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<document id="7E4AF09809F5165838974EC143DCF4A3" ID-DOI="10.1111/zoj.12112" ID-ISSN="0024-4082" ID-Zenodo-Dep="10273730" ID-ZooBank="7BA11142-CBC1-4026-A578-EBAB6D2B6C0C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630107647828" checkinUser="marcus" docAuthor="Mendoza Marroquín, Jorge I." docDate="2014" docId="03E087B45649FFFAFC444BD5B007167E" docLanguage="en" docName="zoj12112.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 170 (4)" docSource="http://dx.doi.org/10.1111/zoj.12112" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Hemirrhagus benzaa Mendoza Marroquín 2014, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="670" masterDocId="FFD9FFCC566AFFDEFFD84973B208116D" masterDocTitle="Taxonomic revision of Hemirrhagus Simon, 1903 (Araneae: Theraphosidae, Theraphosinae), with description of five new species from Mexico" masterLastPageNumber="689" masterPageNumber="634" pageNumber="669" updateTime="1701826367839" updateUser="ExternalLinkService">
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<mods:title id="F4613F63EDB76DBF8C8CE67832B3FFF9">Taxonomic revision of Hemirrhagus Simon, 1903 (Araneae: Theraphosidae, Theraphosinae), with description of five new species from Mexico</mods:title>
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<treatment id="03E087B45649FFFAFC444BD5B007167E" LSID="urn:lsid:plazi:treatment:03E087B45649FFFAFC444BD5B007167E" httpUri="http://treatment.plazi.org/id/03E087B45649FFFAFC444BD5B007167E" lastPageId="36" lastPageNumber="670" pageId="35" pageNumber="669">
<subSubSection id="C35365295649FFFDFC444BD5B71013D6" box="[924,1304,678,699]" pageId="35" pageNumber="669" type="nomenclature">
<paragraph id="8BF636A25649FFFDFC444BD5B71013D6" blockId="35.[877,1351,678,732]" box="[924,1304,678,699]" pageId="35" pageNumber="669">
<heading id="D0BE81CE5649FFFDFC444BD5B71013D6" box="[924,1304,678,699]" centered="true" fontSize="9" level="2" pageId="35" pageNumber="669" reason="2">
<taxonomicName id="4C494D215649FFFDFC444BD5B6B813D6" ID-CoL="3KSDY" authority="Mendoza Marroquín, 2014" authorityName="Mendoza Marroquín" authorityYear="2014" box="[924,1200,678,699]" class="Arachnida" family="Theraphosidae" genus="Hemirrhagus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="686" phylum="Arthropoda" rank="species" species="benzaa" status="sp. nov.">HEMIRRHAGUS BENZAA</taxonomicName>
<emphasis id="B93DEAB05649FFFDFB614BD9B71013D6" bold="true" box="[1209,1304,678,699]" pageId="35" pageNumber="669">
<taxonomicNameLabel id="A20E57CB5649FFFDFB614BD9B71013D6" box="[1209,1304,678,699]" pageId="35" pageNumber="669" rank="species">SP. NOV.</taxonomicNameLabel>
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<subSubSection id="C35365295649FFFAFCB54BB5B007167E" lastPageId="36" lastPageNumber="670" pageId="35" pageNumber="669" type="description">
<paragraph id="8BF636A25649FFFDFCB54BB5B74F13B1" blockId="35.[877,1351,678,732]" box="[877,1351,710,732]" pageId="35" pageNumber="669">
(
<figureCitation id="13722A275649FFFDFCAD4BB5B1D913B6" box="[885,977,710,731]" captionStart="Figure 1" captionStartId="3.[144,223,1334,1353]" captionTargetBox="[303,1262,196,1304]" captionTargetId="figure-144@3.[303,1263,196,1304]" captionTargetPageId="3" captionText="Figure 1. AH, JL, Hemirrhagus spp. habitus; I, Hemirrhagus papalotl egg-sac. A, Hemirrhagus chilango, male CNAN 4253, in life; B, C, Hemirrhagus eros, B, female, C, male; D, E, Hemirrhagus ocellatus, D, male CNAN 4457, in life, E, female; F, Hemirrhagus benzaa sp. nov., female; GI, H. papalotl, G, male CNAN 4461, in life, H, female CNAN 4461, in life, I, female CNAN 4459 with fixed hammock egg-sac, in life; J, K, Hemirrhagus stygius, J, male CNAN 4462, in life, K, female CNAN 4462, in life; L, Hemirrhagus franckei sp. nov., holotype male CNAN-T0776, in life. Photos: A, C, D, E, GK: J. Mendoza; B, F, L: E. Goyer." figureDoi="http://doi.org/10.5281/zenodo.10273732" httpUri="https://zenodo.org/record/10273732/files/figure.png" pageId="35" pageNumber="669">FIGS 1F</figureCitation>
,
<figureCitation id="13722A275649FFFDFC394BB5B1F413B6" box="[993,1020,710,731]" captionStart="Figure 2" captionStartId="4.[164,243,1137,1156]" captionTargetBox="[323,1283,196,1107]" captionTargetId="figure-206@4.[323,1283,196,1107]" captionTargetPageId="4" captionText="Figure 2. AC, Hemirrhagus spp. habitus; DK, habitat. A, B, Hemirrhagus embolulatus sp. nov., A, holotype male CNAN-T0774, in life, B, paratype female CNAN-T0775, in life; C, Hemirrhagus valdezi sp. nov., paratype female CNAN-T0780, in life; D, Cueva de la Peña Blanca entrance, type locality of Hemirrhagus ocellatus; E, 4 km west of Puerto de la Soledad, pine forest, type locality of Hemirrhagus franckei sp. nov.; F, Bosque del ajusco, pine forest, locality of Hemirrhagus chilango; G, Valle de los fantasmas, oak forest on the road to Cueva de la Laguna, type locality of Hemirrhagus elliotti; H, Cueva de los Potrerillos entrance, type locality of Hemirrhagus stygius; I, Paradero Puerta del Sol, pine forest, locality of Hemirrhagus benzaa sp. nov.; J, Omiltemi, tropical forest-pine forest ecotone, type locality of H. embolulatus sp. nov.; K, El punto, type locality of Hemirrhagus eros. Photos: A: M. Gamache; B, E, I, K: E. Goyer; C: L. Vazquez; D, F, J: J. Mendoza; G, H: J. Cruz." figureDoi="http://doi.org/10.5281/zenodo.10273736" httpUri="https://zenodo.org/record/10273736/files/figure.png" pageId="35" pageNumber="669">2I</figureCitation>
,
<figureCitation id="13722A275649FFFDFBD34BB5B65C13B6" box="[1035,1108,710,731]" captionStart="Figure 33" captionStartId="35.[144,223,1765,1784]" captionTargetBox="[303,1264,892,1734]" captionTargetId="figure-306@35.[303,1264,892,1735]" captionTargetPageId="35" captionText="Figure 33. Hemirrhagus benzaa sp. nov., male holotype AMNH. A, carapace, dorsal view. B, prosoma, ventral view showing the retrolateral projection of the coxa (RP) on coxae of all legs (arrows show this feature only on coxa of third and fourth legs). C, ocular tubercle, dorsal view. D, labial and maxillary cuspules. E, opisthosoma, dorsa view. F, cheliceral teeth, ventral view. G, tibial apophyses, prolateral view. H, tibial apophyses, ventral view. I, metatarsus I, prolateral view. Scale bars = 2 mm (A, B, E, I), 1 mm (C, D, FH)." figureDoi="http://doi.org/10.5281/zenodo.10273824" httpUri="https://zenodo.org/record/10273824/files/figure.png" pageId="35" pageNumber="669">33AI</figureCitation>
,
<figureCitation id="13722A275649FFFDFBBA4BB5B6BE13B6" box="[1122,1206,710,731]" captionStart="Figure 34" captionStartId="36.[164,243,738,757]" captionTargetBox="[388,1213,207,702]" captionTargetId="figure-65@36.[387,1219,196,708]" captionTargetPageId="36" captionText="Figure 34. Hemirrhagus benzaa sp. nov., male holotype AMNH, right palpal bulb: A, prolateral view. B, retrolateral view. C, dorsal view. D, ventral view. Abbreviations: SA, subapical keel; VG, ventral groove. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10273826" httpUri="https://zenodo.org/record/10273826/files/figure.png" pageId="35" pageNumber="669">34AD</figureCitation>
,
<figureCitation id="13722A275649FFFDFB1C4BB5B71B13B1" box="[1220,1299,710,732]" captionStart="Figure 35" captionStartId="36.[164,243,1578,1597]" captionTargetBox="[323,1283,861,1549]" captionTargetId="figure-104@36.[323,1283,861,1549]" captionTargetPageId="36" captionText="Figure 35. Hemirrhagus benzaa sp. nov., female paratype AMNH. A, carapace, dorsal view. B, prosoma, ventral view showing the retrolateral projection of the coxa (RP) on coxae of all legs (arrows show this feature only on coxa of third and fourth legs). C, opisthosoma, dorsal view. D, ocular tubercle, dorsal view. E, labial and maxillary cuspules. F, spermathecae, dorsal view. Scale bars = 2 mm (AC), 1 mm (DF)." figureDoi="http://doi.org/10.5281/zenodo.10273828" httpUri="https://zenodo.org/record/10273828/files/figure.png" pageId="35" pageNumber="669">35AF</figureCitation>
,
<figureCitation id="13722A275649FFFDFAFA4BB5B73713B1" box="[1314,1343,710,732]" captionStart="Figure 49" captionStartId="52.[164,243,1007,1026]" captionTargetBox="[172,1436,200,970]" captionTargetId="figure-347@52.[164,1444,195,977]" captionTargetPageId="52" captionText="Figure 49. Known distribution of Hemirrhagus mitchelli (Gertsch, 1982), Hemirrhagus puebla (Gertsch, 1982), Hemirrhagus ocellatus Pérez-Miles &amp; Locht, 2003, Hemirrhagus gertschi Pérez-Miles &amp; Locht, 2003, Hemirrhagus valdezi sp. nov., Hemirrhagus pernix (Ausserer, 1875), Hemirrhagus franckei sp. nov., Hemirrhagus reddelli (Gertsch, 1973), Hemirrhagus benzaa sp. nov., and Hemirrhagus guichi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10273864" httpUri="https://zenodo.org/record/10273864/files/figure.png" pageId="35" pageNumber="669">49</figureCitation>
)
</paragraph>
<paragraph id="8BF636A25649FFFDFCFE4B9EB7851252" blockId="35.[806,1421,748,832]" pageId="35" pageNumber="669">
<emphasis id="B93DEAB05649FFFDFCFE4B9EB1C5126F" box="[806,973,749,770]" italics="true" pageId="35" pageNumber="669">Type material:</emphasis>
<materialsCitation id="3B213CFF5649FFFDFC014B9EB7851252" collectingDate="1985-04" collectionCode="AMNH" collectorName="de Juarez &amp; C. M. Bogert" country="Mexico" county="Llano" location="Sierra de Juarez" municipality="Flores" pageId="35" pageNumber="669" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Oaxaca" typeStatus="holotype">
<typeStatus id="54F288005649FFFDFC014B9EB637126E" box="[985,1087,749,771]" pageId="35" pageNumber="669" type="holotype">Holotype</typeStatus>
<specimenCount id="9D4FFD2B5649FFFDFB934B9FB66C126F" box="[1099,1124,748,770]" pageId="35" pageNumber="669" type="male"></specimenCount>
and
<typeStatus id="54F288005649FFFDFB7D4B9EB703126F" box="[1189,1291,749,770]" pageId="35" pageNumber="669" type="paratype">paratype</typeStatus>
<specimenCount id="9D4FFD2B5649FFFDFACE4B9FB723126F" box="[1302,1323,748,770]" pageId="35" pageNumber="669" type="female"></specimenCount>
<collectionCode id="ED58AE675649FFFDFAED4B9EB782126F" box="[1333,1418,749,770]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="35" pageNumber="669">AMNH</collectionCode>
,
<collectingCountry id="F35E76325649FFFDFCFE4A78B19B124D" box="[806,915,779,800]" name="Mexico" pageId="35" pageNumber="669">MEXICO</collectingCountry>
:
<collectingRegion id="498DF8405649FFFDFC474A78B1FE124D" box="[927,1014,779,800]" country="Mexico" name="Oaxaca" pageId="35" pageNumber="669">Oaxaca</collectingRegion>
,
<quantity id="4CB19B475649FFFDFBDB4A78B633124C" box="[1027,1083,779,801]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="35" pageNumber="669" unit="km" value="1.0">1 km</quantity>
north of
<collectingCounty id="62974E2E5649FFFDFB714A78B6E2124C" box="[1193,1258,779,801]" pageId="35" pageNumber="669">Llano</collectingCounty>
de las
<collectingMunicipality id="6B92ACD85649FFFDFAE74A78B781124C" box="[1343,1417,779,801]" pageId="35" pageNumber="669">Flores</collectingMunicipality>
,
<location id="8E9660795649FFFDFCFE4A59B1EB122D" LSID="urn:lsid:plazi:treatment:03E087B45649FFFAFC444BD5B007167E:8E9660795649FFFDFCFE4A59B1EB122D" box="[806,995,810,832]" country="Mexico" county="Llano" municipality="Flores" name="Sierra de Juarez" pageId="35" pageNumber="669" stateProvince="Oaxaca">
Sierra
<collectorName id="26BC53745649FFFDFCAC4A59B1EB122D" box="[884,995,810,832]" pageId="35" pageNumber="669">de Juárez</collectorName>
</location>
<date id="FFF710625649FFFDFC324A59B634122D" box="[1002,1084,810,832]" pageId="35" pageNumber="669" value="1985-04">
<collectingDate id="EFB3E98A5649FFFDFC324A59B634122D" box="[1002,1084,810,832]" pageId="35" pageNumber="669" value="1985-04">iv.1985</collectingDate>
</date>
, col.
<collectorName id="26BC53745649FFFDFBAC4A59B7001252" box="[1140,1288,810,831]" pageId="35" pageNumber="669">C. M. Bogert</collectorName>
;
<specimenCount id="9D4FFD2B5649FFFDFACB4A59B7851252" box="[1299,1421,810,831]" pageId="35" pageNumber="669" type="female" typeStatus="paratype">1 paratype</specimenCount>
</materialsCitation>
</paragraph>
<caption id="DF36662A5649FFFDFF484F96B0A41603" ID-DOI="http://doi.org/10.5281/zenodo.10273824" ID-Zenodo-Dep="10273824" httpUri="https://zenodo.org/record/10273824/files/figure.png" pageId="35" pageNumber="669" startId="35.[144,223,1765,1784]" targetBox="[303,1264,892,1734]" targetPageId="35">
<paragraph id="8BF636A25649FFFDFF484F96B0A41603" blockId="35.[144,1422,1765,1902]" pageId="35" pageNumber="669">
<emphasis id="B93DEAB05649FFFDFF484F96B0621794" bold="true" box="[144,618,1765,1785]" pageId="35" pageNumber="669">
Figure 33.
<taxonomicName id="4C494D215649FFFDFEC84F96B0001795" authority="Mendoza Marroquín, 2014" authorityName="Mendoza Marroquín" authorityYear="2014" box="[272,520,1765,1784]" class="Arachnida" family="Theraphosidae" genus="Hemirrhagus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="669" phylum="Arthropoda" rank="species" species="benzaa" status="sp. nov.">
<emphasis id="B93DEAB05649FFFDFEC84F96B0001795" bold="true" box="[272,520,1765,1784]" italics="true" pageId="35" pageNumber="669">Hemirrhagus benzaa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57CB5649FFFDFDC94F95B0621794" box="[529,618,1766,1785]" pageId="35" pageNumber="669" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype AMNH. A, carapace, dorsal view. B, prosoma, ventral view showing the retrolateral projection of the coxa (RP) on coxae of all legs (arrows show this feature only on coxa of third and fourth legs). C, ocular tubercle, dorsal view. D, labial and maxillary cuspules. E, opisthosoma, dorsa view. F, cheliceral teeth, ventral view. G, tibial apophyses, prolateral view. H, tibial apophyses, ventral view. I, metatarsus I, prolateral view. Scale bars = 2 mm (A, B, E, I), 1 mm (C, D, FH).
</paragraph>
</caption>
<caption id="DF36662A564EFFFAFF7C4B91B742127E" ID-DOI="http://doi.org/10.5281/zenodo.10273826" ID-Zenodo-Dep="10273826" httpUri="https://zenodo.org/record/10273826/files/figure.png" pageId="36" pageNumber="670" startId="36.[164,243,738,757]" targetBox="[388,1213,207,702]" targetPageId="36">
<paragraph id="8BF636A2564EFFFAFF7C4B91B742127E" blockId="36.[164,1441,738,787]" pageId="36" pageNumber="670">
<emphasis id="B93DEAB0564EFFFAFF7C4B91B073139B" bold="true" box="[164,635,738,758]" pageId="36" pageNumber="670">
Figure 34.
<taxonomicName id="4C494D21564EFFFAFEFC4B91B0131398" authority="Mendoza Marroquín, 2014" authorityName="Mendoza Marroquín" authorityYear="2014" box="[292,539,738,757]" class="Arachnida" family="Theraphosidae" genus="Hemirrhagus" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="670" phylum="Arthropoda" rank="species" species="benzaa" status="sp. nov.">
<emphasis id="B93DEAB0564EFFFAFEFC4B91B0131398" bold="true" box="[292,539,738,757]" italics="true" pageId="36" pageNumber="670">Hemirrhagus benzaa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57CB564EFFFAFDFB4B90B073139B" box="[547,635,739,758]" pageId="36" pageNumber="670" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype AMNH, right palpal bulb: A, prolateral view. B, retrolateral view. C, dorsal view. D, ventral view. Abbreviations: SA, subapical keel; VG, ventral groove. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="DF36662A564EFFFAFF7C4F59B18F17F8" ID-DOI="http://doi.org/10.5281/zenodo.10273828" ID-Zenodo-Dep="10273828" httpUri="https://zenodo.org/record/10273828/files/figure.png" pageId="36" pageNumber="670" startId="36.[164,243,1578,1597]" targetBox="[323,1283,861,1549]" targetPageId="36">
<paragraph id="8BF636A2564EFFFAFF7C4F59B18F17F8" blockId="36.[164,1442,1578,1686]" pageId="36" pageNumber="670">
<emphasis id="B93DEAB0564EFFFAFF7C4F59B0711753" bold="true" box="[164,633,1578,1598]" pageId="36" pageNumber="670">
Figure 35.
<taxonomicName id="4C494D21564EFFFAFEFC4F59B0121750" authority="Mendoza Marroquín, 2014" authorityName="Mendoza Marroquín" authorityYear="2014" box="[292,538,1578,1597]" class="Arachnida" family="Theraphosidae" genus="Hemirrhagus" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="670" phylum="Arthropoda" rank="species" species="benzaa" status="sp. nov.">
<emphasis id="B93DEAB0564EFFFAFEFC4F59B0121750" bold="true" box="[292,538,1578,1597]" italics="true" pageId="36" pageNumber="670">Hemirrhagus benzaa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57CB564EFFFAFDFA4F58B0711753" box="[546,633,1579,1598]" pageId="36" pageNumber="670" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female paratype AMNH. A, carapace, dorsal view. B, prosoma, ventral view showing the retrolateral projection of the coxa (RP) on coxae of all legs (arrows show this feature only on coxa of third and fourth legs). C, opisthosoma, dorsal view. D, ocular tubercle, dorsal view. E, labial and maxillary cuspules. F, spermathecae, dorsal view. Scale bars = 2 mm (AC), 1 mm (DF).
</paragraph>
</caption>
<paragraph id="8BF636A2564EFFFAFF7C4FADB007167E" blockId="36.[164,778,1758,1811]" pageId="36" pageNumber="670">
<materialsCitation id="3B213CFF564EFFFAFF7C4FADB007167E" collectingDate="1962-04" collectionCode="AMNH" collectorName="W. S. Miller." country="Mexico" location="Cerro Pelon" pageId="36" pageNumber="670" specimenCount="1" specimenCount-immature="1" stateProvince="Oaxaca">
♀ and
<specimenCount id="9D4FFD2B564EFFFAFF2F4FADB3881799" box="[247,384,1758,1780]" pageId="36" pageNumber="670" type="immature">1 immature</specimenCount>
<collectionCode id="ED58AE67564EFFFAFE514FACB3D61799" box="[393,478,1759,1780]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="36" pageNumber="670">AMNH</collectionCode>
,
<collectingCountry id="F35E7632564EFFFAFE334FADB050179E" box="[491,600,1758,1779]" name="Mexico" pageId="36" pageNumber="670">MEXICO</collectingCountry>
:
<collectingRegion id="498DF840564EFFFAFDBD4FADB0B4179E" box="[613,700,1758,1779]" country="Mexico" name="Oaxaca" pageId="36" pageNumber="670">Oaxaca</collectingRegion>
,
<location id="8E966079564EFFFAFD124FADB2EC167E" LSID="urn:lsid:plazi:treatment:03E087B45649FFFAFC444BD5B007167E:8E966079564EFFFAFD124FADB2EC167E" country="Mexico" name="Cerro Pelon" pageId="36" pageNumber="670" stateProvince="Oaxaca">Cerro Pelón</location>
<date id="FFF71062564EFFFAFF354F8EB348167E" box="[237,320,1789,1811]" pageId="36" pageNumber="670" value="1962-04">
<collectingDate id="EFB3E98A564EFFFAFF354F8EB348167E" box="[237,320,1789,1811]" pageId="36" pageNumber="670" value="1962-04">iv.1962</collectingDate>
</date>
, col.
<collectorName id="26BC5374564EFFFAFEA64F8EB007167E" box="[382,527,1789,1811]" pageId="36" pageNumber="670">W. S. Miller.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>