderm Research and Diversity in Latin America: 543 - 658. Springer-Verlag, Berlin. https: // doi. org / 10.1007 / 978 - 3 - 642 - 20051 - 9" type="book chapter" year="2013">
|
||
Solís-Marín
|
||
<emphasis id="B9358608532DFFE2FD8098B0FDACC2C8" box="[559,610,1745,1771]" italics="true" pageId="15" pageNumber="26">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD027EB532DFFE2FD1798B3FBDFC2C8" author="Granja-Fernandez R. & Herrero-Perezrul M. D. & Lopez-Perez R. A. & Hernandez L. & Rodriguez-Zaragoza F. A. & Jones R. W. & Pineda-Lopez R." box="[696,1041,1745,1771]" pageId="15" pageNumber="26" pagination="101 - 145" refId="ref15937" refString="Granja-Fernandez R., Herrero-Perezrul M. D., Lopez-Perez R. A., Hernandez L., Rodriguez-Zaragoza F. A., Jones R. W. & Pineda-Lopez R. 2014. Ophiuroidea (Echinodermata) from coral reefs in the Mexican Pacific. Zookeys 406: 101 - 145. https: // doi. org / 10.3897 / zookeys. 406.6306" type="journal article" year="2014">
|
||
Granja-Fernández
|
||
<emphasis id="B9358608532DFFE2FC3E98B0FC0AC2C8" box="[913,964,1745,1771]" italics="true" pageId="15" pageNumber="26">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
). From intertidal to
|
||
<quantity id="4CB9F7FF532DFFE2FB5A98B3FAE1C2C8" box="[1269,1327,1745,1771]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="26" unit="m" value="20.0">20 m</quantity>
|
||
depth (
|
||
<bibRefCitation id="EFD027EB532DFFE2FF6A9896FE2DC32D" author="Solis-Marin F. A. & Alvarado J. J. & Abreu-Perez M. & Aguilera O. & Alio J. & Bacallado-Aranega J. J. & Williams S. M." box="[197,483,1780,1807]" pageId="15" pageNumber="26" pagination="543 - 658" refId="ref17532" refString="Solis-Marin F. A., Alvarado J. J., Abreu-Perez M., Aguilera O., Alio J., Bacallado-Aranega J. J. & Williams S. M. 2013. Appendix. In: Alvarado-Barrientos J. J. & Solis-Marin F. A. (eds) Echinoderm Research and Diversity in Latin America: 543 - 658. Springer-Verlag, Berlin. https: // doi. org / 10.1007 / 978 - 3 - 642 - 20051 - 9" type="book chapter" year="2013">
|
||
Solís-Marín
|
||
<emphasis id="B9358608532DFFE2FEF39897FE5CC32D" box="[348,402,1780,1806]" italics="true" pageId="15" pageNumber="26">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
). Found under rocks and in rubble, coral, sand, algae, and tide pools (
|
||
<bibRefCitation id="EFD027EB532DFFE2FA829896FF35C312" author="Maluf L. Y." pageId="15" pageNumber="26" pagination="1 - 242" refId="ref17113" refString="Maluf L. Y. 1988. Composition and distribution of the central Eastern Pacific echinoderms. Natural History Museum of Los Angeles County, Technical Report 2: 1 - 242." type="book chapter" year="1988">Maluf 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD027EB532DFFE2FEA89975FDAAC312" author="Granja-Fernandez R. & Herrero-Perezrul M. D. & Lopez-Perez R. A. & Hernandez L. & Rodriguez-Zaragoza F. A. & Jones R. W. & Pineda-Lopez R." box="[263,612,1815,1842]" pageId="15" pageNumber="26" pagination="101 - 145" refId="ref15937" refString="Granja-Fernandez R., Herrero-Perezrul M. D., Lopez-Perez R. A., Hernandez L., Rodriguez-Zaragoza F. A., Jones R. W. & Pineda-Lopez R. 2014. Ophiuroidea (Echinodermata) from coral reefs in the Mexican Pacific. Zookeys 406: 101 - 145. https: // doi. org / 10.3897 / zookeys. 406.6306" type="journal article" year="2014">
|
||
Granja-Fernández
|
||
<emphasis id="B9358608532DFFE2FE4D997AFDD8C312" box="[482,534,1815,1841]" italics="true" pageId="15" pageNumber="26">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35B0991532DFFFDFF12993BFAB9C710" lastPageId="16" lastPageNumber="27" pageId="15" pageNumber="26" type="discussion">
|
||
<paragraph id="8BFE5A1A532DFFE2FF12993BFEE1C350" blockId="15.[189,303,1881,1907]" box="[189,303,1881,1907]" pageId="15" pageNumber="26">
|
||
<heading id="D0B6ED76532DFFE2FF12993BFEE1C350" bold="true" box="[189,303,1881,1907]" fontSize="11" level="3" pageId="15" pageNumber="26" reason="3">
|
||
<emphasis id="B9358608532DFFE2FF12993BFEE1C350" bold="true" box="[189,303,1881,1907]" pageId="15" pageNumber="26">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A532DFFFDFF1299E5FAB9C564" blockId="15.[189,1399,1927,2025]" lastBlockId="16.[189,1399,265,327]" lastPageId="16" lastPageNumber="27" pageId="15" pageNumber="26">
|
||
<bibRefCitation id="EFD027EB532DFFE2FF1299E5FEA7C381" author="Lutken C. F." box="[189,361,1927,1954]" pageId="15" pageNumber="26" pagination="75 - 169" refId="ref16788" refString="Lutken C. F. 1859. Additamenta ad Historiam Ophiuridarum. Beskrivelser af de fra Vestindien og Central Amerikas Vestkyst hidtil beckjendte Slangestjerner. Kongelige Danske Videnskabernes Selskab Biologiske Skrifter 2: 75 - 169." type="journal article" year="1859">Lütken (1859)</bibRefCitation>
|
||
did not designate any specific type material in his description of
|
||
<taxonomicName id="4C412199532DFFE2FB3C99EAFAF1C382" authorityName="Lutken" authorityYear="1859" box="[1171,1343,1928,1954]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="15" pageNumber="26" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9358608532DFFE2FB3C99EAFB66C381" box="[1171,1192,1928,1954]" italics="true" pageId="15" pageNumber="26">O</emphasis>
|
||
.
|
||
<emphasis id="B9358608532DFFE2FB1499EAFAF1C382" box="[1211,1343,1928,1953]" italics="true" pageId="15" pageNumber="26">panamense</emphasis>
|
||
</taxonomicName>
|
||
, but according to the International Code of Zoological Nomenclature (
|
||
<bibRefCitation id="EFD027EB532DFFE2FC6B99C9FB9EC3E6" author="ICZN & International Commission on Zoological Nomenclature" box="[964,1104,1963,1989]" pageId="15" pageNumber="26" refId="ref16522" refString="ICZN, International Commission on Zoological Nomenclature. 1999. International Code of Zoological Nomenclature. Available from https: // www. iczn. org / the-code / the-international-code-of-zoological-nomenclature / the-code-online / [accessed 1 Feb. 2020]." type="url" year="1999">ICZN 1999</bibRefCitation>
|
||
), Article 73.1.2, the only specimen reported in the original description (NHMD-107679) must be treated as
|
||
<typeStatus id="54FAE4B8532DFFE2FBC399ACFB1DC3CB" box="[1132,1235,1998,2024]" pageId="15" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
by monotypy. The description provided by
|
||
<bibRefCitation id="EFD027EB5332FFFDFDA29F6BFD7BC507" author="Lutken C. F." box="[525,693,265,292]" pageId="16" pageNumber="27" pagination="75 - 169" refId="ref16788" refString="Lutken C. F. 1859. Additamenta ad Historiam Ophiuridarum. Beskrivelser af de fra Vestindien og Central Amerikas Vestkyst hidtil beckjendte Slangestjerner. Kongelige Danske Videnskabernes Selskab Biologiske Skrifter 2: 75 - 169." type="journal article" year="1859">Lütken (1859)</bibRefCitation>
|
||
is mostly based on the comparison of this species with other
|
||
<taxonomicName id="4C4121995332FFFDFF129F4FFE9FC564" authorityName="Muller & Troschel" authorityYear="1840" box="[189,337,301,327]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="27" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B93586085332FFFDFF129F4FFE9FC564" box="[189,337,301,327]" italics="true" pageId="16" pageNumber="27">Ophioderma</emphasis>
|
||
</taxonomicName>
|
||
from the Atlantic; therefore, a redescription of the
|
||
<typeStatus id="54FAE4B85332FFFDFC329F4FFBCAC564" box="[925,1028,301,327]" pageId="16" pageNumber="27" type="holotype">holotype</typeStatus>
|
||
is provided in the present work.
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5332FFFDFF129F12FB21C642" blockId="16.[189,1399,367,609]" pageId="16" pageNumber="27">
|
||
We observed that the
|
||
<typeStatus id="54FAE4B85332FFFDFE659F0DFDFAC5AA" box="[458,564,367,393]" pageId="16" pageNumber="27" type="holotype">holotype</typeStatus>
|
||
, as well as adult voucher specimens, can have some 'fragmented' or 'broken' dorsal arm plates (
|
||
<figureCitation id="137A469F5332FFFDFE519FF1FD95C58D" box="[510,603,403,430]" captionStart="Fig" captionStartId="9.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@9.[189,1398,291,1855]" captionTargetPageId="9" captionText="Fig. 5. A–D. Ophioderma hendleri sp. nov. A–B. In vivo coloration in natural habitat. C–D. In vivo coloration of collected specimens. – E–J. Ophioderma panamense Lütken, 1859, holotype (NHMD- 107679, DD = 21.9 mm). E. Dorsal disc. F. Ventral disc. G. Mouth. H. Dorsal arm. I. Ventral arm. J. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326625" httpUri="https://zenodo.org/record/4326625/files/figure.png" pageId="16" pageNumber="27">Fig. 5H</figureCitation>
|
||
) indicating that they might have been crushed by moving between rocks and these disruptions are the result of injury (
|
||
<bibRefCitation id="EFD027EB5332FFFDFC9C9FD5FBDCC5F1" author="Ziesenhenne F. C." box="[819,1042,439,466]" pageId="16" pageNumber="27" pagination="185 - 201" refId="ref18074" refString="Ziesenhenne F. C. 1955. A review of the genus Ophioderma Muller and Troschel. In: Allan Hancock Foundation for Scientific Research (eds) Essays in the Natural Sciences in Honor of Captain Allan Hancock: 185 - 201. University of Southern California Press, Los Angeles." type="book chapter" year="1955">Ziesenhenne 1955</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD027EB5332FFFDFB8E9FD5FACFC5F1" author="Stohr S. & Weber A. A. & Boissin E. & Chenuil A." box="[1057,1281,439,466]" pageId="16" pageNumber="27" pagination="1 - 37" refId="ref17759" refString="Stohr S., Weber A. A., Boissin E. & Chenuil A. 2020 b. Resolving the Ophioderma longicauda (Echinodermata: Ophiuroidea) cryptic species complex: five sisters, three of them new. European Journal of Taxonomy 600: 1 - 37. https: // doi. org / 10.5852 / ejt. 2020.600" type="journal article" year="2020">
|
||
Stöhr
|
||
<emphasis id="B93586085332FFFDFBC49FDAFB6FC5F2" box="[1131,1185,439,465]" italics="true" pageId="16" pageNumber="27">et al</emphasis>
|
||
. 2020b
|
||
</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFD027EB5332FFFDFAB79FD5FEC5C5D6" author="Hendler G." pageId="16" pageNumber="27" pagination="189 - 311" refId="ref16304" refString="Hendler G. 2018. Armed to the teeth: a new paradigm for the buccal skeleton of brittle stars (Echinodermata: Ophiuroidea). Contributions in Science 526: 189 - 311." type="journal article" year="2018">Hendler (2018)</bibRefCitation>
|
||
observed that 'fragmented' dorsal arm plates in
|
||
<taxonomicName id="4C4121995332FFFDFCF99FB9FBCFC5D6" authorityName="Lutken" authorityYear="1859" box="[854,1025,475,502]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="27" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B93586085332FFFDFCF99FB9FCA5C5D6" box="[854,875,475,501]" italics="true" pageId="16" pageNumber="27">O</emphasis>
|
||
.
|
||
<emphasis id="B93586085332FFFDFCD29FBEFBCFC5D6" box="[893,1025,476,501]" italics="true" pageId="16" pageNumber="27">panamense</emphasis>
|
||
</taxonomicName>
|
||
frequently occur on basal arm segments of adult specimens, but are rare on distal arm segments of adults and juveniles. It is important to emphasize that these fragmentations on the dorsal arm plates are a result of mechanical damage and are not an inherent feature of the species as occurs in
|
||
<emphasis id="B93586085332FFFDFC9E9C25FBF8C642" box="[817,1078,583,609]" italics="true" pageId="16" pageNumber="27">
|
||
<taxonomicName id="4C4121995332FFFDFC9E9C25FC01C642" authorityName="Pineda-Enriquez, Solis-Marin, Hooker & Laguarda-Figueras" authorityYear="2013" box="[817,975,583,609]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="27" phylum="Echinodermata" rank="species" species="peruanum">O. peruanum</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4121995332FFFDFC759C25FBF8C642" authorityName="Lyman" authorityYear="1860" box="[986,1078,583,609]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="27" phylum="Echinodermata" rank="species" species="teres">O. teres</taxonomicName>
|
||
</emphasis>
|
||
or
|
||
<taxonomicName id="4C4121995332FFFDFBF29C25FB26C642" authorityName="Hendler" authorityYear="1996" box="[1117,1256,583,609]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="27" phylum="Echinodermata" rank="species" species="vansyoci">
|
||
<emphasis id="B93586085332FFFDFBF29C25FB26C642" box="[1117,1256,583,609]" italics="true" pageId="16" pageNumber="27">O. vansyoci</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFE5A1A5332FFFDFF129CEBFAB9C710" blockId="16.[189,1399,649,820]" pageId="16" pageNumber="27">
|
||
Along with
|
||
<taxonomicName id="4C4121995332FFFDFEE79CE8FE03C680" authority="Granja-Fernández & Pineda-Enríquez & Solís-Marín & Laguarda-Figueras, 2020" authorityName="Granja-Fernández & Pineda-Enríquez & Solís-Marín & Laguarda-Figueras" authorityYear="2020" box="[328,461,649,676]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="27" phylum="Echinodermata" rank="species" species="hendleri" status="sp. nov.">
|
||
<emphasis id="B93586085332FFFDFEE79CE8FE03C680" box="[328,461,649,676]" italics="true" pageId="16" pageNumber="27">O. hendleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2063B735332FFFDFE7B9CE8FDFEC687" box="[468,560,650,676]" pageId="16" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C4121995332FFFDFD949CE8FD2CC680" authorityName="Lutken" authorityYear="1859" box="[571,738,650,676]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="27" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B93586085332FFFDFD949CE8FD2CC680" box="[571,738,650,676]" italics="true" pageId="16" pageNumber="27">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
is one of the most conspicuous and abundant ophiuroids in the Eastern Pacific, but it has often been confused with
|
||
<taxonomicName id="4C4121995332FFFDFCDC9CCFFC1EC6E4" authorityName="Lyman" authorityYear="1860" box="[883,976,685,711]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="27" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B93586085332FFFDFCDC9CCFFC1EC6E4" box="[883,976,685,711]" italics="true" pageId="16" pageNumber="27">O. teres</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4121995332FFFDFBA49CCFFB78C6E4" authorityName=", Lutken" authorityYear="1856" box="[1035,1206,685,711]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="27" phylum="Echinodermata" rank="species" species="variegatum">
|
||
<emphasis id="B93586085332FFFDFBA49CCFFB78C6E4" box="[1035,1206,685,711]" italics="true" pageId="16" pageNumber="27">O. variegatum</emphasis>
|
||
</taxonomicName>
|
||
(see Discussion section). Due to its commonness,
|
||
<taxonomicName id="4C4121995332FFFDFDEC9CB3FD27C6C8" authorityName="Lutken" authorityYear="1859" box="[579,745,721,748]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="27" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B93586085332FFFDFDEC9CB3FD96C6C8" box="[579,600,721,747]" italics="true" pageId="16" pageNumber="27">O</emphasis>
|
||
.
|
||
<emphasis id="B93586085332FFFDFDCA9CB0FD27C6C8" box="[613,745,722,747]" italics="true" pageId="16" pageNumber="27">panamense</emphasis>
|
||
</taxonomicName>
|
||
is one of the most studied and commented species of the Eastern Pacific (
|
||
<bibRefCitation id="EFD027EB5332FFFDFED19C97FDD6C733" author="Lyman T." box="[382,536,757,784]" pageId="16" pageNumber="27" pagination="1 - 200" refId="ref16993" refString="Lyman T. 1865. Ophiuridae and Astrophytidae. Illustrated Catalogue of the Museum of Comparative Zoology at Harvard College 1: 1 - 200." type="journal article" year="1865">Lyman 1865</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD027EB5332FFFDFD8A9C97FD78C72C" author="Verrill A. E." box="[549,694,757,783]" pageId="16" pageNumber="27" pagination="251 - 322" refId="ref17963" refString="Verrill A. E. 1867. V. Notes on the Radiata in the Museum of Yale College, with descriptions of new genera and species. No. 2. Notes on the echinoderms of Panama and West Coast of America, with descriptions of new genera and species. Transactions of the Connecticut Academy of Arts and Sciences 1 (2): 251 - 322." type="journal article" year="1867">Verrill 1867</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD027EB5332FFFDFD6B9C94FC87C72C" author="Ives J. E." box="[708,841,757,784]" pageId="16" pageNumber="27" pagination="76 - 77" refId="ref16563" refString="Ives J. E. 1889 a. Variation in Ophiura panamensis and Ophiura teres. Proceedings of the Academy of Natural Sciences of Philadelphia 41: 76 - 77." type="journal article" year="1889">Ives 1889a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD027EB5332FFFDFCF99C97FBE9C72C" author="McClendon J. F." box="[854,1063,757,783]" pageId="16" pageNumber="27" pagination="33 - 64" refId="ref17283" refString="McClendon J. F. 1909. XXV. The ophiurans of the San Diego region. University of California Publications in Zoology 6 (3): 33 - 64." type="journal article" year="1909">McClendon 1909</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD027EB5332FFFDFB9B9C97FB1BC72C" author="Nielsen E." box="[1076,1237,757,783]" pageId="16" pageNumber="27" pagination="241 - 346" refId="ref17315" refString="Nielsen E. 1932. Ophiurans from the Gulf of Panama, California and the Strait of Georgia. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening 91: 241 - 346." type="journal article" year="1932">Nielsen 1932</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD027EB5332FFFDFB4D9C97FF35C717" author="Ziesenhenne F. C." pageId="16" pageNumber="27" pagination="185 - 201" refId="ref18074" refString="Ziesenhenne F. C. 1955. A review of the genus Ophioderma Muller and Troschel. In: Allan Hancock Foundation for Scientific Research (eds) Essays in the Natural Sciences in Honor of Captain Allan Hancock: 185 - 201. University of Southern California Press, Los Angeles." type="book chapter" year="1955">Ziesenhenne 1955</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD027EB5332FFFDFEAA9D7BFD95C710" author="Granja-Fernandez R. & Herrero-Perezrul M. D. & Lopez-Perez R. A. & Hernandez L. & Rodriguez-Zaragoza F. A. & Jones R. W. & Pineda-Lopez R." box="[261,603,793,819]" pageId="16" pageNumber="27" pagination="101 - 145" refId="ref15937" refString="Granja-Fernandez R., Herrero-Perezrul M. D., Lopez-Perez R. A., Hernandez L., Rodriguez-Zaragoza F. A., Jones R. W. & Pineda-Lopez R. 2014. Ophiuroidea (Echinodermata) from coral reefs in the Mexican Pacific. Zookeys 406: 101 - 145. https: // doi. org / 10.3897 / zookeys. 406.6306" type="journal article" year="2014">
|
||
Granja-Fernández
|
||
<emphasis id="B93586085332FFFDFE719D78FDC1C710" box="[478,527,793,819]" italics="true" pageId="16" pageNumber="27">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
), but many of these studies may be based on misidentified specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |