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<document id="3259B143B0B86FF0FA26066425EB374E" ID-DOI="10.26107/RBZ-2022-0023" ID-ISSN="2345-7600" ID-Zenodo-Dep="7174433" ID-ZooBank="E4F49C16-DB5E-4593-AB8E-03D8859A59D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665290687137" checkinUser="felipe" docAuthor="Páll-Gergely, Barna &amp; Hunyadi, András" docDate="2022" docId="03B887D72308266AADC132F122A4FA12" docLanguage="en" docName="RafflesBZool.70.417–439.pdf" docOrigin="Raffles Bulletin of Zoology 70" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Clostophis charybdis Páll-Gergely &amp; Hunyadi 2022, new species" docType="treatment" docVersion="4" lastPageNumber="434" masterDocId="FF81FFAF2305267BAE6F347A2327FFA7" masterDocTitle="New and little-known species of Clostophis Benson, 1860 from Southeast Asia (Gastropoda: Eupulmonata: Hypselostomatidae)" masterLastPageNumber="439" masterPageNumber="417" pageNumber="430" updateTime="1699379781661" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="907E9D8E852F0AE70AABFDDC140172E3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="65E7714E6079C28F39DB0BC7BA72E9CD">
<mods:title id="9FBB4F185494B019C77136E4A9C21C47">New and little-known species of Clostophis Benson, 1860 from Southeast Asia (Gastropoda: Eupulmonata: Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name id="414854CFCD40AC6485163B42494AC414" type="personal">
<mods:role id="6577749E198A9022392733116076B3A0">
<mods:roleTerm id="CBCD5B9A711EC060F4DE7A8A2DDB91FA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C5351236188AA21CF43DE2938E4FCEE2">Páll-Gergely, Barna</mods:namePart>
</mods:name>
<mods:name id="DB13ABC6D7AA7AD1485943360A76897F" type="personal">
<mods:role id="9963EBCC4DE0572A3EEEE40E3D7B2B81">
<mods:roleTerm id="7AE640C6CFE8CC39345BFE2EEBDA29C2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="14A5D5BD634FCAEC3F61D03A4D81B2A3">Hunyadi, András</mods:namePart>
</mods:name>
<mods:typeOfResource id="443DC4CA1BEF489BF0575AAE82AA11B8">text</mods:typeOfResource>
<mods:relatedItem id="52C9A64997F632E32FA7DF8AE723378E" type="host">
<mods:titleInfo id="B45CD17F9751003821B7ADDFFBF9238C">
<mods:title id="4A5AA4D9B38331CBD8AA96F3EBBFE544">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="D522151B0D4D46C48690A80141CC89B7">
<mods:date id="8FDF94155DA0F3E6D4EF68BF49409056">2022</mods:date>
<mods:detail id="C77018BF37C1EE0E00C1EC2FACA872CF" type="pubDate">
<mods:number id="9BB5B3D172A639405C4B76765B8561EB">2022-09-09</mods:number>
</mods:detail>
<mods:detail id="00BCA5319FDEB919E27557AC6F209907" type="volume">
<mods:number id="839357D22E99B2E49977E0E1C45790FA">70</mods:number>
</mods:detail>
<mods:extent id="4CC302E359A40CAC9D0A05D3446AEA62" unit="page">
<mods:start id="58AFF05E47635802CA80531462DB1EB7">417</mods:start>
<mods:end id="E4629257D323225633545D8B2AF6E69A">439</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9B87A1A74F539812AD2AB59B538FCB37">journal article</mods:classification>
<mods:identifier id="2A9279CA307842E61050B55E340A87C0" type="DOI">10.26107/RBZ-2022-0023</mods:identifier>
<mods:identifier id="7654C707F8E68EE1F82C36B121C5B747" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="E0896542ED9BCD5523335B14B161CA89" type="Zenodo-Dep">7174433</mods:identifier>
<mods:identifier id="AF16FF1214C99F98E250148A93A46871" type="ZooBank">E4F49C16-DB5E-4593-AB8E-03D8859A59D2</mods:identifier>
</mods:mods>
<treatment id="03B887D72308266AADC132F122A4FA12" ID-DOI="http://doi.org/10.5281/zenodo.7171135" ID-Zenodo-Dep="7171135" LSID="urn:lsid:plazi:treatment:03B887D72308266AADC132F122A4FA12" httpUri="http://treatment.plazi.org/id/03B887D72308266AADC132F122A4FA12" lastPageId="17" lastPageNumber="434" pageId="13" pageNumber="430">
<subSubSection id="C30B654A23082676ADC132F1260DF904" box="[942,1322,1675,1700]" pageId="13" pageNumber="430" type="nomenclature">
<paragraph id="8BAE36C123082676ADC132F1260DF904" blockId="13.[942,1322,1675,1732]" box="[942,1322,1675,1700]" pageId="13" pageNumber="430">
<heading id="D0E681AD23082676ADC132F1260DF904" bold="true" box="[942,1322,1675,1700]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="430" reason="2">
<emphasis id="B965EAD323082676ADC132F1260DF904" bold="true" box="[942,1322,1675,1700]" pageId="13" pageNumber="430">
<taxonomicName id="4C114D4223082676ADC132F127B1F904" ID-CoL="ceb0ed2d-87d8-4ea3-9b45-3c1ea13ec4c4" authority="Páll-Gergely &amp; Hunyadi, 2022" authorityName="Páll-Gergely &amp; Hunyadi" authorityYear="2022" box="[942,1174,1675,1699]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="430" phylum="Mollusca" rank="species" species="charybdis" status="sp. nov.">
<emphasis id="B965EAD323082676ADC132F127B1F904" bold="true" box="[942,1174,1675,1699]" italics="true" pageId="13" pageNumber="430">Clostophis charybdis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25657A823082676AACA32F6260DF904" box="[1189,1322,1675,1700]" pageId="13" pageNumber="430" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30B654A23082676AA4D32D12791F963" box="[1058,1206,1707,1732]" pageId="13" pageNumber="430" type="description">
<paragraph id="8BAE36C123082676AA4D32D12791F963" blockId="13.[942,1322,1675,1732]" box="[1058,1206,1707,1732]" pageId="13" pageNumber="430">
(
<figureCitation id="132A2A4423082676AA4532D127A2F963" box="[1066,1157,1707,1732]" captionStart="Fig" captionStartId="15.[129,164,2093,2115]" captionTargetBox="[129,1435,189,2077]" captionTargetId="figure-10@15.[129,1435,189,2077]" captionTargetPageId="15" captionText="Fig. 13. Clostophis charybdis, new species (holotype, HNHM 105327)." figureDoi="http://doi.org/10.5281/zenodo.7174461" httpUri="https://zenodo.org/record/7174461/files/figure.png" pageId="13" pageNumber="430">Figs. 13</figureCitation>
,
<figureCitation id="132A2A4423082676AAFD32D6278AF963" box="[1170,1197,1708,1732]" captionStart="Fig" captionStartId="16.[151,186,2093,2115]" captionTargetBox="[151,1457,189,2077]" captionTargetId="figure-8@16.[151,1457,189,2077]" captionTargetPageId="16" captionText="Fig. 14. Clostophis charybdis, new species (paratype, coll. HA)." figureDoi="http://doi.org/10.5281/zenodo.7174463" httpUri="https://zenodo.org/record/7174463/files/figure.png" pageId="13" pageNumber="430">14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30B654A23082676AD42329620D1F7E3" pageId="13" pageNumber="430" type="materials_examined">
<paragraph id="8BAE36C123082676AD42329620D1F7E3" blockId="13.[811,1452,1771,2116]" pageId="13" pageNumber="430">
<emphasis id="B965EAD323082676AD42329620FDF8A4" bold="true" box="[813,986,1771,1796]" pageId="13" pageNumber="430">Type material.</emphasis>
<materialsCitation id="3B793C9C23082676AD8B3291268FF823" collectingDate="2020-02-17" collectionCode="HNHM" collectorName="Di Chi Keo Leng &amp; A. Hunyadi" country="Vietnam" county="Th am Khuyen Cave" elevation="375" latitude="21.938334" location="Province" longLatPrecision="12" longitude="106.397" municipality="Di Chi Keo Leng" pageId="13" pageNumber="430" specimenCode="HNHM 105327" specimenCount="1" typeStatus="holotype">
<typeStatus id="54AA886323082676AD8B3291276EF8A4" box="[996,1097,1771,1795]" pageId="13" pageNumber="430" type="holotype">Holotype</typeStatus>
(SH:
<quantity id="4CE99B2423082676AAFF329627D0F8A4" box="[1168,1271,1772,1796]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" pageId="13" pageNumber="430" unit="mm" value="1.18">1.18 mm</quantity>
, SW:
<quantity id="4CE99B2423082676AB5032962683F8A3" box="[1343,1444,1772,1796]" metricMagnitude="-3" metricUnit="m" metricValue="1.12" pageId="13" pageNumber="430" unit="mm" value="1.12">1.12 mm</quantity>
) (
<specimenCode id="DBB79EBA23082676AD5B337620C4F884" box="[820,995,1803,1828]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="13" pageNumber="430">HNHM 105327</specimenCode>
),
<collectingCountry id="F306765123082676AD9833712771F884" box="[1015,1110,1803,1827]" name="Vietnam" pageId="13" pageNumber="430">Vietnam</collectingCountry>
, Lạng SƠn
<location id="8ECE601A23082676AABB33712611F884" LSID="urn:lsid:plazi:treatment:03B887D72308266AADC132F122A4FA12:8ECE601A23082676AABB33712611F884" box="[1236,1334,1803,1827]" country="Vietnam" county="Th am Khuyen Cave" latitude="21.938334" longLatPrecision="12" longitude="106.397" municipality="Di Chi Keo Leng" name="Province" pageId="13" pageNumber="430">Province</location>
,
<location id="8ECE601A23082676AB2D33712680F884" LSID="urn:lsid:plazi:treatment:03B887D72308266AADC132F122A4FA12:8ECE601A23082676AB2D33712680F884" box="[1346,1447,1803,1827]" country="Vietnam" county="Th am Khuyen Cave" latitude="21.938334" longLatPrecision="12" longitude="106.397" municipality="Di Chi Keo Leng" name="Binh Gia" pageId="13" pageNumber="430">Bình Gia</location>
, eastern edge of the village,
<collectorName id="26E4531723082676AA2133512623F8E3" box="[1102,1284,1835,1860]" pageId="13" pageNumber="430">
<collectingMunicipality id="6BCAACBB23082676AA2133512623F8E3" box="[1102,1284,1835,1860]" pageId="13" pageNumber="430">Di Chí Kéo Lèng</collectingMunicipality>
</collectorName>
(
<collectingCounty id="62CF4E4D23082676AB7E3356204EF8C3" pageId="13" pageNumber="430">Thẩm Khuyên Cave</collectingCounty>
), rock wall;
<geoCoordinate id="EE25500623082676AD933330275BF8C4" box="[1020,1148,1866,1891]" degrees="21" direction="north" minutes="56.30" orientation="latitude" pageId="13" pageNumber="430" precision="9" value="21.938334">21°56.30′N</geoCoordinate>
,
<geoCoordinate id="EE25500623082676AAE433302630F8C4" box="[1163,1303,1866,1891]" degrees="106" direction="east" minutes="23.82" orientation="longitude" pageId="13" pageNumber="430" precision="9" value="106.397">106°23.82′E</geoCoordinate>
,
<elevation id="003CD1F223082676AB4A3331268CF8C4" box="[1317,1451,1867,1891]" metricMagnitude="2" metricUnit="m" metricValue="3.75" pageId="13" pageNumber="430" unit="m" value="375.0">
<quantity id="4CE99B2423082676AB4A33312649F8C4" box="[1317,1390,1867,1891]" metricMagnitude="2" metricUnit="m" metricValue="3.75" pageId="13" pageNumber="430" unit="m" value="375.0">375 m</quantity>
a.s.l.
</elevation>
(locality code: 2020/48), coll.
<collectorName id="26E4531723082676AA09331627C4F824" box="[1126,1251,1899,1924]" pageId="13" pageNumber="430">A. Hunyadi</collectorName>
,
<date id="FFAF100123082676AA833316268FF823" box="[1260,1448,1899,1924]" pageId="13" pageNumber="430" value="2020-02-17">
<collectingDate id="EFEBE9E923082676AA833316268FF823" box="[1260,1448,1899,1924]" pageId="13" pageNumber="430" value="2020-02-17">17 February 2020</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B793C9C23082676AD4433F626A5F804" box="[811,1410,1931,1956]" collectingDate="2020-02-17" collectionCode="HNHM" collectorName="Di Chi Keo Leng &amp; A. Hunyadi" country="Vietnam" county="Th am Khuyen Cave" elevation="375" latitude="21.938334" location="Province" longLatPrecision="12" longitude="106.397" municipality="Di Chi Keo Leng" pageId="13" pageNumber="430" specimenCount="1" typeStatus="paratype">
<typeStatus id="54AA886323082676AD4433F620AAF803" box="[811,909,1932,1956]" pageId="13" pageNumber="430" type="paratype">Paratype</typeStatus>
: 1 shell, coll. HA, same data as for holotype
</materialsCitation>
;
<materialsCitation id="3B793C9C23082676ABE033F620D5F7E3" collectingDate="2020-02-18" collectorName="Province, B &amp; A. Hunyadi" country="Vietnam" elevation="241390" latitude="21.945467" location="District" longLatPrecision="1" longitude="106.32412" municipality="Long Dong" pageId="13" pageNumber="430" specimenCount="30" typeStatus="paratype">
<specimenCount id="9D17FD4823082676ABE033F620B1F863" pageId="13" pageNumber="430" type="generic" typeStatus="paratypes">30 paratypes</specimenCount>
, coll. HA,
<collectingCountry id="F306765123082676AA6433D1274DF864" box="[1035,1130,1963,1987]" name="Vietnam" pageId="13" pageNumber="430">Vietnam</collectingCountry>
, Lạng SƠn
<collectorName id="26E4531723082676AA8B33D12646F861" box="[1252,1377,1963,1990]" pageId="13" pageNumber="430">Province, B</collectorName>
ắc SƠn
<location id="8ECE601A23082676AD4233B120A7F844" LSID="urn:lsid:plazi:treatment:03B887D72308266AADC132F122A4FA12:8ECE601A23082676AD4233B120A7F844" box="[813,896,1995,2019]" country="Vietnam" latitude="21.945467" longLatPrecision="1" longitude="106.32412" municipality="Long Dong" name="District" pageId="13" pageNumber="430">District</location>
,
<collectingMunicipality id="6BCAACBB23082676ADE233B6272AF843" box="[909,1037,1996,2020]" pageId="13" pageNumber="430">Long Dong</collectingMunicipality>
,
<quantity id="4CE99B2423082676AA7533B62740F844" box="[1050,1127,1995,2020]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="13" pageNumber="430" unit="km" value="3.8">3.8 km</quantity>
north from junction of roads no. QL1B (main road) and
<elevation id="003CD1F223082676AA3733962628F7A4" box="[1112,1295,2027,2052]" metricMagnitude="5" metricUnit="m" metricValue="2.4139" pageId="13" pageNumber="430" unit="m" value="241390.0">
<quantity id="4CE99B2423082676AA37339627F3F7A4" box="[1112,1236,2028,2052]" metricMagnitude="5" metricUnit="m" metricValue="2.4139" pageId="13" pageNumber="430" unit="m" value="241390.0">241, 390 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="EE25500623082676AB7233902680F7A4" box="[1309,1447,2026,2051]" degrees="21" direction="north" minutes="56.728" orientation="latitude" pageId="13" pageNumber="430" precision="1" value="21.945467">21°56.728′N</geoCoordinate>
,
<geoCoordinate id="EE25500623082676AD423C7020E6F784" box="[813,961,2058,2083]" degrees="106" direction="east" minutes="19.447" orientation="longitude" pageId="13" pageNumber="430" precision="1" value="106.32412">106°19.447′E</geoCoordinate>
(locality code: 2020/51), coll.
<collectorName id="26E4531723082676AB4C3C762680F784" box="[1315,1447,2059,2084]" pageId="13" pageNumber="430">A. Hunyadi</collectorName>
,
<date id="FFAF100123082676AD423C5620D5F7E3" box="[813,1010,2091,2116]" pageId="13" pageNumber="430" value="2020-02-18">
<collectingDate id="EFEBE9E923082676AD423C5620D5F7E3" box="[813,1010,2091,2116]" pageId="13" pageNumber="430" value="2020-02-18">18 February 2020</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF6E6649230B2675AEF83C57274DF7E4" ID-DOI="http://doi.org/10.5281/zenodo.7174459" ID-Zenodo-Dep="7174459" box="[151,1130,2093,2115]" httpUri="https://zenodo.org/record/7174459/files/figure.png" pageId="14" pageNumber="431" startId="14.[151,186,2093,2115]" targetBox="[151,1457,189,2078]" targetPageId="14">
<paragraph id="8BAE36C1230B2675AEF83C57274DF7E4" blockId="14.[151,1130,2093,2115]" box="[151,1130,2093,2115]" pageId="14" pageNumber="431">
Fig. 12.
<taxonomicName id="4C114D42230B2675AE823C572297F7E5" authority="Páll-Gergely &amp; Hunyadi, 2022" authorityName="Páll-Gergely &amp; Hunyadi" authorityYear="2022" box="[237,432,2093,2114]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="431" phylum="Mollusca" rank="species" species="candidus" status="sp. nov.">
<emphasis id="B965EAD3230B2675AE823C572297F7E5" box="[237,432,2093,2114]" italics="true" pageId="14" pageNumber="431">Clostophis candidus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25657A8230B2675AFD13C542111F7E4" box="[446,566,2093,2115]" pageId="14" pageNumber="431" rank="species">new species</taxonomicNameLabel>
. A, paratype (coll. HA); B–H, holotype, HNHM 105326.
</paragraph>
</caption>
<caption id="DF6E6649230A2674AEEE3C572018F7E5" ID-DOI="http://doi.org/10.5281/zenodo.7174461" ID-Zenodo-Dep="7174461" box="[129,831,2092,2115]" httpUri="https://zenodo.org/record/7174461/files/figure.png" pageId="15" pageNumber="432" startId="15.[129,164,2093,2115]" targetBox="[129,1435,189,2077]" targetPageId="15">
<paragraph id="8BAE36C1230A2674AEEE3C572018F7E5" blockId="15.[129,831,2092,2115]" box="[129,831,2092,2115]" pageId="15" pageNumber="432">
Fig. 13.
<taxonomicName id="4C114D42230A2674AEB83C572285F7E5" authority="Páll-Gergely &amp; Hunyadi, 2022" authorityName="Páll-Gergely &amp; Hunyadi" authorityYear="2022" box="[215,418,2093,2114]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="432" phylum="Mollusca" rank="species" species="charybdis" status="sp. nov.">
<emphasis id="B965EAD3230A2674AEB83C572285F7E5" box="[215,418,2093,2114]" italics="true" pageId="15" pageNumber="432">Clostophis charybdis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25657A8230A2674AFDF3C572102F7E4" box="[432,549,2093,2115]" pageId="15" pageNumber="432" rank="species">new species</taxonomicNameLabel>
(holotype, HNHM 105327).
</paragraph>
</caption>
<caption id="DF6E66492315266BAEF83C572028F7E5" ID-DOI="http://doi.org/10.5281/zenodo.7174463" ID-Zenodo-Dep="7174463" box="[151,783,2093,2115]" httpUri="https://zenodo.org/record/7174463/files/figure.png" pageId="16" pageNumber="433" startId="16.[151,186,2093,2115]" targetBox="[151,1457,189,2077]" targetPageId="16">
<paragraph id="8BAE36C12315266BAEF83C572028F7E5" blockId="16.[151,783,2093,2115]" box="[151,783,2093,2115]" pageId="16" pageNumber="433">
Fig. 14.
<taxonomicName id="4C114D422315266BAE823C57229FF7E5" authority="Páll-Gergely &amp; Hunyadi, 2022" authorityName="Páll-Gergely &amp; Hunyadi" authorityYear="2022" box="[237,440,2093,2114]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="433" phylum="Mollusca" rank="species" species="charybdis" status="sp. nov.">
<emphasis id="B965EAD32315266BAE823C57229FF7E5" box="[237,440,2093,2114]" italics="true" pageId="16" pageNumber="433">Clostophis charybdis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25657A82315266BAFA93C57211CF7E4" box="[454,571,2093,2115]" pageId="16" pageNumber="433" rank="species">new species</taxonomicNameLabel>
(paratype, coll. HA).
</paragraph>
</caption>
<subSubSection id="C30B654A2314266AAE1E34C72145FE92" pageId="17" pageNumber="434" type="diagnosis">
<paragraph id="8BAE36C12314266AAE1E34C72145FE92" blockId="17.[113,753,189,309]" pageId="17" pageNumber="434">
<emphasis id="B965EAD32314266AAE1E34C723CEFF72" bold="true" box="[113,233,189,213]" pageId="17" pageNumber="434">Diagnosis.</emphasis>
A concave conical
<taxonomicName id="4C114D422314266AAFA034C721B9FF72" box="[463,670,189,213]" pageId="17" pageNumber="434">
<emphasis id="B965EAD32314266AAFA034C72163FF72" box="[463,580,189,213]" italics="true" pageId="17" pageNumber="434">Clostophis</emphasis>
species
</taxonomicName>
with a wide umbilicus, a well-developed, elevated parietal, and a shorter, pointed palatal tooth situated slightly upper (more apically) than where the parietal tooth points.
</paragraph>
</subSubSection>
<subSubSection id="C30B654A2314266AAE1E352723E2FB92" pageId="17" pageNumber="434" type="description">
<paragraph id="8BAE36C12314266AAE1E3527225DFC72" blockId="17.[112,755,349,981]" pageId="17" pageNumber="434">
<emphasis id="B965EAD32314266AAE1E35272224FED2" bold="true" box="[113,259,349,373]" pageId="17" pageNumber="434">Description.</emphasis>
Shell concave conical, widest at its base, slightly higher than wide to slightly wider than high, body whorl conspicuously wider than penultimate whorl. Whorls very slightly, bluntly shouldered. Protoconch consisting of 1.25–1.5 whorls, finely pitted/granular, and spirally striated preceding the first teleoconch whorl. Entire shell consisting of 4.5–4.75 whorls, with relatively strong, irregular radial growth lines and equally strong, dense, equidistant spiral striae. Body whorl with few, ca. 16–18 spiral striae from apertural view. Umbilicus regularly growing, wide, shell width is ca. 2.5 times wider than umbilicus. Aperture oblique to shell axis, oval to rectangular with blunt sinulus area. Sinulus large, only slightly smaller than other side of aperture, and well separated due to elevated parietal tooth and palatal tooth. Parietal callus rather blunt, not smeared onto penultimate whorl. Peristome expanded, not reflected. Parietal tooth elevated straight, nearly reaching parietal callus. Parietal wall with a low but pointed tooth that is positioned slightly upper (more apically) compared to where the parietal tooth points.
</paragraph>
<paragraph id="8BAE36C12314266AAE1E378723E2FB92" blockId="17.[113,752,1021,1077]" pageId="17" pageNumber="434">
<emphasis id="B965EAD32314266AAE1E378722A5FBB2" bold="true" box="[113,386,1021,1045]" pageId="17" pageNumber="434">Measurements (in mm).</emphasis>
SH = 1.02–1.21, SW = 1.03–1.19 (n = 6).
</paragraph>
</subSubSection>
<subSubSection id="C30B654A2314266AAE1E302723D8FA92" pageId="17" pageNumber="434" type="diagnosis">
<paragraph id="8BAE36C12314266AAE1E302723D8FA92" blockId="17.[112,754,1117,1333]" pageId="17" pageNumber="434">
<emphasis id="B965EAD32314266AAE1E302722A6FBD2" bold="true" box="[113,385,1117,1141]" pageId="17" pageNumber="434">Differential diagnosis.</emphasis>
Among the
<taxonomicName id="4C114D422314266AAC76302721C8FBD2" box="[537,751,1117,1141]" pageId="17" pageNumber="434">
<emphasis id="B965EAD32314266AAC76302721B5FBD2" box="[537,658,1117,1141]" italics="true" pageId="17" pageNumber="434">Clostophis</emphasis>
species
</taxonomicName>
possessing two parietal teeth,
<taxonomicName id="4C114D422314266AAFA03007229DFB12" authority="PallGergely &amp; Vermeulen, 2020" authorityName="PallGergely &amp; Vermeulen" authorityYear="2020" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="434" phylum="Mollusca" rank="species" species="incurvus">
<emphasis id="B965EAD32314266AAFA030072196FB32" box="[463,689,1149,1173]" italics="true" pageId="17" pageNumber="434">Clostophis incurvus</emphasis>
PállGergely &amp; Vermeulen, 2020
</taxonomicName>
has a conical shell with a narrower umbilicus, and
<taxonomicName id="4C114D422314266AAF0030C7220BFB52" authority="Pall-Gergely &amp; Hunyadi, 2020" authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2020" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="434" phylum="Mollusca" rank="species" species="platytrochus">
<emphasis id="B965EAD32314266AAF0030C72142FB72" box="[367,613,1213,1237]" italics="true" pageId="17" pageNumber="434">Clostophis platytrochus</emphasis>
<bibRefCitation id="EF804B302314266AAC0430C7220BFB52" author="Pall-Gergely B &amp; Hunyadi A &amp; Grego J &amp; Reischutz A &amp; Buczko K &amp; Vermeulen JJ" pageId="17" pageNumber="434" pagination="350 - 368" refId="ref6941" refString="Pall-Gergely B, Hunyadi A, Grego J, Reischutz A, Buczko K &amp; Vermeulen JJ (2020) Clostophis Benson, 1860, is not a monotypic diplommatinid but a speciose hypselostomatid (Gastropoda: Eupulmonata), with descriptions of six new species. Raffles Bulletin of Zoology, 68: 350 - 368." type="journal article" year="2020">Páll-Gergely &amp; Hunyadi, 2020</bibRefCitation>
</taxonomicName>
also has a more conical shell and a more pronounced oblique aperture with a more basally-situated palatal tooth.
</paragraph>
</subSubSection>
<subSubSection id="C30B654A2314266AAE1E312722A4FA12" pageId="17" pageNumber="434" type="etymology">
<paragraph id="8BAE36C12314266AAE1E312722A4FA12" blockId="17.[113,753,1373,1461]" pageId="17" pageNumber="434">
<emphasis id="B965EAD32314266AAE1E312723D1FAD2" bold="true" box="[113,246,1373,1397]" pageId="17" pageNumber="434">Etymology.</emphasis>
Named after Charybdis, the large whirlpool (or sea monster) of the Greek mythology, referring to the whirlpool-like umbilicus.
</paragraph>
</subSubSection>
</treatment>
</document>
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