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<document id="095D5CE1A94EF7629CCED58FF622D648" ID-CLB-Dataset="4006" ID-DOI="10.11646/zootaxa.5052.4.9" ID-GBIF-Dataset="146052d6-ccc2-48f3-b059-3757ce97ba4e" ID-ISSN="1175-5326" ID-Zenodo-Dep="5577387" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634626752609" checkinUser="plazi" docAuthor="Suzuki, Yuya &amp; Tanikawa, Akio" docDate="2021" docId="03B887DCFFE1315A70892721FD3EFC1F" docLanguage="en" docName="zootaxa.5052.4.9.pdf" docOrigin="Zootaxa 5052 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Araneus matsumotoi Suzuki &amp; Tanikawa 2021, new species" docType="treatment" docUuid="5F005F73-B712-49AC-ACE0-3B1C17A03077" docUuidSource="ZooBank" docVersion="5" lastPageNumber="595" masterDocId="FF81FFA4FFE0315E701E2272FFCFFFF4" masterDocTitle="A new species of the genus Araneus (Araneae: Araneidae) from Japan" masterLastPageNumber="596" masterPageNumber="591" pageNumber="592" updateTime="1699460224084" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="A8EDE352F561C1072039162AC3060045" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D69FDABE775E957298CCBBB8BA5216F0">
<mods:title id="61FFEA0A311F3F09D84F708EC5F2D452">A new species of the genus Araneus (Araneae: Araneidae) from Japan</mods:title>
</mods:titleInfo>
<mods:name id="F04093DFB9064742758AFA4F88C5BEA3" type="personal">
<mods:role id="B42E78F18B880285E7DF2A7DC2C0003C">
<mods:roleTerm id="2679DCAD3D365D3F073A561450A50943">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="91BE7A101280FD52E7A4B85210954CCF">Suzuki, Yuya</mods:namePart>
<mods:affiliation id="68AF69FF2C9E89EFAD14FDA33E01E00E">Graduate School of Life and Environmental Sciences, University of Tsukuba, 1 - 1 - 1, Tennodai, Tsukuba-shi, Ibaraki 305 - 8572 Japan. The United Graduate School of Agricultural Sciences, Kagoshima University, 1 - 21 - 24, Korimoto, Kagoshima-shi, Kagoshima 890 - 0065 Japan.</mods:affiliation>
</mods:name>
<mods:name id="EB25EDB453107EE1266818E9F39D493C" type="personal">
<mods:role id="B52CE92501A0ABCD38ABD2A3E64FFB78">
<mods:roleTerm id="7135BA70CAEB630965FB36991849D3AE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0CFDE4B34FCE66ED05C2222737C66A3F">Tanikawa, Akio</mods:namePart>
<mods:affiliation id="7EF0B5E80438D7D2161ABA3E990D3413">Laboratory of Biodiversity Science, School of Agriculture and Life Science, The University of Tokyo, 1 - 1 - 1, Yayoi, Bunkyo-ku, Tokyo 113 - 8657 Japan.</mods:affiliation>
</mods:name>
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<mods:title id="48D725BD788AF19B07290861B2F72F6C">Zootaxa</mods:title>
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<mods:part id="79A9FBD1D2E658B4537D967554184C75">
<mods:date id="05575FB903DE66DA55A7C9D462283F06">2021</mods:date>
<mods:detail id="BC09695C801E31D64B52F97C46A9245E" type="pubDate">
<mods:number id="C34D4321E0AE896FA6D3F3CE7EA912DE">2021-10-18</mods:number>
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<mods:detail id="21E76F258B95ED0681219A013A24C00A" type="volume">
<mods:number id="B56B0838956A7A373F872C5DC56F90AE">5052</mods:number>
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<mods:number id="ABF48C9E4B3B48DF7F8D7E0E58D0BEF4">4</mods:number>
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<mods:classification id="CE9E352CFC89D6423219993180638A35">journal article</mods:classification>
<mods:identifier id="A7528661CDB32971DF630D610C162367" type="CLB-Dataset">4006</mods:identifier>
<mods:identifier id="840DA80F49194435D2812801C01007F8" type="DOI">10.11646/zootaxa.5052.4.9</mods:identifier>
<mods:identifier id="E1284CC4D7B5EAD6AC870B6050023DFE" type="GBIF-Dataset">146052d6-ccc2-48f3-b059-3757ce97ba4e</mods:identifier>
<mods:identifier id="7951551417826221E7B04C493A9BFB32" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D342CE4278EFFAFD872FD266297DEECF" type="Zenodo-Dep">5577387</mods:identifier>
<mods:identifier id="03E4BF6C1D863482D6D4AFA15115F27E" type="ZooBank">B97CDD9C-9ADC-47B1-8020-CF710B4282A1</mods:identifier>
</mods:mods>
<treatment id="03B887DCFFE1315A70892721FD3EFC1F" ID-DOI="http://doi.org/10.5281/zenodo.5577393" ID-GBIF-Taxon="188994763" ID-Zenodo-Dep="5577393" LSID="urn:lsid:zoobank.org:act:5F005F73-B712-49AC-ACE0-3B1C17A03077" httpUri="http://treatment.plazi.org/id/03B887DCFFE1315A70892721FD3EFC1F" lastPageId="4" lastPageNumber="595" pageId="1" pageNumber="592">
<subSubSection id="C30B6541FFE1315F70892721FDE5FA99" box="[151,554,1363,1390]" pageId="1" pageNumber="592" type="nomenclature">
<paragraph id="8BAE36CAFFE1315F70892721FDE5FA99" blockId="1.[151,1437,1292,1965]" box="[151,554,1363,1390]" pageId="1" pageNumber="592">
<heading id="D0E681A6FFE1315F70892721FDE5FA99" bold="true" box="[151,554,1363,1390]" fontSize="11" level="1" pageId="1" pageNumber="592" reason="1">
<emphasis id="B965EAD8FFE1315F70892721FDE5FA99" bold="true" box="[151,554,1363,1390]" pageId="1" pageNumber="592">
<taxonomicName id="4C114D49FFE1315F70892721FE5CFA99" ID-CoL="93QPF" authority="Suzuki &amp; Tanikawa, 2021" authorityName="Suzuki &amp; Tanikawa" authorityYear="2021" box="[151,403,1363,1389]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="matsumotoi" status="sp. nov.">
<emphasis id="B965EAD8FFE1315F70892721FE5CFA99" bold="true" box="[151,403,1363,1389]" italics="true" pageId="1" pageNumber="592">Araneus matsumotoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657A3FFE1315F71842726FDE5FA99" box="[410,554,1363,1390]" pageId="1" pageNumber="592" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30B6541FFE1315F7089270BFEC7FA65" box="[151,264,1401,1425]" pageId="1" pageNumber="592" type="description">
<paragraph id="8BAE36CAFFE1315F7089270BFEC7FA65" blockId="1.[151,1437,1292,1965]" box="[151,264,1401,1425]" pageId="1" pageNumber="592">
<figureCitation id="132A2A4FFFE1315F7089270BFEC7FA65" box="[151,264,1401,1425]" captionStart-0="FIGURES 28" captionStart-1="FIGURES 915" captionStartId-0="2.[151,264,1598,1622]" captionStartId-1="3.[151,264,1751,1775]" captionTargetBox-0="[230,1374,193,1574]" captionTargetBox-1="[189,1402,650,1712]" captionTargetId-0="figure-22@2.[213,1374,181,1574]" captionTargetId-1="figure-243@3.[177,1410,632,1728]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 28. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 2, 4, 5) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 3, 68); habitus (2, 3), male palp (4, 5) and female genitalia (68). 2, Habitus of male, dorsal view. 3, Habitus of female, dorsal view. 4, Male left palp, prolateral view. 5, Same, anterior view. 6, Epigynum, ventral view. 7, Same, lateral view; 8, basal part of epigynum, ventral view. Arrows in Figs. 4 and 5 indicate a sharp appendix on the posterior margin of subterminal apophysis. Scale bars, 47, 1.0 mm; 8, 0.5 mm." captionText-1="FIGURES 915. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 9, 1112) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 10, 1315). 9, Apophysis on prolateral side of tibia II. 10, Projection on ventral side of coxa II. 11, Male left palp, prolateral view. 12, Same, apical part, anterior view.13, Epigynum, ventral view.14, Same, posterior view.15, Internal genitalia, dorsal view. Arrows in Figs. 1112 and 13, 14 indicate a sharp appendix on the posterior margin of subterminal apophysis (arrows 1, 3), bent tip of conductor (arrows 2, 4) and copulatory opening (arrows 5, 6), respectively. Abbreviations: c, conductor; fd, fertilization duct; lp, lateral plate; ma, median apophysis; mp, median plate; pc, paracymbium; s, spermatheca; st, subterminal apophysis; ta, terminal apophysis. Scale bars, 811, 1.0 mm; 1214, 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5577391" figureDoi-1="http://doi.org/10.5281/zenodo.5577397" httpUri-0="https://zenodo.org/record/5577391/files/figure.png" httpUri-1="https://zenodo.org/record/5577397/files/figure.png" pageId="1" pageNumber="592">Figs. 215</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30B6541FFE1315F708927EFFD05F859" pageId="1" pageNumber="592" type="materials_examined">
<paragraph id="8BAE36CAFFE1315F708927EFFB5DF90D" blockId="1.[151,1437,1292,1965]" pageId="1" pageNumber="592">
<emphasis id="B965EAD8FFE1315F708927EFFED4FA41" bold="true" box="[151,283,1437,1461]" inLineHeading="true" pageId="1" pageNumber="592" reason="1">Type series.</emphasis>
<materialsCitation id="3B793C97FFE1315F713F27EFFC8DFA2D" ID-GBIF-Occurrence="3386857304" collectingDate="2019-05-29" collectingDateMax="2019-09-25" collectingDateMin="2019-05-29" collectionCode="R" collectorName="Matsumoto" location="Nara-shi" municipality="Takahata-cho" pageId="1" pageNumber="592" specimenCount="1" stateProvince="Nara" typeStatus="holotype">
<typeStatus id="54AA8868FFE1315F713F27EFFE47FA41" box="[289,392,1437,1461]" pageId="1" pageNumber="592" type="holotype">Holotype</typeStatus>
: male (NSMT-Ar 20952),
<collectingMunicipality id="6BCAACB0FFE1315F728327EFFCE0FA41" box="[669,815,1437,1461]" pageId="1" pageNumber="592">Takahata-cho</collectingMunicipality>
,
<location id="8ECE6011FFE1315F732727EFFC57FA41" LSID="urn:lsid:plazi:treatment:03B887DCFFE1315A70892721FD3EFC1F:8ECE6011FFE1315F732727EFFC57FA41" box="[825,920,1437,1461]" municipality="Takahata-cho" name="Nara-shi" pageId="1" pageNumber="592" stateProvince="Nara">Nara-shi</location>
,
<collectingRegion id="49D5F828FFE1315F73BF27ECFBDEFA41" box="[929,1041,1437,1461]" country="Japan" name="Nara" pageId="1" pageNumber="592">Nara Pref.</collectingRegion>
, collected as a juvenile on
<date id="FFAF100AFFE1315F753427EFFA53FA41" box="[1322,1436,1437,1461]" pageId="1" pageNumber="592" value="2019-05-29">
<collectingDate id="EFEBE9E2FFE1315F753427EFFA53FA41" box="[1322,1436,1437,1461]" pageId="1" pageNumber="592" value="2019-05-29">29-V-2019</collectingDate>
</date>
and matured on
<date id="FFAF100AFFE1315F715227B3FE05FA2D" box="[332,458,1473,1497]" pageId="1" pageNumber="592" value="2019-09-25">
<collectingDate id="EFEBE9E2FFE1315F715227B3FE05FA2D" box="[332,458,1473,1497]" pageId="1" pageNumber="592" value="2019-09-25">25-IX-2019</collectingDate>
</date>
under rearing,
<collectionCode id="ED00AE0FFFE1315F726B27B0FD45FA2D" box="[629,650,1474,1497]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="592">R</collectionCode>
.
<collectorName id="26E4531CFFE1315F728927B0FCDFFA2D" box="[663,784,1474,1497]" pageId="1" pageNumber="592">Matsumoto</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B793C97FFE1315F735427B0FD29FA09" ID-GBIF-Occurrence="3386857301" collectingDate="1990-08-18" collectorName="S. Tazoe" location="Senboku-shi" municipality="Lake Tazawa" pageId="1" pageNumber="592" specimenCount="1" specimenCount-female="1" stateProvince="Akita" typeStatus="paratype">
<typeStatus id="54AA8868FFE1315F735427B0FC79FA2E" box="[842,950,1474,1498]" pageId="1" pageNumber="592" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D17FD43FFE1315F73DD27B3FBECFA2D" box="[963,1059,1473,1497]" pageId="1" pageNumber="592" type="female">1 female</specimenCount>
(NSMT-Ar 20953),
<collectingMunicipality id="6BCAACB0FFE1315F751827B3FA57FA2D" box="[1286,1432,1473,1497]" pageId="1" pageNumber="592">Lake Tazawa</collectingMunicipality>
,
<location id="8ECE6011FFE1315F70892797FED0FA09" LSID="urn:lsid:plazi:treatment:03B887DCFFE1315A70892721FD3EFC1F:8ECE6011FFE1315F70892797FED0FA09" box="[151,287,1509,1533]" municipality="Lake Tazawa" name="Senboku-shi" pageId="1" pageNumber="592" stateProvince="Akita">Senboku-shi</location>
,
<collectingRegion id="49D5F828FFE1315F71352797FE68FA09" box="[299,423,1509,1533]" country="Japan" name="Akita" pageId="1" pageNumber="592">Akita Pref.</collectingRegion>
,
<date id="FFAF100AFFE1315F71AA2797FD89FA09" box="[436,582,1509,1533]" pageId="1" pageNumber="592" value="1990-08-18">
<collectingDate id="EFEBE9E2FFE1315F71AA2797FD89FA09" box="[436,582,1509,1533]" pageId="1" pageNumber="592" value="1990-08-18">18-VIII-1990</collectingDate>
</date>
,
<collectorName id="26E4531CFFE1315F724A2797FD7EFA09" box="[596,689,1509,1533]" pageId="1" pageNumber="592">S. Tazoe</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B793C97FFE1315F72EA2797FB9CF979" ID-GBIF-Occurrence="3386857305" collectingDate="2019-05-29" collectingDateMax="2020-07-15" collectingDateMin="2019-05-29" collectionCode="R" collectorName="Matsumoto" location="Nara-shi" municipality="Takahata-cho" pageId="1" pageNumber="592" specimenCount="10" specimenCount-female="4" specimenCount-male="6" stateProvince="Nara" typeStatus="paratype">
<specimenCount id="9D17FD43FFE1315F72EA2797FC91FA09" box="[756,862,1509,1533]" pageId="1" pageNumber="592" type="female">2 females</specimenCount>
and
<specimenCount id="9D17FD43FFE1315F73862797FC3FFA09" box="[920,1008,1509,1533]" pageId="1" pageNumber="592" type="male">3 males</specimenCount>
, same locality as the holotype [
<specimenCount id="9D17FD43FFE1315F754C2797FA53FA09" box="[1362,1436,1509,1533]" pageId="1" pageNumber="592" type="male">1 male</specimenCount>
(NSMT-Ar 20954), collected as a subadult on
<date id="FFAF100AFFE1315F7292247BFD3DF9D5" box="[652,754,1545,1569]" pageId="1" pageNumber="592" value="2020-05-02">
<collectingDate id="EFEBE9E2FFE1315F7292247BFD3DF9D5" box="[652,754,1545,1569]" pageId="1" pageNumber="592" value="2020-05-02">2-V-2020</collectingDate>
</date>
and matured on
<date id="FFAF100AFFE1315F73B6247BFBD4F9D5" box="[936,1051,1545,1569]" pageId="1" pageNumber="592" value="2020-05-25">
<collectingDate id="EFEBE9E2FFE1315F73B6247BFBD4F9D5" box="[936,1051,1545,1569]" pageId="1" pageNumber="592" value="2020-05-25">25-V-2020</collectingDate>
</date>
under rearing;
<specimenCount id="9D17FD43FFE1315F74DF247BFAD0F9D5" box="[1217,1311,1545,1569]" pageId="1" pageNumber="592" type="female">1 female</specimenCount>
(NSMT-Ar 20955), collected as a juvenile on
<date id="FFAF100AFFE1315F71E2245FFDB5F9B1" box="[508,634,1581,1605]" pageId="1" pageNumber="592" value="2020-06-20">
<collectingDate id="EFEBE9E2FFE1315F71E2245FFDB5F9B1" box="[508,634,1581,1605]" pageId="1" pageNumber="592" value="2020-06-20">20-VI-2020</collectingDate>
</date>
and matured on
<date id="FFAF100AFFE1315F7336245FFC69F9B1" box="[808,934,1581,1605]" pageId="1" pageNumber="592" value="2020-06-24">
<collectingDate id="EFEBE9E2FFE1315F7336245FFC69F9B1" box="[808,934,1581,1605]" pageId="1" pageNumber="592" value="2020-06-24">24-VI-2020</collectingDate>
</date>
under rearing, scape of epigynum incompletely formed;
<specimenCount id="9D17FD43FFE1315F70E82423FE83F99D" box="[246,332,1617,1641]" pageId="1" pageNumber="592" type="male">2 males</specimenCount>
(NSMT-Ar 20956-20957), collected as juveniles on
<date id="FFAF100AFFE1315F73872423FBC4F99D" box="[921,1035,1617,1641]" pageId="1" pageNumber="592" value="2019-05-29">
<collectingDate id="EFEBE9E2FFE1315F73872423FBC4F99D" box="[921,1035,1617,1641]" pageId="1" pageNumber="592" value="2019-05-29">29-V-2019</collectingDate>
</date>
and matured on
<collectingDate id="EFEBE9E2FFE1315F74D32423FF3EF979" isEnumeration="true" pageId="1" pageNumber="592">
<date id="FFAF100AFFE1315F74D32423FAF2F99D" box="[1229,1341,1617,1641]" pageId="1" pageNumber="592" value="2020-06-03">3-VI-2020</date>
and
<date id="FFAF100AFFE1315F75672423FF3EF979" pageId="1" pageNumber="592" value="2020-06-17">17- VI-2020</date>
</collectingDate>
under rearing;
<specimenCount id="9D17FD43FFE1315F71812407FE30F979" box="[415,511,1653,1677]" pageId="1" pageNumber="592" type="female">1 female</specimenCount>
(NSMT-Ar 20958),
<date id="FFAF100AFFE1315F72FD2407FCBFF979" box="[739,880,1653,1677]" pageId="1" pageNumber="592" value="2020-07-15">
<collectingDate id="EFEBE9E2FFE1315F72FD2407FCBFF979" box="[739,880,1653,1677]" pageId="1" pageNumber="592" value="2020-07-15">15-VII-2020</collectingDate>
</date>
],
<collectionCode id="ED00AE0FFFE1315F739D2404FC57F979" box="[899,920,1654,1677]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="592">R</collectionCode>
.
<collectorName id="26E4531CFFE1315F73B82404FBD0F979" box="[934,1055,1654,1677]" pageId="1" pageNumber="592">Matsumoto</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B793C97FFE1315F747F2407FECDF921" ID-GBIF-Occurrence="3386857307" collectingDate="2019-07-14" collectingDateMax="2020-06-17" collectingDateMin="2019-07-14" collectorName="Y. Obae" location="Tottori-shi" municipality="Amedaki" pageId="1" pageNumber="592" specimenCount="1" specimenCount-female="1" stateProvince="Tottori" typeStatus="paratype">
<specimenCount id="9D17FD43FFE1315F747F2407FB0EF979" box="[1121,1217,1653,1677]" pageId="1" pageNumber="592" type="female">1 female</specimenCount>
(NSMT-Ar 20959),
<collectingMunicipality id="6BCAACB0FFE1315F708924EBFF33F945" box="[151,252,1689,1713]" pageId="1" pageNumber="592">Amedaki</collectingMunicipality>
,
<location id="8ECE6011FFE1315F711824EBFEB6F945" LSID="urn:lsid:plazi:treatment:03B887DCFFE1315A70892721FD3EFC1F:8ECE6011FFE1315F711824EBFEB6F945" box="[262,377,1689,1713]" municipality="Amedaki" name="Tottori-shi" pageId="1" pageNumber="592" stateProvince="Tottori">Tottori-shi</location>
,
<collectingRegion id="49D5F828FFE1315F719D24EBFDC7F945" box="[387,520,1689,1713]" country="Japan" name="Tottori" pageId="1" pageNumber="592">Tottori Pref.</collectingRegion>
, collected as a juvenile on
<date id="FFAF100AFFE1315F733824EBFC60F945" box="[806,943,1689,1713]" pageId="1" pageNumber="592" value="2019-07-14">
<collectingDate id="EFEBE9E2FFE1315F733824EBFC60F945" box="[806,943,1689,1713]" pageId="1" pageNumber="592" value="2019-07-14">14-VII-2019</collectingDate>
</date>
, and matured on
<date id="FFAF100AFFE1315F747824EBFB2BF945" box="[1126,1252,1689,1713]" pageId="1" pageNumber="592" value="2020-06-17">
<collectingDate id="EFEBE9E2FFE1315F747824EBFB2BF945" box="[1126,1252,1689,1713]" pageId="1" pageNumber="592" value="2020-06-17">17-VI-2020</collectingDate>
</date>
under rearing,
<collectorName id="26E4531CFFE1315F759824E8FF00F921" pageId="1" pageNumber="592">Y. Obae</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B793C97FFE1315F711024CFFB5DF90D" ID-GBIF-Occurrence="3386857306" collectingDate="2018-08-25" collectingDateMax="2019-01-31" collectingDateMin="2018-08-25" collectorName="K. Terada" county="Kitazato" location="Tsushima-shi" municipality="Izuhara-machi" pageId="1" pageNumber="592" specimenCount="1" specimenCount-male="1" stateProvince="Nagasaki" typeStatus="paratype">
<specimenCount id="9D17FD43FFE1315F711024CFFE98F921" box="[270,343,1725,1749]" pageId="1" pageNumber="592" type="male">1 male</specimenCount>
(NSMT-Ar 20960; COI haplotype: AMT01),
<collectingCounty id="62CF4E46FFE1315F735424CFFC68F921" box="[842,935,1725,1749]" pageId="1" pageNumber="592">Kitazato</collectingCounty>
,
<collectingMunicipality id="6BCAACB0FFE1315F73AD24CFFB9FF921" box="[947,1104,1725,1749]" pageId="1" pageNumber="592">Izuhara-machi</collectingMunicipality>
,
<location id="8ECE6011FFE1315F744524CFFB23F921" LSID="urn:lsid:plazi:treatment:03B887DCFFE1315A70892721FD3EFC1F:8ECE6011FFE1315F744524CFFB23F921" box="[1115,1260,1725,1749]" county="Kitazato" municipality="Izuhara-machi" name="Tsushima-shi" pageId="1" pageNumber="592" stateProvince="Nagasaki">Tsushima-shi</location>
,
<collectingRegion id="49D5F828FFE1315F74E924CFFA56F921" box="[1271,1433,1725,1749]" country="Japan" name="Nagasaki" pageId="1" pageNumber="592">Nagasaki Pref.</collectingRegion>
, collected as a juvenile on
<date id="FFAF100AFFE1315F71B52493FDF4F90D" box="[427,571,1761,1785]" pageId="1" pageNumber="592" value="2018-08-25">
<collectingDate id="EFEBE9E2FFE1315F71B52493FDF4F90D" box="[427,571,1761,1785]" pageId="1" pageNumber="592" value="2018-08-25">25-VIII-2018</collectingDate>
</date>
and matured on
<date id="FFAF100AFFE1315F72F02493FC96F90D" box="[750,857,1761,1785]" pageId="1" pageNumber="592" value="2019-01-31">
<collectingDate id="EFEBE9E2FFE1315F72F02493FC96F90D" box="[750,857,1761,1785]" pageId="1" pageNumber="592" value="2019-01-31">31-I-2019</collectingDate>
</date>
under rearing,
<collectorName id="26E4531CFFE1315F73E22490FBABF90D" box="[1020,1124,1761,1785]" pageId="1" pageNumber="592">K. Terada</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BAE36CAFFE1315F70D92577FD05F859" blockId="1.[151,1437,1292,1965]" pageId="1" pageNumber="592">
<emphasis id="B965EAD8FFE1315F70D92577FDFCF8E9" bold="true" box="[199,563,1797,1821]" inLineHeading="true" pageId="1" pageNumber="592" reason="1">Additional specimens examined.</emphasis>
<materialsCitation id="3B793C97FFE1315F72292577FC41F8B5" ID-GBIF-Occurrence="3386857302" collectingDate="2021-05-03" collectingDateMax="2021-05-23" collectingDateMin="2021-05-03" collectionCode="R" collectorName="Serita" county="Sagayama" location="Nagaoka-gun" municipality="Otoyo-cho" pageId="1" pageNumber="592" specimenCount="1" specimenCount-female="1" stateProvince="Kochi">
<specimenCount id="9D17FD43FFE1315F72292577FD5AF8E9" box="[567,661,1797,1821]" pageId="1" pageNumber="592" type="female">1 female</specimenCount>
,
<collectingCounty id="62CF4E46FFE1315F72802577FCC1F8E9" box="[670,782,1797,1821]" pageId="1" pageNumber="592">Sagayama</collectingCounty>
,
<collectingMunicipality id="6BCAACB0FFE1315F73092574FC43F8D4" box="[791,908,1798,1824]" pageId="1" pageNumber="592">Ôtoyo-cho</collectingMunicipality>
,
<location id="8ECE6011FFE1315F738B2577FBE8F8E9" LSID="urn:lsid:plazi:treatment:03B887DCFFE1315A70892721FD3EFC1F:8ECE6011FFE1315F738B2577FBE8F8E9" box="[917,1063,1797,1821]" county="Sagayama" municipality="Otoyo-cho" name="Nagaoka-gun" pageId="1" pageNumber="592" stateProvince="Kochi">Nagaoka-gun</location>
,
<collectingRegion id="49D5F828FFE1315F742E2577FB62F8E9" box="[1072,1197,1797,1821]" country="Japan" name="Kochi" pageId="1" pageNumber="592">Kochi Pref.</collectingRegion>
, collected as a juvenile on
<date id="FFAF100AFFE1315F70A5255BFEEFF8B5" box="[187,288,1833,1857]" pageId="1" pageNumber="592" value="2021-05-03">
<collectingDate id="EFEBE9E2FFE1315F70A5255BFEEFF8B5" box="[187,288,1833,1857]" pageId="1" pageNumber="592" value="2021-05-03">3-V-2021</collectingDate>
</date>
and matured on
<date id="FFAF100AFFE1315F71C1255BFD9DF8B5" box="[479,594,1833,1857]" pageId="1" pageNumber="592" value="2021-05-23">
<collectingDate id="EFEBE9E2FFE1315F71C1255BFD9DF8B5" box="[479,594,1833,1857]" pageId="1" pageNumber="592" value="2021-05-23">23-V-2021</collectingDate>
</date>
under rearing,
<collectionCode id="ED00AE0FFFE1315F72E22558FCDEF8B5" box="[764,785,1834,1857]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="592">R</collectionCode>
.
<collectorName id="26E4531CFFE1315F7300255BFC93F8B5" box="[798,860,1833,1857]" pageId="1" pageNumber="592">Serita</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B793C97FFE1315F7388255BFE25F87D" ID-GBIF-Occurrence="3386857303" collectingDate="2020-07-19" collectingDateMax="2021-05-04" collectingDateMin="2020-07-19" collectorName="K. Nishiura" location="Sekimoto-cho" municipality="Ogawa" pageId="1" pageNumber="592" specimenCount="1" specimenCount-male="1" stateProvince="Ibaraki">
<specimenCount id="9D17FD43FFE1315F7388255BFC2EF8B5" box="[918,993,1833,1857]" pageId="1" pageNumber="592" type="male">1 male</specimenCount>
(both palps were incompletely formed),
<collectingMunicipality id="6BCAACB0FFE1315F7089253FFF2AF891" box="[151,229,1869,1893]" pageId="1" pageNumber="592">Ogawa</collectingMunicipality>
,
<location id="8ECE6011FFE1315F70EC253FFE45F891" LSID="urn:lsid:plazi:treatment:03B887DCFFE1315A70892721FD3EFC1F:8ECE6011FFE1315F70EC253FFE45F891" box="[242,394,1869,1893]" municipality="Ogawa" name="Sekimoto-cho" pageId="1" pageNumber="592" stateProvince="Ibaraki">Sekimoto-cho</location>
, Kita-ibaraki-shi,
<collectingRegion id="49D5F828FFE1315F724E253FFD14F891" box="[592,731,1869,1893]" country="Japan" name="Ibaraki" pageId="1" pageNumber="592">Ibaraki Pref.</collectingRegion>
, collected as juvenile on
<date id="FFAF100AFFE1315F73ED253FFBB5F891" box="[1011,1146,1869,1893]" pageId="1" pageNumber="592" value="2020-07-19">
<collectingDate id="EFEBE9E2FFE1315F73ED253FFBB5F891" box="[1011,1146,1869,1893]" pageId="1" pageNumber="592" value="2020-07-19">19-VII-2020</collectingDate>
</date>
and matured on
<date id="FFAF100AFFE1315F7529253FFA53F891" box="[1335,1436,1869,1893]" pageId="1" pageNumber="592" value="2021-05-04">
<collectingDate id="EFEBE9E2FFE1315F7529253FFA53F891" box="[1335,1436,1869,1893]" pageId="1" pageNumber="592" value="2021-05-04">4-V-2021</collectingDate>
</date>
under rearing,
<collectorName id="26E4531CFFE1315F71272500FE76F87D" box="[313,441,1905,1929]" pageId="1" pageNumber="592">K. Nishiura</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B793C97FFE1315F71ED2503FD05F859" ID-GBIF-Occurrence="3386857308" collectingDate="2020-11-04" collectingDateMax="2021-04-16" collectingDateMin="2020-11-04" collectorName="Y. Suzuki" location="Tottori-shi" municipality="Amedaki" pageId="1" pageNumber="592" specimenCount="1" specimenCount-male="1" stateProvince="Tottori">
<specimenCount id="9D17FD43FFE1315F71ED2503FD8FF87D" box="[499,576,1905,1929]" pageId="1" pageNumber="592" type="male">1 male</specimenCount>
,
<collectingMunicipality id="6BCAACB0FFE1315F72522503FD7EF87D" box="[588,689,1905,1929]" pageId="1" pageNumber="592">Amedaki</collectingMunicipality>
,
<location id="8ECE6011FFE1315F72A02503FCFEF87D" LSID="urn:lsid:plazi:treatment:03B887DCFFE1315A70892721FD3EFC1F:8ECE6011FFE1315F72A02503FCFEF87D" box="[702,817,1905,1929]" municipality="Amedaki" name="Tottori-shi" pageId="1" pageNumber="592" stateProvince="Tottori">Tottori-shi</location>
,
<collectingRegion id="49D5F828FFE1315F73232503FC09F87D" box="[829,966,1905,1929]" country="Japan" name="Tottori" pageId="1" pageNumber="592">Tottori Pref.</collectingRegion>
, collected as a juvenile on
<date id="FFAF100AFFE1315F74EB2503FAA6F87D" box="[1269,1385,1905,1929]" pageId="1" pageNumber="592" value="2020-11-04">
<collectingDate id="EFEBE9E2FFE1315F74EB2503FAA6F87D" box="[1269,1385,1905,1929]" pageId="1" pageNumber="592" value="2020-11-04">4-XI-2020</collectingDate>
</date>
, and matured on
<date id="FFAF100AFFE1315F710925E7FE5DF859" box="[279,402,1941,1965]" pageId="1" pageNumber="592" value="2021-04-16">
<collectingDate id="EFEBE9E2FFE1315F710925E7FE5DF859" box="[279,402,1941,1965]" pageId="1" pageNumber="592" value="2021-04-16">16-IV-2021</collectingDate>
</date>
under rearing,
<collectorName id="26E4531CFFE1315F722A25E4FD54F859" box="[564,667,1941,1965]" pageId="1" pageNumber="592">Y. Suzuki</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF6E6642FFE2315C7089244CFCDDF912" ID-DOI="http://doi.org/10.5281/zenodo.5577391" ID-Zenodo-Dep="5577391" httpUri="https://zenodo.org/record/5577391/files/figure.png" pageId="2" pageNumber="593" startId="2.[151,264,1598,1622]" targetBox="[230,1374,193,1574]" targetPageId="2">
<paragraph id="8BAE36CAFFE2315C7089244CFCDDF912" blockId="2.[151,1437,1598,1766]" pageId="2" pageNumber="593">
<emphasis id="B965EAD8FFE2315C7089244CFEF9F9A2" bold="true" box="[151,310,1598,1622]" pageId="2" pageNumber="593">FIGURES 28</emphasis>
.
<taxonomicName id="4C114D49FFE2315C715A244DFDD9F9A2" authority="Suzuki &amp; Tanikawa, 2021" authorityName="Suzuki &amp; Tanikawa" authorityYear="2021" box="[324,534,1599,1622]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="matsumotoi" status="sp. nov.">
<emphasis id="B965EAD8FFE2315C715A244DFDD9F9A2" box="[324,534,1599,1622]" italics="true" pageId="2" pageNumber="593">Araneus matsumotoi</emphasis>
</taxonomicName>
<emphasis id="B965EAD8FFE2315C7200244CFD96F9A2" bold="true" box="[542,601,1598,1622]" pageId="2" pageNumber="593">
<taxonomicNameLabel id="A25657A3FFE2315C7200244CFD96F9A2" box="[542,601,1598,1622]" pageId="2" pageNumber="593" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male holotype (NSMT-Ar 20952; 2, 4, 5) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 3, 68); habitus (2, 3), male palp (4, 5) and female genitalia (68). 2, Habitus of male, dorsal view. 3, Habitus of female, dorsal view. 4, Male left palp, prolateral view. 5, Same, anterior view. 6, Epigynum, ventral view. 7, Same, lateral view; 8, basal part of epigynum, ventral view. Arrows in Figs. 4 and 5 indicate a sharp appendix on the posterior margin of subterminal apophysis. Scale bars, 47, 1.0 mm; 8, 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C30B6541FFE2315C70D92564FEACF8A6" pageId="2" pageNumber="593" type="etymology">
<paragraph id="8BAE36CAFFE2315C70D92564FEACF8A6" blockId="2.[151,1437,1814,2019]" pageId="2" pageNumber="593">
<emphasis id="B965EAD8FFE2315C70D92564FE87F8DA" bold="true" box="[199,328,1814,1838]" inLineHeading="true" pageId="2" pageNumber="593" reason="1">Etymology.</emphasis>
The specific name is dedicated to Rikio Matsumoto, who provided us with many invaluable specimens of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C30B6541FFE2315D70D9252CFC22FDE3" lastPageId="3" lastPageNumber="594" pageId="2" pageNumber="593" type="diagnosis">
<paragraph id="8BAE36CAFFE2315D70D9252CFC22FDE3" blockId="2.[151,1437,1814,2019]" lastBlockId="3.[151,1437,151,607]" lastPageId="3" lastPageNumber="594" pageId="2" pageNumber="593">
<emphasis id="B965EAD8FFE2315C70D9252CFEF3F882" bold="true" box="[199,316,1886,1910]" inLineHeading="true" pageId="2" pageNumber="593" reason="1">Diagnosis.</emphasis>
The new species resembles
<taxonomicName id="4C114D49FFE2315C7273252DFC6DF882" authority="(L. Koch 1878)" baseAuthorityName="L. Koch" baseAuthorityYear="1878" box="[621,930,1886,1910]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="seminiger">
<emphasis id="B965EAD8FFE2315C7273252DFD3BF882" box="[621,756,1887,1910]" italics="true" pageId="2" pageNumber="593">A. seminiger</emphasis>
(L.
<bibRefCitation id="EF804B3BFFE2315C733C252CFC55F882" author="Koch, L." box="[802,922,1886,1910]" pageId="2" pageNumber="593" pagination="735 - 798" refId="ref3219" refString="Koch, L. (1878) Japanesische Arachniden und Myriapoden. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 27, 735 - 798." type="journal article" year="1878">Koch 1878</bibRefCitation>
)
</taxonomicName>
in appearance (also see
<bibRefCitation id="EF804B3BFFE2315C74B6252CFADDF882" author="Ono, H." box="[1192,1298,1886,1910]" pageId="2" pageNumber="593" pagination="51 - 60" refId="ref3315" refString="Ono, H. (2002) New and remarkable spiders of the families Liphistiidae, Argyronetidae, Pisauridae, Theridiidae and Araneidae (Arachnida) from Japan. Bulletin of the National Museum of Nature and Science, Series A, 28, 51 - 60." type="journal article" year="2002">Ono 2002</bibRefCitation>
&amp;
<bibRefCitation id="EF804B3BFFE2315C752B252CFF1FF86E" author="Tanikawa, A." pageId="2" pageNumber="593" pagination="63 - 86" refId="ref3446" refString="Tanikawa, A. (2001) Twelve new species and one newly recorded species of the spider genus Araneus (Araneae: Araneidae) from Japan. Acta Arachnologica, 50, 63 - 86. https: // doi. org / 10.2476 / asjaa. 50.63" type="journal article" year="2001">Tanikawa 2001</bibRefCitation>
), but can clearly be distinguished by the combination of the following body color and marking characters: the carapace of the new species is uniformly covered with gray hairs when living, whereas it is reddish brown in
<taxonomicName id="4C114D49FFE2315C74C425D5FAADF84A" baseAuthorityName="L. Koch" baseAuthorityYear="1878" box="[1242,1378,1959,1982]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="seminiger">
<emphasis id="B965EAD8FFE2315C74C425D5FAADF84A" box="[1242,1378,1959,1982]" italics="true" pageId="2" pageNumber="593">A. seminiger</emphasis>
</taxonomicName>
; legs and abdomen of the fresh specimens are dark green, similar to green moss (Bryophyta), whereas those of
<taxonomicName id="4C114D49FFE2315C74EE25B9FAB9F816" baseAuthorityName="L. Koch" baseAuthorityYear="1878" box="[1264,1398,1995,2018]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="seminiger">
<emphasis id="B965EAD8FFE2315C74EE25B9FAB9F816" box="[1264,1398,1995,2018]" italics="true" pageId="2" pageNumber="593">A. seminiger</emphasis>
</taxonomicName>
are whitish green, resembling
<taxonomicName id="4C114D49FFE3315D71AC22EAFDF7FF5B" authorityName="A.Massalongo" authorityYear="1860" box="[434,568,152,175]" class="Lecanoromycetes" family="Parmeliaceae" genus="Parmotrema" kingdom="Fungi" order="Lecanorales" pageId="3" pageNumber="594" phylum="Ascomycota" rank="genus">
<emphasis id="B965EAD8FFE3315D71AC22EAFDF7FF5B" box="[434,568,152,175]" italics="true" pageId="3" pageNumber="594">Parmotrema</emphasis>
</taxonomicName>
lichen (
<taxonomicName id="4C114D49FFE3315D729322E5FCEDFF5B" box="[653,802,151,175]" class="Lecanoromycetes" family="Parmeliaceae" kingdom="Fungi" order="Lecanorales" pageId="3" pageNumber="594" phylum="Ascomycota" rank="family">Parmeliaceae</taxonomicName>
); a light green chevron marking edged with black is present between the shoulder humps, whereas it is absent in
<taxonomicName id="4C114D49FFE3315D72D622CEFC80FF27" baseAuthorityName="L. Koch" baseAuthorityYear="1878" box="[712,847,188,211]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="seminiger">
<emphasis id="B965EAD8FFE3315D72D622CEFC80FF27" box="[712,847,188,211]" italics="true" pageId="3" pageNumber="594">A. seminiger</emphasis>
</taxonomicName>
(
<figureCitation id="132A2A4FFFE3315D734022C9FC0BFF20" box="[862,964,187,212]" captionStart="FIGURES 28" captionStartId="2.[151,264,1598,1622]" captionTargetBox="[230,1374,193,1574]" captionTargetId="figure-22@2.[213,1374,181,1574]" captionTargetPageId="2" captionText="FIGURES 28. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 2, 4, 5) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 3, 68); habitus (2, 3), male palp (4, 5) and female genitalia (68). 2, Habitus of male, dorsal view. 3, Habitus of female, dorsal view. 4, Male left palp, prolateral view. 5, Same, anterior view. 6, Epigynum, ventral view. 7, Same, lateral view; 8, basal part of epigynum, ventral view. Arrows in Figs. 4 and 5 indicate a sharp appendix on the posterior margin of subterminal apophysis. Scale bars, 47, 1.0 mm; 8, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5577391" httpUri="https://zenodo.org/record/5577391/files/figure.png" pageId="3" pageNumber="594">Figs. 2, 3</figureCitation>
). The new species can also be differentiated from
<taxonomicName id="4C114D49FFE3315D70CB2292FE90FF03" baseAuthorityName="L. Koch" baseAuthorityYear="1878" box="[213,351,224,247]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="seminiger">
<emphasis id="B965EAD8FFE3315D70CB2292FE90FF03" box="[213,351,224,247]" italics="true" pageId="3" pageNumber="594">A. seminiger</emphasis>
</taxonomicName>
based on the following characters: a wide unsclerotized area on the tip of subterminal apophysis (unsclerotized area is very narrow in
<taxonomicName id="4C114D49FFE3315D72262376FD0CFEEF" baseAuthorityName="L. Koch" baseAuthorityYear="1878" box="[568,707,260,283]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="seminiger">
<emphasis id="B965EAD8FFE3315D72262376FD0CFEEF" box="[568,707,260,283]" italics="true" pageId="3" pageNumber="594">A. seminiger</emphasis>
</taxonomicName>
); a thin sclerotized appendix on the posterior side of subterminal apophysis (
<figureCitation id="132A2A4FFFE3315D710A2355FE4FFECB" box="[276,384,295,320]" captionStart="FIGURES 28" captionStartId="2.[151,264,1598,1622]" captionTargetBox="[230,1374,193,1574]" captionTargetId="figure-22@2.[213,1374,181,1574]" captionTargetPageId="2" captionText="FIGURES 28. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 2, 4, 5) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 3, 68); habitus (2, 3), male palp (4, 5) and female genitalia (68). 2, Habitus of male, dorsal view. 3, Habitus of female, dorsal view. 4, Male left palp, prolateral view. 5, Same, anterior view. 6, Epigynum, ventral view. 7, Same, lateral view; 8, basal part of epigynum, ventral view. Arrows in Figs. 4 and 5 indicate a sharp appendix on the posterior margin of subterminal apophysis. Scale bars, 47, 1.0 mm; 8, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5577391" httpUri="https://zenodo.org/record/5577391/files/figure.png" pageId="3" pageNumber="594">Figs. 4, 5</figureCitation>
;
<figureCitation id="132A2A4FFFE3315D7191235AFE18FEB4" box="[399,471,296,320]" captionStart="FIGURES 915" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[189,1402,650,1712]" captionTargetId="figure-243@3.[177,1410,632,1728]" captionTargetPageId="3" captionText="FIGURES 915. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 9, 1112) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 10, 1315). 9, Apophysis on prolateral side of tibia II. 10, Projection on ventral side of coxa II. 11, Male left palp, prolateral view. 12, Same, apical part, anterior view.13, Epigynum, ventral view.14, Same, posterior view.15, Internal genitalia, dorsal view. Arrows in Figs. 1112 and 13, 14 indicate a sharp appendix on the posterior margin of subterminal apophysis (arrows 1, 3), bent tip of conductor (arrows 2, 4) and copulatory opening (arrows 5, 6), respectively. Abbreviations: c, conductor; fd, fertilization duct; lp, lateral plate; ma, median apophysis; mp, median plate; pc, paracymbium; s, spermatheca; st, subterminal apophysis; ta, terminal apophysis. Scale bars, 811, 1.0 mm; 1214, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5577397" httpUri="https://zenodo.org/record/5577397/files/figure.png" pageId="3" pageNumber="594">11, 12</figureCitation>
, arrows; the appendix is absent in
<taxonomicName id="4C114D49FFE3315D7343235AFC28FECB" baseAuthorityName="L. Koch" baseAuthorityYear="1878" box="[861,999,296,319]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="seminiger">
<emphasis id="B965EAD8FFE3315D7343235AFC28FECB" box="[861,999,296,319]" italics="true" pageId="3" pageNumber="594">A. seminiger</emphasis>
</taxonomicName>
); a longer slender epigynal scape, with outwardly directed copulatory openings (
<figureCitation id="132A2A4FFFE3315D72572339FD61FE97" box="[585,686,331,355]" captionStart="FIGURES 28" captionStartId="2.[151,264,1598,1622]" captionTargetBox="[230,1374,193,1574]" captionTargetId="figure-22@2.[213,1374,181,1574]" captionTargetPageId="2" captionText="FIGURES 28. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 2, 4, 5) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 3, 68); habitus (2, 3), male palp (4, 5) and female genitalia (68). 2, Habitus of male, dorsal view. 3, Habitus of female, dorsal view. 4, Male left palp, prolateral view. 5, Same, anterior view. 6, Epigynum, ventral view. 7, Same, lateral view; 8, basal part of epigynum, ventral view. Arrows in Figs. 4 and 5 indicate a sharp appendix on the posterior margin of subterminal apophysis. Scale bars, 47, 1.0 mm; 8, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5577391" httpUri="https://zenodo.org/record/5577391/files/figure.png" pageId="3" pageNumber="594">Figs. 68</figureCitation>
,
<figureCitation id="132A2A4FFFE3315D72A7233EFD32FE90" box="[697,765,332,356]" captionStart="FIGURES 915" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[189,1402,650,1712]" captionTargetId="figure-243@3.[177,1410,632,1728]" captionTargetPageId="3" captionText="FIGURES 915. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 9, 1112) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 10, 1315). 9, Apophysis on prolateral side of tibia II. 10, Projection on ventral side of coxa II. 11, Male left palp, prolateral view. 12, Same, apical part, anterior view.13, Epigynum, ventral view.14, Same, posterior view.15, Internal genitalia, dorsal view. Arrows in Figs. 1112 and 13, 14 indicate a sharp appendix on the posterior margin of subterminal apophysis (arrows 1, 3), bent tip of conductor (arrows 2, 4) and copulatory opening (arrows 5, 6), respectively. Abbreviations: c, conductor; fd, fertilization duct; lp, lateral plate; ma, median apophysis; mp, median plate; pc, paracymbium; s, spermatheca; st, subterminal apophysis; ta, terminal apophysis. Scale bars, 811, 1.0 mm; 1214, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5577397" httpUri="https://zenodo.org/record/5577397/files/figure.png" pageId="3" pageNumber="594">13, 14</figureCitation>
; epigynal scape of
<taxonomicName id="4C114D49FFE3315D73DB233EFB85FE97" baseAuthorityName="L. Koch" baseAuthorityYear="1878" box="[965,1098,332,355]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="seminiger">
<emphasis id="B965EAD8FFE3315D73DB233EFB85FE97" box="[965,1098,332,355]" italics="true" pageId="3" pageNumber="594">A. seminiger</emphasis>
</taxonomicName>
is shorter and wider than that of the new species, and copulatory openings of
<taxonomicName id="4C114D49FFE3315D726F2302FD38FE73" baseAuthorityName="L. Koch" baseAuthorityYear="1878" box="[625,759,368,391]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="seminiger">
<emphasis id="B965EAD8FFE3315D726F2302FD38FE73" box="[625,759,368,391]" italics="true" pageId="3" pageNumber="594">A. seminiger</emphasis>
</taxonomicName>
is directed ventrally). In terms of general appearance (i.e., body coloration and markings), the new species can be distinguished from the morphologically related species
<taxonomicName id="4C114D49FFE3315D751A23E6FEF0FE24" authority="(L. Koch 1878)" baseAuthorityName="L. Koch" baseAuthorityYear="1878" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="ventricosus">
<emphasis id="B965EAD8FFE3315D751A23E6FA53FE5F" box="[1284,1436,404,427]" italics="true" pageId="3" pageNumber="594">A. ventricosus</emphasis>
(L.
<bibRefCitation id="EF804B3BFFE3315D70A023C5FEF7FE24" author="Koch, L." box="[190,312,439,464]" pageId="3" pageNumber="594" pagination="735 - 798" refId="ref3219" refString="Koch, L. (1878) Japanesische Arachniden und Myriapoden. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 27, 735 - 798." type="journal article" year="1878">Koch 1878</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C114D49FFE3315D715523CAFD4FFE3B" authority="(Uyemura 1961)" baseAuthorityName="Uyemura" baseAuthorityYear="1961" box="[331,640,439,464]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="macacus">
<emphasis id="B965EAD8FFE3315D715523CAFE09FE3B" box="[331,454,440,463]" italics="true" pageId="3" pageNumber="594">A. macacus</emphasis>
(
<bibRefCitation id="EF804B3BFFE3315D71CA23CAFDB7FE3B" author="Uyemura, T." box="[468,632,439,464]" pageId="3" pageNumber="594" refId="ref3619" refString="Uyemura, T. (1961) Studies on the variation, lineage and distribution of the Japanese spider, Araneus ventricosus (L. Koch) (s. lat.). Arachnological Society of East Asia, Osaka, 116 pp." type="book" year="1961">Uyemura 1961</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C114D49FFE3315D729223CAFC0EFE3B" authority="Yaginuma 1960" authorityName="Yaginuma" authorityYear="1960" box="[652,961,439,463]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="uyemurai">
<emphasis id="B965EAD8FFE3315D729223CAFCC0FE3B" box="[652,783,440,463]" italics="true" pageId="3" pageNumber="594">A. uyemurai</emphasis>
<bibRefCitation id="EF804B3BFFE3315D730823C5FC0EFE3B" author="Yaginuma, T." box="[790,961,439,463]" pageId="3" pageNumber="594" refId="ref3664" refString="Yaginuma, T. (1960) Spiders of Japan in colour. Hoikusha, Osaka, 186 pp." type="book" year="1960">Yaginuma 1960</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C114D49FFE3315D73E723CAFAC8FE24" authority="Tanikawa 2020" authorityName="Tanikawa" authorityYear="2020" box="[1017,1287,439,464]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="reizan">
<emphasis id="B965EAD8FFE3315D73E723CAFB96FE3B" box="[1017,1113,440,463]" italics="true" pageId="3" pageNumber="594">A. reizan</emphasis>
Tanikawa 2020
</taxonomicName>
. Although the epigynum of the new species is similar to that of
<taxonomicName id="4C114D49FFE3315D728023AEFD32FE07" authorityName="Tanikawa" authorityYear="2020" box="[670,765,476,499]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="reizan">
<emphasis id="B965EAD8FFE3315D728023AEFD32FE07" box="[670,765,476,499]" italics="true" pageId="3" pageNumber="594">A. reizan</emphasis>
</taxonomicName>
, it differs with respect to a deeper invagination on the posterior side of median plate (
<figureCitation id="132A2A4FFFE3315D7163238DFE06FDEC" box="[381,457,511,536]" captionStart="FIGURES 28" captionStartId="2.[151,264,1598,1622]" captionTargetBox="[230,1374,193,1574]" captionTargetId="figure-22@2.[213,1374,181,1574]" captionTargetPageId="2" captionText="FIGURES 28. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 2, 4, 5) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 3, 68); habitus (2, 3), male palp (4, 5) and female genitalia (68). 2, Habitus of male, dorsal view. 3, Habitus of female, dorsal view. 4, Male left palp, prolateral view. 5, Same, anterior view. 6, Epigynum, ventral view. 7, Same, lateral view; 8, basal part of epigynum, ventral view. Arrows in Figs. 4 and 5 indicate a sharp appendix on the posterior margin of subterminal apophysis. Scale bars, 47, 1.0 mm; 8, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5577391" httpUri="https://zenodo.org/record/5577391/files/figure.png" pageId="3" pageNumber="594">Figs. 8</figureCitation>
,
<figureCitation id="132A2A4FFFE3315D71CA2072FDD6FDEC" box="[468,537,512,536]" captionStart="FIGURES 915" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[189,1402,650,1712]" captionTargetId="figure-243@3.[177,1410,632,1728]" captionTargetPageId="3" captionText="FIGURES 915. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 9, 1112) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 10, 1315). 9, Apophysis on prolateral side of tibia II. 10, Projection on ventral side of coxa II. 11, Male left palp, prolateral view. 12, Same, apical part, anterior view.13, Epigynum, ventral view.14, Same, posterior view.15, Internal genitalia, dorsal view. Arrows in Figs. 1112 and 13, 14 indicate a sharp appendix on the posterior margin of subterminal apophysis (arrows 1, 3), bent tip of conductor (arrows 2, 4) and copulatory opening (arrows 5, 6), respectively. Abbreviations: c, conductor; fd, fertilization duct; lp, lateral plate; ma, median apophysis; mp, median plate; pc, paracymbium; s, spermatheca; st, subterminal apophysis; ta, terminal apophysis. Scale bars, 811, 1.0 mm; 1214, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5577397" httpUri="https://zenodo.org/record/5577397/files/figure.png" pageId="3" pageNumber="594">13, 14</figureCitation>
;
<taxonomicName id="4C114D49FFE3315D723B2072FD4AFDE3" authorityName="Tanikawa" authorityYear="2020" box="[549,645,512,535]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="reizan">
<emphasis id="B965EAD8FFE3315D723B2072FD4AFDE3" box="[549,645,512,535]" italics="true" pageId="3" pageNumber="594">A. reizan</emphasis>
</taxonomicName>
has a very shallow invagination).
</paragraph>
</subSubSection>
<subSubSection id="C30B6541FFE3315A70D92051FB33FC8B" lastPageId="4" lastPageNumber="595" pageId="3" pageNumber="594" type="description">
<paragraph id="8BAE36CAFFE3315D70D92051FF04FDAB" blockId="3.[151,1437,151,607]" pageId="3" pageNumber="594">
<emphasis id="B965EAD8FFE3315D70D92051FE9DFDCF" bold="true" box="[199,338,547,571]" inLineHeading="true" pageId="3" pageNumber="594" reason="1">Description.</emphasis>
Based on the
<typeStatus id="54AA8868FFE3315D71F12051FD83FDCF" box="[495,588,547,571]" pageId="3" pageNumber="594" type="holotype">holotype</typeStatus>
male (NSMT-Ar 20952) and
<typeStatus id="54AA8868FFE3315D738F2056FC22FDC8" box="[913,1005,548,572]" pageId="3" pageNumber="594" type="paratype">paratype</typeStatus>
<specimenCount id="9D17FD43FFE3315D73EB2056FB9CFDCF" box="[1013,1107,547,572]" pageId="3" pageNumber="594" type="female">1 female</specimenCount>
(NSMT-Ar 20958) from
<collectingRegion id="49D5F828FFE3315D75762056FF04FDAB" country="Japan" name="Nara" pageId="3" pageNumber="594">Nara Pref.</collectingRegion>
</paragraph>
<caption id="DF6E6642FFE3315D708924A5FB2EF833" ID-DOI="http://doi.org/10.5281/zenodo.5577397" ID-Zenodo-Dep="5577397" httpUri="https://zenodo.org/record/5577397/files/figure.png" pageId="3" pageNumber="594" startId="3.[151,264,1751,1775]" targetBox="[189,1402,650,1712]" targetPageId="3">
<paragraph id="8BAE36CAFFE3315D708924A5FB2EF833" blockId="3.[151,1437,1751,1992]" pageId="3" pageNumber="594">
<emphasis id="B965EAD8FFE3315D708924A5FE85F904" bold="true" box="[151,330,1751,1776]" pageId="3" pageNumber="594">FIGURES 915.</emphasis>
<taxonomicName id="4C114D49FFE3315D714D24AAFDE8F91B" authority="Suzuki &amp; Tanikawa, 2021" authorityName="Suzuki &amp; Tanikawa" authorityYear="2021" box="[339,551,1752,1775]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="matsumotoi" status="sp. nov.">
<emphasis id="B965EAD8FFE3315D714D24AAFDE8F91B" box="[339,551,1752,1775]" italics="true" pageId="3" pageNumber="594">Araneus matsumotoi</emphasis>
</taxonomicName>
<emphasis id="B965EAD8FFE3315D722E24AAFDA3F904" bold="true" box="[560,620,1752,1776]" pageId="3" pageNumber="594">
<taxonomicNameLabel id="A25657A3FFE3315D722E24AAFDA3F904" box="[560,620,1752,1776]" pageId="3" pageNumber="594" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male holotype (NSMT-Ar 20952; 9, 1112) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 10, 1315). 9, Apophysis on prolateral side of tibia II. 10, Projection on ventral side of coxa II. 11, Male left palp, prolateral view. 12, Same, apical part, anterior view.13, Epigynum, ventral view.14, Same, posterior view.15, Internal genitalia, dorsal view. Arrows in Figs. 1112 and 13, 14 indicate a sharp appendix on the posterior margin of subterminal apophysis (arrows 1, 3), bent tip of conductor (arrows 2, 4) and copulatory opening (arrows 5, 6), respectively. Abbreviations: c, conductor; fd, fertilization duct; lp, lateral plate; ma, median apophysis; mp, median plate; pc, paracymbium; s, spermatheca; st, subterminal apophysis; ta, terminal apophysis. Scale bars, 811, 1.0 mm; 1214, 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BAE36CAFFE4315A70D922EAFA4DFE73" blockId="4.[151,1437,151,1003]" pageId="4" pageNumber="595">
Measurements and morphology (
<figureCitation id="132A2A4FFFE4315A723B22E5FD55FF5B" box="[549,666,151,175]" captionStart-0="FIGURES 28" captionStart-1="FIGURES 915" captionStartId-0="2.[151,264,1598,1622]" captionStartId-1="3.[151,264,1751,1775]" captionTargetBox-0="[230,1374,193,1574]" captionTargetBox-1="[189,1402,650,1712]" captionTargetId-0="figure-22@2.[213,1374,181,1574]" captionTargetId-1="figure-243@3.[177,1410,632,1728]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 28. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 2, 4, 5) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 3, 68); habitus (2, 3), male palp (4, 5) and female genitalia (68). 2, Habitus of male, dorsal view. 3, Habitus of female, dorsal view. 4, Male left palp, prolateral view. 5, Same, anterior view. 6, Epigynum, ventral view. 7, Same, lateral view; 8, basal part of epigynum, ventral view. Arrows in Figs. 4 and 5 indicate a sharp appendix on the posterior margin of subterminal apophysis. Scale bars, 47, 1.0 mm; 8, 0.5 mm." captionText-1="FIGURES 915. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 9, 1112) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 10, 1315). 9, Apophysis on prolateral side of tibia II. 10, Projection on ventral side of coxa II. 11, Male left palp, prolateral view. 12, Same, apical part, anterior view.13, Epigynum, ventral view.14, Same, posterior view.15, Internal genitalia, dorsal view. Arrows in Figs. 1112 and 13, 14 indicate a sharp appendix on the posterior margin of subterminal apophysis (arrows 1, 3), bent tip of conductor (arrows 2, 4) and copulatory opening (arrows 5, 6), respectively. Abbreviations: c, conductor; fd, fertilization duct; lp, lateral plate; ma, median apophysis; mp, median plate; pc, paracymbium; s, spermatheca; st, subterminal apophysis; ta, terminal apophysis. Scale bars, 811, 1.0 mm; 1214, 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5577391" figureDoi-1="http://doi.org/10.5281/zenodo.5577397" httpUri-0="https://zenodo.org/record/5577391/files/figure.png" httpUri-1="https://zenodo.org/record/5577397/files/figure.png" pageId="4" pageNumber="595">Figs. 215</figureCitation>
).
<specimenCount id="9D17FD43FFE4315A72B522EAFD0DFF44" box="[683,706,152,176]" pageId="4" pageNumber="595" type="male"></specimenCount>
/
<specimenCount id="9D17FD43FFE4315A72D322EAFD12FF44" box="[717,733,152,176]" pageId="4" pageNumber="595" type="female"></specimenCount>
. Body 12.30/17.95 long. Carapace 6.75/7.77 long; 5.34/6.43 wide. Eye size: AME 0.32/0.33; ALE 0.24/0.23; PME 0.22/0.25; PLE 0.19/0.22. Length of legs = [femur + patella + tibia + metatarsus + tarsus = total]. I, 6.82 + 2.65 + 6.75 + 6.20 + 2.18 = 24.60 / 8.47 + 3.58 + 7.72 + 7.69 + 2.93 = 30.39; II, 6.67 + 2.60 + 5.93 + 5.60 + 2.01 = 22.81 / 7.69 + 3.40 + 6.97 + 6.12 + 2.27 = 26.45; III, 4.80 + 1.73 + 3.08 + 2.92 + 1.47 = 14.00 / 5.64 + 2.37 + 3.74 + 3.14 + 1.56 = 16.45; IV, 6.90 + 2.40 + 5.03 + 4.86 + 1.69 = 20.88 / 7.62 + 3.35 + 5.90 + 5.37 + 1.65 = 23.89. Apophysis with a long pedestal developed on prolateral side of male tibia II (
<figureCitation id="132A2A4FFFE4315A74482339FB5AFE90" box="[1110,1173,331,356]" captionStart="FIGURES 915" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[189,1402,650,1712]" captionTargetId="figure-243@3.[177,1410,632,1728]" captionTargetPageId="3" captionText="FIGURES 915. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 9, 1112) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 10, 1315). 9, Apophysis on prolateral side of tibia II. 10, Projection on ventral side of coxa II. 11, Male left palp, prolateral view. 12, Same, apical part, anterior view.13, Epigynum, ventral view.14, Same, posterior view.15, Internal genitalia, dorsal view. Arrows in Figs. 1112 and 13, 14 indicate a sharp appendix on the posterior margin of subterminal apophysis (arrows 1, 3), bent tip of conductor (arrows 2, 4) and copulatory opening (arrows 5, 6), respectively. Abbreviations: c, conductor; fd, fertilization duct; lp, lateral plate; ma, median apophysis; mp, median plate; pc, paracymbium; s, spermatheca; st, subterminal apophysis; ta, terminal apophysis. Scale bars, 811, 1.0 mm; 1214, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5577397" httpUri="https://zenodo.org/record/5577397/files/figure.png" pageId="4" pageNumber="595">Fig. 9</figureCitation>
) and sharp projection on ventral side of coxa II (
<figureCitation id="132A2A4FFFE4315A718E231DFE2FFE7C" box="[400,480,367,392]" captionStart="FIGURES 915" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[189,1402,650,1712]" captionTargetId="figure-243@3.[177,1410,632,1728]" captionTargetPageId="3" captionText="FIGURES 915. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 9, 1112) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 10, 1315). 9, Apophysis on prolateral side of tibia II. 10, Projection on ventral side of coxa II. 11, Male left palp, prolateral view. 12, Same, apical part, anterior view.13, Epigynum, ventral view.14, Same, posterior view.15, Internal genitalia, dorsal view. Arrows in Figs. 1112 and 13, 14 indicate a sharp appendix on the posterior margin of subterminal apophysis (arrows 1, 3), bent tip of conductor (arrows 2, 4) and copulatory opening (arrows 5, 6), respectively. Abbreviations: c, conductor; fd, fertilization duct; lp, lateral plate; ma, median apophysis; mp, median plate; pc, paracymbium; s, spermatheca; st, subterminal apophysis; ta, terminal apophysis. Scale bars, 811, 1.0 mm; 1214, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5577397" httpUri="https://zenodo.org/record/5577397/files/figure.png" pageId="4" pageNumber="595">Fig. 10</figureCitation>
). Abdomen 6.43/10.51 long; 5.77/9.96 wide. A pair of shoulder humps well developed.
</paragraph>
<paragraph id="8BAE36CAFFE4315A70D923E1FA9EFDAB" blockId="4.[151,1437,151,1003]" pageId="4" pageNumber="595">
Coloration and markings (
<figureCitation id="132A2A4FFFE4315A71F923E1FD9DFE58" box="[487,594,403,428]" captionStart="FIGURES 28" captionStartId="2.[151,264,1598,1622]" captionTargetBox="[230,1374,193,1574]" captionTargetId="figure-22@2.[213,1374,181,1574]" captionTargetPageId="2" captionText="FIGURES 28. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 2, 4, 5) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 3, 68); habitus (2, 3), male palp (4, 5) and female genitalia (68). 2, Habitus of male, dorsal view. 3, Habitus of female, dorsal view. 4, Male left palp, prolateral view. 5, Same, anterior view. 6, Epigynum, ventral view. 7, Same, lateral view; 8, basal part of epigynum, ventral view. Arrows in Figs. 4 and 5 indicate a sharp appendix on the posterior margin of subterminal apophysis. Scale bars, 47, 1.0 mm; 8, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5577391" httpUri="https://zenodo.org/record/5577391/files/figure.png" pageId="4" pageNumber="595">Figs. 2, 3</figureCitation>
). Coloration: when living/specimens fixed in ethanol, respectively. Carapace gray/dark yellowish brown with black markings and spots on cervical grooves, redial furrows, and fovea. Sternum pale green/light yellowish brown. Legs pale green with dark green ring bands/light yellowish brown with dark brown ring bands; ventral side of coxae black/dark yellowish brown. Dorsum of abdomen dark green/light green and yellowish brown with black folium; internal area of folium darker; green chevron marking edged with black between humps; two pairs of large sigilla on dorsum; ventral side of abdomen light yellowish brown with pair of light yellowish spots.
</paragraph>
<paragraph id="8BAE36CAFFE4315A70D92019FE01FCAF" blockId="4.[151,1437,151,1003]" pageId="4" pageNumber="595">
Genital organs. Male palp (
<figureCitation id="132A2A4FFFE4315A71EC2019FD94FD77" box="[498,603,619,644]" captionStart="FIGURES 28" captionStartId="2.[151,264,1598,1622]" captionTargetBox="[230,1374,193,1574]" captionTargetId="figure-22@2.[213,1374,181,1574]" captionTargetPageId="2" captionText="FIGURES 28. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 2, 4, 5) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 3, 68); habitus (2, 3), male palp (4, 5) and female genitalia (68). 2, Habitus of male, dorsal view. 3, Habitus of female, dorsal view. 4, Male left palp, prolateral view. 5, Same, anterior view. 6, Epigynum, ventral view. 7, Same, lateral view; 8, basal part of epigynum, ventral view. Arrows in Figs. 4 and 5 indicate a sharp appendix on the posterior margin of subterminal apophysis. Scale bars, 47, 1.0 mm; 8, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5577391" httpUri="https://zenodo.org/record/5577391/files/figure.png" pageId="4" pageNumber="595">Figs. 4, 5</figureCitation>
,
<figureCitation id="132A2A4FFFE4315A7277201EFD7FFD70" box="[617,688,620,644]" captionStart="FIGURES 915" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[189,1402,650,1712]" captionTargetId="figure-243@3.[177,1410,632,1728]" captionTargetPageId="3" captionText="FIGURES 915. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 9, 1112) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 10, 1315). 9, Apophysis on prolateral side of tibia II. 10, Projection on ventral side of coxa II. 11, Male left palp, prolateral view. 12, Same, apical part, anterior view.13, Epigynum, ventral view.14, Same, posterior view.15, Internal genitalia, dorsal view. Arrows in Figs. 1112 and 13, 14 indicate a sharp appendix on the posterior margin of subterminal apophysis (arrows 1, 3), bent tip of conductor (arrows 2, 4) and copulatory opening (arrows 5, 6), respectively. Abbreviations: c, conductor; fd, fertilization duct; lp, lateral plate; ma, median apophysis; mp, median plate; pc, paracymbium; s, spermatheca; st, subterminal apophysis; ta, terminal apophysis. Scale bars, 811, 1.0 mm; 1214, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5577397" httpUri="https://zenodo.org/record/5577397/files/figure.png" pageId="4" pageNumber="595">11, 12</figureCitation>
): tip of terminal apophysis sclerotized and sharp; conductor strongly sclerotized with bent tip; subterminal apophysis with wide unsclerotized area and sharp thin sclerotized appendix on posterior margin; embolus hidden under subterminal apophysis and conductor; median apophysis transverse with prolateral tip pointed and retrolateral tip truncated; paracymbium hook-like with blunt tip. Female genitalia (
<figureCitation id="132A2A4FFFE4315A752E20A5FA58FD1B" box="[1328,1431,727,751]" captionStart="FIGURES 28" captionStartId="2.[151,264,1598,1622]" captionTargetBox="[230,1374,193,1574]" captionTargetId="figure-22@2.[213,1374,181,1574]" captionTargetPageId="2" captionText="FIGURES 28. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 2, 4, 5) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 3, 68); habitus (2, 3), male palp (4, 5) and female genitalia (68). 2, Habitus of male, dorsal view. 3, Habitus of female, dorsal view. 4, Male left palp, prolateral view. 5, Same, anterior view. 6, Epigynum, ventral view. 7, Same, lateral view; 8, basal part of epigynum, ventral view. Arrows in Figs. 4 and 5 indicate a sharp appendix on the posterior margin of subterminal apophysis. Scale bars, 47, 1.0 mm; 8, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5577391" httpUri="https://zenodo.org/record/5577391/files/figure.png" pageId="4" pageNumber="595">Figs. 68</figureCitation>
,
<figureCitation id="132A2A4FFFE4315A70892089FF11FCE7" box="[151,222,763,787]" captionStart="FIGURES 915" captionStartId="3.[151,264,1751,1775]" captionTargetBox="[189,1402,650,1712]" captionTargetId="figure-243@3.[177,1410,632,1728]" captionTargetPageId="3" captionText="FIGURES 915. Araneus matsumotoi n. sp., male holotype (NSMT-Ar 20952; 9, 1112) and one of the female paratypes from Nara Pref. (NSMT-Ar 20958; 10, 1315). 9, Apophysis on prolateral side of tibia II. 10, Projection on ventral side of coxa II. 11, Male left palp, prolateral view. 12, Same, apical part, anterior view.13, Epigynum, ventral view.14, Same, posterior view.15, Internal genitalia, dorsal view. Arrows in Figs. 1112 and 13, 14 indicate a sharp appendix on the posterior margin of subterminal apophysis (arrows 1, 3), bent tip of conductor (arrows 2, 4) and copulatory opening (arrows 5, 6), respectively. Abbreviations: c, conductor; fd, fertilization duct; lp, lateral plate; ma, median apophysis; mp, median plate; pc, paracymbium; s, spermatheca; st, subterminal apophysis; ta, terminal apophysis. Scale bars, 811, 1.0 mm; 1214, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5577397" httpUri="https://zenodo.org/record/5577397/files/figure.png" pageId="4" pageNumber="595">1315</figureCitation>
): scape slender with apical half sclerotized and basal half wrinkled; median plate rounded with deep invagination on posterior edge, looking like inverted heart shaped from ventral view; lateral plates strongly sclerotized; copulatory openings outwardly directed.
</paragraph>
<paragraph id="8BAE36CAFFE4315A70D92115FB33FC8B" blockId="4.[151,1437,151,1003]" box="[199,1276,871,896]" pageId="4" pageNumber="595">
<emphasis id="B965EAD8FFE4315A70D92115FDF0FC8B" box="[199,575,871,896]" inLineHeading="true" pageId="4" pageNumber="595" reason="1">
<emphasis id="B965EAD8FFE4315A70D92115FEFDFC8B" bold="true" box="[199,306,871,895]" pageId="4" pageNumber="595">Variation</emphasis>
(
<specimenCount id="9D17FD43FFE4315A715C2115FE5BFC8B" box="[322,404,871,895]" pageId="4" pageNumber="595" type="male">5 males</specimenCount>
and
<specimenCount id="9D17FD43FFE4315A71D6211AFDFBFC8B" box="[456,564,871,896]" pageId="4" pageNumber="595" type="female">3 females</specimenCount>
).
</emphasis>
Male body length 11.2412.30; female body length 14.3017.95.
</paragraph>
</subSubSection>
<subSubSection id="C30B6541FFE4315A70D921F9FD3EFC1F" pageId="4" pageNumber="595" type="distribution">
<paragraph id="8BAE36CAFFE4315A70D921F9FC49FC57" blockId="4.[151,1437,151,1003]" box="[199,902,907,931]" pageId="4" pageNumber="595">
<emphasis id="B965EAD8FFE4315A70D921F9FE95FC57" bold="true" box="[199,346,907,931]" inLineHeading="true" pageId="4" pageNumber="595" reason="1">Distribution.</emphasis>
The Honshu, Shikoku and Tsushima Islands,
<collectingCountry id="F306765AFFE4315A735F21FEFC4EFC57" box="[833,897,908,931]" name="Japan" pageId="4" pageNumber="595">Japan</collectingCountry>
.
</paragraph>
<paragraph id="8BAE36CAFFE4315A70D921DDFD3EFC1F" blockId="4.[151,1437,151,1003]" pageId="4" pageNumber="595">
<emphasis id="B965EAD8FFE4315A70D921DDFEE9FC33" bold="true" box="[199,294,943,967]" inLineHeading="true" pageId="4" pageNumber="595" reason="1">Habitat.</emphasis>
The new species inhabits deciduous and evergreen forests, in which specimens are often found in green moss on the bark and trunks of trees during the daytime.
</paragraph>
</subSubSection>
</treatment>
</document>