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<document id="C82DED3E27392ACC923406412052F788" ID-CLB-Dataset="2421" ID-DOI="10.11646/phytotaxa.532.1.9" ID-GBIF-Dataset="caf7281b-193e-4f5e-8292-6874b9b8bd30" ID-ISSN="1179-3163" ID-Zenodo-Dep="5900972" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643101474042" checkinUser="plazi" docAuthor="Luna, Efraín De" docDate="2022" docId="03E087F9FFB0FF99FF0A66980CF9DAC6" docLanguage="en" docName="phytotaxa.532.1.9.pdf" docOrigin="Phytotaxa 532 (1)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Braunia plicata Jaeger" docType="treatment" docVersion="5" lastPageNumber="106" masterDocId="FFD9FF81FFB1FF9CFF8266600867D812" masterDocTitle="Lectotypification, taxonomy and distribution of Braunia plicata (Bryopsida, Hedwigiaceae) in South America and Mexico" masterLastPageNumber="106" masterPageNumber="101" pageNumber="102" updateTime="1699304083390" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CB9246AA3446DC3613DAEFC45E2CA1C2">Lectotypification, taxonomy and distribution of Braunia plicata (Bryopsida, Hedwigiaceae) in South America and Mexico</mods:title>
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<treatment id="03E087F9FFB0FF99FF0A66980CF9DAC6" ID-DOI="http://doi.org/10.5281/zenodo.5903280" ID-GBIF-Taxon="192601310" ID-Zenodo-Dep="5903280" LSID="urn:lsid:plazi:treatment:03E087F9FFB0FF99FF0A66980CF9DAC6" httpUri="http://treatment.plazi.org/id/03E087F9FFB0FF99FF0A66980CF9DAC6" lastPageId="5" lastPageNumber="106" pageId="1" pageNumber="102">
<subSubSection id="C3536564FFB0FF9DFF0A66980CD3D91C" box="[136,1204,247,270]" pageId="1" pageNumber="102" type="nomenclature">
<paragraph id="8BF636EFFFB0FF9DFF0A66980CD3D91C" blockId="1.[136,1452,247,666]" box="[136,1204,247,270]" pageId="1" pageNumber="102">
<taxonomicName id="4C494D6CFFB0FF9DFF0A669809D5D91F" ID-CoL="N8B4" authority="(Mitt.) Jaeger" baseAuthorityName="Mitt." box="[136,434,247,270]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="1" pageNumber="102" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB0FF9DFF0A66980943D91F" bold="true" box="[136,292,247,269]" italics="true" pageId="1" pageNumber="102">Braunia plicata</emphasis>
(Mitt.) Jaeger,
</taxonomicName>
Ber. St. Gall. Naturw. Ges. 1874-75: 171. (Ad. 2: 87). 1876. (
<figureCitation id="13722A6AFFB0FF9DFB8166980C38D91F" box="[1027,1119,248,270]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[136,229,2018,2040]" captionStartId-1="2.[136,229,1342,1364]" captionStartId-2="3.[136,229,1279,1301]" captionStartId-3="4.[136,229,1510,1532]" captionTargetBox-0="[186,1404,688,1990]" captionTargetBox-1="[164,1405,196,1305]" captionTargetBox-2="[160,1431,612,1242]" captionTargetBox-3="[155,1433,330,1482]" captionTargetId-0="figure-177@1.[183,1404,688,1990]" captionTargetId-1="figure-29@2.[151,1436,190,1318]" captionTargetId-2="figure-224@3.[151,1436,605,1255]" captionTargetId-3="figure-93@4.[151,1436,317,1485]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURE 1. Lectotype of Braunia plicata. A) Specimen Bridges s. n., 1846, at BM (herb. Hooker). B) Detail of lectotype, showing one capsule. C) Detail of leaves with hyaline apex. D) Leaf apex hyaline and dentate. (Photographs by E. De Luna)." captionText-1="FIGURE 2. Jameson syntype of Braunia plicata. AB) Vegetative leaves in the Jameson syntype. C) Perichaetial leaf in the Jameson syntype. DE) Vegetative leaves characteristic of Braunia cirrhifolia. F. Perichaetial leaf of B. cirrhifolia.AC from Jameson s.n. (syntype of B. plicata, NY-herb. Mitten). DF from Jameson 1 (BM), type for B. cirrhifolia." captionText-2="FIGURE 3. Jameson syntype of Braunia plicata. AB) Leaf cells in the apex and upper leaf, in the Jameson syntype. CD) Leaf cells in the apex and upper leaf, characteristic of B. cirrhifolia. AB from Jameson s.n. (syntype of B. plicata, NY-herb. Mitten). CD from Jameson 1 (BM), type for B. cirrhifolia." captionText-3="FIGURE 4. Gametophytic and sporophytic features of Braunia plicata. AC, IJ) Vegetative leaves. DE) Perichaetial leaves. F, K) Capsules. G) Upper leaf cells. H) Apical leaf cells.AH from lectotype Bridges s.n. (BM-herb-Hooker). IK from Herzog 3703 (JE)." figureDoi-0="http://doi.org/10.5281/zenodo.5900974" figureDoi-1="http://doi.org/10.5281/zenodo.5900976" figureDoi-2="http://doi.org/10.5281/zenodo.5900978" figureDoi-3="http://doi.org/10.5281/zenodo.5900982" httpUri-0="https://zenodo.org/record/5900974/files/figure.png" httpUri-1="https://zenodo.org/record/5900976/files/figure.png" httpUri-2="https://zenodo.org/record/5900978/files/figure.png" httpUri-3="https://zenodo.org/record/5900982/files/figure.png" pageId="1" pageNumber="102">Figs. 14</figureCitation>
, 1525).
</paragraph>
</subSubSection>
<subSubSection id="C3536564FFB0FF9DFF0A677C0929D944" pageId="1" pageNumber="102" type="reference_group">
<paragraph id="8BF636EFFFB0FF9DFF0A677C0929D944" blockId="1.[136,1452,247,666]" pageId="1" pageNumber="102">
Basionym:
<taxonomicName id="4C494D6CFFB0FF9DFF74677C0AEAD923" authority="Mitt., J. Linn. Soc., Bot." authorityName="Mitt., J. Linn. Soc., Bot." box="[246,653,284,306]" class="Bryopsida" family="Hedwigiaceae" genus="Hedwigia" kingdom="Plantae" order="Leucodontales" pageId="1" pageNumber="102" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB0FF9DFF74677C09FAD923" box="[246,413,284,305]" italics="true" pageId="1" pageNumber="102">Hedwigia plicata</emphasis>
Mitt., J. Linn. Soc., Bot.
</taxonomicName>
12: 406-407. 1869. Protologue. “Andes Quitensis,
<emphasis id="B93DEAFDFFB0FF9DFBF8677C0CA9D923" box="[1146,1230,284,305]" italics="true" pageId="1" pageNumber="102">Jameson</emphasis>
. In
<collectingCountry id="F35E767FFFB0FF9DFB77677C0D59D920" box="[1269,1342,284,306]" name="Bolivia" pageId="1" pageNumber="102">Bolivia</collectingCountry>
,
<emphasis id="B93DEAFDFFB0FF9DFACA677C0DF4D923" box="[1352,1427,284,305]" italics="true" pageId="1" pageNumber="102">Bridges</emphasis>
in herb. Hooker”.
</paragraph>
</subSubSection>
<subSubSection id="C3536564FFB0FF9DFF0A67040B24D98C" pageId="1" pageNumber="102" type="materials_examined">
<paragraph id="8BF636EFFFB0FF9DFF0A67040B24D98C" blockId="1.[136,1452,247,666]" pageId="1" pageNumber="102">
<typeStatus id="54F2884DFFB0FF9DFF0A6704088FD96B" box="[136,232,356,377]" pageId="1" pageNumber="102" type="lectotype">Lectotype</typeStatus>
(designated here):—
<collectingCountry id="F35E767FFFB0FF9DFE3667040A7CD968" box="[436,539,356,378]" name="Bolivia" pageId="1" pageNumber="102">BOLIVIA</collectingCountry>
. 1846,
<emphasis id="B93DEAFDFFB0FF9DFDE167040ABFD96B" box="[611,728,356,377]" italics="true" pageId="1" pageNumber="102">Bridges s. n</emphasis>
. (
<typeStatus id="54F2884DFFB0FF9DFD6E67040B36D96B" box="[748,849,356,377]" pageId="1" pageNumber="102" type="lectotype">Lectotype</typeStatus>
: BM! (herb. Hooker-BM) [bc.BM000960770!]; isolectotypes, BM! (herb. Wilson), NY (2)! [bc.NY00913932, bc.NY00913953]).
</paragraph>
</subSubSection>
<subSubSection id="C3536564FFB0FF9DFF0A67CC0A9ED9F4" pageId="1" pageNumber="102" type="reference_group">
<paragraph id="8BF636EFFFB0FF9DFF0A67CC0A9ED9F4" blockId="1.[136,1452,247,666]" pageId="1" pageNumber="102">
<taxonomicName id="4C494D6CFFB0FF9DFF0A67CC0A08D9D0" authority="Card., Rev. Bryol." authorityName="Card., Rev. Bryol." box="[136,623,428,450]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="1" pageNumber="102" phylum="Bryophyta" rank="variety" species="plicata" variety="canescens">
<emphasis id="B93DEAFDFFB0FF9DFF0A67CC0946D9D3" box="[136,289,428,449]" italics="true" pageId="1" pageNumber="102">Braunia plicata</emphasis>
var.
<emphasis id="B93DEAFDFFB0FF9DFED167CC09D3D9D3" box="[339,436,428,449]" italics="true" pageId="1" pageNumber="102">canescens</emphasis>
Card., Rev. Bryol.
</taxonomicName>
38: 38. 1911. Protologue. “Etat de
<collectingRegion id="498DF80DFFB0FF9DFC4E67CC0C7AD9D0" box="[972,1053,428,450]" country="Mexico" name="Hidalgo" pageId="1" pageNumber="102">Hidalgo</collectingRegion>
: Cuyamaloya, rochers (
<emphasis id="B93DEAFDFFB0FF9DFA8767CC0D34D9D3" box="[1285,1363,428,449]" italics="true" pageId="1" pageNumber="102">Pringle,</emphasis>
1908,
<emphasis id="B93DEAFDFFB0FF9DFA1A67CC089ED9F7" italics="true" pageId="1" pageNumber="102">nº 10627</emphasis>
). Etat de
<collectingRegion id="498DF80DFFB0FF9DFED567B009C3D9F4" box="[343,420,464,486]" country="Mexico" name="Mexico" pageId="1" pageNumber="102">Mexico</collectingRegion>
: Toluca (
<emphasis id="B93DEAFDFFB0FF9DFE7967B00A2ED9F7" box="[507,585,464,485]" italics="true" pageId="1" pageNumber="102">Pringle,</emphasis>
1908,
<emphasis id="B93DEAFDFFB0FF9DFD0967B00A86D9F7" box="[651,737,464,485]" italics="true" pageId="1" pageNumber="102">nº 15171</emphasis>
)”.
</paragraph>
</subSubSection>
<subSubSection id="C3536564FFB0FF9DFF0A67940AD3DA88" pageId="1" pageNumber="102" type="materials_examined">
<paragraph id="8BF636EFFFB0FF9DFF0A67940AD3DA88" blockId="1.[136,1452,247,666]" pageId="1" pageNumber="102">
<typeStatus id="54F2884DFFB0FF9DFF0A6794088FDA1B" box="[136,232,500,521]" pageId="1" pageNumber="102" type="lectotype">Lectotype</typeStatus>
(designated here):—
<collectingCountry id="F35E767FFFB0FF9DFE2D67940A75DA18" box="[431,530,500,522]" name="Mexico" pageId="1" pageNumber="102">MEXICO</collectingCountry>
. Toluca,
<date id="FFF7102FFFB0FF9DFDE767940ABBDA1B" box="[613,732,500,521]" pageId="1" pageNumber="102" value="1908-08-04">Aug 4, 1908</date>
,
<emphasis id="B93DEAFDFFB0FF9DFD6767940B09DA1B" box="[741,878,500,521]" italics="true" pageId="1" pageNumber="102">Pringle 15171</emphasis>
(
<typeStatus id="54F2884DFFB0FF9DFCF867940BBBDA1B" box="[890,988,500,521]" pageId="1" pageNumber="102" type="lectotype">Lectotype</typeStatus>
P (herb. Cardot) [bc.PC0701219]; isolectotypes, NY! [bc.NY00792519]; S!, FH!) [
<emphasis id="B93DEAFDFFB0FF9DFD9B64780A0BDA3F" box="[537,620,536,557]" italics="true" pageId="1" pageNumber="102">proparte</emphasis>
, the specimen at FH contains mostly plants of
<taxonomicName id="4C494D6CFFB0FF9DFBD764780CB7DA3F" box="[1109,1232,536,557]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="1" pageNumber="102" phylum="Bryophyta" rank="species" species="andrieuxi">
<emphasis id="B93DEAFDFFB0FF9DFBD764780CB7DA3F" box="[1109,1232,536,557]" italics="true" pageId="1" pageNumber="102">B. andrieuxi</emphasis>
</taxonomicName>
];
<collectingCountry id="F35E767FFFB0FF9DFB6B64780D2CDA3F" box="[1257,1355,536,557]" name="Mexico" pageId="1" pageNumber="102">MEXICO</collectingCountry>
.
<collectingRegion id="498DF80DFFB0FF9DFADA64780DCFDA3C" box="[1368,1448,536,558]" country="Mexico" name="Hidalgo" pageId="1" pageNumber="102">Hidalgo</collectingRegion>
, Cuyamaloya,
<emphasis id="B93DEAFDFFB0FF9DFEC5645C09B2DA43" box="[327,469,572,593]" italics="true" pageId="1" pageNumber="102">Pringle 10627</emphasis>
(BM (2)! [bc. BM000960771, bc.BM000960772], E (2) [bc.E00108810, bc.E00246841], H! [bc. H3301459], JE (2)! [bc.JE04000337, bc.JE04000338], MO! [bc.MO-407560], NY! [bc.NY00792521], P(2) [bc.PC0147304, bc.PC0701217], S!, VT [bc.UVMVT031831],
<collectingCountry id="F35E767FFFB0FF9DFDFC64E40AC3DA88" box="[638,676,644,666]" name="United States of America" pageId="1" pageNumber="102">US</collectingCountry>
!).
</paragraph>
</subSubSection>
<caption id="DF366667FFB0FF9DFF0A61820CDCD009" ID-DOI="http://doi.org/10.5281/zenodo.5900974" ID-Zenodo-Dep="5900974" httpUri="https://zenodo.org/record/5900974/files/figure.png" pageId="1" pageNumber="102" startId="1.[136,229,2018,2040]" targetBox="[186,1404,688,1990]" targetPageId="1">
<paragraph id="8BF636EFFFB0FF9DFF0A61820CDCD009" blockId="1.[136,1452,2018,2076]" pageId="1" pageNumber="102">
<emphasis id="B93DEAFDFFB0FF9DFF0A61820899DFE5" bold="true" box="[136,254,2018,2040]" pageId="1" pageNumber="102">FIGURE 1.</emphasis>
Lectotype of
<taxonomicName id="4C494D6CFFB0FF9DFE0461830A79DFE5" box="[390,542,2018,2040]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="1" pageNumber="102" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB0FF9DFE0461830A79DFE5" box="[390,542,2018,2040]" italics="true" pageId="1" pageNumber="102">Braunia plicata</emphasis>
</taxonomicName>
.
<emphasis id="B93DEAFDFFB0FF9DFDA861820A23DFEA" bold="true" box="[554,580,2018,2040]" pageId="1" pageNumber="102">A)</emphasis>
Specimen
<emphasis id="B93DEAFDFFB0FF9DFD3261820B4EDFEA" box="[688,809,2018,2040]" italics="true" pageId="1" pageNumber="102">Bridges s. n.</emphasis>
, 1846, at BM (herb. Hooker).
<emphasis id="B93DEAFDFFB0FF9DFBD061820C0EDFEA" bold="true" box="[1106,1129,2018,2040]" pageId="1" pageNumber="102">B)</emphasis>
Detail of lectotype, showing one capsule.
<emphasis id="B93DEAFDFFB0FF9DFF5E6E660892D00E" bold="true" box="[220,245,2054,2076]" pageId="1" pageNumber="102">C)</emphasis>
Detail of leaves with hyaline apex.
<emphasis id="B93DEAFDFFB0FF9DFDCD6E660A0FD00E" bold="true" box="[591,616,2054,2076]" pageId="1" pageNumber="102">D)</emphasis>
Leaf apex hyaline and dentate. (Photographs by E. De Luna).
</paragraph>
</caption>
<caption id="DF366667FFB3FF9EFF0A635E0BC2DD8E" ID-DOI="http://doi.org/10.5281/zenodo.5900976" ID-Zenodo-Dep="5900976" httpUri="https://zenodo.org/record/5900976/files/figure.png" pageId="2" pageNumber="103" startId="2.[136,229,1342,1364]" targetBox="[164,1405,196,1305]" targetPageId="2">
<paragraph id="8BF636EFFFB3FF9EFF0A635E0BC2DD8E" blockId="2.[136,1452,1342,1436]" pageId="2" pageNumber="103">
<emphasis id="B93DEAFDFFB3FF9EFF0A635E0898DD41" bold="true" box="[136,255,1342,1364]" pageId="2" pageNumber="103">FIGURE 2.</emphasis>
Jameson syntype of
<taxonomicName id="4C494D6CFFB3FF9EFE4D635F0A0FDD41" box="[463,616,1342,1364]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="2" pageNumber="103" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB3FF9EFE4D635F0A0FDD41" box="[463,616,1342,1364]" italics="true" pageId="2" pageNumber="103">Braunia plicata</emphasis>
</taxonomicName>
.
<emphasis id="B93DEAFDFFB3FF9EFDF7635E0ACCDD46" bold="true" box="[629,683,1342,1364]" pageId="2" pageNumber="103">AB)</emphasis>
Vegetative leaves in the Jameson syntype.
<emphasis id="B93DEAFDFFB3FF9EFBD6635E0C0ADD46" bold="true" box="[1108,1133,1342,1364]" pageId="2" pageNumber="103">C)</emphasis>
Perichaetial leaf in the Jameson syntype.
<emphasis id="B93DEAFDFFB3FF9EFF5C63020974DD6A" bold="true" box="[222,275,1378,1400]" pageId="2" pageNumber="103">DE)</emphasis>
Vegetative leaves characteristic of
<taxonomicName id="4C494D6CFFB3FF9EFDE263030B72DD65" box="[608,789,1378,1400]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="2" pageNumber="103" phylum="Bryophyta" rank="species" species="cirrhifolia">
<emphasis id="B93DEAFDFFB3FF9EFDE263030B72DD65" box="[608,789,1378,1400]" italics="true" pageId="2" pageNumber="103">Braunia cirrhifolia</emphasis>
</taxonomicName>
.
<emphasis id="B93DEAFDFFB3FF9EFCA263020B55DD65" bold="true" box="[800,818,1378,1399]" pageId="2" pageNumber="103">F.</emphasis>
Perichaetial leaf of
<taxonomicName id="4C494D6CFFB3FF9EFC6D63030C0DDD65" box="[1007,1130,1378,1400]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="2" pageNumber="103" phylum="Bryophyta" rank="species" species="cirrhifolia">
<emphasis id="B93DEAFDFFB3FF9EFC6D63030C0DDD65" box="[1007,1130,1378,1400]" italics="true" pageId="2" pageNumber="103">B. cirrhifolia</emphasis>
</taxonomicName>
. AC from
<emphasis id="B93DEAFDFFB3FF9EFB5863030D33DD6A" box="[1242,1364,1379,1400]" italics="true" pageId="2" pageNumber="103">Jameson s.n.</emphasis>
(syntype of
<taxonomicName id="4C494D6CFFB3FF9EFF2063E70967DD89" box="[162,256,1414,1436]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="2" pageNumber="103" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB3FF9EFF2063E70967DD89" box="[162,256,1414,1436]" italics="true" pageId="2" pageNumber="103">B. plicata</emphasis>
</taxonomicName>
, NY-herb. Mitten). DF from
<emphasis id="B93DEAFDFFB3FF9EFDA163E70AEEDD89" box="[547,649,1414,1436]" italics="true" pageId="2" pageNumber="103">Jameson 1</emphasis>
(BM), type for
<taxonomicName id="4C494D6CFFB3FF9EFCA063E70BF8DD89" box="[802,927,1414,1436]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="2" pageNumber="103" phylum="Bryophyta" rank="species" species="cirrhifolia">
<emphasis id="B93DEAFDFFB3FF9EFCA063E70BF8DD89" box="[802,927,1414,1436]" italics="true" pageId="2" pageNumber="103">B. cirrhifolia</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C3536564FFB3FF9FFF0A63AA0CB3DF8C" lastPageId="3" lastPageNumber="104" pageId="2" pageNumber="103" type="discussion">
<paragraph id="8BF636EFFFB3FF9EFF0A63AA0A51DF5F" blockId="2.[136,1452,1482,2085]" pageId="2" pageNumber="103">
<emphasis id="B93DEAFDFFB3FF9EFF0A63AA09E8DDF7" bold="true" box="[136,399,1482,1509]" pageId="2" pageNumber="103">
<emphasis id="B93DEAFDFFB3FF9EFF0A63AA09EFDDF7" bold="true" box="[136,392,1482,1509]" italics="true" pageId="2" pageNumber="103">Nomenclatural note 1</emphasis>
.
</emphasis>
The selection of a specimen collected by Bridges from
<collectingCountry id="F35E767FFFB3FF9EFB9C63AB0C17DDF7" box="[1054,1136,1483,1509]" name="Bolivia" pageId="2" pageNumber="103">Bolivia</collectingCountry>
in 1846 as
<typeStatus id="54F2884DFFB3FF9EFB7863AB0D06DDF7" box="[1274,1377,1483,1509]" pageId="2" pageNumber="103" type="lectotype">lectotype</typeStatus>
for
<taxonomicName id="4C494D6CFFB3FF9EFA1163AB08B1DE1B" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="2" pageNumber="103" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB3FF9EFA1163AB08B1DE1B" italics="true" pageId="2" pageNumber="103">B. plicata</emphasis>
</taxonomicName>
conforms strictly to the species concept in the original description (
<bibRefCitation id="EFD84B1EFFB3FF9EFC5A638F0C0EDE18" author="Mitten, W." box="[984,1129,1519,1546]" pageId="2" pageNumber="103" pagination="1 - 659" refId="ref3887" refString="Mitten, W. (1869) Musci Austro-Americani. Journal of the Linnean Society, Botany 12: 1 - 659. https: // doi. org / 10.1111 / j. 1095 - 8339.1871. tb 00633. x" type="journal article" year="1869">Mitten 1869</bibRefCitation>
). The original description of
<taxonomicName id="4C494D6CFFB3FF9EFF0A6073092BDE3F" box="[136,332,1555,1581]" class="Bryopsida" family="Hedwigiaceae" genus="Hedwigia" kingdom="Plantae" order="Leucodontales" pageId="2" pageNumber="103" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB3FF9EFF0A6073092BDE3F" box="[136,332,1555,1581]" italics="true" pageId="2" pageNumber="103">Hedwigia plicata</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFD84B1EFFB3FF9EFEF660730A75DE3C" author="Mitten, W." box="[372,530,1555,1582]" pageId="2" pageNumber="103" pagination="1 - 659" refId="ref3887" refString="Mitten, W. (1869) Musci Austro-Americani. Journal of the Linnean Society, Botany 12: 1 - 659. https: // doi. org / 10.1111 / j. 1095 - 8339.1871. tb 00633. x" type="journal article" year="1869">Mitten (1869)</bibRefCitation>
reads: “Folia erecto-patentia, imbricata, latissime ovalia, concava, cuspide brevi interdum albescente terminata, quater plicata, margine late reflexo apice serrulato, … “. These leaf features constitute the core of the species concept of
<emphasis id="B93DEAFDFFB3FF9EFD85603B0A1CDE67" box="[519,635,1627,1653]" italics="true" pageId="2" pageNumber="103">
<taxonomicName id="4C494D6CFFB3FF9EFD85603B0A10DE67" box="[519,631,1627,1653]" class="Bryopsida" family="Hedwigiaceae" genus="Hedwigia" kingdom="Plantae" order="Leucodontales" pageId="2" pageNumber="103" phylum="Bryophyta" rank="species" species="plicata">B. plicata</taxonomicName>
,
</emphasis>
as recognized here. Only the plants in Bridges specimens display leaves widely ovate, leaf acumen hyaline, and leaf margin at apex serrulate to dentate (
<figureCitation id="13722A6AFFB3FF9EFB98601F0C3ADE8B" box="[1050,1117,1663,1689]" captionStart="FIGURE 1" captionStartId="1.[136,229,2018,2040]" captionTargetBox="[186,1404,688,1990]" captionTargetId="figure-177@1.[183,1404,688,1990]" captionTargetPageId="1" captionText="FIGURE 1. Lectotype of Braunia plicata. A) Specimen Bridges s. n., 1846, at BM (herb. Hooker). B) Detail of lectotype, showing one capsule. C) Detail of leaves with hyaline apex. D) Leaf apex hyaline and dentate. (Photographs by E. De Luna)." figureDoi="http://doi.org/10.5281/zenodo.5900974" httpUri="https://zenodo.org/record/5900974/files/figure.png" pageId="2" pageNumber="103">Fig. 1</figureCitation>
). I have seen
<specimenCount id="9D4FFD66FFB3FF9EFA82601F0DCBDE8B" box="[1280,1452,1663,1689]" pageId="2" pageNumber="103" type="generic">four specimens</specimenCount>
of the Bridges collection. There are two duplicates at BM (
<emphasis id="B93DEAFDFFB3FF9EFCB260C30BA7DEAE" box="[816,960,1699,1725]" italics="true" pageId="2" pageNumber="103">Bridges s. n.</emphasis>
BM-herb. Hooker;
<emphasis id="B93DEAFDFFB3FF9EFB2760C30D52DEAE" box="[1189,1333,1699,1725]" italics="true" pageId="2" pageNumber="103">Bridges s. n.</emphasis>
BM-herb. Wilson) and two duplicates at NY (
<emphasis id="B93DEAFDFFB3FF9EFD9C60A70ACADEF2" box="[542,685,1735,1761]" italics="true" pageId="2" pageNumber="103">Bridges s. n.</emphasis>
NY-herb. Mitten). These
<specimenCount id="9D4FFD66FFB3FF9EFC5660A70C1BDEF3" box="[980,1148,1735,1761]" pageId="2" pageNumber="103" type="generic">two specimens</specimenCount>
at NY consist of only few stems glued on a card (bc.NY00913932, bc.NY00913953). The specimen at BM-herb Hooker (bc.BM000960770) is selected here as
<typeStatus id="54F2884DFFB3FF9EFEBC616F09C2DF3B" box="[318,421,1807,1833]" pageId="2" pageNumber="103" type="lectotype">lectotype</typeStatus>
(
<figureCitation id="13722A6AFFB3FF9EFE31616F0993DF3B" box="[435,500,1807,1833]" captionStart="FIGURE 1" captionStartId="1.[136,229,2018,2040]" captionTargetBox="[186,1404,688,1990]" captionTargetId="figure-177@1.[183,1404,688,1990]" captionTargetPageId="1" captionText="FIGURE 1. Lectotype of Braunia plicata. A) Specimen Bridges s. n., 1846, at BM (herb. Hooker). B) Detail of lectotype, showing one capsule. C) Detail of leaves with hyaline apex. D) Leaf apex hyaline and dentate. (Photographs by E. De Luna)." figureDoi="http://doi.org/10.5281/zenodo.5900974" httpUri="https://zenodo.org/record/5900974/files/figure.png" pageId="2" pageNumber="103">Fig. 1</figureCitation>
), because the type citation makes clear reference to a particular specimen as follows: “In
<collectingCountry id="F35E767FFFB3FF9EFF316153096FDF5F" box="[179,264,1843,1869]" name="Bolivia" pageId="2" pageNumber="103">Bolivia</collectingCountry>
,
<emphasis id="B93DEAFDFFB3FF9EFE916153090DDF5F" box="[275,362,1843,1869]" italics="true" pageId="2" pageNumber="103">Bridges</emphasis>
in herb. Hooker”.
</paragraph>
<paragraph id="8BF636EFFFB3FF9FFF3F61360A2FDA57" blockId="2.[136,1452,1482,2085]" lastBlockId="3.[136,1452,158,581]" lastPageId="3" lastPageNumber="104" pageId="2" pageNumber="103">
<emphasis id="B93DEAFDFFB3FF9EFF3F613609A3DF63" bold="true" box="[189,452,1878,1905]" italics="true" pageId="2" pageNumber="103">Nomenclatural note 2.</emphasis>
<bibRefCitation id="EFD84B1EFFB3FF9EFE4F61370A0CDF60" author="Mitten, W." box="[461,619,1879,1906]" pageId="2" pageNumber="103" pagination="1 - 659" refId="ref3887" refString="Mitten, W. (1869) Musci Austro-Americani. Journal of the Linnean Society, Botany 12: 1 - 659. https: // doi. org / 10.1111 / j. 1095 - 8339.1871. tb 00633. x" type="journal article" year="1869">Mitten (1869)</bibRefCitation>
cited jointly in the protologue of
<taxonomicName id="4C494D6CFFB3FF9EFC7261370C38DF63" box="[1008,1119,1879,1905]" class="Bryopsida" family="Hedwigiaceae" genus="Hedwigia" kingdom="Plantae" order="Leucodontales" pageId="2" pageNumber="103" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB3FF9EFC7261370C38DF63" box="[1008,1119,1879,1905]" italics="true" pageId="2" pageNumber="103">B. plicata</emphasis>
</taxonomicName>
a specimen from
<collectingCountry id="F35E767FFFB3FF9EFAAC61370DECDF63" box="[1326,1419,1879,1905]" name="Ecuador" pageId="2" pageNumber="103">Ecuador</collectingCountry>
as “Andes quitenses,
<emphasis id="B93DEAFDFFB3FF9EFED8611B09DBDF86" box="[346,444,1915,1940]" italics="true" pageId="2" pageNumber="103">Jameson</emphasis>
”. This
<typeStatus id="54F2884DFFB3FF9EFD8D611B0A01DF87" box="[527,614,1915,1941]" pageId="2" pageNumber="103" type="syntype">syntype</typeStatus>
is difficult to discern among several specimens collected by Jameson, due to incomplete collection data. Two duplicates at NY-herb. Mitten with herbarium labels “Andes Quitensis,
<emphasis id="B93DEAFDFFB3FF9EFACB61FF08D8DFCF" italics="true" pageId="2" pageNumber="103">Jameson s.n.”</emphasis>
, no year, are very likely this
<typeStatus id="54F2884DFFB3FF9EFE7B61A30A37DFCF" box="[505,592,1987,2013]" pageId="2" pageNumber="103" type="syntype">syntype</typeStatus>
(bc.NY0001244170, bc.NY0001244171). Plants in these Jameson specimens do not display the features of
<taxonomicName id="4C494D6CFFB3FF9EFE6761870A32D013" box="[485,597,2023,2049]" class="Bryopsida" family="Hedwigiaceae" genus="Hedwigia" kingdom="Plantae" order="Leucodontales" pageId="2" pageNumber="103" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB3FF9EFE6761870A32D013" box="[485,597,2023,2049]" italics="true" pageId="2" pageNumber="103">B. plicata</emphasis>
</taxonomicName>
, such as obovate leaves, revolute leaf margins, hyaline leaf acumens, and dentate leaf apices. Instead, these Jameson
<typeStatus id="54F2884DFFB3FF9EFDED6E6B0AB6D037" box="[623,721,2059,2085]" pageId="2" pageNumber="103" type="syntype">syntypes</typeStatus>
have the features of
<taxonomicName id="4C494D6CFFB3FF9EFC3C6E6B0C36D037" box="[958,1105,2059,2085]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="2" pageNumber="103" phylum="Bryophyta" rank="species" species="cirrhifolia">
<emphasis id="B93DEAFDFFB3FF9EFC3C6E6B0C36D037" box="[958,1105,2059,2085]" italics="true" pageId="2" pageNumber="103">B. cirrhifolia</emphasis>
</taxonomicName>
, namely, leaves are long ovate lanceolate, leaf margins reflexed at the base, leaf acumens are concolorous, and the leaf apex is entire (
<figureCitation id="13722A6AFFB2FF9FFA9466FE0D3BD8AA" box="[1302,1372,158,184]" captionStart="FIGURE 2" captionStartId="2.[136,229,1342,1364]" captionTargetBox="[164,1405,196,1305]" captionTargetId="figure-29@2.[151,1436,190,1318]" captionTargetPageId="2" captionText="FIGURE 2. Jameson syntype of Braunia plicata. AB) Vegetative leaves in the Jameson syntype. C) Perichaetial leaf in the Jameson syntype. DE) Vegetative leaves characteristic of Braunia cirrhifolia. F. Perichaetial leaf of B. cirrhifolia.AC from Jameson s.n. (syntype of B. plicata, NY-herb. Mitten). DF from Jameson 1 (BM), type for B. cirrhifolia." figureDoi="http://doi.org/10.5281/zenodo.5900976" httpUri="https://zenodo.org/record/5900976/files/figure.png" pageId="3" pageNumber="104">Fig. 2</figureCitation>
). Also, the upper leaf cells are long and sinuous, just like in
<taxonomicName id="4C494D6CFFB2FF9FFD6966A30BE7D8CE" box="[747,896,194,220]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="cirrhifolia">
<emphasis id="B93DEAFDFFB2FF9FFD6966A30BE7D8CE" box="[747,896,194,220]" italics="true" pageId="3" pageNumber="104">B. cirrhifolia</emphasis>
</taxonomicName>
(
<figureCitation id="13722A6AFFB2FF9FFC1266A20BB0D8CE" box="[912,983,194,220]" captionStart="FIGURE 3" captionStartId="3.[136,229,1279,1301]" captionTargetBox="[160,1431,612,1242]" captionTargetId="figure-224@3.[151,1436,605,1255]" captionTargetPageId="3" captionText="FIGURE 3. Jameson syntype of Braunia plicata. AB) Leaf cells in the apex and upper leaf, in the Jameson syntype. CD) Leaf cells in the apex and upper leaf, characteristic of B. cirrhifolia. AB from Jameson s.n. (syntype of B. plicata, NY-herb. Mitten). CD from Jameson 1 (BM), type for B. cirrhifolia." figureDoi="http://doi.org/10.5281/zenodo.5900978" httpUri="https://zenodo.org/record/5900978/files/figure.png" pageId="3" pageNumber="104">Fig. 3</figureCitation>
). Additionally, I have seen the following Jameson specimens from
<collectingCountry id="F35E767FFFB2FF9FFE2B66860A6FD912" box="[425,520,230,256]" name="Ecuador" pageId="3" pageNumber="104">Ecuador</collectingCountry>
, which are also
<taxonomicName id="4C494D6CFFB2FF9FFD3F66870B28D912" box="[701,847,230,256]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="cirrhifolia">
<emphasis id="B93DEAFDFFB2FF9FFD3F66870B28D912" box="[701,847,230,256]" italics="true" pageId="3" pageNumber="104">B. cirrhifolia</emphasis>
</taxonomicName>
(ordered chronologically): Quito,
<emphasis id="B93DEAFDFFB2FF9FFB5066870D0BD912" box="[1234,1388,230,256]" italics="true" pageId="3" pageNumber="104">Jameson 135,</emphasis>
1845 (BM!-herb. Wilson, BM!-herb. Hooker, G!),
<emphasis id="B93DEAFDFFB2FF9FFD07676B0B71D936" box="[645,790,267,292]" italics="true" pageId="3" pageNumber="104">Jameson s.n.</emphasis>
, 1846 (BM!- herb. Wilson, BM!-herb. Hooker, BM!-herb. Dixon),
<emphasis id="B93DEAFDFFB2FF9FFF64674F091AD95A" box="[230,381,303,328]" italics="true" pageId="3" pageNumber="104">Jameson s.n.,</emphasis>
1847 (BM!- herb. Dixon),
<emphasis id="B93DEAFDFFB2FF9FFD30674F0B4DD95A" box="[690,810,302,328]" italics="true" pageId="3" pageNumber="104">Jameson 1</emphasis>
, 1847 (BM!-herb. Hooker, BR!, MICH!, NY!),
<emphasis id="B93DEAFDFFB2FF9FFACB674F08C3D97E" italics="true" pageId="3" pageNumber="104">Jameson 25</emphasis>
, 1847 (BM!-herb. Lyon),
<emphasis id="B93DEAFDFFB2FF9FFE4867330A34D97E" box="[458,595,339,364]" italics="true" pageId="3" pageNumber="104">Jameson s.n</emphasis>
., 1848 (G!), and
<emphasis id="B93DEAFDFFB2FF9FFC9467330BF8D97E" box="[790,927,339,364]" italics="true" pageId="3" pageNumber="104">Jameson s.n</emphasis>
. 1851 (G!). Moreover, there are images online of a specimen at P (bc.PC0701220) labeled “
<emphasis id="B93DEAFDFFB2FF9FFD0467170B77D982" box="[646,784,375,400]" italics="true" pageId="3" pageNumber="104">Jameson s.n</emphasis>
. Quito, Mitten Herbarium, NYBG”. This specimen is very likely
<taxonomicName id="4C494D6CFFB2FF9FFF4D67FB0905D9A6" box="[207,354,410,436]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="cirrhifolia">
<emphasis id="B93DEAFDFFB2FF9FFF4D67FB0905D9A6" box="[207,354,410,436]" italics="true" pageId="3" pageNumber="104">B. cirrhifolia</emphasis>
</taxonomicName>
since it consists of one very large stem (over
<quantity id="4CB19B0AFFB2FF9FFCEA67FA0BC7D9A7" box="[872,928,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="104" unit="cm" value="4.0">4 cm</quantity>
long), leaves with long concolorous acumens, and sporophytes with a very long seta (
<quantity id="4CB19B0AFFB2FF9FFDC767DE0AE4D9CB" box="[581,643,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="104" unit="cm" value="2.0">2 cm</quantity>
). In conclusion, all Jameson specimens I have seen actually correspond to
<taxonomicName id="4C494D6CFFB2FF9FFF246783095ED9EE" box="[166,313,482,508]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="cirrhifolia">
<emphasis id="B93DEAFDFFB2FF9FFF246783095ED9EE" box="[166,313,482,508]" italics="true" pageId="3" pageNumber="104">B. cirrhifolia</emphasis>
</taxonomicName>
. Therefore, the Jameson
<typeStatus id="54F2884DFFB2FF9FFDDA67830AC8D9EF" box="[600,687,483,509]" pageId="3" pageNumber="104" type="syntype">syntype</typeStatus>
cited in the protologue cannot possibly be selected as
<typeStatus id="54F2884DFFB2FF9FFA9E67820DE4D9EE" box="[1308,1411,482,508]" pageId="3" pageNumber="104" type="lectotype">lectotype</typeStatus>
for
<taxonomicName id="4C494D6CFFB2FF9FFF0A64670891DA32" box="[136,246,518,544]" class="Bryopsida" family="Hedwigiaceae" genus="Hedwigia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB2FF9FFF0A64670891DA32" box="[136,246,518,544]" italics="true" pageId="3" pageNumber="104">B. plicata</emphasis>
</taxonomicName>
. The identity of the
<typeStatus id="54F2884DFFB2FF9FFE5C64670A52DA33" box="[478,565,519,545]" pageId="3" pageNumber="104" type="syntype">syntype</typeStatus>
“Andes Quitensis,
<emphasis id="B93DEAFDFFB2FF9FFC9064670BCEDA32" box="[786,937,519,544]" italics="true" pageId="3" pageNumber="104">Jameson s.n.,</emphasis>
no year, NY!-herb. Mitten (bc.NY01244170, bc.NY01244171) is
<taxonomicName id="4C494D6CFFB2FF9FFEEE644B0A25DA56" box="[364,578,554,580]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="cirrhifolia">
<emphasis id="B93DEAFDFFB2FF9FFEEE644B0A25DA56" box="[364,578,554,580]" italics="true" pageId="3" pageNumber="104">Braunia cirrhifolia</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF366667FFB2FF9FFF0A629F0A6DDD4F" ID-DOI="http://doi.org/10.5281/zenodo.5900978" ID-Zenodo-Dep="5900978" httpUri="https://zenodo.org/record/5900978/files/figure.png" pageId="3" pageNumber="104" startId="3.[136,229,1279,1301]" targetBox="[160,1431,612,1242]" targetPageId="3">
<paragraph id="8BF636EFFFB2FF9FFF0A629F0A6DDD4F" blockId="3.[136,1452,1279,1374]" pageId="3" pageNumber="104">
<emphasis id="B93DEAFDFFB2FF9FFF0A629F0898DD06" bold="true" box="[136,255,1279,1301]" pageId="3" pageNumber="104">FIGURE 3.</emphasis>
Jameson syntype of
<taxonomicName id="4C494D6CFFB2FF9FFE4E63600A02DD06" box="[460,613,1279,1301]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB2FF9FFE4E63600A02DD06" box="[460,613,1279,1301]" italics="true" pageId="3" pageNumber="104">Braunia plicata</emphasis>
</taxonomicName>
.
<emphasis id="B93DEAFDFFB2FF9FFDF0629F0AC0DD07" bold="true" box="[626,679,1279,1301]" pageId="3" pageNumber="104">AB)</emphasis>
Leaf cells in the apex and upper leaf, in the Jameson syntype.
<emphasis id="B93DEAFDFFB2FF9FFA8D629F0D22DD07" bold="true" box="[1295,1349,1279,1301]" pageId="3" pageNumber="104">CD)</emphasis>
Leaf cells in the apex and upper leaf, characteristic of
<taxonomicName id="4C494D6CFFB2FF9FFDBA63440AD1DD2A" box="[568,694,1315,1337]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="cirrhifolia">
<emphasis id="B93DEAFDFFB2FF9FFDBA63440AD1DD2A" box="[568,694,1315,1337]" italics="true" pageId="3" pageNumber="104">B. cirrhifolia</emphasis>
</taxonomicName>
. AB from
<emphasis id="B93DEAFDFFB2FF9FFCAF63440BCDDD2B" box="[813,938,1316,1337]" italics="true" pageId="3" pageNumber="104">Jameson s.n.</emphasis>
(syntype of
<taxonomicName id="4C494D6CFFB2FF9FFBA463440CE2DD2A" box="[1062,1157,1315,1337]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB2FF9FFBA463440CE2DD2A" box="[1062,1157,1315,1337]" italics="true" pageId="3" pageNumber="104">B. plicata</emphasis>
</taxonomicName>
, NY-herb. Mitten). CD from
<emphasis id="B93DEAFDFFB2FF9FFF0A63280889DD4F" box="[136,238,1352,1373]" italics="true" pageId="3" pageNumber="104">Jameson 1</emphasis>
(BM), type for
<taxonomicName id="4C494D6CFFB2FF9FFE0563280A64DD4E" box="[391,515,1351,1373]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="cirrhifolia">
<emphasis id="B93DEAFDFFB2FF9FFE0563280A64DD4E" box="[391,515,1351,1373]" italics="true" pageId="3" pageNumber="104">B. cirrhifolia</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BF636EFFFB2FF9FFF3F63EB0CB1DE48" blockId="3.[136,1452,1419,2058]" pageId="3" pageNumber="104">
<emphasis id="B93DEAFDFFB2FF9FFF3F63EB09D8DDB4" bold="true" box="[189,447,1419,1446]" italics="true" pageId="3" pageNumber="104">Nomenclatural note 3.</emphasis>
<bibRefCitation id="EFD84B1EFFB2FF9FFE4963EC0A00DDB4" author="Cardot, J." box="[459,615,1420,1446]" pageId="3" pageNumber="104" pagination="33 - 43" refId="ref3734" refString="Cardot, J. (1911) Diagnoses preliminaires de mousses mexicaines (9 e article). Revue Bryologique 38: 33 - 43." type="journal article" year="1911">Cardot (1911)</bibRefCitation>
cited
<specimenCount id="9D4FFD66FFB2FF9FFD2863EC0B28DDB4" box="[682,847,1420,1446]" pageId="3" pageNumber="104" type="generic">two specimens</specimenCount>
in the description of
<taxonomicName id="4C494D6CFFB2FF9FFBB863ED0DCCDDB4" authority=": Pringle" authorityName="Pringle" box="[1082,1451,1420,1446]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="variety" species="plicata" variety="canescens">
<emphasis id="B93DEAFDFFB2FF9FFBB863ED0CC1DDB4" box="[1082,1190,1420,1446]" italics="true" pageId="3" pageNumber="104">B. plicata</emphasis>
var
<emphasis id="B93DEAFDFFB2FF9FFB5463ED0DCCDDB4" box="[1238,1451,1420,1446]" italics="true" pageId="3" pageNumber="104">canescens: Pringle</emphasis>
</taxonomicName>
<emphasis id="B93DEAFDFFB2FF9FFF0A63D008A9DDD8" box="[136,206,1456,1482]" italics="true" pageId="3" pageNumber="104">15171</emphasis>
and
<emphasis id="B93DEAFDFFB2FF9FFE8663D009C2DDD8" box="[260,421,1456,1482]" italics="true" pageId="3" pageNumber="104">Pringle 10627</emphasis>
. I have seen several duplicates of both collections. The specimen
<emphasis id="B93DEAFDFFB2FF9FFB0C63D00D48DDD8" box="[1166,1327,1456,1482]" italics="true" pageId="3" pageNumber="104">Pringle 10627</emphasis>
should not be considered as
<typeStatus id="54F2884DFFB2FF9FFEC863B409D6DDFC" box="[330,433,1492,1518]" pageId="3" pageNumber="104" type="lectotype">lectotype</typeStatus>
since most duplicates consist of mixtures of different
<taxonomicName id="4C494D6CFFB2FF9FFB9763B50CAEDDFC" box="[1045,1225,1492,1518]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="undetermined">
<emphasis id="B93DEAFDFFB2FF9FFB9763B50C16DDFC" box="[1045,1137,1493,1518]" italics="true" pageId="3" pageNumber="104">Braunia</emphasis>
species
</taxonomicName>
(see Nomenclatural note 4). Only the selection of
<emphasis id="B93DEAFDFFB2FF9FFE6A63980AEBDE00" box="[488,652,1528,1554]" italics="true" pageId="3" pageNumber="104">Pringle 15171</emphasis>
(P-herb. Cardot) as the
<typeStatus id="54F2884DFFB2FF9FFC2B63980C77DE00" box="[937,1040,1528,1554]" pageId="3" pageNumber="104" type="lectotype">lectotype</typeStatus>
preserves the identity of this taxon without confusion. The duplicates
<emphasis id="B93DEAFDFFB2FF9FFD8D607C0AD6DE24" box="[527,689,1564,1590]" italics="true" pageId="3" pageNumber="104">Pringle 15171</emphasis>
at NY! and S! are good isolectotypes. But the specimen of
<emphasis id="B93DEAFDFFB2FF9FFAD5607C08A9DE48" italics="true" pageId="3" pageNumber="104">Pringle 15171</emphasis>
at FH contains mostly plants of
<taxonomicName id="4C494D6CFFB2FF9FFDB960210AAADE48" authorityName="Lorentz" authorityYear="1864" box="[571,717,1600,1626]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="andrieuxii">
<emphasis id="B93DEAFDFFB2FF9FFDB960210AAADE48" box="[571,717,1600,1626]" italics="true" pageId="3" pageNumber="104">B. andrieuxii</emphasis>
</taxonomicName>
, which I separated in an envelope with a note.
</paragraph>
<paragraph id="8BF636EFFFB2FF9FFF3F60030CB3DF8C" blockId="3.[136,1452,1419,2058]" pageId="3" pageNumber="104">
<emphasis id="B93DEAFDFFB2FF9FFF3F600309A3DE6C" box="[189,452,1635,1662]" italics="true" pageId="3" pageNumber="104">
<emphasis id="B93DEAFDFFB2FF9FFF3F600309DADE6C" bold="true" box="[189,445,1635,1662]" italics="true" pageId="3" pageNumber="104">Nomenclatural note 4</emphasis>
.
</emphasis>
The
<typeStatus id="54F2884DFFB2FF9FFD8860040A06DE6C" box="[522,609,1636,1662]" pageId="3" pageNumber="104" type="syntype">syntype</typeStatus>
specimen
<emphasis id="B93DEAFDFFB2FF9FFD5F60040BE7DE6C" box="[733,896,1636,1662]" italics="true" pageId="3" pageNumber="104">Pringle 10627</emphasis>
of
<taxonomicName id="4C494D6CFFB2FF9FFC2B60050CD8DE6C" box="[937,1215,1636,1662]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="variety" species="plicata" variety="canescens">
<emphasis id="B93DEAFDFFB2FF9FFC2B60050C7FDE6C" box="[937,1048,1636,1662]" italics="true" pageId="3" pageNumber="104">B. plicata</emphasis>
var
<emphasis id="B93DEAFDFFB2FF9FFBCF60050CD8DE6C" box="[1101,1215,1637,1662]" italics="true" pageId="3" pageNumber="104">canescens</emphasis>
</taxonomicName>
is problematic since most duplicates consist of mixtures of up to three species:
<taxonomicName id="4C494D6CFFB2FF9FFC9E60E90BEEDEB0" box="[796,905,1672,1698]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB2FF9FFC9E60E90BEEDEB0" box="[796,905,1672,1698]" italics="true" pageId="3" pageNumber="104">B. plicata</emphasis>
</taxonomicName>
,
<emphasis id="B93DEAFDFFB2FF9FFC1A60E90C56DEB0" box="[920,1073,1672,1698]" italics="true" pageId="3" pageNumber="104">
<taxonomicName id="4C494D6CFFB2FF9FFC1A60E90C4ADEB0" authorityName="Lorentz" authorityYear="1864" box="[920,1069,1672,1698]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="andrieuxii">B. andrieuxii</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C494D6CFFB2FF9FFBEB60E90C83DEB0" box="[1129,1252,1672,1698]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="secunda">
<emphasis id="B93DEAFDFFB2FF9FFBEB60E90C83DEB0" box="[1129,1252,1672,1698]" italics="true" pageId="3" pageNumber="104">B. secunda</emphasis>
</taxonomicName>
. Notably,
<bibRefCitation id="EFD84B1EFFB2FF9FFADB60E808B4DED5" author="Theriot, E." pageId="3" pageNumber="104" pagination="1 - 29" refId="ref3929" refString="Theriot, E. (1926) Mexican mosses collected by Brother Arsene Brouard. Smithsonian Miscellaneous Collections 78: 1 - 29." type="journal article" year="1926">Thériot (1926)</bibRefCitation>
had already warned that not all duplicates of
<emphasis id="B93DEAFDFFB2FF9FFD5860CC0B1BDED4" box="[730,892,1708,1734]" italics="true" pageId="3" pageNumber="104">Pringle 10627</emphasis>
were really
<taxonomicName id="4C494D6CFFB2FF9FFB8860CD0D7ADED4" box="[1034,1309,1708,1734]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="variety" species="plicata" variety="canescens">
<emphasis id="B93DEAFDFFB2FF9FFB8860CD0C1FDED4" box="[1034,1144,1708,1734]" italics="true" pageId="3" pageNumber="104">B. plicata</emphasis>
var
<emphasis id="B93DEAFDFFB2FF9FFB2960CD0D7ADED4" box="[1195,1309,1709,1734]" italics="true" pageId="3" pageNumber="104">canescens</emphasis>
</taxonomicName>
. Only seven duplicates of
<emphasis id="B93DEAFDFFB2FF9FFE9E60B009DBDEF8" box="[284,444,1744,1770]" italics="true" pageId="3" pageNumber="104">Pringle 10627</emphasis>
(BM!, H!, JE!, MO!, NY! [bc.00792521], S!,
<collectingCountry id="F35E767FFFB2FF9FFC3860B00B83DEF8" box="[954,996,1744,1770]" name="United States of America" pageId="3" pageNumber="104">US</collectingCountry>
!) are authentic
<taxonomicName id="4C494D6CFFB2FF9FFB0860B10C91DEF8" box="[1162,1270,1744,1770]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB2FF9FFB0860B10C91DEF8" box="[1162,1270,1744,1770]" italics="true" pageId="3" pageNumber="104">B. plicata</emphasis>
</taxonomicName>
. But I have seen other eight duplicates, which are misidentified;
<specimenCount id="9D4FFD66FFB2FF9FFD1560940B5ADF1C" box="[663,829,1780,1806]" pageId="3" pageNumber="104" type="generic">five specimens</specimenCount>
(FH!, G!, NSW!, S!, B!) are
<taxonomicName id="4C494D6CFFB2FF9FFBFC60950D68DF1C" authorityName="Lorentz" authorityYear="1864" box="[1150,1295,1780,1806]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="andrieuxii">
<emphasis id="B93DEAFDFFB2FF9FFBFC60950D68DF1C" box="[1150,1295,1780,1806]" italics="true" pageId="3" pageNumber="104">B. andrieuxii</emphasis>
</taxonomicName>
, and the other three duplicates (MICH!, NY!, JE! [bc.JE04000338]) are
<taxonomicName id="4C494D6CFFB2FF9FFC9561790BF4DF20" box="[791,915,1816,1842]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="secunda">
<emphasis id="B93DEAFDFFB2FF9FFC9561790BF4DF20" box="[791,915,1816,1842]" italics="true" pageId="3" pageNumber="104">B. secunda</emphasis>
</taxonomicName>
. Additionally, one of the duplicates of
<emphasis id="B93DEAFDFFB2FF9FFAD5617808A9DF44" italics="true" pageId="3" pageNumber="104">Pringle 10627</emphasis>
at P (bc.PC0147304) is certainly not genuine
<taxonomicName id="4C494D6CFFB2FF9FFD4D615D0B5BDF44" box="[719,828,1852,1878]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB2FF9FFD4D615D0B5BDF44" box="[719,828,1852,1878]" italics="true" pageId="3" pageNumber="104">B. plicata</emphasis>
</taxonomicName>
. The images available at the MNHN web site show leaf features that correspond most likely to
<taxonomicName id="4C494D6CFFB2FF9FFDC361010AABDF68" box="[577,716,1888,1914]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="andrieuxi">
<emphasis id="B93DEAFDFFB2FF9FFDC361010AABDF68" box="[577,716,1888,1914]" italics="true" pageId="3" pageNumber="104">B. andrieuxi</emphasis>
</taxonomicName>
. The images online in the same web site for two other duplicates (bc.PC0701217, bc.PC0701218) show plants, with hyaline leaf apices, which are likely
<taxonomicName id="4C494D6CFFB2FF9FFBE361E50CA9DF8C" box="[1121,1230,1924,1950]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="3" pageNumber="104" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB2FF9FFBE361E50CA9DF8C" box="[1121,1230,1924,1950]" italics="true" pageId="3" pageNumber="104">B. plicata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3536564FFB2FF98FF3F61C80AF4DF0E" lastPageId="4" lastPageNumber="105" pageId="3" pageNumber="104" type="description">
<paragraph id="8BF636EFFFB2FF98FF3F61C80B17D936" blockId="3.[136,1452,1419,2058]" lastBlockId="4.[136,1452,158,293]" lastPageId="4" lastPageNumber="105" pageId="3" pageNumber="104">
Plants in dense mats; main stems sympodial, plagiotropic, 3(45)
<quantity id="4CB19B0AFFB2FF9FFC6261C80C7BDFD0" box="[992,1052,1960,1987]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="104" unit="cm" value="7.0">7 cm</quantity>
long, closely branched;
<emphasis id="B93DEAFDFFB2FF9FFAC761C80A07DFF4" italics="true" pageId="3" pageNumber="104">branches elongated, ending in obtuse, rosulate tips</emphasis>
; flagelliform branches common; pseudoparaphylia foliose, with papillose cells. Leaves ovate or broadly ovate,
<quantity id="4CB19B0AFFB2FF9FFDA961900AD5D018" box="[555,690,2032,2058]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="3" pageNumber="104" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.92.2 mm</quantity>
long,
<quantity id="4CB19B0AFFB2FF9FFD7B61900BE7D018" box="[761,896,2032,2058]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="3" pageNumber="104" unit="mm" value="0.95" valueMax="1.1" valueMin="0.8">0.81.1 mm</quantity>
wide, concave, imbricate, spreading when moist,
<emphasis id="B93DEAFDFFB5FF98FF0A66FE095ED8AA" box="[136,313,158,184]" italics="true" pageId="4" pageNumber="105">strongly plicate</emphasis>
; margins recurved to acumen;
<emphasis id="B93DEAFDFFB5FF98FD2666FE0C8DD8AA" box="[676,1258,158,184]" italics="true" pageId="4" pageNumber="105">high base, widest point at or above middle of leaf</emphasis>
; apex cuspidate,
<emphasis id="B93DEAFDFFB5FF98FF0A66A2099ED8CE" box="[136,505,194,220]" italics="true" pageId="4" pageNumber="105">hyaline, margins sharply dentate</emphasis>
; apical leaf cells homogeneous,
<emphasis id="B93DEAFDFFB5FF98FCE966A20CE8D8CE" box="[875,1167,194,220]" italics="true" pageId="4" pageNumber="105">elongated, narrow elliptic</emphasis>
(57:1), 1828 µm long, not sinuose, with marginal papillae; upper and median leaf cells 1418 µm long, sub-quadrate to short elliptic (2:1), walls sinuose, papillae 2-3, low, on side walls, overarching lumen.
</paragraph>
<caption id="DF366667FFB5FF98FF0A63860DEDDE32" ID-DOI="http://doi.org/10.5281/zenodo.5900982" ID-Zenodo-Dep="5900982" httpUri="https://zenodo.org/record/5900982/files/figure.png" pageId="4" pageNumber="105" startId="4.[136,229,1510,1532]" targetBox="[155,1433,330,1482]" targetPageId="4">
<paragraph id="8BF636EFFFB5FF98FF0A63860DEDDE32" blockId="4.[136,1451,1510,1568]" pageId="4" pageNumber="105">
<emphasis id="B93DEAFDFFB5FF98FF0A63860898DDE9" bold="true" box="[136,255,1510,1532]" pageId="4" pageNumber="105">FIGURE 4.</emphasis>
Gametophytic and sporophytic features of
<taxonomicName id="4C494D6CFFB5FF98FD2863870B23DDE9" box="[682,836,1510,1532]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="4" pageNumber="105" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB5FF98FD2863870B23DDE9" box="[682,836,1510,1532]" italics="true" pageId="4" pageNumber="105">Braunia plicata</emphasis>
</taxonomicName>
.
<emphasis id="B93DEAFDFFB5FF98FCD363860BDFDDEE" bold="true" box="[849,952,1510,1532]" pageId="4" pageNumber="105">AC, IJ)</emphasis>
Vegetative leaves.
<emphasis id="B93DEAFDFFB5FF98FBF463860CCBDDEE" bold="true" box="[1142,1196,1510,1532]" pageId="4" pageNumber="105">DE)</emphasis>
Perichaetial leaves.
<emphasis id="B93DEAFDFFB5FF98FAF463860DCCDDEE" bold="true" box="[1398,1451,1510,1532]" pageId="4" pageNumber="105">F, K)</emphasis>
Capsules.
<emphasis id="B93DEAFDFFB5FF98FF69606A0962DE32" bold="true" box="[235,261,1546,1568]" pageId="4" pageNumber="105">G)</emphasis>
Upper leaf cells.
<emphasis id="B93DEAFDFFB5FF98FE2D606A09A5DE0D" bold="true" box="[431,450,1546,1567]" pageId="4" pageNumber="105">H</emphasis>
) Apical leaf cells. AH from lectotype
<emphasis id="B93DEAFDFFB5FF98FCBF606A0BC8DE32" box="[829,943,1546,1568]" italics="true" pageId="4" pageNumber="105">Bridges s.n.</emphasis>
(BM-herb-Hooker). IK from
<emphasis id="B93DEAFDFFB5FF98FB5A606B0D32DE0D" box="[1240,1365,1546,1568]" italics="true" pageId="4" pageNumber="105">Herzog 3703</emphasis>
(JE).
</paragraph>
</caption>
<paragraph id="8BF636EFFFB5FF98FF3F602E0AF4DF0E" blockId="4.[136,1452,1614,2073]" pageId="4" pageNumber="105">
Autoicous. Archegonia medial on short vaginula (
<quantity id="4CB19B0AFFB5FF98FD79602E0BEDDE7B" box="[763,906,1614,1641]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="4" pageNumber="105" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.41.6 mm</quantity>
); perichaetial leaves narrow lanceolate,
<quantity id="4CB19B0AFFB5FF98FAD5602F08D3DE9F" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="4" pageNumber="105" unit="mm" value="2.65" valueMax="2.9" valueMin="2.4">2.42.9 mm</quantity>
long,
<emphasis id="B93DEAFDFFB5FF98FE8060120B7DDE9E" box="[258,794,1650,1677]" italics="true" pageId="4" pageNumber="105">less than twice the length of vegetative leaves</emphasis>
, covering
<quantity id="4CB19B0AFFB5FF98FC1B60130BB9DE9F" box="[921,990,1651,1677]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="105" unit="mm" value="1.0">1 mm</quantity>
of seta, acuminate, plicate, perichaetial paraphyses biseriate at base, immersed. Setae 7 (910)
<quantity id="4CB19B0AFFB5FF98FC9260F70B04DEA3" box="[784,867,1687,1713]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="105" unit="mm" value="12.0">12 mm</quantity>
long; capsule exserted, necks
<quantity id="4CB19B0AFFB5FF98FB4B60F70D32DEA3" box="[1225,1365,1687,1713]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="4" pageNumber="105" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.30.4 mm</quantity>
, broad, gradually attenuated, urns
<quantity id="4CB19B0AFFB5FF98FE3360DA0A5FDEC7" box="[433,568,1722,1749]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.5" metricValueMin="1.7" pageId="4" pageNumber="105" unit="mm" value="2.1" valueMax="2.5" valueMin="1.7">1.72.5 mm</quantity>
long, narrowly elliptic, mouth narrow, nearly 1/2 as wide as urn, capsule walls smooth, exothecial cells rectangular; stomates superficial; opercula base conical, short oblique rostrate. Calyptrae cucullate, large. Spores 2931 µm in diameter.
</paragraph>
</subSubSection>
<subSubSection id="C3536564FFB5FF98FF3F61460A07DF77" pageId="4" pageNumber="105" type="biology_ecology">
<paragraph id="8BF636EFFFB5FF98FF3F61460A07DF77" blockId="4.[136,1452,1614,2073]" pageId="4" pageNumber="105">
<emphasis id="B93DEAFDFFB5FF98FF3F61460947DF52" bold="true" box="[189,288,1830,1856]" pageId="4" pageNumber="105">Habitat.</emphasis>
Commonly grows on trees, but also colonizes andesitic rocks in exposed habitats and within forests at middle to high elevations (
<quantity id="4CB19B0AFFB5FF98FE31612A0A34DF77" box="[435,595,1866,1893]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.7" metricValueMin="1.5" pageId="4" pageNumber="105" unit="m" value="2100.0" valueMax="2700.0" valueMin="1500.0">15002700 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C3536564FFB5FF98FF3F610E0A3ADF9A" box="[189,605,1902,1929]" pageId="4" pageNumber="105" type="distribution">
<paragraph id="8BF636EFFFB5FF98FF3F610E0A3ADF9A" blockId="4.[136,1452,1614,2073]" box="[189,605,1902,1929]" pageId="4" pageNumber="105">
<emphasis id="B93DEAFDFFB5FF98FF3F610E0928DF9A" bold="true" box="[189,335,1902,1928]" pageId="4" pageNumber="105">Distribution</emphasis>
.
<collectingCountry id="F35E767FFFB5FF98FEDC610E09D4DF9A" box="[350,435,1902,1928]" name="Bolivia" pageId="4" pageNumber="105">Bolivia</collectingCountry>
,
<collectingCountry id="F35E767FFFB5FF98FE3C610F0993DF9B" box="[446,500,1903,1929]" name="Peru" pageId="4" pageNumber="105">Peru</collectingCountry>
,
<collectingCountry id="F35E767FFFB5FF98FE7D610E0A3FDF9A" box="[511,600,1902,1928]" name="Mexico" pageId="4" pageNumber="105">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3536564FFB5FF99FF3F61F20882D9CA" lastPageId="5" lastPageNumber="106" pageId="4" pageNumber="105" type="materials_examined">
<paragraph id="8BF636EFFFB5FF98FF3F61F20DF6D00B" blockId="4.[136,1452,1614,2073]" pageId="4" pageNumber="105">
<emphasis id="B93DEAFDFFB5FF98FF3F61F209DADFBE" bold="true" box="[189,445,1938,1964]" pageId="4" pageNumber="105">Specimens examined</emphasis>
:—
<collectingCountry id="F35E767FFFB5FF98FE5D61F30A3FDFBF" box="[479,600,1939,1965]" name="Bolivia" pageId="4" pageNumber="105">BOLIVIA</collectingCountry>
.
<emphasis id="B93DEAFDFFB5FF98FDE861F20A99DFBE" box="[618,766,1938,1964]" italics="true" pageId="4" pageNumber="105">Bridges s.n.,</emphasis>
1846 (BM!-herb. Hooker, BM!-herb. Wilson, NY!-herb. Hooker, NY!-herb. Wilson);
<collectingRegion id="498DF80DFFB5FF98FE4C61D70A7BDFC3" box="[462,540,1975,2001]" country="Bolivia" name="La Paz" pageId="4" pageNumber="105">La Paz</collectingRegion>
: Saavedra, NW of Chuma,
<emphasis id="B93DEAFDFFB5FF98FCD561D70B97DFC3" box="[855,1008,1975,2001]" italics="true" pageId="4" pageNumber="105">Lewis 79-947</emphasis>
(F!);
<collectingRegion id="498DF80DFFB5FF98FBB161D60CA2DFC2" box="[1075,1221,1974,2000]" country="Bolivia" name="Cochabamba" pageId="4" pageNumber="105">Cochabamba</collectingRegion>
: Iscaypata, cerca de Quillacollo,
<emphasis id="B93DEAFDFFB5FF98FE9461BB09DCDFE7" box="[278,443,2011,2037]" italics="true" pageId="4" pageNumber="105">De Luna 2094</emphasis>
(DUKE!, LPB!); in market at
<collectingRegion id="498DF80DFFB5FF98FC9D61BA0BD3DFE6" box="[799,948,2010,2036]" country="Bolivia" name="Cochabamba" pageId="4" pageNumber="105">Cochabamba</collectingRegion>
,
<emphasis id="B93DEAFDFFB5FF98FC4361BB0C1DDFE7" box="[961,1146,2011,2037]" italics="true" pageId="4" pageNumber="105">Hermann 25280</emphasis>
(NY!);
<collectingRegion id="498DF80DFFB5FF98FB5A61BB0D31DFE7" box="[1240,1366,2011,2037]" country="Bolivia" name="Santa Cruz" pageId="4" pageNumber="105">Santa Cruz</collectingRegion>
: In der Cordillere von
<collectingRegion id="498DF80DFFB5FF98FEB0619F09D7D00B" box="[306,432,2047,2073]" country="Bolivia" name="Santa Cruz" pageId="4" pageNumber="105">Santa Cruz</collectingRegion>
,
<quantity id="4CB19B0AFFB5FF98FE39619E0A6CD00A" box="[443,523,2046,2072]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="105" unit="m" value="1600.0">1600m</quantity>
,
<emphasis id="B93DEAFDFFB5FF98FD95619F0ACFD00B" box="[535,680,2047,2073]" italics="true" pageId="4" pageNumber="105">Herzog 3482</emphasis>
(JE!, S!); Comarapa und Abra de
<taxonomicName id="4C494D6CFFB5FF98FBA5619E0C83D00A" authority=", Herzog" authorityName="Herzog" box="[1063,1252,2046,2072]" class="Insecta" family="Staphylinidae" genus="Catalina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="genus">
Catalina,
<emphasis id="B93DEAFDFFB5FF98FB10619F0C83D00A" box="[1170,1252,2047,2072]" italics="true" pageId="4" pageNumber="105">Herzog</emphasis>
</taxonomicName>
<emphasis id="B93DEAFDFFB5FF98FB6E619F0D43D00B" box="[1260,1316,2047,2073]" italics="true" pageId="4" pageNumber="105">3703</emphasis>
(JE!, S!).
</paragraph>
<paragraph id="8BF636EFFFB4FF99FF3F66FF0D59D8AA" blockId="5.[136,1452,158,725]" box="[189,1342,158,185]" pageId="5" pageNumber="106">
<collectingCountry id="F35E767FFFB4FF99FF5966FF0942D8AB" box="[219,293,159,185]" name="Peru" pageId="5" pageNumber="106">PERU</collectingCountry>
.
<collectingRegion id="498DF80DFFB4FF99FEAD66FE0912D8AA" box="[303,373,158,184]" country="Peru" name="Cusco" pageId="5" pageNumber="106">Cusco</collectingRegion>
: Calca, Pisac,
<emphasis id="B93DEAFDFFB4FF99FD9966FF0AF5D8AA" box="[539,658,158,184]" italics="true" pageId="5" pageNumber="106">Marin 622</emphasis>
(F!); Laguna Urpicancha, near Huacarpay,
<emphasis id="B93DEAFDFFB4FF99FBF866FE0C9FD8AA" box="[1146,1272,158,184]" italics="true" pageId="5" pageNumber="106">Inohue 828</emphasis>
(JE!).
</paragraph>
<paragraph id="8BF636EFFFB4FF99FF3F66A20882D9CA" blockId="5.[136,1452,158,725]" pageId="5" pageNumber="106">
<collectingCountry id="F35E767FFFB4FF99FF5966A2092BD8CE" box="[219,332,194,220]" name="Mexico" pageId="5" pageNumber="106">MEXICO</collectingCountry>
.
<collectingRegion id="498DF80DFFB4FF99FED166A20A66D8CE" box="[339,513,194,220]" country="Mexico" name="Aguascalientes" pageId="5" pageNumber="106">Aguascalientes</collectingRegion>
: San José de Gracia, La Congoja,
<emphasis id="B93DEAFDFFB4FF99FCF166A20B90D8CE" box="[883,1015,194,220]" italics="true" pageId="5" pageNumber="106">Villalobos 8</emphasis>
(MEXU! NY!);
<collectingRegion id="498DF80DFFB4FF99FB2D66A20D69D8CE" box="[1199,1294,194,220]" country="Mexico" name="Hidalgo" pageId="5" pageNumber="106">Hidalgo</collectingRegion>
: Cuyamaloya,
<emphasis id="B93DEAFDFFB4FF99FF0A6686094CD912" box="[136,299,230,256]" italics="true" pageId="5" pageNumber="106">Pringle 10627</emphasis>
(BM!, H!, JE!, MO!, NY!, S!,
<collectingCountry id="F35E767FFFB4FF99FD1066860AD9D912" box="[658,702,230,256]" name="United States of America" pageId="5" pageNumber="106">US</collectingCountry>
!); ridge above Real del Monte, near Pachuca,
<emphasis id="B93DEAFDFFB4FF99FB4966860D27D912" box="[1227,1344,230,256]" italics="true" pageId="5" pageNumber="106">Sharp 797</emphasis>
(MICH!,
<collectingCountry id="F35E767FFFB4FF99FF0A676A08D3D936" box="[136,180,266,292]" name="United States of America" pageId="5" pageNumber="106">US</collectingCountry>
!); S of San Juan del Rio,
<emphasis id="B93DEAFDFFB4FF99FE55676A0AFBD936" box="[471,668,266,293]" italics="true" pageId="5" pageNumber="106">Rauh &amp; Dull 122</emphasis>
(H!); Estado de México: Toluca,
<emphasis id="B93DEAFDFFB4FF99FB9E676A0CD8D936" box="[1052,1215,266,292]" italics="true" pageId="5" pageNumber="106">Pringle 15171</emphasis>
(FH! proparte, NY!, S!); Cerro Gordo,
<emphasis id="B93DEAFDFFB4FF99FEDB674E0A62D95A" box="[345,517,302,328]" italics="true" pageId="5" pageNumber="106">Cárdenas 2200</emphasis>
(DUKE!); San Martín de las Pirámides, Cerro Gordo,
<emphasis id="B93DEAFDFFB4FF99FBF4674E0C98D95A" box="[1142,1279,302,328]" italics="true" pageId="5" pageNumber="106">Allen 12943</emphasis>
(MO!); Nuevo León:
<quantity id="4CB19B0AFFB4FF99FF5267320964D97E" box="[208,259,338,364]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="5" pageNumber="106" unit="mi" value="3.0">3 mi</quantity>
NW of Picacho Onofre,
<emphasis id="B93DEAFDFFB4FF99FD9A67330AB8D97E" box="[536,735,338,364]" italics="true" pageId="5" pageNumber="106">Nesom &amp;Wells 34</emphasis>
(DUKE!); Cerro del Viejo,
<quantity id="4CB19B0AFFB4FF99FB9567320C3FD97E" box="[1047,1112,338,365]" metricMagnitude="4" metricUnit="m" metricValue="2.414016" pageId="5" pageNumber="106" unit="mi" value="15.0">15 mi</quantity>
W Dulces Nombres,
<emphasis id="B93DEAFDFFB4FF99FAC56733096DD982" italics="true" pageId="5" pageNumber="106">Meyer &amp; Rogers 149</emphasis>
(G!, NY!); Querétaro: cerro El Pinguical, cerca de Pinal de Amoles,
<emphasis id="B93DEAFDFFB4FF99FB9C67170CC8D982" box="[1054,1199,374,401]" italics="true" pageId="5" pageNumber="106">Ocampo 620</emphasis>
(XAL); Tlaxcala: Mt. Tlacuapanga,
<emphasis id="B93DEAFDFFB4FF99FEA667FA09F0D9A6" box="[292,407,410,436]" italics="true" pageId="5" pageNumber="106">Sharp 433</emphasis>
(MICH!); cerca de Villareal, al NE de Huamantla,
<emphasis id="B93DEAFDFFB4FF99FC4E67FB0C0DD9A6" box="[972,1130,410,436]" italics="true" pageId="5" pageNumber="106">De Luna 2646</emphasis>
(XAL!),
<emphasis id="B93DEAFDFFB4FF99FB5167FA0D6CD9A6" box="[1235,1291,410,436]" italics="true" pageId="5" pageNumber="106">2718</emphasis>
(XAL!),
<emphasis id="B93DEAFDFFB4FF99FAF167FA0DCCD9A6" box="[1395,1451,410,436]" italics="true" pageId="5" pageNumber="106">2720</emphasis>
(XAL!).
</paragraph>
</subSubSection>
<subSubSection id="C3536564FFB4FF99FF3F67820CF9DAC6" pageId="5" pageNumber="106" type="discussion">
<paragraph id="8BF636EFFFB4FF99FF3F67820CF9DAC6" blockId="5.[136,1452,158,725]" pageId="5" pageNumber="106">
<emphasis id="B93DEAFDFFB4FF99FF3F67820925D9EE" bold="true" box="[189,322,482,508]" italics="true" pageId="5" pageNumber="106">Discussion.</emphasis>
This is strictly the species concept from that of
<bibRefCitation id="EFD84B1EFFB4FF99FCC767820B85D9EF" author="Mitten, W." box="[837,994,482,509]" pageId="5" pageNumber="106" pagination="1 - 659" refId="ref3887" refString="Mitten, W. (1869) Musci Austro-Americani. Journal of the Linnean Society, Botany 12: 1 - 659. https: // doi. org / 10.1111 / j. 1095 - 8339.1871. tb 00633. x" type="journal article" year="1869">Mitten (1869)</bibRefCitation>
. Morphological characters distinguishing
<taxonomicName id="4C494D6CFFB4FF99FF0A64670890DA32" box="[136,247,518,544]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="5" pageNumber="106" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB4FF99FF0A64670890DA32" box="[136,247,518,544]" italics="true" pageId="5" pageNumber="106">B. plicata</emphasis>
</taxonomicName>
are the leaves imbricate, strongly plicate, and concave (
<figureCitation id="13722A6AFFB4FF99FCFB64660BB5DA32" box="[889,978,518,544]" captionStart="FIGURE 1" captionStartId="1.[136,229,2018,2040]" captionTargetBox="[186,1404,688,1990]" captionTargetId="figure-177@1.[183,1404,688,1990]" captionTargetPageId="1" captionText="FIGURE 1. Lectotype of Braunia plicata. A) Specimen Bridges s. n., 1846, at BM (herb. Hooker). B) Detail of lectotype, showing one capsule. C) Detail of leaves with hyaline apex. D) Leaf apex hyaline and dentate. (Photographs by E. De Luna)." figureDoi="http://doi.org/10.5281/zenodo.5900974" httpUri="https://zenodo.org/record/5900974/files/figure.png" pageId="5" pageNumber="106">Fig. 1C</figureCitation>
). The leaf apex is cuspidate, hyaline, and dentate (
<figureCitation id="13722A6AFFB4FF99FF6F644A0933DA56" box="[237,340,554,580]" captionStart="FIGURE 1" captionStartId="1.[136,229,2018,2040]" captionTargetBox="[186,1404,688,1990]" captionTargetId="figure-177@1.[183,1404,688,1990]" captionTargetPageId="1" captionText="FIGURE 1. Lectotype of Braunia plicata. A) Specimen Bridges s. n., 1846, at BM (herb. Hooker). B) Detail of lectotype, showing one capsule. C) Detail of leaves with hyaline apex. D) Leaf apex hyaline and dentate. (Photographs by E. De Luna)." figureDoi="http://doi.org/10.5281/zenodo.5900974" httpUri="https://zenodo.org/record/5900974/files/figure.png" pageId="5" pageNumber="106">Figs. 1D</figureCitation>
,
<figureCitation id="13722A6AFFB4FF99FEE7644A0997DA56" box="[357,496,554,580]" captionStart="FIGURE 4" captionStartId="4.[136,229,1510,1532]" captionTargetBox="[155,1433,330,1482]" captionTargetId="figure-93@4.[151,1436,317,1485]" captionTargetPageId="4" captionText="FIGURE 4. Gametophytic and sporophytic features of Braunia plicata. AC, IJ) Vegetative leaves. DE) Perichaetial leaves. F, K) Capsules. G) Upper leaf cells. H) Apical leaf cells.AH from lectotype Bridges s.n. (BM-herb-Hooker). IK from Herzog 3703 (JE)." figureDoi="http://doi.org/10.5281/zenodo.5900982" httpUri="https://zenodo.org/record/5900982/files/figure.png" pageId="5" pageNumber="106">4AC, 4IJ</figureCitation>
). Shoots are often dark-green to brownish, especially darker green along the leaf plications, which contrasts sharply with the hyaline leaf acumens (
<figureCitation id="13722A6AFFB4FF99FC0A642E0C61DA7A" box="[904,1030,590,616]" captionStart="FIGURE 1" captionStartId="1.[136,229,2018,2040]" captionTargetBox="[186,1404,688,1990]" captionTargetId="figure-177@1.[183,1404,688,1990]" captionTargetPageId="1" captionText="FIGURE 1. Lectotype of Braunia plicata. A) Specimen Bridges s. n., 1846, at BM (herb. Hooker). B) Detail of lectotype, showing one capsule. C) Detail of leaves with hyaline apex. D) Leaf apex hyaline and dentate. (Photographs by E. De Luna)." figureDoi="http://doi.org/10.5281/zenodo.5900974" httpUri="https://zenodo.org/record/5900974/files/figure.png" pageId="5" pageNumber="106">Fig. 1AC</figureCitation>
).
<taxonomicName id="4C494D6CFFB4FF99FB9E642F0CA8DA7A" box="[1052,1231,590,616]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="5" pageNumber="106" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB4FF99FB9E642F0CA8DA7A" box="[1052,1231,590,616]" italics="true" pageId="5" pageNumber="106">Braunia plicata</emphasis>
</taxonomicName>
is mostly montane Andean. Among the South American species with hyaline leaf acumens, the dentate apex and the narrowly elongated apical cells (
<figureCitation id="13722A6AFFB4FF99FE9564F6091BDAA2" box="[279,380,662,688]" captionStart="FIGURE 1" captionStartId="1.[136,229,2018,2040]" captionTargetBox="[186,1404,688,1990]" captionTargetId="figure-177@1.[183,1404,688,1990]" captionTargetPageId="1" captionText="FIGURE 1. Lectotype of Braunia plicata. A) Specimen Bridges s. n., 1846, at BM (herb. Hooker). B) Detail of lectotype, showing one capsule. C) Detail of leaves with hyaline apex. D) Leaf apex hyaline and dentate. (Photographs by E. De Luna)." figureDoi="http://doi.org/10.5281/zenodo.5900974" httpUri="https://zenodo.org/record/5900974/files/figure.png" pageId="5" pageNumber="106">Figs. 1A</figureCitation>
,
<figureCitation id="13722A6AFFB4FF99FE0B64F609CBDAA2" box="[393,428,662,688]" captionStart="FIGURE 4" captionStartId="4.[136,229,1510,1532]" captionTargetBox="[155,1433,330,1482]" captionTargetId="figure-93@4.[151,1436,317,1485]" captionTargetPageId="4" captionText="FIGURE 4. Gametophytic and sporophytic features of Braunia plicata. AC, IJ) Vegetative leaves. DE) Perichaetial leaves. F, K) Capsules. G) Upper leaf cells. H) Apical leaf cells.AH from lectotype Bridges s.n. (BM-herb-Hooker). IK from Herzog 3703 (JE)." figureDoi="http://doi.org/10.5281/zenodo.5900982" httpUri="https://zenodo.org/record/5900982/files/figure.png" pageId="5" pageNumber="106">4H</figureCitation>
) separate
<taxonomicName id="4C494D6CFFB4FF99FDA364F70AE8DAA2" box="[545,655,662,688]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="5" pageNumber="106" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB4FF99FDA364F70AE8DAA2" box="[545,655,662,688]" italics="true" pageId="5" pageNumber="106">B. plicata</emphasis>
</taxonomicName>
from
<emphasis id="B93DEAFDFFB4FF99FD5464F70B83DAA2" box="[726,996,663,688]" italics="true" pageId="5" pageNumber="106">
<taxonomicName id="4C494D6CFFB4FF99FD5464F70B22DAA2" box="[726,837,663,688]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="5" pageNumber="106" phylum="Bryophyta" rank="species" species="incana">B. incana</taxonomicName>
,
<taxonomicName id="4C494D6CFFB4FF99FCD064F70B83DAA2" box="[850,996,663,688]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="5" pageNumber="106" phylum="Bryophyta" rank="species" species="canescens">B. canescens</taxonomicName>
</emphasis>
and
<emphasis id="B93DEAFDFFB4FF99FB9E64F70CD0DAA2" box="[1052,1207,662,688]" italics="true" pageId="5" pageNumber="106">
<taxonomicName id="4C494D6CFFB4FF99FB9E64F70CD3DAA2" box="[1052,1204,662,688]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="5" pageNumber="106" phylum="Bryophyta" rank="species" species="subincana">B. subincana</taxonomicName>
.
</emphasis>
The following key is proposed to help in distinguishing
<taxonomicName id="4C494D6CFFB4FF99FD8E64DB0A1EDAC6" box="[524,633,698,724]" class="Bryopsida" family="Hedwigiaceae" genus="Braunia" kingdom="Plantae" order="Leucodontales" pageId="5" pageNumber="106" phylum="Bryophyta" rank="species" species="plicata">
<emphasis id="B93DEAFDFFB4FF99FD8E64DB0A1EDAC6" box="[524,633,698,724]" italics="true" pageId="5" pageNumber="106">B. plicata</emphasis>
</taxonomicName>
from the other three species with hyaline apices.
</paragraph>
</subSubSection>
</treatment>
</document>