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<document id="910F6573B4EB4DF8FB68DD771592BB82" ID-DOI="10.11646/zootaxa.3872.3.1" ID-GBIF-Dataset="47ff6515-876a-4d83-b631-64004e51c683" ID-ISSN="1175-5326" ID-Zenodo-Dep="230974" ID-ZooBank="72ACE4FF-9A41-4D26-A201-01E020439899" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460926869511" checkinUser="plazi" docAuthor="Pessacq, Pablo" docDate="2014" docId="03D3FD33FFB2FFFF4E8F71C3F474FB44" docLanguage="en" docName="zt03872p234.pdf" docOrigin="Zootaxa 3872 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epipleoneura kaxuriana Machado 1985" docType="treatment" docVersion="10" lastPageNumber="221" masterDocId="FFEA854BFFA2FFEB4E18773AF60FFFC8" masterDocTitle="Synopsis of Epipleoneura (Zygoptera, Coenagrionidae, “ Protoneuridae ”), with emphasis on its Brazilian species" masterLastPageNumber="234" masterPageNumber="201" pageNumber="217" updateTime="1698362651403" updateUser="plazi">
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<mods:title id="25614DFF243E1E3D69E592192230CECD">Synopsis of Epipleoneura (Zygoptera, Coenagrionidae, “ Protoneuridae ”), with emphasis on its Brazilian species</mods:title>
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<mods:namePart id="EDEBF68674D49384A39D5F6FBE8B6A39">Pessacq, Pablo</mods:namePart>
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<mods:date id="1E8B32698FAE91B6D9B9CB68C0BC21F4">2014</mods:date>
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<treatment id="03D3FD33FFB2FFFF4E8F71C3F474FB44" ID-DOI="http://doi.org/10.5281/zenodo.5296822" ID-GBIF-Taxon="119590007" ID-Zenodo-Dep="5296822" LSID="urn:lsid:plazi:treatment:03D3FD33FFB2FFFF4E8F71C3F474FB44" httpUri="http://treatment.plazi.org/id/03D3FD33FFB2FFFF4E8F71C3F474FB44" lastPageId="20" lastPageNumber="221" pageId="16" pageNumber="217">
<subSubSection id="C3601FAEFFB2FFFB4E8F71C3F745F8FD" pageId="16" pageNumber="217" type="nomenclature">
<paragraph id="8BC54C25FFB2FFFB4E8F71C3F489F8DB" blockId="16.[151,646,1785,1845]" box="[151,646,1785,1811]" pageId="16" pageNumber="217">
<heading id="D08DFB49FFB2FFFB4E8F71C3F489F8DB" bold="true" box="[151,646,1785,1811]" fontSize="11" level="1" pageId="16" pageNumber="217" reason="1">
<taxonomicName id="4C7A37A6FFB2FFFB4E8F71C3F489F8DB" ID-CoL="6G6VB" authority="Machado, 1985" authorityName="Machado" authorityYear="1985" box="[151,646,1785,1811]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="kaxuriana">
<emphasis id="B90E9037FFB2FFFB4E8F71C3F489F8DB" bold="true" box="[151,646,1785,1811]" pageId="16" pageNumber="217">
<emphasis id="B90E9037FFB2FFFB4E8F71C3F7CFF8DB" bold="true" box="[151,448,1785,1811]" italics="true" pageId="16" pageNumber="217">Epipleoneura kaxuriana</emphasis>
<bibRefCitation id="EFEB31D4FFB2FFFB4FDF71C3F489F8DB" author="Machado" box="[455,646,1785,1811]" pageId="16" pageNumber="217" refString="Machado, A. B. M. (1985) Studies on Neotropical Protoneuridae. 5. Redescription of the type of Phasmoneura ephippigera (Selys, 1886) (Zygoptera, Protoneuridae). Odonatologica, 14, 363 - 368." type="journal article" year="1985">Machado, 1985</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC54C25FFB2FFFB4E8F7026F745F8FD" blockId="16.[151,646,1785,1845]" box="[151,330,1820,1845]" pageId="16" pageNumber="217">
(
<figureCitation id="134150A0FFB2FFFB4E877026F6E3F8FC" box="[159,236,1820,1845]" captionStart="FIGURES 1 3" captionStartId="5.[151,264,1656,1678]" captionTargetBox="[154,1428,189,1639]" captionTargetId="figure@5.[143,1429,189,1639]" captionTargetPageId="5" captionText="FIGURES 1 3. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. AH: apical hook. DB: cercus dorsal branch. VB: cercus ventral branch. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth. TS: triangular swollen structure." httpUri="https://zenodo.org/record/230975/files/figure.png" pageId="16" pageNumber="217">Figs. 1</figureCitation>
,
<figureCitation id="134150A0FFB2FFFB4EE27026F716F8FC" box="[250,281,1820,1844]" captionStart="FIGURES 28 47" captionStartId="17.[151,264,1969,1991]" captionTargetBox="[233,1339,231,1909]" captionTargetId="figure@17.[212,1378,204,1943]" captionTargetPageId="17" captionText="FIGURES 28 47. Genital ligula, lateral and ectal view. Figures 29, 33, 47 modified from Garrison et al. (2010). PLP: posterolateral projection." httpUri="https://zenodo.org/record/230984/files/figure.png" pageId="16" pageNumber="217">40</figureCitation>
,
<figureCitation id="134150A0FFB2FFFB4F3D7026F74DF8FD" box="[293,322,1820,1845]" captionStart="FIGURES 53 66" captionStartId="19.[151,264,1880,1902]" captionTargetBox="[166,1421,208,1847]" captionTargetId="figure@19.[151,1436,193,1858]" captionTargetPageId="19" captionText="FIGURES 53 66. Female pronotum in lateral, dorsal, antero-lateral, anterior postero-lateral and posterior view. Figures 57, 64 and 65 modified from Garrison et al. (2010), Rácenis (1960) and Machado (2005) respectively." httpUri="https://zenodo.org/record/230986/files/figure.png" pageId="16" pageNumber="217">54</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3601FAEFFB2FFF94E8F705EF786FAFE" lastPageId="18" lastPageNumber="219" pageId="16" pageNumber="217" type="reference_group">
<paragraph id="8BC54C25FFB2FFFB4E8F705EF7C7F83C" blockId="16.[151,1436,1892,2036]" pageId="16" pageNumber="217">
<treatmentCitationGroup id="AB6A6B0BFFB2FFFB4E8F705EF7C7F83C" infered="true" pageId="16" pageNumber="217">
<taxonomicName id="4C7A37A6FFB2FFFB4E8F705EF451F8B2" ID-CoL="6G6VB" authority="Machado 1985: 14" authorityName="Machado" authorityPageNumber="14" authorityYear="1985" box="[151,606,1892,1914]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="kaxuriana">
<emphasis id="B90E9037FFB2FFFB4E8F705EF780F8B2" box="[151,399,1892,1914]" italics="true" pageId="16" pageNumber="217">Epipleoneura kaxuriana</emphasis>
<treatmentCitation id="0ADB6A34FFB2FFFB4F8E705EF451F8B2" author="Machado" box="[406,606,1892,1914]" page="14" pageId="16" pageNumber="217" year="1985">
<bibRefCitation id="EFEB31D4FFB2FFFB4F8E705EF43AF8B2" author="Machado" box="[406,565,1892,1914]" pageId="16" pageNumber="217" refString="Machado, A. B. M. (1985) Studies on Neotropical Protoneuridae. 5. Redescription of the type of Phasmoneura ephippigera (Selys, 1886) (Zygoptera, Protoneuridae). Odonatologica, 14, 363 - 368." type="journal article" year="1985">
<emphasis id="B90E9037FFB2FFFB4F8E705EF43AF8B2" bold="true" box="[406,565,1892,1914]" pageId="16" pageNumber="217">Machado 1985</emphasis>
</bibRefCitation>
: 14
</treatmentCitation>
</taxonomicName>
, 17, 28, 29 (new species, description of male, scanning of cercus and epiproct in posterior and lateral view, detail of epiproct,
<typeStatus id="54C1F287FFB2FFFB4C9470B9F4B7F850" box="[652,696,1923,1944]" pageId="16" pageNumber="217">type</typeStatus>
material at ABMM).
<treatmentCitation id="0ADB6A34FFB2FFFB4D8C70B8F26FF850" author="Lencioni" box="[916,1120,1922,1944]" page="177" pageId="16" pageNumber="217" year="2005">
<bibRefCitation id="EFEB31D4FFB2FFFB4D8C70B8F224F850" author="Lencioni" box="[916,1067,1922,1944]" pageId="16" pageNumber="217" refString="Lencioni, F. A. A. (2005) Damselflies of Brazil. An Illustrated Identification Guide. 1 - Non-Coenagrionidae Families. All Print Editora, Sao Paulo, 324 pp." type="book" year="2005">
<emphasis id="B90E9037FFB2FFFB4D8C70B8F224F850" bold="true" box="[916,1067,1922,1944]" pageId="16" pageNumber="217">Lencioni 2005</emphasis>
</bibRefCitation>
: 177
</treatmentCitation>
, figs. 107 ab (references,
<typeStatus id="54C1F287FFB2FFFB4B6870B9F393F850" box="[1392,1436,1923,1944]" pageId="16" pageNumber="217">type</typeStatus>
locality, distribution, described stages, illustrations of male S
<quantity id="4C82E1C0FFB2FFFB4D2D709BF565F87F" box="[821,874,1953,1975]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="217" unit="in" value="10.0">10 in</quantity>
posterior and lateral view).
<treatmentCitation id="0ADB6A34FFB2FFFB4A93709BF397F87F" author="Garrison" box="[1163,1432,1952,1975]" page="352" pageId="16" pageNumber="217" year="2010">
<bibRefCitation id="EFEB31D4FFB2FFFB4A93709BF36DF87F" author="Garrison" box="[1163,1378,1952,1975]" pageId="16" pageNumber="217" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2010) Damselfly genera of the New World. An Illustrated and Annotated Key to the Zygoptera. The Johns Hopkins University Press, Baltimore, 490 pp." type="book" year="2010">
<emphasis id="B90E9037FFB2FFFB4A93709BF36DF87F" bold="true" box="[1163,1378,1952,1975]" pageId="16" pageNumber="217">
Garrison
<emphasis id="B90E9037FFB2FFFB4AEC7098F327F87F" bold="true" box="[1268,1320,1952,1975]" italics="true" pageId="16" pageNumber="217">et al.</emphasis>
2010
</emphasis>
</bibRefCitation>
: 352
</treatmentCitation>
, 355 (included in list of species, illustrations of epiproct in posterior view).
<treatmentCitation id="0ADB6A34FFB2FFFB4DFA70FAF2C1F81E" author="Pessacq" box="[994,1230,1983,2006]" page="4" pageId="16" pageNumber="217" year="2012">
<bibRefCitation id="EFEB31D4FFB2FFFB4DFA70FAF2BDF81E" author="Pessacq" box="[994,1202,1983,2006]" pageId="16" pageNumber="217" refString="Pessacq, P., Santos, T. C. &amp; Costa, JM. (2012) Checklist and updated distribution of Protoneuridae from Brazil. International Journal of Odonatology, 15 (1), 1 - 15. http: // dx. doi. org / 10.1080 / 13887890.2012.672158" type="journal article" year="2012">
<emphasis id="B90E9037FFB2FFFB4DFA70FAF2BDF81E" bold="true" box="[994,1202,1983,2006]" pageId="16" pageNumber="217">
Pessacq
<emphasis id="B90E9037FFB2FFFB4A5870FAF279F81E" bold="true" box="[1088,1142,1983,2006]" italics="true" pageId="16" pageNumber="217">et al.</emphasis>
2012
</emphasis>
</bibRefCitation>
: 4
</treatmentCitation>
(included in list of Brazilian
<taxonomicName id="4C7A37A6FFB2FFFB4F3270E4F7B2F83C" ID-CoL="F5S" box="[298,445,2014,2036]" class="Insecta" family="Protoneuridae" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="217" phylum="Arthropoda" rank="family">Protoneuridae</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<caption id="DF051CADFFB3FFFA4E8F708BF745F82D" httpUri="https://zenodo.org/record/230984/files/figure.png" pageId="17" pageNumber="218" targetBox="[233,1339,231,1909]" targetPageId="17">
<paragraph id="8BC54C25FFB3FFFA4E8F708BF745F82D" blockId="17.[151,1436,1969,2021]" pageId="17" pageNumber="218">
<emphasis id="B90E9037FFB3FFFA4E8F708BF75BF80F" bold="true" box="[151,340,1969,1991]" pageId="17" pageNumber="218">FIGURES 2847.</emphasis>
Genital ligula, lateral and ectal view. Figures 29, 33, 47 modified from Garrison
<emphasis id="B90E9037FFB3FFFA4A9F7088F2BBF80F" box="[1159,1204,1969,1991]" italics="true" pageId="17" pageNumber="218">et al</emphasis>
. (2010). PLP: posterolateral projection.
</paragraph>
</caption>
<caption id="DF051CADFFB0FFF94E8F7355F331FB4D" box="[151,1342,1135,1157]" httpUri="https://zenodo.org/record/230985/files/figure.png" pageId="18" pageNumber="219" targetBox="[359,1226,206,1082]" targetPageId="18">
<paragraph id="8BC54C25FFB0FFF94E8F7355F331FB4D" blockId="18.[151,1342,1135,1157]" box="[151,1342,1135,1157]" pageId="18" pageNumber="219">
<emphasis id="B90E9037FFB0FFF94E8F7355F75BFB4D" bold="true" box="[151,340,1135,1157]" pageId="18" pageNumber="219">FIGURES 4852.</emphasis>
Genital ligula, a: lateral view; b: ectal view. Figures 48 and 51 modified from De Marmels (1989).
</paragraph>
</caption>
<paragraph id="8BC54C25FFB0FFF94E8F738BF786FAFE" blockId="18.[151,1437,1201,2018]" pageId="18" pageNumber="219">
<emphasis id="B90E9037FFB0FFF94E8F738BF796FB02" bold="true" box="[151,409,1201,1226]" pageId="18" pageNumber="219">Specimens examined.</emphasis>
<specimenCount id="9D7C87ACFFB0FFF94FBA7388F7C7FB02" box="[418,456,1202,1226]" pageId="18" pageNumber="219" type="generic">4♂</specimenCount>
, 2♀.
<specimenCount id="9D7C87ACFFB0FFF94C127388F420FB02" box="[522,559,1202,1226]" pageId="18" pageNumber="219" type="generic">3♂</specimenCount>
,
<collectingCountry id="F36D0CB5FFB0FFF94C26738BF48AFB02" box="[574,645,1201,1226]" name="Brazil" pageId="18" pageNumber="219">Brazil</collectingCountry>
, Amazonas State, small stream within the forest, Igarapé, km 3040 from Road Itacoatiara to Manaus, (
<geoCoordinate id="EE4E2AE2FFB0FFF94C3673EFF4BCFB26" box="[558,691,1237,1262]" direction="south" orientation="latitude" pageId="18" pageNumber="219" precision="15" value="-3.1055555">03º0620”S</geoCoordinate>
,
<geoCoordinate id="EE4E2AE2FFB0FFF94CDA73EFF55DFB26" box="[706,850,1237,1262]" direction="west" orientation="longitude" pageId="18" pageNumber="219" precision="15" value="-60.0225">60º0121”W</geoCoordinate>
), leg. N.D. Santos,
<date id="FFC46AE5FFB0FFF94A2073ECF2BEFB26" box="[1080,1201,1237,1262]" pageId="18" pageNumber="219" value="1959-10-27">27 x 1959</date>
.
<specimenCount id="9D7C87ACFFB0FFF94AA773ECF2E2FB25" box="[1215,1261,1238,1262]" pageId="18" pageNumber="219" type="generic">1 ♂</specimenCount>
, same data but Manaus, Colonia Santo Antonio, Igarapé do Alberto (Colonia Santo Antonio:
<geoCoordinate id="EE4E2AE2FFB0FFF94DE773C3F28BFADA" box="[1023,1156,1273,1298]" direction="south" orientation="latitude" pageId="18" pageNumber="219" precision="15" value="-3.0402777">03º0225”S</geoCoordinate>
,
<geoCoordinate id="EE4E2AE2FFB0FFF94A9773C3F310FADA" box="[1167,1311,1273,1298]" direction="west" orientation="longitude" pageId="18" pageNumber="219" precision="15" value="-60.021946">60º0119”W</geoCoordinate>
), leg. M.C. de Melo,
<date id="FFC46AE5FFB0FFF94F197224F78AFAFE" box="[257,389,1309,1334]" pageId="18" pageNumber="219" value="1960-07-29">29 vii 1960</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3601FAEFFB0FFF94EDF727BF49DF99E" pageId="18" pageNumber="219" type="description">
<paragraph id="8BC54C25FFB0FFF94EDF727BF49DF99E" blockId="18.[151,1437,1201,2018]" pageId="18" pageNumber="219">
<emphasis id="B90E9037FFB0FFF94EDF727BF7F5FA92" bold="true" box="[199,506,1345,1370]" pageId="18" pageNumber="219">Female description. Head</emphasis>
: Dorsally metallic green; antennifer dorsum black, frons yellow; antefrons yellow with lateral and dorsal margins black; postclypeus black, anteclypeus basally brown, distally yellow; labrum basally black, distally yellow; genae yellow; venter of head black, with two narrow yellow stripes bordering eyes; labrum and maxilla yellow.
<emphasis id="B90E9037FFB0FFF94FCE7297F420FA0E" bold="true" box="[470,559,1453,1478]" pageId="18" pageNumber="219">Thorax</emphasis>
: Prothorax dorsally metallic green, becoming yellow laterally, posterior lobe (
<figureCitation id="134150A0FFB0FFF94E8772EBF6FEFA22" box="[159,241,1489,1514]" captionStart="FIGURES 53 66" captionStartId="19.[151,264,1880,1902]" captionTargetBox="[166,1421,208,1847]" captionTargetId="figure@19.[151,1436,193,1858]" captionTargetPageId="19" captionText="FIGURES 53 66. Female pronotum in lateral, dorsal, antero-lateral, anterior postero-lateral and posterior view. Figures 57, 64 and 65 modified from Garrison et al. (2010), Rácenis (1960) and Machado (2005) respectively." httpUri="https://zenodo.org/record/230986/files/figure.png" pageId="18" pageNumber="219">Fig. 54</figureCitation>
) erect, with a slightly produced medial lobe and an antero-medial depression. Mesothorax metallic green, metathorax yellow, with a posterior metallic green area at the metepisternum; pterothoracic venter and legs light yellow; apex of femora light brown.
<emphasis id="B90E9037FFB0FFF94C367123F4ACF9FA" bold="true" box="[558,675,1561,1586]" pageId="18" pageNumber="219">Abdomen</emphasis>
: S12 dorsally metallic green, sides light brown, S37 dorsally dark brown, sides light brown, S810 dark brown.
</paragraph>
</subSubSection>
<subSubSection id="C3601FAEFFB0FFF94EDF715BF419F852" pageId="18" pageNumber="219" type="diagnosis">
<paragraph id="8BC54C25FFB0FFF94EDF715BF419F852" blockId="18.[151,1437,1201,2018]" pageId="18" pageNumber="219">
<emphasis id="B90E9037FFB0FFF94EDF715BF74EF9B2" bold="true" box="[199,321,1633,1658]" pageId="18" pageNumber="219">Diagnosis.</emphasis>
Male cercus resembles that described for
<taxonomicName id="4C7A37A6FFB0FFF94D3B7158F5DCF9B2" box="[803,979,1634,1658]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="capilliformis">
<emphasis id="B90E9037FFB0FFF94D3B7158F5DCF9B2" box="[803,979,1634,1658]" italics="true" pageId="18" pageNumber="219">E. capilliformis</emphasis>
</taxonomicName>
. The cercus possesses a well-developed inner-basal branch with its apex directed posteriorly, not reaching midline of S10. The genital ligula, cercus, and epiproct structure resemble those of
<emphasis id="B90E9037FFB0FFF94C287190F512F909" box="[560,797,1706,1730]" italics="true" pageId="18" pageNumber="219">
<taxonomicName id="4C7A37A6FFB0FFF94C287190F4CFF90A" box="[560,704,1706,1730]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="machadoi">E. machadoi</taxonomicName>
,
<taxonomicName id="4C7A37A6FFB0FFF94CD37190F512F909" box="[715,797,1706,1729]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="ottoi">E. ottoi</taxonomicName>
</emphasis>
and
<taxonomicName id="4C7A37A6FFB0FFF94D4C7190F5DCF90A" box="[852,979,1706,1730]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="westfalli">
<emphasis id="B90E9037FFB0FFF94D4C7190F5DCF90A" box="[852,979,1706,1730]" italics="true" pageId="18" pageNumber="219">E. westfalli</emphasis>
</taxonomicName>
but differ from all of them by the cercus inner-basal branch shape and size. Additionally, the base of the epiproct branches is not separated by a triangular swollen structure (TS,
<figureCitation id="134150A0FFB0FFF94F8F71CBF7D7F8C2" box="[407,472,1777,1802]" captionStart="FIGURES 1 3" captionStartId="5.[151,264,1656,1678]" captionTargetBox="[154,1428,189,1639]" captionTargetId="figure@5.[143,1429,189,1639]" captionTargetPageId="5" captionText="FIGURES 1 3. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. AH: apical hook. DB: cercus dorsal branch. VB: cercus ventral branch. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth. TS: triangular swollen structure." httpUri="https://zenodo.org/record/230975/files/figure.png" pageId="18" pageNumber="219">Fig. 2</figureCitation>
b) in
<taxonomicName id="4C7A37A6FFB0FFF94C0C71C8F4AAF8C2" box="[532,677,1778,1802]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="kaxuriana">
<emphasis id="B90E9037FFB0FFF94C0C71C8F4AAF8C2" box="[532,677,1778,1802]" italics="true" pageId="18" pageNumber="219">E. kaxuriana</emphasis>
</taxonomicName>
, which is present in the other species. See under
<taxonomicName id="4C7A37A6FFB0FFF94AD671C8F32EF8C1" box="[1230,1313,1778,1801]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="ottoi">
<emphasis id="B90E9037FFB0FFF94AD671C8F32EF8C1" box="[1230,1313,1778,1801]" italics="true" pageId="18" pageNumber="219">E. ottoi</emphasis>
</taxonomicName>
for further discussion. Segment 3 of genital ligula (
<figureCitation id="134150A0FFB0FFF94C78702FF4BCF8E6" box="[608,691,1813,1838]" captionStart="FIGURES 28 47" captionStartId="17.[151,264,1969,1991]" captionTargetBox="[233,1339,231,1909]" captionTargetId="figure@17.[212,1378,204,1943]" captionTargetPageId="17" captionText="FIGURES 28 47. Genital ligula, lateral and ectal view. Figures 29, 33, 47 modified from Garrison et al. (2010). PLP: posterolateral projection." httpUri="https://zenodo.org/record/230984/files/figure.png" pageId="18" pageNumber="219">Fig. 40</figureCitation>
) is slightly narrowed medially, its sides are slightly elevated, and the apex posses two small lobes, folded entally and with two small spines at the sides. The postero-lateral projections are widely rounded in lateral view, each with a short peduncle, barely visible, and a small rounded lobe basal to postero-lateral projection.
</paragraph>
</subSubSection>
<subSubSection id="C3601FAEFFB0FFFF4EDF709FF474FB44" lastPageId="20" lastPageNumber="221" pageId="18" pageNumber="219" type="distribution">
<paragraph id="8BC54C25FFB0FFF94EDF709FF20FF82A" blockId="18.[151,1437,1201,2018]" pageId="18" pageNumber="219">
<emphasis id="B90E9037FFB0FFF94EDF709FF76FF876" bold="true" box="[199,352,1957,1982]" pageId="18" pageNumber="219">Distribution.</emphasis>
Northern
<collectingCountry id="F36D0CB5FFB0FFF94FC2709FF411F876" box="[474,542,1957,1982]" name="Brazil" pageId="18" pageNumber="219">Brazil</collectingCountry>
in the Amazonas region, Amazonas, Pará, and Roraima States (
<typeStatus id="54C1F287FFB0FFF94B14709CF331F876" box="[1292,1342,1958,1982]" pageId="18" pageNumber="219">type</typeStatus>
locality Oriximiná, Porto Trombetas, Igarapé Saracazinho,
<geoCoordinate id="EE4E2AE2FFB0FFF94CCA70F3F559F82A" box="[722,854,1993,2018]" direction="south" orientation="latitude" pageId="18" pageNumber="219" precision="15" value="-1.7652777">01º4555”S</geoCoordinate>
,
<geoCoordinate id="EE4E2AE2FFB0FFF94D7A70F3F5FBF82A" box="[866,1012,1993,2018]" direction="west" orientation="longitude" pageId="18" pageNumber="219" precision="15" value="-55.866665">55º5160”W</geoCoordinate>
).
</paragraph>
<caption id="DF051CADFFB1FFF84E8F7062F253F844" httpUri="https://zenodo.org/record/230986/files/figure.png" pageId="19" pageNumber="220" targetBox="[166,1421,208,1847]" targetPageId="19">
<paragraph id="8BC54C25FFB1FFF84E8F7062F253F844" blockId="19.[151,1435,1880,1933]" pageId="19" pageNumber="220">
<emphasis id="B90E9037FFB1FFF84E8F7062F75BF8A6" bold="true" box="[151,340,1880,1902]" pageId="19" pageNumber="220">FIGURES 5366.</emphasis>
Female pronotum in lateral, dorsal, antero-lateral, anterior postero-lateral and posterior view. Figures 57, 64 and 65 modified from Garrison
<emphasis id="B90E9037FFB1FFF84FC17042F409F844" box="[473,518,1910,1933]" italics="true" pageId="19" pageNumber="220">et al</emphasis>
. (2010), Rácenis (1960) and Machado (2005) respectively.
</paragraph>
</caption>
<caption id="DF051CADFFB6FFFF4E8F7462F584FCA6" box="[151,907,856,878]" pageId="20" pageNumber="221">
<paragraph id="8BC54C25FFB6FFFF4E8F7462F584FCA6" blockId="20.[151,907,856,878]" box="[151,907,856,878]" pageId="20" pageNumber="221">
<emphasis id="B90E9037FFB6FFFF4E8F7462F75CFCA6" bold="true" box="[151,339,856,878]" pageId="20" pageNumber="221">FIGURES 6769.</emphasis>
Male epiproct, posterior view, morphological variation.
</paragraph>
</caption>
<paragraph id="8BC54C25FFB6FFFF4EDF74A1F474FB44" blockId="20.[151,1437,923,1164]" pageId="20" pageNumber="221">
<emphasis id="B90E9037FFB6FFFF4EDF74A1F71EFC7C" bold="true" box="[199,273,923,948]" pageId="20" pageNumber="221">Notes.</emphasis>
I have assigned a female to
<taxonomicName id="4C7A37A6FFB6FFFF4C4274A6F4E3FC7B" box="[602,748,923,947]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="kaxuriana">
<emphasis id="B90E9037FFB6FFFF4C4274A6F4E3FC7B" box="[602,748,923,947]" italics="true" pageId="20" pageNumber="221">E. kaxuriana</emphasis>
</taxonomicName>
since it was taken together with males of this species and it matches no other female known to me. In the collection of R.W. Garrison (pers. com.) there is a female taken in tandem with a male of
<taxonomicName id="4C7A37A6FFB6FFFF4FB874DEF43CFC33" box="[416,563,995,1019]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="kaxuriana">
<emphasis id="B90E9037FFB6FFFF4FB874DEF43CFC33" box="[416,563,995,1019]" italics="true" pageId="20" pageNumber="221">E. kaxuriana</emphasis>
</taxonomicName>
that agrees with the description herein presented. Machado (1980) mentions variation in thorax coloration, from metallic green to reddish metallic. One male and one female close to
<emphasis id="B90E9037FFB6FFFF4B9C7332F700FB8B" italics="true" pageId="20" pageNumber="221">
<taxonomicName id="4C7A37A6FFB6FFFF4B9C7332F703FB8B" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="kaxuriana">E. kaxuriana</taxonomicName>
,
</emphasis>
from the federal district in central
<collectingCountry id="F36D0CB5FFB6FFFF4C867311F4EDFB8C" box="[670,738,1067,1092]" name="Brazil" pageId="20" pageNumber="221">Brazil</collectingCountry>
differ slightly by epiproct and female prothorax morphology, but due to the lack of more material and the bad state of preservation of this pair, I have refrained from describing them pending acquisition of more material.
</paragraph>
</subSubSection>
</treatment>
</document>