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<mods:title id="314868016E0D03EF27A9965366339BAD">Checklist and typification of Parnassia (Celastraceae)</mods:title>
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<treatment id="03E01B0FFFD9FFE5FF76F8AFFC17FD72" LSID="urn:lsid:plazi:treatment:03E01B0FFFD9FFE5FF76F8AFFC17FD72" httpUri="http://treatment.plazi.org/id/03E01B0FFFD9FFE5FF76F8AFFC17FD72" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<subSubSection id="C353F992FFD9FFE4FF76F8AFFD21F8CE" box="[136,699,1814,1841]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8BF6AA19FFD9FFE4FF76F8AFFD21F8CE" blockId="8.[136,699,1814,1841]" box="[136,699,1814,1841]" pageId="8" pageNumber="9">
<heading id="D0BE1D75FFD9FFE4FF76F8AFFD21F8CE" box="[136,699,1814,1841]" fontSize="11" level="2" pageId="8" pageNumber="9" reason="3">
<treatmentCitationGroup id="AB598D37FFD9FFE4FF76F8AFFD21F8CE" box="[136,699,1814,1841]" pageId="8" pageNumber="9">
<treatmentCitation id="0AE88C08FFD9FFE4FF76F8AFFD21F8CE" author="Royle, J. F." box="[136,699,1814,1841]" page=")" pageId="8" pageNumber="9" year="1835">
<taxonomicName id="4C49D19AFFD9FFE4FF76F8AFFD21F8CE" ID-CoL="4DQ7Q" authority="Wallich ex Royle (1835: 227)" authorityName="Wallich ex Royle" authorityPageNumber="227" authorityYear="1835" box="[136,699,1814,1841]" class="Magnoliopsida" family="Parnassiaceae" genus="Parnassia" kingdom="Plantae" order="Celastrales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="nubicola">
<emphasis id="B93D760BFFD9FFE4FF76F8AFFEFDF8CE" bold="true" box="[136,359,1814,1841]" italics="true" pageId="8" pageNumber="9">Parnassia nubicola</emphasis>
Wallich ex
<bibRefCitation id="EFD8D7E8FFD9FFE4FE13F8A1FD21F8CE" author="Royle, J. F." box="[493,699,1816,1841]" pageId="8" pageNumber="9" pagination="225 - 227" refId="ref17115" refString="Royle, J. F. (1835) Saxifrageae. In: Royle, J. F. (Ed.) Illustrations of the botany and other branches of the natural history of the Himalayan Mountains, and of the Flora of Cashmere. William Allen, London, pp. 225 - 227 http: // doi. org / 10.5962 / bhl. title. 449" type="book chapter" year="1835">Royle (1835: 227)</bibRefCitation>
</taxonomicName>
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</heading>
</paragraph>
</subSubSection>
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<paragraph id="8BF6AA19FFD9FFE4FF76F887FA36F8AC" blockId="8.[136,1452,1854,2093]" box="[136,1452,1854,1875]" pageId="8" pageNumber="9">
<collectionCode id="ED5832DCFFD9FFE4FF76F887FF5FF8AC" box="[136,197,1854,1875]" pageId="8" pageNumber="9">TYPE</collectionCode>
:—
<collectingCountry id="F35EEA89FFD9FFE4FF1FF887FEABF8AC" box="[225,305,1854,1875]" name="Nepal" pageId="8" pageNumber="9">NEPAL</collectingCountry>
. Gossain-than (Gossain-kund),
<emphasis id="B93D760BFFD9FFE4FD9BF887FCA1F8AC" box="[613,827,1854,1875]" italics="true" pageId="8" pageNumber="9">Wallich Cat. n. 1246a</emphasis>
(
<typeStatus id="54F214BBFFD9FFE4FCB4F887FC3DF8AC" box="[842,935,1854,1875]" pageId="8" pageNumber="9" type="lectotype">lectotype</typeStatus>
:
<collectionCode id="ED5832DCFFD9FFE4FC4FF887FC53F8AC" box="[945,969,1854,1875]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="8" pageNumber="9" type="Herbarium">W</collectionCode>
1889-0305721, designated by
<bibRefCitation id="EFD8D7E8FFD9FFE4FB0EF887FA36F8AC" author="Schonbeck-Temesy, E." box="[1264,1452,1854,1875]" pageId="8" pageNumber="9" pagination="1 - 4" refId="ref17269" refString="Schonbeck-Temesy, E. (1966) Parnassiaceae. In: Rechinger, K. H. (Ed.) Flora Iranica. Graz, Austria, pp. 1 - 4." type="book chapter" year="1966">Schönbeck-Temesy</bibRefCitation>
</paragraph>
<paragraph id="8BF6AA19FFD9FFE4FF43F8DBFA36F888" blockId="8.[136,1452,1854,2093]" box="[189,1452,1890,1911]" pageId="8" pageNumber="9">
1966;
<materialsCitation id="3B21A044FFD9FFE4FEFBF8DBFA3DF888" box="[261,1447,1890,1911]" collectionCode="E, GZU, NY, BM, L" pageId="8" pageNumber="9" specimenCode="E00301173, E00301174, GZU000100103, NY00186147, BM000947005, BM000521923, L0281535" specimenCount="1" typeStatus="isolectotype">
<typeStatus id="54F214BBFFD9FFE4FEFBF8DBFE1CF888" box="[261,390,1890,1911]" pageId="8" pageNumber="9" type="isolectotype">isolectotypes</typeStatus>
:
<specimenCode id="DBEF0262FFD9FFE4FE64F8DBFD91F888" box="[410,523,1890,1911]" collectionCode="E" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="8" pageNumber="9" type="Herbarium">E00301173</specimenCode>
,
<specimenCode id="DBEF0262FFD9FFE4FDE1F8DBFD15F888" box="[543,655,1890,1911]" collectionCode="E" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="8" pageNumber="9" type="Herbarium">E00301174</specimenCode>
,
<specimenCode id="DBEF0262FFD9FFE4FD5AF8DBFCDEF888" box="[676,836,1890,1911]" collectionCode="GZU" country="Austria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15750" lsid="urn:lsid:biocol.org:col:15750" name="Karl-Franzens-Universität Graz" pageId="8" pageNumber="9" type="Herbarium">GZU000100103</specimenCode>
,
<specimenCode id="DBEF0262FFD9FFE4FCA6F8DBFC47F888" box="[856,989,1890,1911]" collectionCode="NY" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="8" pageNumber="9" type="Herbarium">NY00186147</specimenCode>
,
<specimenCode id="DBEF0262FFD9FFE4FC0CF8DBFB1CF888" box="[1010,1158,1890,1911]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="8" pageNumber="9">BM000947005</specimenCode>
,
<specimenCode id="DBEF0262FFD9FFE4FB64F8DBFAB4F888" box="[1178,1326,1890,1911]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="8" pageNumber="9">BM000521923</specimenCode>
,
<specimenCode id="DBEF0262FFD9FFE4FABDF8DBFA3DF888" box="[1347,1447,1890,1911]" collectionCode="L" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="8" pageNumber="9" type="Herbarium">L0281535</specimenCode>
</materialsCitation>
,
</paragraph>
<paragraph id="8BF6AA19FFD9FFE4FF43F83FFA36F7D2" blockId="8.[136,1452,1854,2093]" pageId="8" pageNumber="9">
<collectionCode id="ED5832DCFFD9FFE4FF43F83FFF54F864" box="[189,206,1926,1947]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="8" pageNumber="9" type="Herbarium">K</collectionCode>
000739481,
<collectionCode id="ED5832DCFFD9FFE4FEB9F83FFEC2F864" box="[327,344,1926,1947]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="8" pageNumber="9" type="Herbarium">K</collectionCode>
000739482)
<taxonomicName id="4C49D19AFFD9FFE4FF76F813FDEFF840" authority="Drude (1875: 316)" authorityName="Drude" authorityPageNumber="316" authorityYear="1875" box="[136,629,1962,1983]" class="Magnoliopsida" family="Parnassiaceae" genus="Parnassia" kingdom="Plantae" order="Celastrales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="variety" species="nubicola" variety="cordata">
<emphasis id="B93D760BFFD9FFE4FF76F813FED9F840" box="[136,323,1962,1983]" italics="true" pageId="8" pageNumber="9">Parnassia nubicola</emphasis>
var.
<emphasis id="B93D760BFFD9FFE4FE8DF813FE27F840" box="[371,445,1962,1983]" italics="true" pageId="8" pageNumber="9">cordata</emphasis>
<bibRefCitation id="EFD8D7E8FFD9FFE4FE3CF813FDEFF840" author="Drude, O." box="[450,629,1962,1983]" pageId="8" pageNumber="9" pagination="239 - 324" refId="ref14013" refString="Drude, O. (1875) Uber die Bluthengestaltung und die Verwandtschaftsverhaltnisse des Genus Parnassia, nebst einer systematischen Revision seiner Arten. Linnaea 39: 239 - 324. Available from: www. biodiversitylibrary. org / item / 10885 (accessed 1 June 2017)" type="journal article" year="1875">Drude (1875: 316)</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C49D19AFFD9FFE4FD77F813FB0EF840" authority="(Drude) Jien ex T. C. Ku (1987: 38)" authorityName="Jien ex T. C. Ku" authorityPageNumber="38" authorityYear="1987" baseAuthorityName="Drude" box="[649,1172,1961,1983]" class="Magnoliopsida" family="Parnassiaceae" genus="Parnassia" kingdom="Plantae" order="Celastrales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="cordata">
<emphasis id="B93D760BFFD9FFE4FD77F813FCA6F840" bold="true" box="[649,828,1961,1983]" italics="true" pageId="8" pageNumber="9">Parnassia cordata</emphasis>
(Drude) Jien ex T.C.
<bibRefCitation id="EFD8D7E8FFD9FFE4FBF2F813FB0EF840" author="Ku, T. C." box="[1036,1172,1962,1983]" pageId="8" pageNumber="9" pagination="1 - 61" refId="ref15603" refString="Ku, T. C. (1987) A revision of the genus Parnassia (Saxifragaceae) in China. Bulletin of Botanical Research (Harbin) 7 (1): 1 - 61." type="journal article" year="1987">Ku (1987: 38)</bibRefCitation>
</taxonomicName>
<collectionCode id="ED5832DCFFD9FFE4FF76F877FF5DF81C" box="[136,199,1998,2019]" pageId="8" pageNumber="9">TYPE</collectionCode>
:—
<collectingCountry id="F35EEA89FFD9FFE4FF1DF877FEB0F81C" box="[227,298,1998,2019]" name="India" pageId="8" pageNumber="9">INDIA</collectingCountry>
. Kumaon, Blinkworth s.n.,
<emphasis id="B93D760BFFD9FFE4FDCCF877FC99F81C" box="[562,771,1998,2019]" italics="true" pageId="8" pageNumber="9">Wallich Cat. n. 1246b</emphasis>
(
<typeStatus id="54F214BBFFD9FFE4FCEEF877FCF7F81C" box="[784,877,1998,2019]" pageId="8" pageNumber="9" type="lectotype">lectotype</typeStatus>
:
<collectionCode id="ED5832DCFFD9FFE4FC88F877FC1EF81C" box="[886,900,1998,2019]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="8" pageNumber="9" type="Herbarium">L</collectionCode>
0281535 designated here by marker II) Notes:
<taxonomicName id="4C49D19AFFD9FFE4FF29F849FE3FF7F6" authorityName="Jien ex T. C. Ku" authorityYear="1987" baseAuthorityName="Drude" box="[215,421,2032,2057]" class="Magnoliopsida" family="Parnassiaceae" genus="Parnassia" kingdom="Plantae" order="Celastrales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="cordata">
<emphasis id="B93D760BFFD9FFE4FF29F849FE3FF7F6" box="[215,421,2032,2057]" italics="true" pageId="8" pageNumber="9">Parnassia cordata</emphasis>
</taxonomicName>
was an accepted name in the latest treatment by
<bibRefCitation id="EFD8D7E8FFD9FFE4FC3FF849FB26F7F6" author="Ku, T. C. &amp; Hultgard, U. M." box="[961,1212,2032,2057]" pageId="8" pageNumber="9" pagination="95 - 118" refId="ref15749" refString="Ku, T. C. &amp; Hultgard, U. M. (2001) Parnassia. In: Wu, Z. Y. &amp; Raven, P. H. (Eds.) Flora of China, vol. 8. Missouri Botanical Garden Press, St. Louis and Science Press, Beijing, pp. 95 - 118." type="book chapter" year="2001">Ku &amp; Hultgard (2001)</bibRefCitation>
, but the type material was related to that of
<taxonomicName id="4C49D19AFFD9FFE4FE89F7ADFE6EF7D2" authorityName="Wallich ex Royle" authorityYear="1835" box="[375,500,2068,2093]" class="Magnoliopsida" family="Parnassiaceae" genus="Parnassia" kingdom="Plantae" order="Celastrales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="nubicola">
<emphasis id="B93D760BFFD9FFE4FE89F7ADFE6EF7D2" box="[375,500,2068,2093]" italics="true" pageId="8" pageNumber="9">P. nubicola</emphasis>
</taxonomicName>
, so we arranged it here.
<taxonomicName id="4C49D19AFFD9FFE4FCFDF7ADFC47F7D2" authorityName="Wallich ex Royle" authorityYear="1835" box="[771,989,2068,2093]" class="Magnoliopsida" family="Parnassiaceae" genus="Parnassia" kingdom="Plantae" order="Celastrales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="nubicola">
<emphasis id="B93D760BFFD9FFE4FCFDF7ADFC47F7D2" box="[771,989,2068,2093]" italics="true" pageId="8" pageNumber="9">Parnassia nubicola</emphasis>
</taxonomicName>
was formally described based on
<emphasis id="B93D760BFFD9FFE4FAA9F7ADFA36F7D2" box="[1367,1452,2068,2093]" italics="true" pageId="8" pageNumber="9">Wallich</emphasis>
</paragraph>
<paragraph id="8BF6AA19FFD8FFE5FF76FF19FC17FD72" blockId="9.[136,1452,160,653]" pageId="9" pageNumber="10">
<emphasis id="B93D760BFFD8FFE5FF76FF19FE88FF46" box="[136,274,160,185]" italics="true" pageId="9" pageNumber="10">Cat. n. 1246</emphasis>
, which contains two gatherings, one by Wallich from Gossain Than (
<emphasis id="B93D760BFFD8FFE5FBE9FF19FA90FF46" box="[1047,1290,160,185]" italics="true" pageId="9" pageNumber="10">Wallich Cat. n. 1246a</emphasis>
) and the other by Blinkworth from Kumaon (
<emphasis id="B93D760BFFD8FFE5FE10FF7DFD76FF22" box="[494,748,196,221]" italics="true" pageId="9" pageNumber="10">Wallich Cat. n. 1246b</emphasis>
). Several specimens of both gatherings were traced, and we found that both gatherings are mixed in some individuals that should be more accurately determined as
<taxonomicName id="4C49D19AFFD8FFE5FAE6FF51FA3EFEFE" authorityName="Wallich ex Wight &amp; Arnott" authorityYear="1834" box="[1304,1444,232,257]" class="Magnoliopsida" family="Parnassiaceae" genus="Parnassia" kingdom="Plantae" order="Celastrales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="wightiana">
<emphasis id="B93D760BFFD8FFE5FAE6FF51FA3EFEFE" box="[1304,1444,232,257]" italics="true" pageId="9" pageNumber="10">P. wightiana</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFD8D7E8FFD8FFE5FF76FEB5FE29FEDA" author="Schonbeck-Temesy, E." box="[136,435,268,293]" pageId="9" pageNumber="10" pagination="1 - 4" refId="ref17269" refString="Schonbeck-Temesy, E. (1966) Parnassiaceae. In: Rechinger, K. H. (Ed.) Flora Iranica. Graz, Austria, pp. 1 - 4." type="book chapter" year="1966">Schönbeck-Temesy (1966)</bibRefCitation>
selected a sheet of
<emphasis id="B93D760BFFD8FFE5FD78FEB5FCEFFEDA" box="[646,885,268,293]" italics="true" pageId="9" pageNumber="10">Wallich Cat. n. 1246a</emphasis>
from
<collectionCode id="ED5832DCFFD8FFE5FC48FEB5FC4AFEDA" box="[950,976,268,293]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="9" pageNumber="10" type="Herbarium">W</collectionCode>
as the
<typeStatus id="54F214BBFFD8FFE5FBE7FEB5FB19FEDA" box="[1049,1155,268,293]" pageId="9" pageNumber="10" type="lectotype">lectotype</typeStatus>
. Based on these specimens collected from Kumaon, Drude described
<taxonomicName id="4C49D19AFFD8FFE5FD9DFE89FCEBFEB6" authorityName="Drude" authorityYear="1875" box="[611,881,304,329]" class="Magnoliopsida" family="Parnassiaceae" genus="Parnassia" kingdom="Plantae" order="Celastrales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="variety" species="nubicola" variety="cordata">
<emphasis id="B93D760BFFD8FFE5FD9DFE89FD78FEB6" box="[611,738,304,329]" italics="true" pageId="9" pageNumber="10">P. nubicola</emphasis>
var.
<emphasis id="B93D760BFFD8FFE5FCE5FE89FCEBFEB6" box="[795,881,304,329]" italics="true" pageId="9" pageNumber="10">cordata</emphasis>
</taxonomicName>
, but no
<typeStatus id="54F214BBFFD8FFE5FC31FE89FBABFEB6" box="[975,1073,304,329]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
was indicated. In the protologue, he mentioned
<emphasis id="B93D760BFFD8FFE5FED2FEEDFDE8FE6E" italics="true" pageId="9" pageNumber="10">
foliis cordatis
<quantity id="4CB107FCFFD8FFE5FE25FEEDFD90FE92" box="[475,522,340,365]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="10" unit="cm" value="3.0">3cm</quantity>
longis et paulo latioribus petalisque ad basin magis fimbriatis diversa; haec ad Parnassiam Wrightianam propius accedit.
</emphasis>
From this description, we can speculate that Drude had seen the mixed specimens and named the individuals with fimbriate petals and cordate leaves as
<taxonomicName id="4C49D19AFFD8FFE5FBE0FE25FB3FFE4A" authorityName="Drude" authorityYear="1875" box="[1054,1189,412,437]" class="Magnoliopsida" family="Parnassiaceae" genus="Parnassia" kingdom="Plantae" order="Celastrales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="variety" species="nubicola" variety="cordata">
var.
<emphasis id="B93D760BFFD8FFE5FBB1FE25FB3FFE4A" box="[1103,1189,412,437]" italics="true" pageId="9" pageNumber="10">cordata</emphasis>
</taxonomicName>
, and he also noted that these materials may belong to
<taxonomicName id="4C49D19AFFD8FFE5FE19FE79FDEFFE26" authorityName="Wallich ex Wight &amp; Arnott" authorityYear="1834" box="[487,629,448,473]" class="Magnoliopsida" family="Parnassiaceae" genus="Parnassia" kingdom="Plantae" order="Celastrales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="wightiana">
<emphasis id="B93D760BFFD8FFE5FE19FE79FDEFFE26" box="[487,629,448,473]" italics="true" pageId="9" pageNumber="10">P. wightiana</emphasis>
</taxonomicName>
. Eight sheets of
<emphasis id="B93D760BFFD8FFE5FCC7FE79FBA9FE26" box="[825,1075,448,473]" italics="true" pageId="9" pageNumber="10">Wallich Cat. n. 1246b</emphasis>
have been found, in which most of the materials have ovate leaves and subentire petals, thus do not match the descriptions.
<collectionCode id="ED5832DCFFD8FFE5FB65FE5DFB34FE02" box="[1179,1198,484,509]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="9" pageNumber="10" type="Herbarium">A</collectionCode>
scape marked II from
<collectionCode id="ED5832DCFFD8FFE5FF76FDB1FF03FDDE" box="[136,153,520,545]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="9" pageNumber="10" type="Herbarium">L</collectionCode>
0281535 has a cordate leaf and proximally fimbriate petals, which should be determined as
<taxonomicName id="4C49D19AFFD8FFE5FB4DFDB1FADAFDDE" authorityName="Wallich ex Wight &amp; Arnott" authorityYear="1834" box="[1203,1344,520,545]" class="Magnoliopsida" family="Parnassiaceae" genus="Parnassia" kingdom="Plantae" order="Celastrales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="wightiana">
<emphasis id="B93D760BFFD8FFE5FB4DFDB1FADAFDDE" box="[1203,1344,520,545]" italics="true" pageId="9" pageNumber="10">P. wightiana</emphasis>
</taxonomicName>
, matches the description well, and this should be what Drude intended to describe. Since no specimens labeled with Wallichs number have been found from
<collectionCode id="ED5832DCFFD8FFE5FE12FDE9FDA2FD96" box="[492,568,592,617]" country="Germany" httpUri="http://grbio.org/cool/pq74-a94z" name="Universität Göttingen" pageId="9" pageNumber="10" type="Herbarium">GOET</collectionCode>
, where Drude worked when he described this new variety, we designated the individual marked II from
<collectionCode id="ED5832DCFFD8FFE5FE4DFDCDFE5EFD72" box="[435,452,628,653]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="9" pageNumber="10" type="Herbarium">L</collectionCode>
0281535 as the
<typeStatus id="54F214BBFFD8FFE5FD8AFDCDFD41FD72" box="[628,731,628,653]" pageId="9" pageNumber="10" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C49D19AFFD8FFE5FCFEFDCDFC1CFD72" authorityName="Drude" authorityYear="1875" box="[768,902,628,653]" class="Magnoliopsida" family="Parnassiaceae" genus="Parnassia" kingdom="Plantae" order="Celastrales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="variety" species="nubicola" variety="cordata">
var.
<emphasis id="B93D760BFFD8FFE5FCCEFDCDFC1CFD72" box="[816,902,628,653]" italics="true" pageId="9" pageNumber="10">cordata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>