treatments-xml/data/03/9F/5A/039F5A5FFFE9FFECFF20FB0EFD6A39C0.xml

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<mods:title id="B2C133C25A80A9938D02B00BA2A4C017">Trichadenotecnum species from Peninsular Malaysia and Singapore (Insecta: Psocodea: ' Psocoptera': Psocidae)</mods:title>
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<mods:namePart id="FC5E24035753E694672CADF1ED3DABAF">Yoshizawa, Kazunori</mods:namePart>
<mods:affiliation id="C542323568AA3822C7895CD19EC2F9C6">Systematic Entomology, School of Agriculture, Hokkaido University, Sapporo 060 - 8589, Japan.</mods:affiliation>
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<mods:namePart id="A2A59A9D2CB2B7E94D551169B2A70E2B">Lienhard, Charles</mods:namePart>
<mods:affiliation id="7D0EF58CB2C180A4C95B95FB36DE109B">Muséum d'histoire naturelle, C. P. 6434, CH- 1211 Genève 6, Switzerland</mods:affiliation>
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<mods:namePart id="B33930FDD4FC96D9BF19899E5E7F779D">Abd, Idris</mods:namePart>
<mods:affiliation id="2ADCE58304ABCF85E40AF763B1B0B8EE">Center for Insect Systematics, Faculty of Sciences and Technology, Universiti Kebangsaan Malaysia, 43600 Bangi, Malaysia</mods:affiliation>
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<heading id="D0C15C25FFE9FFECFF20FB0EFD483AB5" bold="true" box="[151,705,1169,1195]" fontSize="11" level="1" pageId="22" pageNumber="491" reason="1">
<taxonomicName id="4C3690CAFFE9FFECFF20FB0EFD483AB5" ID-CoL="587CL" authority="Endang &amp; New" authorityName="Endang &amp; New" authorityYear="2005" box="[151,705,1169,1195]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="22" pageNumber="491" phylum="Arthropoda" rank="species" species="cornutum">
<emphasis id="B942375BFFE9FFECFF20FB0EFD483AB5" bold="true" box="[151,705,1169,1195]" pageId="22" pageNumber="491">
<emphasis id="B942375BFFE9FFECFF20FB0EFE733AB5" bold="true" box="[151,506,1169,1195]" italics="true" pageId="22" pageNumber="491">Trichadenotecnum cornutum</emphasis>
Endang &amp; New
</emphasis>
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<paragraph id="8B89EB49FFE9FFECFF20FB2BFEA43AD3" blockId="22.[151,705,1169,1229]" box="[151,301,1204,1229]" pageId="22" pageNumber="491">
(
<figureCitation id="130DF7CCFFE9FFECFF28FB2BFF723AD3" box="[159,251,1204,1229]" captionStart="FIGURE 1.1" captionStartId="2.[151,250,1786,1808]" captionTargetBox="[151,1435,196,1746]" captionTargetId="figure-94@2.[151,1436,193,1765]" captionTargetPageId="2" captionText="FIGURE 1.1. Forewings of Trichadenotecnum spp.: A: T. circularoides, female; B: T. cinnamonum, male; C: ditto, female; D: T. imrum, male; E: T.pardus, female; F: T. yatai, holotype male; G: T. kojimai, holotype male; H: T. adika, male; I: T. endangae, holotype male; J: T. hammani, female; K: T. sp.My19, female; L: T. gombakense, female; M: T. malayense, male." figureDoi="http://doi.org/10.5281/zenodo.4921625" httpUri="https://zenodo.org/record/4921625/files/figure.png" pageId="22" pageNumber="491">Figs 1Q</figureCitation>
, 
<figureCitation id="130DF7CCFFE9FFECFEBFFB2BFEAC3AD3" box="[264,293,1204,1229]" captionStart="FIGURE 24" captionStartId="25.[151,250,1151,1173]" captionTargetBox="[155,1422,199,1123]" captionTargetId="figure-311@25.[151,1435,193,1129]" captionTargetPageId="25" captionText="FIGURE 24. Male terminalia of Trichadenotecnum cornutum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view. E: phallosome, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4921674" httpUri="https://zenodo.org/record/4921674/files/figure.png" pageId="22" pageNumber="491">24</figureCitation>
)
</paragraph>
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<subSubSection id="C32CB8C2FFE9FFECFF20FB63FE723B65" pageId="22" pageNumber="491" type="reference_group">
<paragraph id="8B89EB49FFE9FFECFF20FB63FD403B0C" blockId="22.[151,713,1276,1298]" box="[151,713,1276,1298]" pageId="22" pageNumber="491">
<treatmentCitationGroup id="AB26CC67FFE9FFECFF20FB63FD403B0C" box="[151,713,1276,1298]" pageId="22" pageNumber="491">
<treatmentCitation id="0A97CD58FFE9FFECFF20FB63FD4C3B0C" author="Endang, S. K. &amp; New, T. R." box="[151,709,1276,1298]" page="28" pageId="22" pageNumber="491" year="2005">
<taxonomicName id="4C3690CAFFE9FFECFF20FB63FD4C3B0C" ID-CoL="587CL" authority="Endang &amp; New, 2005: 28" authorityName="Endang &amp; New" authorityPageNumber="28" authorityYear="2005" box="[151,709,1276,1298]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="22" pageNumber="491" phylum="Arthropoda" rank="species" species="cornutum">
<emphasis id="B942375BFFE9FFECFF20FB63FE323B0C" box="[151,443,1276,1298]" italics="true" pageId="22" pageNumber="491">Trichadenotecnum cornutum</emphasis>
<bibRefCitation id="EFA796B8FFE9FFECFE76FB63FD4C3B0C" author="Endang, S. K. &amp; New, T. R." box="[449,709,1276,1298]" pageId="22" pageNumber="491" pagination="15 - 38" refId="ref14985" refString="Endang, S. K. &amp; New, T. R. (2005) The genus Trichadenotecnum (Insecta: Psocoptera: Psocidae) in Sumatra, Indonesia, with description of thirteen new species. Records of the Australian Museum, 57, 15 - 38. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.57.2005.1437" type="journal article" year="2005">Endang &amp; New, 2005: 28</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="8B89EB49FFE9FFECFF20FAA1FE723B65" blockId="22.[151,1438,1342,2015]" pageId="22" pageNumber="491">
<emphasis id="B942375BFFE9FFECFF20FAA1FE103B49" bold="true" box="[151,409,1342,1367]" inLineHeading="true" pageId="22" pageNumber="491" reason="1">Specimens examined.</emphasis>
[
<collectingCountry id="F321ABD9FFE9FFECFE1DFAA1FD9F3B49" box="[426,534,1342,1367]" name="Malaysia" pageId="22" pageNumber="491">Malaysia</collectingCountry>
] 
<specimenCount id="9D3020C0FFE9FFECFD90FAA1FDF13B49" box="[551,632,1342,1367]" pageId="22" pageNumber="491" type="male">1 male</specimenCount>
, Endau Rompin, 
<collectingRegion id="49F225ABFFE9FFECFCF4FAA1FC0A3B49" box="[835,899,1342,1367]" country="Malaysia" name="Johor" pageId="22" pageNumber="491">Johor</collectingRegion>
, 
<date id="FF88CD89FFE9FFECFC25FAA1FB8F3B49" box="[914,1030,1342,1367]" pageId="22" pageNumber="491" value="2003-07-09">9.vii.2003</date>
, NT (UKM); 
<materialsCitation id="3B5EE114FFE9FFECFB1DFAA1FE7E3B65" ID-GBIF-Occurrence="3326724325" collectingDate="2003-03-06" collectionCode="UKM" location="Fraser's Hill" pageId="22" pageNumber="491" specimenCount="1" specimenCount-male="1" stateProvince="Pahang">
<specimenCount id="9D3020C0FFE9FFECFB1DFAA1FB723B49" box="[1194,1275,1342,1367]" pageId="22" pageNumber="491" type="male">1 male</specimenCount>
, 
<location id="8EE9BD92FFE9FFECFABEFAA1FA1E3B49" LSID="urn:lsid:plazi:treatment:039F5A5FFFE9FFECFF20FB0EFD6A39C0:8EE9BD92FFE9FFECFABEFAA1FA1E3B49" box="[1289,1431,1342,1367]" name="Fraser's Hill" pageId="22" pageNumber="491" stateProvince="Pahang">Fraser's Hill</location>
, 
<collectingRegion id="49F225ABFFE9FFECFF20FAFDFF653B65" box="[151,236,1378,1403]" country="Malaysia" name="Pahang" pageId="22" pageNumber="491">Pahang</collectingRegion>
, 
<date id="FF88CD89FFE9FFECFF40FAFDFEEC3B65" box="[247,357,1378,1403]" pageId="22" pageNumber="491" value="2003-03-06">
<collectingDate id="EFCC3461FFE9FFECFF40FAFDFEEC3B65" box="[247,357,1378,1403]" pageId="22" pageNumber="491" value="2003-03-06">6.iii.2003</collectingDate>
</date>
, KY (
<collectionCode id="ED27738CFFE9FFECFE1FFAFDFE663B65" box="[424,495,1378,1403]" country="Malaysia" httpUri="http://grbio.org/cool/ashc-3pfs" name="Universiti Kebangsaan Malaysia" pageId="22" pageNumber="491">UKM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32CB8C2FFE9FFECFF70FA19FD6A39C0" pageId="22" pageNumber="491" type="discussion">
<paragraph id="8B89EB49FFE9FFECFF70FA19FEE73918" blockId="22.[151,1438,1342,2015]" pageId="22" pageNumber="491">
<emphasis id="B942375BFFE9FFECFF70FA19FEB23B80" box="[199,315,1414,1439]" inLineHeading="true" pageId="22" pageNumber="491" reason="1">
<emphasis id="B942375BFFE9FFECFF70FA19FEBD3B81" bold="true" box="[199,308,1414,1439]" pageId="22" pageNumber="491">Remarks</emphasis>
.
</emphasis>
This species was originally described from 
<collectingRegion id="49F225ABFFE9FFECFC85FA19FC1B3B81" box="[818,914,1414,1439]" country="Indonesia" name="Sumatera Utara" pageId="22" pageNumber="491">Sumatra</collectingRegion>
, 
<collectingCountry id="F321ABD9FFE9FFECFC28FA19FB853B81" box="[927,1036,1414,1439]" name="Indonesia" pageId="22" pageNumber="491">Indonesia</collectingCountry>
based on a single male and is here recorded from Peninsula 
<collectingCountry id="F321ABD9FFE9FFECFE05FA35FD903BDD" box="[434,537,1450,1475]" name="Malaysia" pageId="22" pageNumber="491">Malaysia</collectingCountry>
for the first time. The specimens examined here are in good agreement with the original description of 
<taxonomicName id="4C3690CAFFE9FFECFE16FA50FDA53BF8" authorityName="Endang &amp; New" authorityYear="2005" box="[417,556,1487,1510]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="22" pageNumber="491" phylum="Arthropoda" rank="species" species="cornutum">
<emphasis id="B942375BFFE9FFECFE16FA50FDA53BF8" box="[417,556,1487,1510]" italics="true" pageId="22" pageNumber="491">T. cornutum</emphasis>
</taxonomicName>
(
<figureCitation id="130DF7CCFFE9FFECFD89FA51FD1F3BF9" box="[574,662,1486,1511]" captionStart="FIGURE 24" captionStartId="25.[151,250,1151,1173]" captionTargetBox="[155,1422,199,1123]" captionTargetId="figure-311@25.[151,1435,193,1129]" captionTargetPageId="25" captionText="FIGURE 24. Male terminalia of Trichadenotecnum cornutum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view. E: phallosome, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4921674" httpUri="https://zenodo.org/record/4921674/files/figure.png" pageId="22" pageNumber="491">Fig. 24</figureCitation>
). However, 
<bibRefCitation id="EFA796B8FFE9FFECFC94FA51FBA73BF9" author="Endang, S. K. &amp; New, T. R." box="[803,1070,1486,1511]" pageId="22" pageNumber="491" pagination="15 - 38" refId="ref14985" refString="Endang, S. K. &amp; New, T. R. (2005) The genus Trichadenotecnum (Insecta: Psocoptera: Psocidae) in Sumatra, Indonesia, with description of thirteen new species. Records of the Australian Museum, 57, 15 - 38. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.57.2005.1437" type="journal article" year="2005">Endang &amp; New (2005)</bibRefCitation>
described the phallosome of 
<taxonomicName id="4C3690CAFFE9FFECFA30FA50FE8B3814" authorityName="Endang &amp; New" authorityYear="2005" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="22" pageNumber="491" phylum="Arthropoda" rank="species" species="cornutum">
<emphasis id="B942375BFFE9FFECFA30FA50FE8B3814" italics="true" pageId="22" pageNumber="491">T. cornutum</emphasis>
</taxonomicName>
as open posteriorly, whereas, in the presently examined specimens, this structure is closed (
<figureCitation id="130DF7CCFFE9FFECFA92FA6DFA193815" box="[1317,1424,1522,1547]" captionStart="FIGURE 24" captionStartId="25.[151,250,1151,1173]" captionTargetBox="[155,1422,199,1123]" captionTargetId="figure-311@25.[151,1435,193,1129]" captionTargetPageId="25" captionText="FIGURE 24. Male terminalia of Trichadenotecnum cornutum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view. E: phallosome, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4921674" httpUri="https://zenodo.org/record/4921674/files/figure.png" pageId="22" pageNumber="491">Fig. 24D</figureCitation>
). However, the figure of the strongly deformed phallosome in the original description clearly shows that it was broken before being illustrated. Apart from the posteromedian part, the phallosomal structure of the present specimens is also in good agreement with the original illustration of 
<emphasis id="B942375BFFE9FFECFC21F9C0FBAD3868" box="[918,1060,1631,1654]" italics="true" pageId="22" pageNumber="491">
<taxonomicName id="4C3690CAFFE9FFECFC21F9C0FBA83868" authorityName="Endang &amp; New" authorityYear="2005" box="[918,1057,1631,1654]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="22" pageNumber="491" phylum="Arthropoda" rank="species" species="cornutum">T. cornutum</taxonomicName>
.
</emphasis>
Therefore, we concluded that the difference between the present specimens and the original description corresponds to an artifact, and that the specimens examined here are conspecific with 
<taxonomicName id="4C3690CAFFE9FFECFD01F938FCB638A0" authorityName="Endang &amp; New" authorityYear="2005" box="[694,831,1703,1726]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="22" pageNumber="491" phylum="Arthropoda" rank="species" species="cornutum">
<emphasis id="B942375BFFE9FFECFD01F938FCB638A0" box="[694,831,1703,1726]" italics="true" pageId="22" pageNumber="491">T. cornutum</emphasis>
</taxonomicName>
. Examination of the complete phallosome showed a very particular, long posteromedian spine, which is uniquely observed in this species within the genus 
<taxonomicName id="4C3690CAFFE9FFECFF20F971FEEE3919" authorityName="Enderlein" authorityYear="1909" box="[151,359,1774,1799]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="22" pageNumber="491" phylum="Arthropoda" rank="genus">
<emphasis id="B942375BFFE9FFECFF20F971FEEE3919" box="[151,359,1774,1799]" italics="true" pageId="22" pageNumber="491">Trichadenotecnum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B89EB49FFE9FFECFF70F88CFD6A39C0" blockId="22.[151,1438,1342,2015]" pageId="22" pageNumber="491">
Because the female of this species remains unknown, identification of its close relatives among the 
<taxonomicName id="4C3690CAFFE9FFECFAEEF88CFA153934" baseAuthorityName="Kolbe" baseAuthorityYear="1880" box="[1369,1436,1811,1834]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="22" pageNumber="491" phylum="Arthropoda" rank="species" species="majus">
<emphasis id="B942375BFFE9FFECFAEEF88CFA153934" box="[1369,1436,1811,1834]" italics="true" pageId="22" pageNumber="491">majus</emphasis>
</taxonomicName>
group is difficult. Presence of the median tongue resembles 
<taxonomicName id="4C3690CAFFE9FFECFCCFF8A7FC573951" baseAuthorityName="Kolbe" baseAuthorityYear="1880" box="[888,990,1848,1871]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="22" pageNumber="491" phylum="Arthropoda" rank="species" species="majus">
<emphasis id="B942375BFFE9FFECFCCFF8A7FC573951" box="[888,990,1848,1871]" italics="true" pageId="22" pageNumber="491">T. majus</emphasis>
</taxonomicName>
and its relatives, but it represents a plesiomorphic condition within the group. 
<taxonomicName id="4C3690CAFFE9FFECFDCBF8C4FC8D396C" authorityName="Endang &amp; New" authorityYear="2005" box="[636,772,1883,1906]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="22" pageNumber="491" phylum="Arthropoda" rank="species" species="cornutum">
<emphasis id="B942375BFFE9FFECFDCBF8C4FC8D396C" box="[636,772,1883,1906]" italics="true" pageId="22" pageNumber="491">T. cornutum</emphasis>
</taxonomicName>
resembles 
<taxonomicName id="4C3690CAFFE9FFECFC33F8C4FC74396C" authorityName="Endang, Thornton &amp; New" authorityYear="2002" box="[900,1021,1883,1906]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="22" pageNumber="491" phylum="Arthropoda" rank="species" species="soenarti">
<emphasis id="B942375BFFE9FFECFC33F8C4FC74396C" box="[900,1021,1883,1906]" italics="true" pageId="22" pageNumber="491">T. soenarti</emphasis>
</taxonomicName>
, for which both sexes are known, in male hypandrial structures (median tongue present) and in the structure of the phallosome (posterior margin strongly extended ventrally). Their close affinity is undoubted so that 
<taxonomicName id="4C3690CAFFE9FFECFC29F83CFBAD39A4" authorityName="Endang &amp; New" authorityYear="2005" box="[926,1060,1955,1978]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="22" pageNumber="491" phylum="Arthropoda" rank="species" species="cornutum">
<emphasis id="B942375BFFE9FFECFC29F83CFBAD39A4" box="[926,1060,1955,1978]" italics="true" pageId="22" pageNumber="491">T. cornutum</emphasis>
</taxonomicName>
is also likely to be closely related to 
<taxonomicName id="4C3690CAFFE9FFECFF03F857FEA539C1" authorityName="Thornton" authorityYear="1961" box="[180,300,1992,2015]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="22" pageNumber="491" phylum="Arthropoda" rank="species" species="apertum">
<emphasis id="B942375BFFE9FFECFF03F857FEA539C1" box="[180,300,1992,2015]" italics="true" pageId="22" pageNumber="491">T. apertum</emphasis>
</taxonomicName>
and its relatives, as well as 
<taxonomicName id="4C3690CAFFE9FFECFDD2F857FD5539C1" authorityName="Endang, Thornton &amp; New" authorityYear="2002" box="[613,732,1992,2015]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="22" pageNumber="491" phylum="Arthropoda" rank="species" species="soenarti">
<emphasis id="B942375BFFE9FFECFDD2F857FD5539C1" box="[613,732,1992,2015]" italics="true" pageId="22" pageNumber="491">T. soenarti</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>