500 lines
69 KiB
XML
500 lines
69 KiB
XML
<document id="E02871ED4A61A5836FFDB731A2489A5A" ID-CLB-Dataset="299704" ID-DOI="10.5281/zenodo.11512288" ID-GBIF-Dataset="c67efb5d-a1aa-431a-b0e2-5352ba10f138" ID-ISSN="0068-547X" ID-Zenodo-Dep="11512288" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1717719284002" checkinUser="felipe" docAuthor="Gosliner, Terrence" docDate="2015" docId="03A36E275638EE76FFD9FCFDAE49FCBE" docLanguage="en" docName="ProcCalifAcadSci.62.6.191-205.pdf" docOrigin="Proceedings of the California Academy of Sciences 62 (6)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Philinopsis buntot Gosliner 2015, sp. nov." docType="treatment" docVersion="5" lastPageNumber="197" masterDocId="FF9A165F5639EE70FFA2FFB7AD7BFFAD" masterDocTitle="Three New Species of Aglajid Cephalaspidean Mollusks from the Tropical Indo-Pacific of the Verde Island Passage" masterLastPageNumber="205" masterPageNumber="191" pageNumber="192" updateTime="1720727630445" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="B3689E4A5A11A3BD2EF2E1F4D963B282">
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<mods:title id="F28583254236C68B9BF7723716E258DA">Three New Species of Aglajid Cephalaspidean Mollusks from the Tropical Indo-Pacific of the Verde Island Passage</mods:title>
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<mods:name id="8B942E7CDDC5349A507983E45A634348" type="personal">
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<mods:namePart id="ACCBADC5B482CB78E1616C79955B5891">Gosliner, Terrence</mods:namePart>
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<mods:typeOfResource id="02246CD41A3A4EB75EFC8748B945F9CB">text</mods:typeOfResource>
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<mods:title id="4A86B367F98BE7A3EDAE04790DB0684D">Proceedings of the California Academy of Sciences</mods:title>
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</mods:titleInfo>
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<mods:part id="4F736BE3D3E01ADC486882EEDE898E71">
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<mods:date id="2159225C235D0B6158D3A8D57C48D625">2015</mods:date>
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<mods:number id="79EA9736302D5175D487807C4BA5F65F">2015-04-15</mods:number>
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<mods:number id="9E801009F49DFC8D9B63DED505FB35C6">62</mods:number>
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<mods:number id="499D8E0CE60105EA1876A479B90E8FE2">6</mods:number>
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<mods:identifier id="B7573D2CE2BA50A59031720095E51751" type="CLB-Dataset">299704</mods:identifier>
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<mods:identifier id="F7D8DC96EA0CD15DF40DB061B5B4D727" type="DOI">10.5281/zenodo.11512288</mods:identifier>
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<treatment id="03A36E275638EE76FFD9FCFDAE49FCBE" ID-DOI="http://doi.org/10.5281/zenodo.12727287" ID-GBIF-Taxon="235040539" ID-Zenodo-Dep="12727287" LSID="urn:lsid:plazi:treatment:03A36E275638EE76FFD9FCFDAE49FCBE" httpUri="http://treatment.plazi.org/id/03A36E275638EE76FFD9FCFDAE49FCBE" lastPageId="6" lastPageNumber="197" pageId="1" pageNumber="192">
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<subSubSection id="C3108CBA5638EE71FFD9FCFDAF74FCCE" box="[123,527,842,867]" pageId="1" pageNumber="192" type="nomenclature">
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<paragraph id="8BB5DF315638EE71FFD9FCFDAF74FCCE" blockId="1.[123,527,842,897]" box="[123,527,842,867]" pageId="1" pageNumber="192">
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<heading id="D0FD685D5638EE71FFD9FCFDAF74FCCE" box="[123,527,842,867]" fontSize="10" level="2" pageId="1" pageNumber="192" reason="4">
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<emphasis id="B97E03235638EE71FFD9FCFDAF74FCCE" bold="true" box="[123,527,842,867]" pageId="1" pageNumber="192">
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<taxonomicName id="4C0AA4B25638EE71FFD9FCFDACD1FCCF" ID-CoL="4G6F5" authority="Gosliner" authorityName="Gosliner" authorityYear="2015" box="[123,426,842,866]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="1" pageNumber="192" phylum="Mollusca" rank="species" species="buntot" status="sp. nov.">
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<emphasis id="B97E03235638EE71FFD9FCFDAC3FFCCF" bold="true" box="[123,324,842,866]" italics="true" pageId="1" pageNumber="192">Philinopsis buntot</emphasis>
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Gosliner
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24DBE585638EE71FE1BFCFCAF74FCCE" box="[441,527,843,867]" pageId="1" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3108CBA5638EE71FFD9FCDCAC1AFC2D" box="[123,353,875,897]" pageId="1" pageNumber="192" type="description">
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<paragraph id="8BB5DF315638EE71FFD9FCDCAC1AFC2D" blockId="1.[123,527,842,897]" box="[123,353,875,897]" pageId="1" pageNumber="192">
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<figureCitation id="1331C3B45638EE71FFD9FCDCAC7BFC2D" box="[123,256,875,897]" captionStart="FIGURE" captionStartId="2.[171,183,1160,1179]" captionTargetBox="[168,1128,216,1139]" captionTargetId="figure-322@2.[168,1128,216,1139]" captionTargetPageId="2" captionText="FIGURE 1. Living animals A–B. Philinopsis aliciae sp. nov., holotype, PNM 41061, Mabini, Philippines. C. Philinopsis buntot sp. nov., holotype, PNM 41060, Mabini, Philippines, ventral view showing markings on foot. D. Philinopsis buntotparatype, CASIZ 185942, Tingloy, Philippines. E. Chelidonura alexisi sp. nov., holotype, PNM 41062, Mabini, Philippines." figureDoi="http://doi.org/10.5281/zenodo.11512292" httpUri="https://zenodo.org/record/11512292/files/figure.png" pageId="1" pageNumber="192">Figures 1C–E</figureCitation>
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,
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<figureCitation id="1331C3B45638EE71FEAFFCDCAC51FC2D" box="[269,298,875,896]" captionStart="FIGURE" captionStartId="3.[171,183,1395,1414]" captionTargetBox="[163,1123,216,1378]" captionTargetId="figure-184@3.[163,1123,216,1378]" captionTargetPageId="3" captionText="FIGURE 2. Shells.A. Philinopsis buntot. sp. nov., paratype, CASIZ 185942, Tingloy, Philippines. B. Philinopsis aliciae sp. nov., holotype, PNM 41061, Mabini, Philippines. C. Chelidonura alexisi sp. nov., holotype, PNM 41062, Mabini, Philippines." figureDoi="http://doi.org/10.5281/zenodo.11512294" httpUri="https://zenodo.org/record/11512294/files/figure.png" pageId="1" pageNumber="192">2A</figureCitation>
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,
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<figureCitation id="1331C3B45638EE71FE94FCDCAC39FC2D" box="[310,322,875,896]" captionStart="FIGURE" captionStartId="4.[555,567,1152,1171]" captionTargetBox="[504,1176,216,1124]" captionTargetId="figure-521@4.[504,1176,216,1141]" captionText="FIGURE 3. Internal anatomy, Philinopsis buntot. sp. nov., paratype, CASIZ 185942, Tingloy, Philippines. A. Central nervous system c-cerebral ganglion, gg–genital ganglion, on–osphradial nerve, pe–pedal ganglion, pl–pleural ganglion, sp–supraintestinal ganglion, su–subintestinal ganglion, v–visceral ganglion, scale = 1 mm. B. Penis, p–penial papilla, pr–prostate, scale = 0.75 mm. C. Posterior reproductive organs, al–albumen gland, am–ampulla, bc–bursa copulatrix, me–membrane gland, mu1–mucous gland large, mu2–mucous gland small, rs–receptaculum seminis,vg–vestibular gland, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12727283" httpUri="https://zenodo.org/record/12727283/files/figure.png" pageId="1" pageNumber="192">3</figureCitation>
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, 4.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3108CBA5638EE71FF09FC2AADC1FB75" pageId="1" pageNumber="192" type="materials_examined">
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<paragraph id="8BB5DF315638EE71FF09FC2AAE53FB3A" blockId="1.[123,1174,925,1725]" pageId="1" pageNumber="192">
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.—
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:
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<specimenCode id="DBAC774A5638EE71FD94FC2AAFC2FC18" box="[566,697,925,949]" collectionCode="PNM" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="1" pageNumber="192">PNM 41060</specimenCode>
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,
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depth
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</elevation>
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, subsampled for molecular study,
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<location id="8ED589EA5638EE71FFD9FC0AAC62FC78" LSID="urn:lsid:plazi:treatment:03A36E275638EE76FFD9FCFDAE49FCBE:8ED589EA5638EE71FFD9FC0AAC62FC78" box="[123,281,957,981]" country="Philippines" county="Luzon" latitude="-13.75994" longLatPrecision="1" longitude="120.83036" municipality="Mabini" name="Anilao Harbor" pageId="1" pageNumber="192" stateProvince="Batangas">Anilao Harbor</location>
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,
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<collectingMunicipality id="6BD1454B5638EE71FE8EFC0AAC02FC78" box="[300,377,957,981]" pageId="1" pageNumber="192">Mabini</collectingMunicipality>
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,
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<collectingRegion id="49CE11D35638EE71FE2EFC09AF23FC78" box="[396,600,957,982]" country="Philippines" name="Batangas" pageId="1" pageNumber="192">Batangas Province</collectingRegion>
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,
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<collectingCounty id="62D4A7BD5638EE71FDC9FC09AFD4FC78" box="[619,687,958,981]" pageId="1" pageNumber="192">Luzon</collectingCounty>
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,
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<collectingCountry id="F31D9FA15638EE71FD63FC0AAE42FC78" box="[705,825,957,981]" name="Philippines" pageId="1" pageNumber="192">Philippines</collectingCountry>
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,
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,
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,
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<collectingDate id="EFF000195638EE71FBD9FC0AAD95FC5B" pageId="1" pageNumber="192" value="2011-04-30">30 April 2011</collectingDate>
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</date>
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,
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<collectorName id="26FFBAE75638EE71FF59FC69AC24FC58" box="[251,351,990,1013]" pageId="1" pageNumber="192">E. Jessup</collectorName>
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</materialsCitation>
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.
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<materialsCitation id="3B62D56C5638EE71FEC9FC69AE83FBBB" ID-GBIF-Occurrence="4905161302" collectingDate="2011-05-24" collectionCode="CASIZ" country="Philippines" county="Luzon" elevation="5" latitude="-13.75994" location="Anilao Harbor" longLatPrecision="1" longitude="120.83036" municipality="Mabini" pageId="1" pageNumber="192" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
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:
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, 185780,
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,
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depth
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</elevation>
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,
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<location id="8ED589EA5638EE71FC0FFC69A93DFC5B" LSID="urn:lsid:plazi:treatment:03A36E275638EE76FFD9FCFDAE49FCBE:8ED589EA5638EE71FC0FFC69A93DFC5B" box="[941,1094,990,1014]" country="Philippines" county="Luzon" latitude="-13.75994" longLatPrecision="1" longitude="120.83036" municipality="Mabini" name="Anilao Harbor" pageId="1" pageNumber="192" stateProvince="Batangas">Anilao Harbor</location>
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,
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<collectingMunicipality id="6BD1454B5638EE71FBF6FC69ADEBFBBB" pageId="1" pageNumber="192">Mabini</collectingMunicipality>
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,
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<collectingRegion id="49CE11D35638EE71FF3FFC49AC1FFBBB" box="[157,356,1022,1046]" country="Philippines" name="Batangas" pageId="1" pageNumber="192">Batangas Province</collectingRegion>
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,
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<collectingCounty id="62D4A7BD5638EE71FED3FC49ACCEFBB8" box="[369,437,1022,1045]" pageId="1" pageNumber="192">Luzon</collectingCounty>
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,
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<collectingCountry id="F31D9FA15638EE71FE60FC49AF41FBBB" box="[450,570,1022,1046]" name="Philippines" pageId="1" pageNumber="192">Philippines</collectingCountry>
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,
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,
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,
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<collectingDate id="EFF000195638EE71FCC9FC49AE83FBBB" box="[875,1016,1022,1046]" pageId="1" pageNumber="192" value="2011-05-24">24 May 2011</collectingDate>
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</date>
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</materialsCitation>
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,
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<collectionCode id="ED1B47F45638EE71FBA7FC49A968FBB8" box="[1029,1043,1022,1045]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="192">T</collectionCode>
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||
. M. Gosliner.
|
||
<materialsCitation id="3B62D56C5638EE71FF00FBA9AEFDFBFA" ID-GBIF-Occurrence="4905161306" collectingDate="2011-05-19" collectionCode="CASIZ" country="Philippines" county="Luzon" elevation="7" latitude="-13.68433" location="Cemetery Beach" longLatPrecision="1" longitude="120.82993" municipality="Tingloy" pageId="1" pageNumber="192" specimenCode="CASIZ 185942" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
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<specimenCode id="DBAC774A5638EE71FF00FBA9AC37FB9B" box="[162,332,1054,1078]" collectionCode="CASIZ" pageId="1" pageNumber="192">CASIZ 185942</specimenCode>
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,
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, dissected,
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|
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depth
|
||
</elevation>
|
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,
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<location id="8ED589EA5638EE71FCA4FBA9AEC6FB9B" LSID="urn:lsid:plazi:treatment:03A36E275638EE76FFD9FCFDAE49FCBE:8ED589EA5638EE71FCA4FBA9AEC6FB9B" box="[774,957,1054,1078]" country="Philippines" county="Luzon" latitude="-13.68433" longLatPrecision="1" longitude="120.82993" municipality="Tingloy" name="Cemetery Beach" pageId="1" pageNumber="192" stateProvince="Batangas">Cemetery Beach</location>
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,
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||
<collectingMunicipality id="6BD1454B5638EE71FC6DFBA9A95AFB9B" box="[975,1057,1054,1078]" pageId="1" pageNumber="192">Tingloy</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49CE11D35638EE71FB96FBA8ADA1FBFA" country="Philippines" name="Batangas" pageId="1" pageNumber="192">Batangas Province</collectingRegion>
|
||
,
|
||
<collectingCounty id="62D4A7BD5638EE71FF48FB88AC55FBFB" box="[234,302,1087,1110]" pageId="1" pageNumber="192">Luzon</collectingCounty>
|
||
,
|
||
<collectingCountry id="F31D9FA15638EE71FE9DFB88ACCCFBFA" box="[319,439,1087,1111]" name="Philippines" pageId="1" pageNumber="192">Philippines</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE3EB9F65638EE71FE65FB88AF3EFBFA" box="[455,581,1087,1111]" degrees="13. 68433" direction="south" orientation="latitude" pageId="1" pageNumber="192" precision="1" value="-13.68433">13.68433°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3EB9F65638EE71FDF7FB88AF9AFBFB" box="[597,737,1087,1111]" degrees="120. 82993" direction="east" orientation="longitude" pageId="1" pageNumber="192" precision="1" value="120.82993">120.82993°E</geoCoordinate>
|
||
,
|
||
<date id="FFB4F9F15638EE71FD50FB88AEFDFBFA" box="[754,902,1087,1111]" pageId="1" pageNumber="192" value="2011-05-19">
|
||
<collectingDate id="EFF000195638EE71FD50FB88AEFDFBFA" box="[754,902,1087,1111]" pageId="1" pageNumber="192" value="2011-05-19">19 May 2011</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B62D56C5638EE71FC34FB88A94CFBFA" ID-GBIF-Occurrence="4905161305" box="[918,1079,1087,1111]" collectionCode="T" collectorName="Gosliner" country="Philippines" pageId="1" pageNumber="192" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
|
||
<collectionCode id="ED1B47F45638EE71FC34FB88AEDFFBFB" box="[918,932,1087,1110]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="192">T</collectionCode>
|
||
. M.
|
||
<collectorName id="26FFBAE75638EE71FC7FFB88A94CFBFA" box="[989,1079,1087,1111]" pageId="1" pageNumber="192">Gosliner</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B62D56C5638EE71FBEAFB88AF0EFB3A" ID-GBIF-Occurrence="4905161303" collectingDate="2011-05-19" collectionCode="CASIZ" country="Philippines" county="Luzon" elevation="7" latitude="-13.68433" location="Cemetery Beach" longLatPrecision="1" longitude="120.82993" municipality="Tingloy" pageId="1" pageNumber="192" specimenCode="CASIZ 185777" specimenCount="2" stateProvince="Batangas" typeStatus="paratype">
|
||
<specimenCode id="DBAC774A5638EE71FBEAFB88ADB0FBDA" collectionCode="CASIZ" pageId="1" pageNumber="192">CASIZ 185777</specimenCode>
|
||
,
|
||
<specimenCount id="9D0C14B85638EE71FF7BFBE8AC03FBDA" box="[217,376,1119,1143]" count="2" pageId="1" pageNumber="192" type="generic">two specimens</specimenCount>
|
||
,
|
||
<elevation id="002738025638EE71FE25FBE8AC88FBDA" box="[391,499,1119,1143]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="192" unit="m" value="7.0">
|
||
<quantity id="4CF272D45638EE71FE25FBE8ACCBFBDB" box="[391,432,1119,1142]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="192" unit="m" value="7.0">7 m</quantity>
|
||
depth
|
||
</elevation>
|
||
,
|
||
<location id="8ED589EA5638EE71FDA0FBE8AFCFFBDA" LSID="urn:lsid:plazi:treatment:03A36E275638EE76FFD9FCFDAE49FCBE:8ED589EA5638EE71FDA0FBE8AFCFFBDA" box="[514,692,1119,1143]" country="Philippines" county="Luzon" latitude="-13.68433" longLatPrecision="1" longitude="120.82993" municipality="Tingloy" name="Cemetery Beach" pageId="1" pageNumber="192" stateProvince="Batangas">Cemetery Beach</location>
|
||
,
|
||
<collectingMunicipality id="6BD1454B5638EE71FD60FBE8AE6FFBDA" box="[706,788,1119,1143]" pageId="1" pageNumber="192">Tingloy</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49CE11D35638EE71FC81FBE8AE90FBDA" box="[803,1003,1119,1143]" country="Philippines" name="Batangas" pageId="1" pageNumber="192">Batangas Province</collectingRegion>
|
||
,
|
||
<collectingCounty id="62D4A7BD5638EE71FC58FBE8A945FBDB" box="[1018,1086,1119,1142]" pageId="1" pageNumber="192">Luzon</collectingCounty>
|
||
,
|
||
<collectingCountry id="F31D9FA15638EE71FBEFFBE8ADC8FB3A" name="Philippines" pageId="1" pageNumber="192">Philippines</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE3EB9F65638EE71FF63FBC8AC44FB3A" box="[193,319,1151,1175]" degrees="13. 68433" direction="south" orientation="latitude" pageId="1" pageNumber="192" precision="1" value="-13.68433">13.68433°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3EB9F65638EE71FEEEFBC8ACA3FB3A" box="[332,472,1151,1175]" degrees="120. 82993" direction="east" orientation="longitude" pageId="1" pageNumber="192" precision="1" value="120.82993">120.82993°E</geoCoordinate>
|
||
,
|
||
<date id="FFB4F9F15638EE71FE45FBC8AF0EFB3A" box="[487,629,1151,1176]" pageId="1" pageNumber="192" value="2011-05-19">
|
||
<collectingDate id="EFF000195638EE71FE45FBC8AF0EFB3A" box="[487,629,1151,1176]" pageId="1" pageNumber="192" value="2011-05-19">19 May 2011</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B62D56C5638EE71FD21FB37AE5AFB3A" ID-GBIF-Occurrence="4905161307" box="[643,801,1151,1175]" collectionCode="T" collectorName="Gosliner" country="Philippines" pageId="1" pageNumber="192" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
|
||
<collectionCode id="ED1B47F45638EE71FD21FB37AFEAFB3A" box="[643,657,1152,1175]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="192">T</collectionCode>
|
||
. M.
|
||
<collectorName id="26FFBAE75638EE71FD67FBC8AE5AFB3A" box="[709,801,1151,1175]" pageId="1" pageNumber="192">Gosliner</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB5DF315638EE71FF09FB28ADC1FB75" blockId="1.[123,1174,925,1725]" pageId="1" pageNumber="192">
|
||
<emphasis id="B97E03235638EE71FF09FB28AC89FB1B" bold="true" box="[171,498,1183,1207]" pageId="1" pageNumber="192">
|
||
<smallCapsWord id="8D5349ED5638EE71FF09FB28AC2DFB1B" baselines="1201,1201" box="[171,342,1183,1207]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="1" pageNumber="192">GEOGRAPHICAL</smallCapsWord>
|
||
<smallCapsWord id="8D5349ED5638EE71FEFCFB14AC89FB1B" baselines="1201" box="[350,498,1187,1206]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="1" pageNumber="192">DISTRIBUTION</smallCapsWord>
|
||
</emphasis>
|
||
.— Thus far, known only from southern Luzon Island,
|
||
<collectingCountry id="F31D9FA15638EE71FBEFFB17ADC8FB75" name="Philippines" pageId="1" pageNumber="192">Philippines</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3108CBA5638EE71FF09FB56A921FAF4" pageId="1" pageNumber="192" type="etymology">
|
||
<paragraph id="8BB5DF315638EE71FF09FB56AC81FAB5" blockId="1.[123,1174,925,1725]" pageId="1" pageNumber="192">
|
||
<smallCapsWord id="8D5349ED5638EE71FF09FB56AC4EFB5A" baselines="1267,1266" box="[171,309,1249,1273]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Etymology" pageId="1" pageNumber="192">
|
||
<emphasis id="B97E03235638EE71FF09FB56AC4EFB5A" bold="true" box="[171,309,1249,1273]" pageId="1" pageNumber="192">ETYMOLOGY</emphasis>
|
||
</smallCapsWord>
|
||
.— The name
|
||
<taxonomicName id="4C0AA4B25638EE71FE76FB56AF58FB55" authorityName="Gosliner" authorityYear="2015" box="[468,547,1248,1273]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="1" pageNumber="192" phylum="Mollusca" rank="species" species="buntot">“buntot</taxonomicName>
|
||
” is the Tagalog word for tail, referring to the distinctive posterior appendage of this species.
|
||
</paragraph>
|
||
<paragraph id="8BB5DF315638EE71FF09FA96A921FAF4" blockId="1.[123,1174,925,1725]" pageId="1" pageNumber="192">
|
||
<emphasis id="B97E03235638EE71FF09FA96AC0EFA95" bold="true" box="[171,373,1313,1337]" pageId="1" pageNumber="192">
|
||
<smallCapsWord id="8D5349ED5638EE71FF09FA96AC6AFA95" baselines="1331,1331" box="[171,273,1313,1337]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Natural" pageId="1" pageNumber="192">NATURAL</smallCapsWord>
|
||
<smallCapsWord id="8D5349ED5638EE71FEB8FA92AC0EFA95" baselines="1331" box="[282,373,1317,1336]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="history" pageId="1" pageNumber="192">HISTORY</smallCapsWord>
|
||
</emphasis>
|
||
.— This species is found crawling on clean sand where it is nocturnally active. It has not been observed diurnally and may remain buried in the sand during the day.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3108CBA5638EE74FF09FAD5AEB6F91E" lastPageId="4" lastPageNumber="195" pageId="1" pageNumber="192" type="description">
|
||
<paragraph id="8BB5DF315638EE71FF09FAD5AE31F9B6" blockId="1.[123,1174,925,1725]" pageId="1" pageNumber="192">
|
||
<smallCapsWord id="8D5349ED5638EE71FF09FAD5AC41FAD5" baselines="1396,1395" box="[171,314,1378,1402]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Description" pageId="1" pageNumber="192">
|
||
<emphasis id="B97E03235638EE71FF09FAD5AC41FAD5" bold="true" box="[171,314,1378,1402]" pageId="1" pageNumber="192">DESCRIPTION</emphasis>
|
||
</smallCapsWord>
|
||
.—
|
||
<emphasis id="B97E03235638EE71FEC0FAD5AF3CFAD7" box="[354,583,1378,1402]" italics="true" pageId="1" pageNumber="192">External morphology</emphasis>
|
||
: The living animals are
|
||
<quantity id="4CF272D45638EE71FCF0FAD5AEB4FAD4" box="[850,975,1378,1402]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7.0" pageId="1" pageNumber="192" unit="mm" value="9.5" valueMax="12.0" valueMin="7.0">7 to 12 mm</quantity>
|
||
in length and
|
||
<quantity id="4CF272D45638EE71FBCFFAD5ADDFFA37" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="1" pageNumber="192" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6 mm</quantity>
|
||
wide. The general body color of the living animal (
|
||
<figureCitation id="1331C3B45638EE71FD68FA35AE39FA37" box="[714,834,1410,1434]" captionStart="FIGURE" captionStartId="2.[171,183,1160,1179]" captionTargetBox="[168,1128,216,1139]" captionTargetId="figure-322@2.[168,1128,216,1139]" captionTargetPageId="2" captionText="FIGURE 1. Living animals A–B. Philinopsis aliciae sp. nov., holotype, PNM 41061, Mabini, Philippines. C. Philinopsis buntot sp. nov., holotype, PNM 41060, Mabini, Philippines, ventral view showing markings on foot. D. Philinopsis buntotparatype, CASIZ 185942, Tingloy, Philippines. E. Chelidonura alexisi sp. nov., holotype, PNM 41062, Mabini, Philippines." figureDoi="http://doi.org/10.5281/zenodo.11512292" httpUri="https://zenodo.org/record/11512292/files/figure.png" pageId="1" pageNumber="192">Figs. 1C–E</figureCitation>
|
||
) is translucent white with varying amounts of reticulated brown pigment on the dorsal surface. In specimens with sparse brown pigment patches of opaque white may also be present. The brown pigment frequently surrounds small yellow spots on the dorsal surface. Larger yellow spots are evident along the margins of the cephalic and posterior shields and on the margins of the parapodia.
|
||
</paragraph>
|
||
<paragraph id="8BB5DF315638EE72FF09F993AF1CFA20" blockId="1.[123,1174,925,1725]" lastBlockId="2.[122,1174,1269,1709]" lastPageId="2" lastPageNumber="193" pageId="1" pageNumber="192">Living animals are elongate, and relatively slender. The anterior end of the cephalic shield is blunt and quadrangular. The cephalic shield is roughly rectangular and terminates posteriorly with a rounded edge. No upturned papilla was observed on the end of the cephalic shield. The posterior shield is slightly rounded anteriorly and terminates in a medial elongate, digitiform posterior projection that is well-elevated from the base of the shield. This appendage is rounded at the apex and is often held upright in actively crawling individuals. The posterior end of the posterior shield is much lower that the area with the projection and two relatively flat lateral posterior lobes are short and simply rounded. The parapodia are relatively short, leaving most of the cephalic and posterior shields visible. The gill is simply plicate consisting of 12 primary folds and is situated ventrally on the right posterior end of the animal.</paragraph>
|
||
<caption id="DF758FB9563BEE72FF09FB3FA9EEFB63" ID-DOI="http://doi.org/10.5281/zenodo.11512292" ID-Zenodo-Dep="11512292" httpUri="https://zenodo.org/record/11512292/files/figure.png" pageId="2" pageNumber="193" startId="2.[171,183,1160,1179]" targetBox="[168,1128,216,1139]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8BB5DF31563BEE72FF09FB3FA9EEFB63" blockId="2.[123,1174,1159,1230]" pageId="2" pageNumber="193">
|
||
<smallCapsWord id="8D5349ED563BEE72FF09FB3FAD93FB34" baselines="1174,1173" box="[171,232,1160,1179]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="2" pageNumber="193">FIGURE</smallCapsWord>
|
||
1. Living animals A–B.
|
||
<taxonomicName id="4C0AA4B2563BEE72FE14FB30AF2EFB37" authority="Gosliner, 2015" authorityName="Gosliner" authorityYear="2015" box="[438,597,1159,1178]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="2" pageNumber="193" phylum="Mollusca" rank="species" species="aliciae" status="sp. nov.">
|
||
<emphasis id="B97E0323563BEE72FE14FB30AF2EFB37" box="[438,597,1159,1178]" italics="true" pageId="2" pageNumber="193">Philinopsis aliciae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DBE58563BEE72FDF8FB3FAFE7FB36" box="[602,668,1160,1179]" pageId="2" pageNumber="193" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, PNM 41061, Mabini, Philippines. C.
|
||
<taxonomicName id="4C0AA4B2563BEE72FB97FB30ADC9FB19" authority="Gosliner, 2015" authorityName="Gosliner" authorityYear="2015" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="2" pageNumber="193" phylum="Mollusca" rank="species" species="buntot" status="sp. nov.">
|
||
<emphasis id="B97E0323563BEE72FB97FB30ADC9FB19" italics="true" pageId="2" pageNumber="193">Philinopsis buntot</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DBE58563BEE72FF15FB16AD81FB19" box="[183,250,1185,1204]" pageId="2" pageNumber="193" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, PNM 41060, Mabini, Philippines, ventral view showing markings on foot. D.
|
||
<taxonomicName id="4C0AA4B2563BEE72FC50FB16AC77FB60" authority=", CASIZ" authorityName="CASIZ" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="2" pageNumber="193" phylum="Mollusca" rank="species" species="buntotparatype">
|
||
<emphasis id="B97E0323563BEE72FC50FB16ADBCFB63" italics="true" pageId="2" pageNumber="193">Philinopsis buntotparatype</emphasis>
|
||
, CASIZ
|
||
</taxonomicName>
|
||
185942, Tingloy, Philippines. E.
|
||
<taxonomicName id="4C0AA4B2563BEE72FD8AFB0DAFAAFB60" authority="Gosliner, 2015" authorityName="Gosliner" authorityYear="2015" box="[552,721,1210,1229]" class="Gastropoda" family="Aglajidae" genus="Chelidonura" kingdom="Animalia" order="Cephalaspidea" pageId="2" pageNumber="193" phylum="Mollusca" rank="species" species="alexisi" status="sp. nov.">
|
||
<emphasis id="B97E0323563BEE72FD8AFB0DAFAAFB60" box="[552,721,1210,1229]" italics="true" pageId="2" pageNumber="193">Chelidonura alexisi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DBE58563BEE72FD77FB0CAE62FB63" box="[725,793,1211,1230]" pageId="2" pageNumber="193" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, PNM 41062, Mabini, Philippines.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB5DF31563BEE72FF09FA22AF9DF9A0" blockId="2.[122,1174,1269,1709]" pageId="2" pageNumber="193">
|
||
<emphasis id="B97E0323563BEE72FF09FA22AD9BFA00" box="[171,224,1429,1453]" italics="true" pageId="2" pageNumber="193">Shell</emphasis>
|
||
(
|
||
<figureCitation id="1331C3B4563BEE72FF50FA22AC30FA03" box="[242,331,1429,1454]" captionStart="FIGURE" captionStartId="3.[171,183,1395,1414]" captionTargetBox="[163,1123,216,1378]" captionTargetId="figure-184@3.[163,1123,216,1378]" captionTargetPageId="3" captionText="FIGURE 2. Shells.A. Philinopsis buntot. sp. nov., paratype, CASIZ 185942, Tingloy, Philippines. B. Philinopsis aliciae sp. nov., holotype, PNM 41061, Mabini, Philippines. C. Chelidonura alexisi sp. nov., holotype, PNM 41062, Mabini, Philippines." figureDoi="http://doi.org/10.5281/zenodo.11512294" httpUri="https://zenodo.org/record/11512294/files/figure.png" pageId="2" pageNumber="193">Fig. 2A</figureCitation>
|
||
): The shell is relatively thickly calcified and consists of a narrow band that occupies the posterior extreme of the animal. There is a membranous periostracum that is slightly more extensive anteriorly than the calcified portion. The area at the base of the shell near the protoconch is more thickly calcified that the rest of the shell.
|
||
</paragraph>
|
||
<paragraph id="8BB5DF31563BEE73FF09F9A1AEA2F9A7" blockId="2.[122,1174,1269,1709]" lastBlockId="3.[123,1174,1491,1707]" lastPageId="3" lastPageNumber="194" pageId="2" pageNumber="193">
|
||
<emphasis id="B97E0323563BEE72FF09F9A1AC26F980" box="[171,349,1558,1581]" italics="true" pageId="2" pageNumber="193">Digestive system</emphasis>
|
||
: The buccal mass is large, highly muscularized and slightly elongate posteriorly and occupies the body cavity for the entire length of the cephalic shield. The buccal bulb entirely lacks any vestige of a radula. There is a large ventral oral gland and a smaller dorsal oral gland. At the posterior end of the buccal mass, near the junction with the crop, is a pair of elongate salivary glands. The crop is large and saccate, wider than the buccal bulb. The crop narrows posteriorly and enters the digestive gland. The intestine emerges from the right side of the digestive gland and terminates near the posterior end of the body near the base of the gill.
|
||
</paragraph>
|
||
<caption id="DF758FB9563AEE73FF09FAC4ADD6FA17" ID-DOI="http://doi.org/10.5281/zenodo.11512294" ID-Zenodo-Dep="11512294" httpUri="https://zenodo.org/record/11512294/files/figure.png" pageId="3" pageNumber="194" startId="3.[171,183,1395,1414]" targetBox="[163,1123,216,1378]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BB5DF31563AEE73FF09FAC4ADD6FA17" blockId="3.[123,1173,1395,1466]" pageId="3" pageNumber="194">
|
||
<smallCapsWord id="8D5349ED563AEE73FF09FAC4AD93FA28" baselines="1409,1409" box="[171,232,1395,1414]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="3" pageNumber="194">FIGURE</smallCapsWord>
|
||
2. Shells.A.
|
||
<emphasis id="B97E0323563AEE73FEFAFAC4AC83FA2B" box="[344,504,1395,1414]" italics="true" pageId="3" pageNumber="194">
|
||
<taxonomicName id="4C0AA4B2563AEE73FEFAFAC4AC8FFA2B" authorityName="Gosliner" authorityYear="2015" box="[344,500,1395,1414]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="3" pageNumber="194" phylum="Mollusca" rank="species" species="buntot">Philinopsis buntot</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
sp. nov., paratype, CASIZ 185942, Tingloy, Philippines. B.
|
||
<taxonomicName id="4C0AA4B2563AEE73FC54FAC4A9EEFA2B" authority="Gosliner, 2015" authorityName="Gosliner" authorityYear="2015" box="[1014,1173,1395,1414]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="3" pageNumber="194" phylum="Mollusca" rank="species" species="aliciae" status="sp. nov.">
|
||
<emphasis id="B97E0323563AEE73FC54FAC4A9EEFA2B" box="[1014,1173,1395,1414]" italics="true" pageId="3" pageNumber="194">Philinopsis aliciae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DBE58563AEE73FFD9FA3AADC6FA0D" box="[123,189,1421,1440]" pageId="3" pageNumber="194" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, PNM 41061, Mabini, Philippines. C.
|
||
<taxonomicName id="4C0AA4B2563AEE73FDF4FA3BAF85FA32" authority="Gosliner, 2015" authorityName="Gosliner" authorityYear="2015" box="[598,766,1420,1439]" class="Gastropoda" family="Aglajidae" genus="Chelidonura" kingdom="Animalia" order="Cephalaspidea" pageId="3" pageNumber="194" phylum="Mollusca" rank="species" species="alexisi" status="sp. nov.">
|
||
<emphasis id="B97E0323563AEE73FDF4FA3BAF85FA32" box="[598,766,1420,1439]" italics="true" pageId="3" pageNumber="194">Chelidonura alexisi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DBE58563AEE73FCA0FA3AAE3DFA0D" box="[770,838,1421,1440]" pageId="3" pageNumber="194" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, PNM 41062, Mabini, Philippines.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB5DF31563AEE74FF09F9A4AC50FD1E" blockId="3.[123,1174,1491,1707]" lastBlockId="4.[123,478,219,1395]" lastPageId="4" lastPageNumber="195" pageId="3" pageNumber="194">
|
||
<emphasis id="B97E0323563AEE73FF09F9A4ACD4F987" box="[171,431,1555,1579]" italics="true" pageId="3" pageNumber="194">Central Nervous System</emphasis>
|
||
(
|
||
<figureCitation id="1331C3B4563AEE73FE1DF9A4AF6DF986" box="[447,534,1555,1579]" captionStart="FIGURE" captionStartId="4.[555,567,1152,1171]" captionTargetBox="[504,1176,216,1124]" captionTargetId="figure-521@4.[504,1176,216,1141]" captionText="FIGURE 3. Internal anatomy, Philinopsis buntot. sp. nov., paratype, CASIZ 185942, Tingloy, Philippines. A. Central nervous system c-cerebral ganglion, gg–genital ganglion, on–osphradial nerve, pe–pedal ganglion, pl–pleural ganglion, sp–supraintestinal ganglion, su–subintestinal ganglion, v–visceral ganglion, scale = 1 mm. B. Penis, p–penial papilla, pr–prostate, scale = 0.75 mm. C. Posterior reproductive organs, al–albumen gland, am–ampulla, bc–bursa copulatrix, me–membrane gland, mu1–mucous gland large, mu2–mucous gland small, rs–receptaculum seminis,vg–vestibular gland, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12727283" httpUri="https://zenodo.org/record/12727283/files/figure.png" pageId="3" pageNumber="194">Fig. 3A</figureCitation>
|
||
): The circumesophageal nerve ring consists of paired cerebral, pedal and pleural ganglia and a single supraintestinal ganglion on the right side. The cerebral and pedal commissures are both elongate with well-separated respective ganglia. Immediately adjacent and posterior to the right pleural ganglion is the supraintestinal ganglion. From its posterior end is the right branch of the visceral loop and the osphradial nerve. The two lateral branches of the visceral loop join posteriorly at the posterior ganglia. The left visceral loop enters the subintestinal ganglion, whereas the right lateral nerve enters the visceral ganglion. The visceral ganglion is larger than the subintestinal ganglion. From the visceral ganglion is the genital nerve, which has a distinct genital ganglion. The buccal ganglia can be seen near the middle of the ventral surface of the buccal mas and are somewhat separated from each other.
|
||
</paragraph>
|
||
<paragraph id="8BB5DF31563DEE74FF09FD0CAEB6F91E" blockId="4.[123,478,219,1395]" lastBlockId="4.[123,1174,1403,1715]" pageId="4" pageNumber="195">
|
||
<emphasis id="B97E0323563DEE74FF09FD0CACEAFD7F" box="[171,401,699,723]" italics="true" pageId="4" pageNumber="195">Reproductive System</emphasis>
|
||
(
|
||
<figureCitation id="1331C3B4563DEE74FE0AFD0CADC2FD5E" captionStart="FIGURE" captionStartId="4.[555,567,1152,1171]" captionTargetBox="[504,1176,216,1124]" captionTargetId="figure-521@4.[504,1176,216,1141]" captionText="FIGURE 3. Internal anatomy, Philinopsis buntot. sp. nov., paratype, CASIZ 185942, Tingloy, Philippines. A. Central nervous system c-cerebral ganglion, gg–genital ganglion, on–osphradial nerve, pe–pedal ganglion, pl–pleural ganglion, sp–supraintestinal ganglion, su–subintestinal ganglion, v–visceral ganglion, scale = 1 mm. B. Penis, p–penial papilla, pr–prostate, scale = 0.75 mm. C. Posterior reproductive organs, al–albumen gland, am–ampulla, bc–bursa copulatrix, me–membrane gland, mu1–mucous gland large, mu2–mucous gland small, rs–receptaculum seminis,vg–vestibular gland, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12727283" httpUri="https://zenodo.org/record/12727283/files/figure.png" pageId="4" pageNumber="195">Figs. 3B–C</figureCitation>
|
||
, 4): The arrangement of reproductive organs is essentially monaulic (as discussed by Gosliner 1994) but with a single branch of the hermaphroditic duct to the albumen and membrane glands (
|
||
<figureCitation id="1331C3B4563DEE74FEB1FC2CAC13FC1E" box="[275,360,923,947]" captionStart="FIGURE" captionStartId="4.[555,567,1152,1171]" captionTargetBox="[504,1176,216,1124]" captionTargetId="figure-521@4.[504,1176,216,1141]" captionText="FIGURE 3. Internal anatomy, Philinopsis buntot. sp. nov., paratype, CASIZ 185942, Tingloy, Philippines. A. Central nervous system c-cerebral ganglion, gg–genital ganglion, on–osphradial nerve, pe–pedal ganglion, pl–pleural ganglion, sp–supraintestinal ganglion, su–subintestinal ganglion, v–visceral ganglion, scale = 1 mm. B. Penis, p–penial papilla, pr–prostate, scale = 0.75 mm. C. Posterior reproductive organs, al–albumen gland, am–ampulla, bc–bursa copulatrix, me–membrane gland, mu1–mucous gland large, mu2–mucous gland small, rs–receptaculum seminis,vg–vestibular gland, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12727283" httpUri="https://zenodo.org/record/12727283/files/figure.png" pageId="4" pageNumber="195">Fig. 3C</figureCitation>
|
||
). From the large ovotestis, which is intermingled with the digestive gland, emerges the convoluted ampulla. The ampulla narrows into the hermaphroditic duct, which curves around the receptaculum seminis and enters the short, coiled albumen and membrane glands by means of a single duct. The larger mucous gland is bilobed with a massive primary lobe and smaller secondary one. The outer margins of both lobes bear a rounded margin with numerous tooth-shaped lobes. The hermaphroditic duct then joins the duct of the receptaculum seminis and continues to the genital atrium, where it joins the duct of the bursa copulatrix. The bursa is large and spherical. Its duct is narrow where it joins the bursa and widens until its widest portion at the genital atrium. The genital atrium is large and muscular. The atrium also has a lobate vestibular gland situated on its ventral surface. From the genital atrium the open, ciliated sperm groove leads to the cephalic penis. The penis (
|
||
<figureCitation id="1331C3B4563DEE74FDDCF9ACAFA7F99E" box="[638,732,1563,1587]" captionStart="FIGURE" captionStartId="4.[555,567,1152,1171]" captionTargetBox="[504,1176,216,1124]" captionTargetId="figure-521@4.[504,1176,216,1141]" captionText="FIGURE 3. Internal anatomy, Philinopsis buntot. sp. nov., paratype, CASIZ 185942, Tingloy, Philippines. A. Central nervous system c-cerebral ganglion, gg–genital ganglion, on–osphradial nerve, pe–pedal ganglion, pl–pleural ganglion, sp–supraintestinal ganglion, su–subintestinal ganglion, v–visceral ganglion, scale = 1 mm. B. Penis, p–penial papilla, pr–prostate, scale = 0.75 mm. C. Posterior reproductive organs, al–albumen gland, am–ampulla, bc–bursa copulatrix, me–membrane gland, mu1–mucous gland large, mu2–mucous gland small, rs–receptaculum seminis,vg–vestibular gland, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12727283" httpUri="https://zenodo.org/record/12727283/files/figure.png" pageId="4" pageNumber="195">Figs. 3B</figureCitation>
|
||
, 4) consists of a penial sac and a thick, curved, irregularly-shaped prostate gland that is joined to the penial sac by a narrow duct. Within the penial sac is an elongate penial papilla (
|
||
<figureCitation id="1331C3B4563DEE74FDF1F9ECAFD4F9DE" box="[595,687,1627,1651]" captionStart="FIGURE" captionStartId="4.[555,567,1152,1171]" captionTargetBox="[504,1176,216,1124]" captionTargetId="figure-521@4.[504,1176,216,1141]" captionText="FIGURE 3. Internal anatomy, Philinopsis buntot. sp. nov., paratype, CASIZ 185942, Tingloy, Philippines. A. Central nervous system c-cerebral ganglion, gg–genital ganglion, on–osphradial nerve, pe–pedal ganglion, pl–pleural ganglion, sp–supraintestinal ganglion, su–subintestinal ganglion, v–visceral ganglion, scale = 1 mm. B. Penis, p–penial papilla, pr–prostate, scale = 0.75 mm. C. Posterior reproductive organs, al–albumen gland, am–ampulla, bc–bursa copulatrix, me–membrane gland, mu1–mucous gland large, mu2–mucous gland small, rs–receptaculum seminis,vg–vestibular gland, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12727283" httpUri="https://zenodo.org/record/12727283/files/figure.png" pageId="4" pageNumber="195">Figs. 3B</figureCitation>
|
||
, 4A). The papilla has a bilobed apex with an expanded outer collar. The collar (Figs. 4A–C) is ornamented with undivided and bifid penial spines as are the inner (Fig. 4D) and outer (Fig. 4E) lobes of the penial papilla.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF758FB9563DEE74FD89FB37AFD6FACD" ID-DOI="http://doi.org/10.5281/zenodo.12727283" ID-Zenodo-Dep="12727283" httpUri="https://zenodo.org/record/12727283/files/figure.png" pageId="4" pageNumber="195" startId="4.[555,567,1152,1171]" targetBox="[504,1176,216,1124]" targetPageId="4">
|
||
<paragraph id="8BB5DF31563DEE74FD89FB37AFD6FACD" blockId="4.[507,1174,1152,1376]" pageId="4" pageNumber="195">
|
||
<subScript id="178EDD74563DEE74FD89FB37A9EDFB3E" attach="right" box="[555,1174,1152,1171]" fontSize="8" pageId="4" pageNumber="195">FIGURE 3. Internal anatomy, Philinopsis buntot. sp. nov., paratype,</subScript>
|
||
<superScript id="7C7F7279563DEE74FE59FB2EA9EEFB01" attach="right" box="[507,1173,1177,1197]" fontSize="8" pageId="4" pageNumber="195">CASIZ 185942, Tingloy, Philippines. A. Central nervous system c-cerebral</superScript>
|
||
<superScript id="7C7F7279563DEE74FE59FB04A9EDFB6B" attach="right" box="[507,1174,1203,1222]" fontSize="8" pageId="4" pageNumber="195">ganglion, gg–genital ganglion, on–osphradial nerve, pe–pedal ganglion,</superScript>
|
||
pl–pleural ganglion, sp–supraintestinal ganglion, su–subintestinal ganglion, v–visceral ganglion, scale = 1 mm. B. Penis, p–penial papilla, pr–prostate, scale = 0.75 mm. C. Posterior reproductive organs, al–albumen gland, am–ampulla, bc–bursa copulatrix, me–membrane gland, mu1–mucous gland large, mu2–mucous gland small, rs–receptaculum seminis,vg–vestibular gland, scale = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF758FB9563CEE75FF09FBD3AEECFB06" ID-DOI="http://doi.org/10.5281/zenodo.11512296" ID-Zenodo-Dep="11512296" httpUri="https://zenodo.org/record/11512296/files/figure.png" pageId="5" pageNumber="196" startId="5.[171,183,1124,1143]" targetBox="[176,1120,216,1111]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8BB5DF31563CEE75FF09FBD3AEECFB06" blockId="5.[123,1174,1124,1195]" pageId="5" pageNumber="196">
|
||
<smallCapsWord id="8D5349ED563CEE75FF09FBD3AD93FBDB" baselines="1138,1138" box="[171,232,1124,1143]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="5" pageNumber="196">FIGURE</smallCapsWord>
|
||
4.
|
||
<emphasis id="B97E0323563CEE75FEA0FBD3ACDFFBDA" box="[258,420,1124,1143]" italics="true" pageId="5" pageNumber="196">
|
||
<taxonomicName id="4C0AA4B2563CEE75FEA0FBD3ACE4FBDA" authorityName="Gosliner" authorityYear="2015" box="[258,415,1124,1143]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="5" pageNumber="196" phylum="Mollusca" rank="species" species="buntot">Philinopsis buntot</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
sp. nov., paratype, CASIZ 185942, Tingloy, Philippines, scanning electron micrographs of penis. A. Entire penial papilla, c-collar. i-inner lobe, o-outer lobe, scale = 150 µm. B. Collar, scale = 80 µm. C. Collar showing detail of spines, scale = 20 µm. D. Inner lobe, scale = 20 µm. E. Outer lobe, 20 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3108CBA563CEE76FF09FB76AE49FCBE" lastPageId="6" lastPageNumber="197" pageId="5" pageNumber="196" type="discussion">
|
||
<paragraph id="8BB5DF31563CEE75FF09FB76ACF9F914" blockId="5.[122,1174,1217,1721]" pageId="5" pageNumber="196">
|
||
<smallCapsWord id="8D5349ED563CEE75FF09FB76AC6DFB7A" baselines="1235,1234" box="[171,278,1217,1241]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Remarks" pageId="5" pageNumber="196">
|
||
<emphasis id="B97E0323563CEE75FF09FB76AC6DFB7A" bold="true" box="[171,278,1217,1241]" pageId="5" pageNumber="196">REMARKS</emphasis>
|
||
</smallCapsWord>
|
||
.— The presence of a quadrangular anterior end of the body, a posterior lobe of the cephalic shield, a large muscular bulb, the single branch of the hermaphroditic duct to the albumen and membrane glands, a bilobed mucous gland are all characteristics of species of
|
||
<taxonomicName id="4C0AA4B2563CEE75FBBFFAB6AC4CFA94" authority="(Rudman, 1972 a)" baseAuthorityName="Rudman" baseAuthorityYear="1972" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="5" pageNumber="196" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97E0323563CEE75FBBFFAB6A9EEFAB4" box="[1053,1173,1281,1305]" italics="true" pageId="5" pageNumber="196">Philinopsis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9BA2C0563CEE75FF20FA96AC54FA94" author="RUDMAN, W. B." box="[130,303,1313,1337]" pageId="5" pageNumber="196" pagination="50 - 62" refId="ref6426" refString="RUDMAN, W. B. 1972 a. On Melanochlamys Cheeseman, 1881, a genus of the Aglajidae (Opisthobranchia: Gastropoda). Pacific Science 26: 50 - 62." type="journal article" year="1972">Rudman, 1972a</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C0AA4B2563CEE75FEE7FA96AF72FA94" authorityName="Gosliner" authorityYear="2015" box="[325,521,1313,1337]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="5" pageNumber="196" phylum="Mollusca" rank="species" species="buntot">
|
||
<emphasis id="B97E0323563CEE75FEE7FA96AF72FA94" box="[325,521,1313,1337]" italics="true" pageId="5" pageNumber="196">Philinopsis buntot</emphasis>
|
||
</taxonomicName>
|
||
lacks the elevated posterior extension of the cephalic shield but has all the other characteristics of
|
||
<taxonomicName id="4C0AA4B2563CEE75FDBBFAF6AFEAFAF4" baseAuthorityName="Rudman" baseAuthorityYear="1972" box="[537,657,1345,1369]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="5" pageNumber="196" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97E0323563CEE75FDBBFAF6AFEAFAF4" box="[537,657,1345,1369]" italics="true" pageId="5" pageNumber="196">Philinopsis</emphasis>
|
||
</taxonomicName>
|
||
. The anatomy of
|
||
<taxonomicName id="4C0AA4B2563CEE75FCF1FAF6AEC8FAF4" authorityName="Gosliner" authorityYear="2015" box="[851,947,1345,1369]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="5" pageNumber="196" phylum="Mollusca" rank="species" species="buntot">
|
||
<emphasis id="B97E0323563CEE75FCF1FAF6AEC8FAF4" box="[851,947,1345,1369]" italics="true" pageId="5" pageNumber="196">P. buntot</emphasis>
|
||
</taxonomicName>
|
||
differs significatntly from other described members of the genus. The most obvious morphological attribute that distinguishes
|
||
<taxonomicName id="4C0AA4B2563CEE75FF70FA36AC4AFA34" authorityName="Gosliner" authorityYear="2015" box="[210,305,1409,1433]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="5" pageNumber="196" phylum="Mollusca" rank="species" species="buntot">
|
||
<emphasis id="B97E0323563CEE75FF70FA36AC4AFA34" box="[210,305,1409,1433]" italics="true" pageId="5" pageNumber="196">P. buntot</emphasis>
|
||
</taxonomicName>
|
||
is the presence of the elongate posteromedial appendage on the posterior shield. The penis is also similar to that described for
|
||
<taxonomicName id="4C0AA4B2563CEE75FDF9FA16AEEFFA14" authority="Gosliner, 2011" authorityName="Gosliner" authorityYear="2011" box="[603,916,1441,1465]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="5" pageNumber="196" phylum="Mollusca" rank="species" species="falciphallus">
|
||
<emphasis id="B97E0323563CEE75FDF9FA16AF89FA14" box="[603,754,1441,1465]" italics="true" pageId="5" pageNumber="196">P. falciphallus</emphasis>
|
||
<bibRefCitation id="EF9BA2C0563CEE75FD5AFA16AEEFFA14" author="GOSLINER, T." box="[760,916,1441,1465]" pageId="5" pageNumber="196" pagination="1 - 24" refId="ref6216" refString="GOSLINER, T. 2011. Six new species of aglajid opisthobranch mollusks from the tropical Indo-Pacific. Zootaxa 2751: 1 - 24." type="journal article" year="2011">Gosliner, 2011</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0AA4B2563CEE75FC64FA16A944FA15" authorityName="Gosliner" authorityYear="2011" box="[966,1087,1441,1464]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="5" pageNumber="196" phylum="Mollusca" rank="species" species="coronata">
|
||
<emphasis id="B97E0323563CEE75FC64FA16A944FA15" box="[966,1087,1441,1464]" italics="true" pageId="5" pageNumber="196">P. coronata</emphasis>
|
||
</taxonomicName>
|
||
Gosliner, 2011. These two species also have a oblong prostate that is connected to the penial sac by means of a thin duct. In all three species, the penial papilla is armed with spines, whereas no other species of
|
||
<taxonomicName id="4C0AA4B2563CEE75FF38F9B6AC69F9B4" baseAuthorityName="Rudman" baseAuthorityYear="1972" box="[154,274,1537,1561]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="5" pageNumber="196" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97E0323563CEE75FF38F9B6AC69F9B4" box="[154,274,1537,1561]" italics="true" pageId="5" pageNumber="196">Philinopsis</emphasis>
|
||
</taxonomicName>
|
||
are known to have an armed penis. In
|
||
<taxonomicName id="4C0AA4B2563CEE75FD67F9B6AE1BF9B4" authorityName="Gosliner" authorityYear="2011" box="[709,864,1537,1561]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="5" pageNumber="196" phylum="Mollusca" rank="species" species="falciphallus">
|
||
<emphasis id="B97E0323563CEE75FD67F9B6AE1BF9B4" box="[709,864,1537,1561]" italics="true" pageId="5" pageNumber="196">P. falciphallus</emphasis>
|
||
</taxonomicName>
|
||
, the base of the penis has a series of spines and a large chitinous sickle-shaped spine near the apex, whereas
|
||
<taxonomicName id="4C0AA4B2563CEE75FC7CF996A92CF995" authorityName="Gosliner" authorityYear="2011" box="[990,1111,1569,1592]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="5" pageNumber="196" phylum="Mollusca" rank="species" species="coronata">
|
||
<emphasis id="B97E0323563CEE75FC7CF996A92CF995" box="[990,1111,1569,1592]" italics="true" pageId="5" pageNumber="196">P. coronata</emphasis>
|
||
</taxonomicName>
|
||
has a ring of spines around the apex of the penial papilla and a ring of larger spines at the base of the penial papilla with a second ring of larger spines above the penial papilla. The arrangement of spines along the margins of the two penial papilla lobes and spines along a basal collar in
|
||
<taxonomicName id="4C0AA4B2563CEE75FB95F936A9EEF934" authorityName="Gosliner" authorityYear="2015" box="[1079,1173,1665,1689]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="5" pageNumber="196" phylum="Mollusca" rank="species" species="buntot">
|
||
<emphasis id="B97E0323563CEE75FB95F936A9EEF934" box="[1079,1173,1665,1689]" italics="true" pageId="5" pageNumber="196">P. buntot</emphasis>
|
||
</taxonomicName>
|
||
is unique to this species.
|
||
</paragraph>
|
||
<paragraph id="8BB5DF31563FEE76FF09FF6CAFF3FE1E" blockId="6.[123,1174,219,787]" pageId="6" pageNumber="197">
|
||
In his review of
|
||
<taxonomicName id="4C0AA4B2563FEE76FECBFF6CAFD0FF5E" authority=", Rudman (1972 a)" authorityName=", Rudman" authorityYear="1972" box="[361,683,219,243]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="197" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97E0323563FEE76FECBFF6CAC9AFF5E" box="[361,481,219,243]" italics="true" pageId="6" pageNumber="197">Philinopsis</emphasis>
|
||
,
|
||
<bibRefCitation id="EF9BA2C0563FEE76FE51FF6CAFD0FF5E" author="RUDMAN, W. B." box="[499,683,219,243]" pageId="6" pageNumber="197" pagination="50 - 62" refId="ref6426" refString="RUDMAN, W. B. 1972 a. On Melanochlamys Cheeseman, 1881, a genus of the Aglajidae (Opisthobranchia: Gastropoda). Pacific Science 26: 50 - 62." type="journal article" year="1972">Rudman (1972a)</bibRefCitation>
|
||
</taxonomicName>
|
||
indicated there were two distinct groups of species. Members of the first group are characterized by having a large quadrangular head, a bulbous buccal mass and a specialized penis with a basal elongate penial papilla and a short prostate (Marcus and
|
||
<bibRefCitation id="EF9BA2C0563FEE76FEAAFE8BACEFFEFE" author="MARCUS, EV & AND ER & MARCUS" box="[264,404,315,339]" pageId="6" pageNumber="197" pagination="1 - 137" refId="ref6300" refString="MARCUS, EV., AND ER. MARCUS. 1967. American opisthobranch mollusks Part I, Tropical American opisthobranchs. Studies in Tropical Oceanography, Miami 6: 1 - 137." type="journal article" year="1967">Marcus 1967</bibRefCitation>
|
||
: fig. 12) or a simple penis in
|
||
<taxonomicName id="4C0AA4B2563FEE76FD71FE8BA9F4FEFE" authority="(Renier, 1807) (Gosliner 1980)" authorityName="Gosliner" authorityYear="1980" baseAuthorityName="Renier" baseAuthorityYear="1807" box="[723,1167,315,339]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="197" phylum="Mollusca" rank="species" species="depicta">
|
||
<emphasis id="B97E0323563FEE76FD71FE8BAE47FEFE" box="[723,828,315,339]" italics="true" pageId="6" pageNumber="197">P. depicta</emphasis>
|
||
(Renier, 1807) (
|
||
<bibRefCitation id="EF9BA2C0563FEE76FC4CFE8CA9FEFEFE" author="GOSLINER, T. M." box="[1006,1157,315,339]" pageId="6" pageNumber="197" pagination="325 - 360" refId="ref6130" refString="GOSLINER, T. M. 1980. Systematics and phylogeny of the Aglajidae (Opisthobranchia: Mollusca). Zoological Journal of the Linnean Society 68: 325 - 360." type="journal article" year="1980">Gosliner 1980</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Members of the second group of species have a rounded head with an elevated bulbous region (that resembles the front end of a Boeing 747) with prominent eyes visible at the base of the bulb, an elongate, tubular buccal bulb and a simple penis.
|
||
</paragraph>
|
||
<paragraph id="8BB5DF31563FEE76FF09FE0BAE49FCBE" blockId="6.[123,1174,219,787]" pageId="6" pageNumber="197">
|
||
In the molecular study by
|
||
<bibRefCitation id="EF9BA2C0563FEE76FE66FE0CAE79FE7E" author="CAMACHO-GARCIA, Y. E. & E. ORNELAS-GATDULA & T. M. GOSLINER & A. VALDES" box="[452,770,443,467]" pageId="6" pageNumber="197" pagination="113 - 126" refId="ref5986" refString="CAMACHO-GARCIA, Y. E., E. ORNELAS-GATDULA, T. M. GOSLINER, AND A. VALDES. 2014. Phylogeny of the family Aglajidae (Pilsbry, 1895) (Opisthobranchia: Cephalaspidea) inferred from mtDNA and nDNA. Molecular Phylogenetics and Evolution 71: 113 - 126." type="journal article" year="2014">Camacho-García et al. (2014)</bibRefCitation>
|
||
(
|
||
<figureCitation id="1331C3B4563FEE76FCB0FE0CAE2BFE7E" box="[786,848,443,467]" captionStart="FIGURE" captionStartId="11.[171,183,1323,1342]" captionTargetBox="[172,1125,216,1306]" captionTargetId="figure-121@11.[172,1125,216,1306]" captionTargetPageId="11" captionText="FIGURE 7. Bayesian maximum credibility tree showing all taxa included for the analysis of the combined H3, 16S, and COI sequence alignments (after Camacho et al. 2014). Posterior probabilities areindicated above each branch (for the analysis without COI third positions on the left, and for the analysis with COI third positions on the right) and bootstrap values areindicated below each branch. Branch lengths are also shown here indicating the proportion of substitutions. Species names are color-coded according to their range, blue forIndo-Pacific, red for Atlantic and green for Eastern Pacific. The phylogenetic positions of Philinopsis buntot sp. nov. and Chelidonura alexisi sp. nov. are indicated by arrows on the tree." figureDoi="http://doi.org/10.5281/zenodo.11512298" httpUri="https://zenodo.org/record/11512298/files/figure.png" pageId="6" pageNumber="197">Fig. 7</figureCitation>
|
||
), these two groups are largely upheld, but two additional groups of taxa being included in the same clade as the taxa with a tubular buccal bulb.
|
||
<taxonomicName id="4C0AA4B2563FEE76FE8EFE4CAFD2FDBE" authority="(Baba, 1949)" baseAuthorityName="Baba" baseAuthorityYear="1949" box="[300,681,507,531]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="197" phylum="Mollusca" rank="species" species="orientalis">
|
||
<emphasis id="B97E0323563FEE76FE8EFE4CAF68FDBE" box="[300,531,507,531]" italics="true" pageId="6" pageNumber="197">Philinopsis orientalis</emphasis>
|
||
(Baba, 1949)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0AA4B2563FEE76FD43FE4BA917FDBE" authority="(Marcus and Marcus, 1976)" baseAuthorityName="Marcus and Marcus" baseAuthorityYear="1976" box="[737,1132,507,531]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="197" phylum="Mollusca" rank="species" species="petra">
|
||
<emphasis id="B97E0323563FEE76FD43FE4BAE4EFDBE" box="[737,821,508,531]" italics="true" pageId="6" pageNumber="197">P. petra</emphasis>
|
||
(Marcus and Marcus, 1976)
|
||
</taxonomicName>
|
||
are sister to the clade of species with the tubular buccal mass.
|
||
<taxonomicName id="4C0AA4B2563FEE76FCA4FDACA9F5FD9E" authority="Gosliner, 2011" authorityName="Gosliner" authorityYear="2011" box="[774,1166,539,563]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="197" phylum="Mollusca" rank="species" species="coronata">
|
||
<emphasis id="B97E0323563FEE76FCA4FDACAE9DFD9E" box="[774,998,539,563]" italics="true" pageId="6" pageNumber="197">Philinopsis coronata</emphasis>
|
||
<bibRefCitation id="EF9BA2C0563FEE76FC52FDACA9F5FD9E" author="GOSLINER, T." box="[1008,1166,539,563]" pageId="6" pageNumber="197" pagination="1 - 24" refId="ref6216" refString="GOSLINER, T. 2011. Six new species of aglajid opisthobranch mollusks from the tropical Indo-Pacific. Zootaxa 2751: 1 - 24." type="journal article" year="2011">Gosliner, 2011</bibRefCitation>
|
||
</taxonomicName>
|
||
, together with
|
||
<taxonomicName id="4C0AA4B2563FEE76FEADFD8BAC16FDFE" authorityName="Gosliner" authorityYear="2015" box="[271,365,571,595]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="197" phylum="Mollusca" rank="species" species="buntot">
|
||
<emphasis id="B97E0323563FEE76FEADFD8BAC16FDFE" box="[271,365,571,595]" italics="true" pageId="6" pageNumber="197">P. buntot</emphasis>
|
||
</taxonomicName>
|
||
(as
|
||
<emphasis id="B97E0323563FEE76FE38FD8BACD1FDFE" box="[410,426,572,595]" italics="true" pageId="6" pageNumber="197">P</emphasis>
|
||
. sp.
|
||
<collectionCode id="ED1B47F4563FEE76FE7FFD8CAF51FDFE" box="[477,554,571,595]" pageId="6" pageNumber="197">CASIZ</collectionCode>
|
||
185779), is sister to the remainder of the members of this clade.
|
||
<taxonomicName id="4C0AA4B2563FEE76FF60FDECACBAFDDE" authorityName="Gosliner" authorityYear="2011" box="[194,449,603,627]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="197" phylum="Mollusca" rank="species" species="falciphallus">
|
||
<emphasis id="B97E0323563FEE76FF60FDECACBAFDDE" box="[194,449,603,627]" italics="true" pageId="6" pageNumber="197">Philinopsis falciphallus</emphasis>
|
||
</taxonomicName>
|
||
, which has very similar morphology to
|
||
<taxonomicName id="4C0AA4B2563FEE76FCD9FDEBAEA0FDDE" authorityName="Gosliner" authorityYear="2015" box="[891,987,603,627]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="197" phylum="Mollusca" rank="species" species="buntot">
|
||
<emphasis id="B97E0323563FEE76FCD9FDEBAEA0FDDE" box="[891,987,603,627]" italics="true" pageId="6" pageNumber="197">P. buntot</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0AA4B2563FEE76FBB1FDEBA9F5FDDE" authorityName="Gosliner" authorityYear="2011" box="[1043,1166,604,627]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="197" phylum="Mollusca" rank="species" species="coronata">
|
||
<emphasis id="B97E0323563FEE76FBB1FDEBA9F5FDDE" box="[1043,1166,604,627]" italics="true" pageId="6" pageNumber="197">P. coronata</emphasis>
|
||
</taxonomicName>
|
||
, surprisingly nests with
|
||
<taxonomicName id="4C0AA4B2563FEE76FED0FDCCAFACFD3E" authority="Bertsch, 1972" authorityName="Bertsch" authorityYear="1972" box="[370,727,635,659]" class="Gastropoda" family="Aglajidae" genus="Aglaja" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="197" phylum="Mollusca" rank="species" species="regiscorona">
|
||
<emphasis id="B97E0323563FEE76FED0FDCCAF45FD3E" box="[370,574,635,659]" italics="true" pageId="6" pageNumber="197">Aglaja regiscorona</emphasis>
|
||
Bertsch, 1972
|
||
</taxonomicName>
|
||
, and several species of
|
||
<taxonomicName id="4C0AA4B2563FEE76FC71FDCCA921FD3E" authorityName="A. Adams" authorityYear="1850" box="[979,1114,635,659]" class="Gastropoda" family="Aglajidae" genus="Chelidonura" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="197" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97E0323563FEE76FC71FDCCA921FD3E" box="[979,1114,635,659]" italics="true" pageId="6" pageNumber="197">Chelidonura</emphasis>
|
||
</taxonomicName>
|
||
. This relationship requires additional study as the COI mitochondrial gene was not able to be amplified for
|
||
<taxonomicName id="4C0AA4B2563FEE76FF03FD0BAC41FD7E" authorityName="Gosliner" authorityYear="2011" box="[161,314,699,723]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="197" phylum="Mollusca" rank="species" species="falciphallus">
|
||
<emphasis id="B97E0323563FEE76FF03FD0BAC41FD7E" box="[161,314,699,723]" italics="true" pageId="6" pageNumber="197">P. falciphallus</emphasis>
|
||
</taxonomicName>
|
||
. The morphological distinctions of
|
||
<taxonomicName id="4C0AA4B2563FEE76FD18FD0BAE63FD7E" authorityName="Gosliner" authorityYear="2015" box="[698,792,699,723]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="197" phylum="Mollusca" rank="species" species="buntot">
|
||
<emphasis id="B97E0323563FEE76FD18FD0BAE63FD7E" box="[698,792,699,723]" italics="true" pageId="6" pageNumber="197">P. buntot</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0AA4B2563FEE76FC84FD0BAEDBFD7E" authorityName="Gosliner" authorityYear="2011" box="[806,928,700,723]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="197" phylum="Mollusca" rank="species" species="coronata">
|
||
<emphasis id="B97E0323563FEE76FC84FD0BAEDBFD7E" box="[806,928,700,723]" italics="true" pageId="6" pageNumber="197">P. coronata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0AA4B2563FEE76FC77FD0BA915FD7E" authorityName="Gosliner" authorityYear="2011" box="[981,1134,699,723]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="197" phylum="Mollusca" rank="species" species="falciphallus">
|
||
<emphasis id="B97E0323563FEE76FC77FD0BA915FD7E" box="[981,1134,699,723]" italics="true" pageId="6" pageNumber="197">P. falciphallus</emphasis>
|
||
</taxonomicName>
|
||
are discussed above, but the molecular data suggesting a sister species relationship between
|
||
<taxonomicName id="4C0AA4B2563FEE76FB94FD6BA9EEFD5E" authorityName="Gosliner" authorityYear="2015" box="[1078,1173,731,755]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="197" phylum="Mollusca" rank="species" species="buntot">
|
||
<emphasis id="B97E0323563FEE76FB94FD6BA9EEFD5E" box="[1078,1173,731,755]" italics="true" pageId="6" pageNumber="197">P. buntot</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0AA4B2563FEE76FF0BFD4BAC59FCBE" authorityName="Gosliner" authorityYear="2011" box="[169,290,764,787]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="197" phylum="Mollusca" rank="species" species="coronata">
|
||
<emphasis id="B97E0323563FEE76FF0BFD4BAC59FCBE" box="[169,290,764,787]" italics="true" pageId="6" pageNumber="197">P. coronata</emphasis>
|
||
</taxonomicName>
|
||
is consistent with their morphological similarity.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |