426 lines
88 KiB
XML
426 lines
88 KiB
XML
<document ID-DOI="10.35929/RSZ.0083" ID-ISSN="0035-418" ID-Zenodo-Dep="7761487" checkinTime="1679514280647" checkinUser="valdenar" docAuthor="Schwendinger, Peter J., Huber, Siegfried, Lehmann-Graber, Christina, Ono, Hirotsugu, Aung, Mu Mu & Hongpadharakiree, Komsan" docDate="2022" docId="03E18D71726C0347B298FB805703FAA1" docLanguage="en" docName="RevSuisseZool.129.2.375-424.pdf" docOrigin="Revue suisse de Zoologie 129 (2)" docSource="http://dx.doi.org/10.35929/rsz.0083" docStyle="DocumentStyle:EA5207814960044D272DF5BE47DFF256.3:RevSuisseZool.2020-.journal_article.1cover" docStyleId="EA5207814960044D272DF5BE47DFF256" docStyleName="RevSuisseZool.2020-.journal_article.1cover" docStyleVersion="3" docTitle="Liphistius platnicki Schwendinger & Huber, sp. nov." docType="treatment" docVersion="3" lastPageNumber="420" masterDocId="FFD8F50972460369B239FFEC5305FF92" masterDocTitle="A taxonomic revision of the Liphistius birmanicus-group (Araneae: Liphistiidae) with the description of five new species" masterLastPageNumber="424" masterPageNumber="375" pageNumber="416" updateTime="1679571001943" updateUser="valdenar">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A taxonomic revision of the Liphistius birmanicus-group (Araneae: Liphistiidae) with the description of five new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Schwendinger, Peter J.</mods:namePart>
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<mods:affiliation>Muséum d’histoire naturelle de Genève, C. P. 6404, CH- 1211 Genève 6, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Huber, Siegfried</mods:namePart>
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<mods:affiliation>Ottenbohlstrasse 12, D- 88690 Mühlhofen, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lehmann-Graber, Christina</mods:namePart>
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<mods:affiliation>Muséum d’histoire naturelle de Genève, C. P. 6404, CH- 1211 Genève 6, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ono, Hirotsugu</mods:namePart>
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<mods:affiliation>Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Aung, Mu Mu</mods:namePart>
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<mods:affiliation>Forest Research Institute, Forest Department, Ministry of Natural Resources and Environmental Conservation, Yezin, Nay Pyi Taw, Myanmar</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hongpadharakiree, Komsan</mods:namePart>
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<mods:affiliation>Sirinart Rajini Mangrove Ecosystem Learning Center, Pranburi, Prachuab Khiri Khan Province, Thailand</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Revue suisse de Zoologie</mods:title>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-10-03</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>129</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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<mods:start>375</mods:start>
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<mods:end>424</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.35929/rsz.0083</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.35929/RSZ.0083</mods:identifier>
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<mods:identifier type="ISSN">0035-418</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7761487</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7761557" ID-Zenodo-Dep="7761557" LSID="urn:lsid:plazi:treatment:03E18D71726C0347B298FB805703FAA1" httpUri="http://treatment.plazi.org/id/03E18D71726C0347B298FB805703FAA1" lastPageId="46" lastPageNumber="420" pageId="42" pageNumber="416">
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<subSubSection box="[161,747,1132,1156]" pageId="42" pageNumber="416" type="nomenclature">
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<paragraph blockId="42.[161,747,1132,1188]" box="[161,747,1132,1156]" pageId="42" pageNumber="416">
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<heading bold="true" box="[161,747,1132,1156]" centered="true" fontSize="10" level="2" pageId="42" pageNumber="416" reason="2">
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<emphasis bold="true" box="[161,747,1132,1156]" pageId="42" pageNumber="416">
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<taxonomicName authority="Schwendinger & Huber" authorityName="Schwendinger & Huber" box="[161,649,1132,1156]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="416" phylum="Arthropoda" rank="species" species="platnicki" status="sp. nov.">
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<emphasis bold="true" box="[161,373,1132,1156]" italics="true" pageId="42" pageNumber="416">Liphistius platnicki</emphasis>
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Schwendinger & Huber
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</taxonomicName>
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,
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<taxonomicNameLabel box="[661,747,1133,1156]" pageId="42" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[361,546,1165,1188]" pageId="42" pageNumber="416" type="description">
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<paragraph blockId="42.[161,747,1132,1188]" box="[361,546,1165,1188]" pageId="42" pageNumber="416">
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<figureCitation box="[361,426,1165,1188]" captionStart="Fig" captionStartId="7.[151,186,1431,1452]" captionTargetBox="[151,1436,257,1395]" captionTargetId="figure-270@7.[151,1436,257,1395]" captionTargetPageId="7" captionText="Fig. 1. Localities of species in the Liphistius birmanicus-group in northern Thailand, eastern Myanmar and southern China. Blue lines indicate major rivers, black lines national borders, grey lines major roads." figureDoi="http://doi.org/10.5281/zenodo.7761489" httpUri="https://zenodo.org/record/7761489/files/figure.png" pageId="42" pageNumber="416">Figs 1</figureCitation>
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,
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<figureCitation box="[440,473,1165,1188]" captionStart="Fig" captionStartId="8.[151,186,1932,1953]" captionTargetBox="[155,1431,783,1902]" captionTargetId="figure-159@8.[151,1436,777,1910]" captionTargetPageId="8" captionText="Fig. 2. Habitus of some species of the Liphistius birmanicus-group. (A) Liphistius tung sp. nov., male holotype.(B) Liphistius pyinoolwin, male (matured 1.X.2014) from the Anisakan Falls; note exuviae of ectoparasitic mites (Ljunghia sp.) on the opisthosoma. (C) Liphistius platnicki sp. nov., female paratype (moulted 10.III.2021) from near Loi Kan Village. (D, F) Liphistius cupreus sp. nov., male paratype from near Kong Nyaung Village. (E) Same species, different male paratype. (G) Same species, female paratype (moulted 13.II.2021). (H-I) Same species, different female paratype (moulted 21.V.2021). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7761491" httpUri="https://zenodo.org/record/7761491/files/figure.png" pageId="42" pageNumber="416">2C</figureCitation>
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,
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<figureCitation box="[484,546,1165,1188]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="42.[151,186,1797,1818]" captionStartId-1="44.[153,188,1912,1933]" captionTargetId-0="figure-9@43.[151,1436,257,2027]" captionTargetId-1="figure-9@45.[161,1428,257,2074]" captionTargetPageId-0="43" captionTargetPageId-1="45" captionText-0="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." captionText-1="Fig. 24. Liphistius platnicki sp. nov., vulval plates of ten females (all from exuviae), dorsal view (A, D, K, O), ventral view (B-C, E, H-J, L-N, P), anterior view (F), and isolated uterus externus (G). (A-B) Moult of 29.I.2021. (C, F) Moult of 10.III.2021 (al- lotype). (D-E) Moult of 25.IV.2020. (G, N) Moult of 14.X.2021. (H) Moult of 14.V.2020. (I) Moult of 20.XI.2021. (J) Moult of 13.X.2021. (K-L) Moult of 2.IV.2021. (M) Moult of 7.I.2021. (O-P) Moult of 3.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm" figureDoi-0="http://doi.org/10.5281/zenodo.7761553" figureDoi-1="http://doi.org/10.5281/zenodo.7761555" httpUri-0="https://zenodo.org/record/7761553/files/figure.png" httpUri-1="https://zenodo.org/record/7761555/files/figure.png" pageId="42" pageNumber="416">23-24</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="42" pageNumber="416" type="materials_examined">
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<paragraph blockId="42.[151,756,1211,1362]" pageId="42" pageNumber="416">
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<materialsCitation pageId="42" pageNumber="416">
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<emphasis bold="true" box="[151,264,1211,1234]" pageId="42" pageNumber="416">Holotype:</emphasis>
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MHNG-ARTO-0028361 (sample MT-14/25); male (matured 1.VIII.2016); Myanmar, Shan State, N of Lashio, near Loi Kan Village (23°05’23”N, 97°47’02”E), 1140 m; 10.VI.2014; leg. P.J. Schwendinger & S. Huber.
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</materialsCitation>
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</paragraph>
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<paragraph blockId="42.[151,756,1386,1664]" pageId="42" pageNumber="416">
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<materialsCitation pageId="42" pageNumber="416">
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<emphasis bold="true" box="[151,278,1386,1409]" pageId="42" pageNumber="416">Paratypes:</emphasis>
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MHNG-ARTO-0028362 to MHNGARTO-0028366, BRCM (sample MT-14/25); 6 males (matured 10.IX.2015; 26.V.2016; 2.VII.2016; 3.VII.2016; 26.VII.2016; 8.VIII.2016); collected together with the holotype.
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</materialsCitation>
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–
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<materialsCitation pageId="42" pageNumber="416">BRCM, MHNGARTO-0028369 to MHNG-ARTO-0028377 (sample MT-14/25); 10 female paratypes (allotype, MHNGARTO-0028377, last moulted 10.III.2021); collected together with the holotype.</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="42" pageNumber="416" type="etymology">
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<paragraph blockId="42.[151,756,1687,1743]" pageId="42" pageNumber="416">
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<emphasis bold="true" box="[151,284,1687,1710]" pageId="42" pageNumber="416">Etymology:</emphasis>
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We name this species in honour of the renowned American arachnologist Norman I. Platnick
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</paragraph>
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<paragraph blockId="42.[831,1436,258,509]" pageId="42" pageNumber="416">
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(1951-2020) who, together with Walter C. Sedgwick, revised and properly defined the genus
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<taxonomicName box="[1329,1436,290,313]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="416" phylum="Arthropoda" rank="genus">
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<emphasis box="[1329,1436,290,313]" italics="true" pageId="42" pageNumber="416">Liphistius</emphasis>
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</taxonomicName>
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in 1984 and described several species (also from Myanmar) in this genus. Initially we wanted to name after him the species that he misidentified as
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<taxonomicName authorityName="Thorell" authorityYear="1897" box="[831,989,421,444]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="416" phylum="Arthropoda" rank="species" species="birmanicus">
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<emphasis box="[831,989,421,444]" italics="true" pageId="42" pageNumber="416">L. birmanicus</emphasis>
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</taxonomicName>
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and that he described and illustrated from both sexes in much detail, but that meanwhile was named
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<taxonomicName authorityName="Xu, Yu, Aung, Yu, Liu, Lwin, Sang & Li" authorityYear="2021" box="[909,1056,486,509]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="416" phylum="Arthropoda" rank="species" species="pyinoolwin">
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<emphasis box="[909,1056,486,509]" italics="true" pageId="42" pageNumber="416">L. pyinoolwin</emphasis>
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</taxonomicName>
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in 2021.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="42" pageNumber="416" type="diagnosis">
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<paragraph blockId="42.[831,1437,533,1272]" pageId="42" pageNumber="416">
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<emphasis bold="true" box="[831,953,533,556]" pageId="42" pageNumber="416">Diagnosis:</emphasis>
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Medium-sized, mostly brown species with indistinctly annulated legs in females (but not in males). Most similar to
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<taxonomicName box="[1137,1249,598,621]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="416" phylum="Arthropoda" rank="species" species="nabang">
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<emphasis box="[1137,1249,598,621]" italics="true" pageId="42" pageNumber="416">L. nabang</emphasis>
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</taxonomicName>
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. Males different by distal contrategular edge in distal view without a prodorsal protrusion (
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<figureCitation box="[1107,1214,663,686]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="42" pageNumber="416">Fig. 23A</figureCitation>
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, E-F, I; present in
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<taxonomicName authority=", Yu et al., 2021" authorityName="Yu" authorityYear="2021" box="[831,1117,696,719]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="416" phylum="Arthropoda" rank="species" species="nabang">
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<emphasis box="[831,941,696,719]" italics="true" pageId="42" pageNumber="416">L. nabang</emphasis>
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,
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<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[954,1117,696,719]" pageId="42" pageNumber="416" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
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Yu
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<emphasis box="[991,1046,696,719]" italics="true" pageId="42" pageNumber="416">et al.</emphasis>
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, 2021
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</bibRefCitation>
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</taxonomicName>
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: fig. 3B) and with a narrowly rounded apex (
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<figureCitation box="[995,1073,729,752]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="42" pageNumber="416">Fig. 23</figureCitation>
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E-G; pointed in
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<taxonomicName authority=", Yu et al., 2021" authorityName="Yu" authorityYear="2021" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="416" phylum="Arthropoda" rank="species" species="nabang">
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||
<emphasis box="[1253,1364,729,752]" italics="true" pageId="42" pageNumber="416">L. nabang</emphasis>
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||
,
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||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." pageId="42" pageNumber="416" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
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||
Yu
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||
<emphasis italics="true" pageId="42" pageNumber="416">et al.</emphasis>
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, 2021
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</bibRefCitation>
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</taxonomicName>
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: fig. 3B); proximal edge of tegulum widely rounded (
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<figureCitation box="[939,1021,794,817]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="42" pageNumber="416">Fig. 23</figureCitation>
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J-L; angular in
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<taxonomicName authority=", Yu et al., 2021" authorityName="Yu" authorityYear="2021" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="416" phylum="Arthropoda" rank="species" species="nabang">
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<emphasis box="[1199,1312,794,817]" italics="true" pageId="42" pageNumber="416">L. nabang</emphasis>
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,
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." pageId="42" pageNumber="416" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[1371,1430,794,817]" italics="true" pageId="42" pageNumber="416">et al.</emphasis>
|
||
, 2021
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
: fig. 3A); paracymbium without retrolateralproximal heel (
|
||
<figureCitation box="[1004,1108,859,882]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="42" pageNumber="416">Fig. 23A</figureCitation>
|
||
; present in
|
||
<taxonomicName authority=", Yu et al., 2021" authorityName="Yu" authorityYear="2021" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="416" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis box="[1245,1358,859,882]" italics="true" pageId="42" pageNumber="416">L. nabang</emphasis>
|
||
,
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." pageId="42" pageNumber="416" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis italics="true" pageId="42" pageNumber="416">et al</emphasis>
|
||
., 2021
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
: fig. 3E). Females distinguished from those of
|
||
<taxonomicName box="[831,940,924,947]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="416" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis box="[831,940,924,947]" italics="true" pageId="42" pageNumber="416">L. nabang</emphasis>
|
||
</taxonomicName>
|
||
by poreplate anteriorly wider than posteriorly (in
|
||
<taxonomicName box="[874,988,956,979]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="416" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis box="[874,988,956,979]" italics="true" pageId="42" pageNumber="416">L. nabang</emphasis>
|
||
</taxonomicName>
|
||
anteriorly narrower than posteriorly or equally wide), its anterior margin more strongly invaginated (in
|
||
<taxonomicName box="[963,1076,1021,1044]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="416" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis box="[963,1076,1021,1044]" italics="true" pageId="42" pageNumber="416">L. nabang</emphasis>
|
||
</taxonomicName>
|
||
slightly invaginated to arched); posterior stalk relatively longer and slightly constricted in anterior portion, without “bulging margins” between poreplate and posterior stalk (in
|
||
<taxonomicName box="[1208,1323,1119,1142]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="416" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis box="[1208,1323,1119,1142]" italics="true" pageId="42" pageNumber="416">L. nabang</emphasis>
|
||
</taxonomicName>
|
||
transition between poreplate and posterior stalk very wide and carrying widely separated “bulging margins”); posterior part of genital atrium not strongly curved ventrad (
|
||
<figureCitation captionStart="Fig" captionStartId="44.[153,188,1912,1933]" captionTargetId="figure-9@45.[161,1428,257,2074]" captionTargetPageId="45" captionText="Fig. 24. Liphistius platnicki sp. nov., vulval plates of ten females (all from exuviae), dorsal view (A, D, K, O), ventral view (B-C, E, H-J, L-N, P), anterior view (F), and isolated uterus externus (G). (A-B) Moult of 29.I.2021. (C, F) Moult of 10.III.2021 (al- lotype). (D-E) Moult of 25.IV.2020. (G, N) Moult of 14.X.2021. (H) Moult of 14.V.2020. (I) Moult of 20.XI.2021. (J) Moult of 13.X.2021. (K-L) Moult of 2.IV.2021. (M) Moult of 7.I.2021. (O-P) Moult of 3.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.7761555" httpUri="https://zenodo.org/record/7761555/files/figure.png" pageId="42" pageNumber="416">Fig. 24</figureCitation>
|
||
cf.
|
||
<bibRefCitation author="Yu K. & Zhang F. & Zhang J. - X." box="[898,1058,1249,1272]" pageId="42" pageNumber="416" pagination="36 - 40" refId="ref38634" refString="Yu K., Zhang F., Zhang J. - X. 2021. First new species of the genus Liphistius Schiodte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30 (1): 36 - 40." type="journal article" year="2021">
|
||
Yu
|
||
<emphasis box="[934,988,1249,1272]" italics="true" pageId="42" pageNumber="416">et al.</emphasis>
|
||
, 2021
|
||
</bibRefCitation>
|
||
: fig. 4).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="46" lastPageNumber="420" pageId="42" pageNumber="416" type="description">
|
||
<paragraph blockId="42.[831,1437,1297,1743]" pageId="42" pageNumber="416">
|
||
<emphasis bold="true" box="[831,1213,1297,1320]" pageId="42" pageNumber="416">Description of male (holotype):</emphasis>
|
||
<emphasis box="[1228,1436,1297,1320]" italics="true" pageId="42" pageNumber="416">Colour in alcohol</emphasis>
|
||
(darker in life): Mostly light brown. Carapace with slightly darkened, large, orchid-shaped pattern on pars thoracica; eye mound black. Chelicerae cream-coloured proximally, light brown distally. Ventral side of palps and legs slightly lighter than dorsal side; most of ventral side of palpal coxae and leg tarsi cream-coloured. Legs and palps mostly light brown, without annulations; palpal tarsus and sclerotised parts of palpal organ dark brown. Membranous cuticle of opisthosoma creamcoloured and finely mottled with grey. Sternites of opisthosoma slightly darker than membranous cuticle; tergite I entirely dark brown, tergites II-VI mostly light yellow-brown, with darker spots laterally and on
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7761553" ID-Zenodo-Dep="7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="42" pageNumber="416" startId="42.[151,186,1797,1818]" targetBox="[156,1430,266,2023]" targetPageId="43">
|
||
<paragraph blockId="42.[151,1437,1797,2077]" pageId="42" pageNumber="416">
|
||
Fig. 23.
|
||
<taxonomicName box="[237,423,1797,1818]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="416" phylum="Arthropoda" rank="species" species="platnicki" status="sp. nov.">
|
||
<emphasis box="[237,423,1797,1818]" italics="true" pageId="42" pageNumber="416">Liphistius platnicki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[431,510,1797,1818]" pageId="42" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="42.[151,1437,1797,2077]" lastBlockId="44.[151,757,258,1859]" lastPageId="44" lastPageNumber="418" pageId="42" pageNumber="416">► posterior margin, tergites VII-X light yellow-brown medially and dark brown laterally.</paragraph>
|
||
<paragraph blockId="44.[151,757,258,1859]" pageId="44" pageNumber="418">
|
||
<emphasis box="[151,366,322,345]" italics="true" pageId="44" pageNumber="418">Setae on carapace</emphasis>
|
||
: Several small setae around eye mound, including one closely spaced enlarged pair side by side behind eye mound; few small setae on lateral margins of carapace and on coxal elevations; 1 enlarged seta on each posterolateral corner; no setae on posterior margin; 2 small setae anterior to fovea.
|
||
</paragraph>
|
||
<paragraph blockId="44.[151,757,258,1859]" pageId="44" pageNumber="418">
|
||
<emphasis box="[151,329,513,536]" italics="true" pageId="44" pageNumber="418">Cheliceral teeth</emphasis>
|
||
: 12 small teeth of different sizes on promargin of each cheliceral groove.
|
||
</paragraph>
|
||
<paragraph blockId="44.[151,757,258,1859]" pageId="44" pageNumber="418">
|
||
<emphasis box="[151,237,577,600]" italics="true" pageId="44" pageNumber="418">Scopula</emphasis>
|
||
: Only distally divided by median stripe on all tarsi; quite weak on tarsus I, only slightly denser on other tarsi; covering distal 5/6 of tarsi I-III, distal 4/5 of tarsus IV.
|
||
</paragraph>
|
||
<paragraph blockId="44.[151,757,258,1859]" pageId="44" pageNumber="418">
|
||
<emphasis box="[151,288,706,729]" italics="true" pageId="44" pageNumber="418">Tarsal claws</emphasis>
|
||
: Paired claws with 3 teeth on tarsus I, 3-4 on tarsus II, 4 on tarsus III, 5-6 on tarsus IV; unpaired claw with 1-2 (mostly 2) denticles on tarsi I-III, 1 and 3 on tarsus IV.
|
||
</paragraph>
|
||
<paragraph blockId="44.[151,757,258,1859]" lastBlockId="44.[831,1437,258,865]" pageId="44" pageNumber="418">
|
||
<emphasis box="[151,201,835,858]" italics="true" pageId="44" pageNumber="418">Palp</emphasis>
|
||
: Tibial apophysis short, triangular and basally wide in ventral view, not set back from distal margin of tibia (
|
||
<figureCitation box="[159,255,900,923]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23C</figureCitation>
|
||
), carrying 4 long apical megaspines, the ventral 3 distinctly longer than the dorsal one; dorsal to apophysis a quite long spine on a slightly elevated base (
|
||
<figureCitation box="[652,729,964,987]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23</figureCitation>
|
||
CD). Palpal tarsus with a pronounced subdistal suture on ventral side (
|
||
<figureCitation box="[293,391,1029,1052]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23A</figureCitation>
|
||
) and with a widely but shallowly invaginated distal margin (
|
||
<figureCitation box="[458,542,1061,1084]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23</figureCitation>
|
||
H-I). Paracymbium moderately deep (
|
||
<figureCitation box="[338,433,1093,1116]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23B</figureCitation>
|
||
), in ventral view clearly longer than wide, with a distinctly conical distal side, without a retrolateral-proximal heel (
|
||
<figureCitation box="[440,540,1158,1181]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23A</figureCitation>
|
||
); cumulus distinctly elevated, unpigmented on its retroventral side, carrying a group of 5 long, strong bristles (
|
||
<figureCitation box="[485,561,1223,1246]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23</figureCitation>
|
||
A-B). Subtegulum without apophysis (
|
||
<figureCitation box="[373,507,1255,1278]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23A, J</figureCitation>
|
||
). Tegulum wide, axeblade-shaped; its distal margin elevated, developed as a short, wide keel with a quite straight margin (
|
||
<figureCitation box="[644,721,1319,1342]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23</figureCitation>
|
||
I-J, M); its proximal edge widely rounded and finely dentate, straight (not bent proximad) and strongly protruding from surface of palpal organ (
|
||
<figureCitation box="[407,483,1416,1439]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23</figureCitation>
|
||
I-J, M). Pigmented bridge between tegulum and contrategulum on retrodorsal side of palpal organ well developed (
|
||
<figureCitation box="[489,579,1481,1504]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23J</figureCitation>
|
||
). Contrategulum with moderately developed, conical proventral process (
|
||
<figureCitation box="[159,270,1545,1568]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23E</figureCitation>
|
||
, showing paratype); essentially only a single strong wrinkle on prolateral surface and a very indistinct proximal ledge on retrodorsal side (
|
||
<figureCitation captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23</figureCitation>
|
||
H-J, see also
|
||
<figureCitation box="[334,463,1642,1665]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23F, K</figureCitation>
|
||
for paratype); distal edge of contrategulum narrow and evenly arched, without protrusion prodorsally; dorsal apex of contrategulum quite long and very narrowly rounded (
|
||
<figureCitation box="[610,704,1739,1762]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23I</figureCitation>
|
||
, see also
|
||
<figureCitation box="[202,277,1772,1795]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23</figureCitation>
|
||
E-G for paratype). Para-embolic plate short, about as long as retroventral edge of embolus complex and not clearly separated from it by an invagination (
|
||
<figureCitation box="[713,882,258,1859]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23A</figureCitation>
|
||
). Membranous prolateral-proximal zone of embolus complex not swollen (
|
||
<figureCitation box="[1076,1185,290,313]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23 A</figureCitation>
|
||
; swollen in
|
||
<taxonomicName box="[1322,1436,290,313]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="418" phylum="Arthropoda" rank="species" species="cupreus" status="sp. nov.">
|
||
<emphasis box="[1322,1436,290,313]" italics="true" pageId="44" pageNumber="418">L. cupreus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[831,921,323,346]" pageId="44" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<figureCitation box="[935,1038,323,346]" captionStart="Fig" captionStartId="38.[152,187,1761,1782]" captionTargetId="figure-9@39.[168,1420,257,2071]" captionTargetPageId="39" captionText="Fig. 21. Liphistius cupreus sp. nov., palps of three males; holotype (A-D, H-J, L-N), paratype matured end VI.2018 (E-G), paratype matured mid VI.2018 (K). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left contrategulum, distal and slightly ventral view. (I) Right contrategulum, same view. (J) Part of left palpal organ, retroventral view; slightly bent proximal edge of tegulum indicated by arrow. (K) Right palpal organ, retrolateral and slightly distal view. (L) Left palpal organ and distal margin of tarsus, dorsal and slightly proximal view. (M) Left palpal organ, retrolateral and slightly distal view. (N) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. Scale lines 1.0 mm (A-B, E-N; C-D). Abbreviation: a - angular prodorsal protrusion at base of distal edge of contrategulum." figureDoi="http://doi.org/10.5281/zenodo.7761547" httpUri="https://zenodo.org/record/7761547/files/figure.png" pageId="44" pageNumber="418">Fig. 21A</figureCitation>
|
||
); embolus proper narrowly divided, its sclerotised part strengthened by 3 longitudinal ribs reaching apex and carrying denticles distally (
|
||
<figureCitation box="[1340,1432,388,411]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23J</figureCitation>
|
||
, see also
|
||
<figureCitation box="[925,1002,421,444]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23</figureCitation>
|
||
E-F, K for paratypes); membranous part of embolus proper distinctly shorter than sclerotised part, at its base a short, weakly pigmented area with only a few longitudinal wrinkles and with a very narrow and concave distal margin; embolic folds short (
|
||
<figureCitation box="[1300,1400,551,574]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="44" pageNumber="418">Fig. 23H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="44.[831,1437,258,865]" pageId="44" pageNumber="418">
|
||
<emphasis box="[831,986,583,606]" italics="true" pageId="44" pageNumber="418">Measurements</emphasis>
|
||
: Total length 13.12; CL 5.45, CW 4.93; opisthosoma 5.56 long, 3.82 wide; eye mound 0.90 long, 0.96 wide; palpal coxa 1.91 long, 1.19 wide; labium 0.40 long, 0.95 wide; sternum 2.70 long, 1.91 wide (0.95 on ventral surface); palp 9.14 long (2.74 + 1.67 + 3.18 + 1.55); leg I 16.10 long (4.41 + 2.23 + 3.34 +3.97 + 2.15); leg II 17.01 long (4.45 + 2.23 + 3.46 + 4.49 + 2.38); leg III 19.12 long (4.77 + 2.23 + 3.78 + 5.52 + 2.82); leg IV 23.48 long (5.80 + 2.38 + 4.77 + 7.03 + 3.50).
|
||
</paragraph>
|
||
<paragraph blockId="44.[831,1437,889,1858]" pageId="44" pageNumber="418">
|
||
<emphasis bold="true" box="[831,1220,889,912]" pageId="44" pageNumber="418">Description of female (allotype):</emphasis>
|
||
<emphasis box="[1233,1437,889,912]" italics="true" pageId="44" pageNumber="418">Colour in alcohol</emphasis>
|
||
(darker in life, as in
|
||
<figureCitation box="[1069,1158,922,945]" captionStart="Fig" captionStartId="8.[151,186,1932,1953]" captionTargetBox="[155,1431,783,1902]" captionTargetId="figure-159@8.[151,1436,777,1910]" captionTargetPageId="8" captionText="Fig. 2. Habitus of some species of the Liphistius birmanicus-group. (A) Liphistius tung sp. nov., male holotype.(B) Liphistius pyinoolwin, male (matured 1.X.2014) from the Anisakan Falls; note exuviae of ectoparasitic mites (Ljunghia sp.) on the opisthosoma. (C) Liphistius platnicki sp. nov., female paratype (moulted 10.III.2021) from near Loi Kan Village. (D, F) Liphistius cupreus sp. nov., male paratype from near Kong Nyaung Village. (E) Same species, different male paratype. (G) Same species, female paratype (moulted 13.II.2021). (H-I) Same species, different female paratype (moulted 21.V.2021). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7761491" httpUri="https://zenodo.org/record/7761491/files/figure.png" pageId="44" pageNumber="418">Fig. 2C</figureCitation>
|
||
): Mostly as in male, but generally darker, distal part of chelicerae much darker. Legs and palps with indistinct annulations, i.e. faint and broken light median rings on all tibiae of legs and palps, plus on metararsi of legs II-IV, and very faint on tarsi II-IV; a faint light proximal ring on all leg metatarsi. Membranous cuticle of opisthosoma more distinctly mottled with grey-brown than in male; tergites II-VI with much larger dark portions, extending from anterior to posterior margins, leaving rather small paramedian pairs of longitudinal bands.
|
||
</paragraph>
|
||
<paragraph blockId="44.[831,1437,889,1858]" pageId="44" pageNumber="418">
|
||
<emphasis box="[831,1034,1281,1304]" italics="true" pageId="44" pageNumber="418">Setae on carapace</emphasis>
|
||
: Slightly more than in the holotype; pair of stiff bristles in front of eye mound much larger; pair of enlarged setae behind eye mound situated behind each other (in male side by side); 5 small setae anterior to fovea.
|
||
</paragraph>
|
||
<paragraph blockId="44.[831,1437,889,1858]" pageId="44" pageNumber="418">
|
||
<emphasis box="[831,1006,1444,1467]" italics="true" pageId="44" pageNumber="418">Cheliceral teeth</emphasis>
|
||
: 12 strong teeth on promargin of each cheliceral groove.
|
||
</paragraph>
|
||
<paragraph blockId="44.[831,1437,889,1858]" pageId="44" pageNumber="418">
|
||
<emphasis box="[831,898,1509,1532]" italics="true" pageId="44" pageNumber="418">Claws</emphasis>
|
||
: Left palpal claw with 2 denticles, right claw with 3 denticles. Paired tarsal claws of legs I-II with 3 teeth, of leg III with 4 teeth, of leg IV with 3-4 teeth; unpaired claws of leg I with 3 denticles, of leg II with 3-4 denticles, of leg III with 2 denticles, of leg IV with 1-2 denticles. All tarsi without scopula.
|
||
</paragraph>
|
||
<paragraph blockId="44.[831,1437,889,1858]" pageId="44" pageNumber="418">
|
||
<taxonomicName authorityName="Olafsen" authorityYear="1772" box="[831,891,1704,1727]" class="Polychaeta" genus="Vulva" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="418" phylum="Annelida" rank="genus">
|
||
<emphasis box="[831,891,1704,1727]" italics="true" pageId="44" pageNumber="418">Vulva</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[907,1033,1704,1727]" captionStart="Fig" captionStartId="44.[153,188,1912,1933]" captionTargetId="figure-9@45.[161,1428,257,2074]" captionTargetPageId="45" captionText="Fig. 24. Liphistius platnicki sp. nov., vulval plates of ten females (all from exuviae), dorsal view (A, D, K, O), ventral view (B-C, E, H-J, L-N, P), anterior view (F), and isolated uterus externus (G). (A-B) Moult of 29.I.2021. (C, F) Moult of 10.III.2021 (al- lotype). (D-E) Moult of 25.IV.2020. (G, N) Moult of 14.X.2021. (H) Moult of 14.V.2020. (I) Moult of 20.XI.2021. (J) Moult of 13.X.2021. (K-L) Moult of 2.IV.2021. (M) Moult of 7.I.2021. (O-P) Moult of 3.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.7761555" httpUri="https://zenodo.org/record/7761555/files/figure.png" pageId="44" pageNumber="418">Fig. 24C, F</figureCitation>
|
||
): Membranous uterus externus with a small but distinct pair of lateral pockets (see
|
||
<figureCitation box="[1305,1399,1737,1760]" captionStart="Fig" captionStartId="44.[153,188,1912,1933]" captionTargetId="figure-9@45.[161,1428,257,2074]" captionTargetPageId="45" captionText="Fig. 24. Liphistius platnicki sp. nov., vulval plates of ten females (all from exuviae), dorsal view (A, D, K, O), ventral view (B-C, E, H-J, L-N, P), anterior view (F), and isolated uterus externus (G). (A-B) Moult of 29.I.2021. (C, F) Moult of 10.III.2021 (al- lotype). (D-E) Moult of 25.IV.2020. (G, N) Moult of 14.X.2021. (H) Moult of 14.V.2020. (I) Moult of 20.XI.2021. (J) Moult of 13.X.2021. (K-L) Moult of 2.IV.2021. (M) Moult of 7.I.2021. (O-P) Moult of 3.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.7761555" httpUri="https://zenodo.org/record/7761555/files/figure.png" pageId="44" pageNumber="418">Fig. 24G</figureCitation>
|
||
for another female). Vulval plate slightly longer than wide, with a few hairs on lateral folds. Poreplate with widely and quite deeply invaginated anterior margin carrying a
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7761555" ID-Zenodo-Dep="7761555" httpUri="https://zenodo.org/record/7761555/files/figure.png" pageId="44" pageNumber="418" startId="44.[153,188,1912,1933]" targetBox="[167,1422,270,2070]" targetPageId="45">
|
||
<paragraph blockId="44.[153,1440,1912,2048]" pageId="44" pageNumber="418">
|
||
Fig. 24.
|
||
<taxonomicName box="[237,422,1912,1933]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="418" phylum="Arthropoda" rank="species" species="platnicki" status="sp. nov.">
|
||
<emphasis box="[237,422,1912,1933]" italics="true" pageId="44" pageNumber="418">Liphistius platnicki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[429,506,1912,1933]" pageId="44" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, vulval plates of ten females (all from exuviae), dorsal view (A, D, K, O), ventral view (B-C, E, H-J, L-N, P), anterior view (F), and isolated uterus externus (G). (A-B) Moult of 29.I.2021. (C, F) Moult of 10.III.2021 (al- lotype). (D-E) Moult of 25.IV.2020. (G, N) Moult of 14.X.2021. (H) Moult of 14.V.2020. (I) Moult of 20.XI.2021. (J) Moult of 13.X.2021. (K-L) Moult of 2.IV.2021. (M) Moult of 7.I.2021. (O-P) Moult of 3.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="44.[153,1440,1912,2048]" box="[816,820,2027,2048]" pageId="44" pageNumber="418">.</paragraph>
|
||
<paragraph blockId="44.[1416,1440,2056,2077]" box="[1416,1440,2056,2077]" pageId="44" pageNumber="418">►</paragraph>
|
||
<paragraph blockId="46.[151,757,258,896]" pageId="46" pageNumber="420">
|
||
fairly large and widely separated pair of vesicle clusters (corresponding to anterior lobes); anterolateral processes on each side of poreplate small, composed of only one vesicle (on one side) or a few vesicles (on other side). CDO small; receptacular cluster racemose, slightly longer than wide, not reaching anterior margin of poreplate, very deep (
|
||
<figureCitation box="[220,320,452,475]" captionStart="Fig" captionStartId="44.[153,188,1912,1933]" captionTargetId="figure-9@45.[161,1428,257,2074]" captionTargetPageId="45" captionText="Fig. 24. Liphistius platnicki sp. nov., vulval plates of ten females (all from exuviae), dorsal view (A, D, K, O), ventral view (B-C, E, H-J, L-N, P), anterior view (F), and isolated uterus externus (G). (A-B) Moult of 29.I.2021. (C, F) Moult of 10.III.2021 (al- lotype). (D-E) Moult of 25.IV.2020. (G, N) Moult of 14.X.2021. (H) Moult of 14.V.2020. (I) Moult of 20.XI.2021. (J) Moult of 13.X.2021. (K-L) Moult of 2.IV.2021. (M) Moult of 7.I.2021. (O-P) Moult of 3.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.7761555" httpUri="https://zenodo.org/record/7761555/files/figure.png" pageId="46" pageNumber="420">Fig. 24F</figureCitation>
|
||
). Posterior stalk quite long, wider than poreplate, with a slight constriction at transition from poreplate to posterior stalk; axe-blade-shaped, anteriorly narrower than posteriorly, with a widely arched posterior margin (
|
||
<figureCitation box="[241,339,581,604]" captionStart="Fig" captionStartId="44.[153,188,1912,1933]" captionTargetId="figure-9@45.[161,1428,257,2074]" captionTargetPageId="45" captionText="Fig. 24. Liphistius platnicki sp. nov., vulval plates of ten females (all from exuviae), dorsal view (A, D, K, O), ventral view (B-C, E, H-J, L-N, P), anterior view (F), and isolated uterus externus (G). (A-B) Moult of 29.I.2021. (C, F) Moult of 10.III.2021 (al- lotype). (D-E) Moult of 25.IV.2020. (G, N) Moult of 14.X.2021. (H) Moult of 14.V.2020. (I) Moult of 20.XI.2021. (J) Moult of 13.X.2021. (K-L) Moult of 2.IV.2021. (M) Moult of 7.I.2021. (O-P) Moult of 3.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.7761555" httpUri="https://zenodo.org/record/7761555/files/figure.png" pageId="46" pageNumber="420">Fig. 24C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="46.[151,757,258,896]" pageId="46" pageNumber="420">
|
||
<emphasis box="[151,306,614,637]" italics="true" pageId="46" pageNumber="420">Measurements</emphasis>
|
||
: Total length 18.56; CL 6.68, CW 5.84; opisthosoma 8.62 long, 6.92 wide; eye mound 0.97 long, 1.00 wide; palpal coxa 2.42 long, 1.55 wide; labium 0.64 long, 1.55 wide; sternum 3.54 long, 2.34 wide (1.43 on ventral surface); palp 10.61 long (3.38 + 2.03 + 2.54 + 2.66); leg I 13.59 long (4.21 + 2.42 + 2.74 + 2.70 + 1.52); leg II 14.06 long (4.21 + 2.46 + 2.74 + 2.98 + 1.67); leg III 15.30 long (4.25 + 2.54 + 2.94 + 3.66 + 1.91); leg IV 21.09 long (5.68 + 2.82 + 4.17 + 5.76 + 2.66).
|
||
</paragraph>
|
||
<paragraph blockId="46.[151,757,920,1396]" lastBlockId="46.[831,1437,258,832]" pageId="46" pageNumber="420">
|
||
<emphasis bold="true" box="[151,269,920,943]" pageId="46" pageNumber="420">Variation:</emphasis>
|
||
For carapace measurements and prefoveal setae counts see
|
||
<tableCitation box="[355,444,953,976]" captionStart="Table 1" captionStartId="46.[151,204,1454,1475]" captionTargetId="graphics-627@46.[151,1436,1545,2074]" captionTargetPageId="46" captionText="Table 1. Carapace measurements (in mm; including data from the literature) and prefoveal setae counts of Liphistius birmanicus group species." httpUri="http://table.plazi.org/id/DF376CEF72680347B2AEFA425282FA71" pageId="46" pageNumber="420" tableUuid="DF376CEF72680347B2AEFA425282FA71">Table 1</tableCitation>
|
||
. All specimens have both AME well developed. In four specimens the prefoveal setae are situated relatively far away from the fovea. Most females have two enlarged bristles behind the eye mound, one specimen has only one. In most males the scopula covers 5/6 of the ventral side of tarsi I-III and 4/5 of tarsus IV, in one male it covers 4/5 of tarsus I. Variation in details of the male palp is given in
|
||
<figureCitation box="[151,234,1211,1234]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="46" pageNumber="420">Fig. 23</figureCitation>
|
||
. None of the males has a prodorsal protrusion at the base of the distal contrategular edge (present in
|
||
<taxonomicName box="[151,266,1276,1299]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="420" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis box="[151,266,1276,1299]" italics="true" pageId="46" pageNumber="420">L. nabang</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[328,446,1276,1299]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="420" phylum="Arthropoda" rank="species" species="cupreus" status="sp. nov.">
|
||
<emphasis box="[328,446,1276,1299]" italics="true" pageId="46" pageNumber="420">L. cupreus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[458,550,1276,1299]" pageId="46" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
|
||
). The apex of the proventral contrategular process is narrowly rounded or obliquely truncate (
|
||
<figureCitation box="[367,465,1341,1364]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="46" pageNumber="420">Fig. 23E</figureCitation>
|
||
) in distal view. All males have a distinctly elevated distal tegular edge which is dentate (
|
||
<figureCitation box="[954,1033,258,281]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="46" pageNumber="420">Fig. 23</figureCitation>
|
||
K-L) or quite smooth (
|
||
<figureCitation box="[1288,1382,258,281]" captionStart="Fig" captionStartId="42.[151,186,1797,1818]" captionTargetId="figure-9@43.[151,1436,257,2027]" captionTargetPageId="43" captionText="Fig. 23. Liphistius platnicki sp. nov., palps of three males; holotype (A-D, H-J, M), paratype matured 26.VII.2019 (E-G), paratype matured 8.VIII.2016 (K-L). (A) Left tarsus and palpal organ, ventral view. (B) Left paracymbium and distal margin of tibia, retrolateral view. (C) Distal part of left tibia, ventral view. (D) Same, retrolateral and slightly proximal view. (E) Left palpal organ, distal view (dorsal side up). (F) Dorsal part of left contrategulum and embolus proper, distal and slightly ventral view. (G) Dorsal part of left contrategulum, distal and slightly dorsal view. (H) Left palpal organ and distal margin of tarsus, prodorsal and slightly proximal view. (I) Same, dorsal and slightly proximal view. (J) Same, retrolateral and slightly distal view. (K) Left palpal organ, retrolateral view. (L) Tegulum and nearby parts of right palpal organ, same view. (M) Part of left palpal organ, retroventral view; elevated distal edge of tegulum indicated by short arrow, straight, salient proximal tegular edge indicated by long arrow. Scale lines 1.0 mm (A-B; C-D; E-M)." figureDoi="http://doi.org/10.5281/zenodo.7761553" httpUri="https://zenodo.org/record/7761553/files/figure.png" pageId="46" pageNumber="420">Fig. 23J</figureCitation>
|
||
). All males examined have three ridges reaching the apex of the sclerotised embolus part. The lightly pigmented area at the base of the membranous embolus part has wrinkles that are more or less strongly inclined, in some specimens being almost horizontal. In small females the annulations (light median rings) on the legs and palps are more pronounced than in the allotype (the largest female examined). Variation in the shape of the vulval plate is shown in
|
||
<figureCitation box="[1032,1116,550,573]" captionStart="Fig" captionStartId="44.[153,188,1912,1933]" captionTargetId="figure-9@45.[161,1428,257,2074]" captionTargetPageId="45" captionText="Fig. 24. Liphistius platnicki sp. nov., vulval plates of ten females (all from exuviae), dorsal view (A, D, K, O), ventral view (B-C, E, H-J, L-N, P), anterior view (F), and isolated uterus externus (G). (A-B) Moult of 29.I.2021. (C, F) Moult of 10.III.2021 (al- lotype). (D-E) Moult of 25.IV.2020. (G, N) Moult of 14.X.2021. (H) Moult of 14.V.2020. (I) Moult of 20.XI.2021. (J) Moult of 13.X.2021. (K-L) Moult of 2.IV.2021. (M) Moult of 7.I.2021. (O-P) Moult of 3.X.2021. Arrows indicating paired lateral pockets of membranous uterus externus. Scale lines 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.7761555" httpUri="https://zenodo.org/record/7761555/files/figure.png" pageId="46" pageNumber="420">Fig. 24</figureCitation>
|
||
: the lateral folds carry no or only very few hairs; the anterior margin of the poreplate is slightly to deeply invaginated, widely concave to straight; the anterior lobes (vesicle clusters) are narrow to wide; the anterolateral processes are small and composed of one to only a few vesicles; the receptacular cluster is small to medium-sized, always racemose, never reaching the anterior poreplate margin; the posterior stalk is axe-blade-shaped to almost elliptical.
|
||
</paragraph>
|
||
<paragraph blockId="46.[831,1436,856,1009]" pageId="46" pageNumber="420">
|
||
<emphasis bold="true" box="[831,996,856,879]" pageId="46" pageNumber="420">Relationships:</emphasis>
|
||
The new species appears most closely related to
|
||
<taxonomicName box="[945,1057,889,912]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="420" phylum="Arthropoda" rank="species" species="nabang">
|
||
<emphasis box="[945,1057,889,912]" italics="true" pageId="46" pageNumber="420">L. nabang</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName box="[1092,1208,889,912]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="420" phylum="Arthropoda" rank="species" species="cupreus" status="sp. nov.">
|
||
<emphasis box="[1092,1208,889,912]" italics="true" pageId="46" pageNumber="420">L. cupreus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1218,1304,889,912]" pageId="46" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
|
||
with which it shares characters explained further below (see Discussion - Relationships). These three species form the most distinct subgroup within the
|
||
<taxonomicName authorityName="Thorell" authorityYear="1897" box="[1234,1354,986,1009]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="420" phylum="Arthropoda" rank="species" species="birmanicus">
|
||
<emphasis box="[1234,1354,986,1009]" italics="true" pageId="46" pageNumber="420">birmanicus</emphasis>
|
||
</taxonomicName>
|
||
-group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="46" pageNumber="420" type="distribution">
|
||
<paragraph blockId="46.[831,1436,1034,1121]" pageId="46" pageNumber="420">
|
||
<emphasis bold="true" box="[831,980,1034,1057]" pageId="46" pageNumber="420">Distribution:</emphasis>
|
||
This species is only known from the type locality near Lashio, about 17 km north of the type locality of
|
||
<taxonomicName box="[947,1058,1098,1121]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="420" phylum="Arthropoda" rank="species" species="cupreus" status="sp. nov.">
|
||
<emphasis box="[947,1058,1098,1121]" italics="true" pageId="46" pageNumber="420">L. cupreus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1065,1148,1098,1121]" pageId="46" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation box="[1162,1225,1098,1121]" captionStart="Fig" captionStartId="7.[151,186,1431,1452]" captionTargetBox="[151,1436,257,1395]" captionTargetId="figure-270@7.[151,1436,257,1395]" captionTargetPageId="7" captionText="Fig. 1. Localities of species in the Liphistius birmanicus-group in northern Thailand, eastern Myanmar and southern China. Blue lines indicate major rivers, black lines national borders, grey lines major roads." figureDoi="http://doi.org/10.5281/zenodo.7761489" httpUri="https://zenodo.org/record/7761489/files/figure.png" pageId="46" pageNumber="420">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="46" pageNumber="420" type="biology_ecology">
|
||
<paragraph blockId="46.[831,1436,1146,1331]" pageId="46" pageNumber="420">
|
||
<emphasis bold="true" box="[831,927,1146,1169]" pageId="46" pageNumber="420">Biology:</emphasis>
|
||
No mature spiders and no egg cases were found at the type locality, but the dates at which males matured in captivity (especially the first date after capture) indicate that this species has a phenology very similar to other congeneric species in Myanmar and northern Thailand.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |