treatments-xml/data/03/9F/5A/039F5A5FFFE3FFE7FF20F97BFED03975.xml

<document id="DB0AF6593E02C928C1516964A5D90E87" ID-CLB-Dataset="5285" ID-DOI="10.11646/zootaxa.3835.4.3" ID-GBIF-Dataset="dbf8de20-9c7a-4b66-9130-615b6b42e84e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4921623" ID-ZooBank="AD845CF3-CB19-4924-891F-330AAE283D07" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623283883149" checkinUser="felipe" docAuthor="Yoshizawa, Kazunori, Lienhard, Charles &amp; Abd, Idris" docDate="2014" docId="039F5A5FFFE3FFE7FF20F97BFED03975" docLanguage="en" docName="zt03835p500.pdf" docOrigin="Zootaxa 3835 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichadenotecnum krucilense Endang, Thornton &amp; New" docType="treatment" docVersion="7" lastPageNumber="498" masterDocId="FFA62227FFFFFFFAFFB7FF9FFF893E1E" masterDocTitle="Trichadenotecnum species from Peninsular Malaysia and Singapore (Insecta: Psocodea: ' Psocoptera': Psocidae)" masterLastPageNumber="500" masterPageNumber="469" pageNumber="497" updateTime="1699009618324" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="42E540DFBFBCEDAB9FFF238811960FE1">Trichadenotecnum species from Peninsular Malaysia and Singapore (Insecta: Psocodea: ' Psocoptera': Psocidae)</mods:title>
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<mods:namePart id="31F2BAC7F5D623B4B4FE1DBFA9F4C76D">Yoshizawa, Kazunori</mods:namePart>
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<mods:namePart id="F51060DBB71B08387BEAC792699F4AE4">Lienhard, Charles</mods:namePart>
<mods:affiliation id="AF6DD3CFBCEB55394724EA991384A602">Muséum d'histoire naturelle, C. P. 6434, CH- 1211 Genève 6, Switzerland</mods:affiliation>
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<mods:namePart id="3D245DEEE5EC69D8876A564197A2112C">Abd, Idris</mods:namePart>
<mods:affiliation id="86242D8B5CE666BF375923853E74BC09">Center for Insect Systematics, Faculty of Sciences and Technology, Universiti Kebangsaan Malaysia, 43600 Bangi, Malaysia</mods:affiliation>
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<heading id="D0C15C25FFE3FFE6FF20F97BFCC738E0" bold="true" box="[151,846,1764,1790]" fontSize="11" level="1" pageId="28" pageNumber="497" reason="1">
<taxonomicName id="4C3690CAFFE3FFE6FF20F97BFCC738E0" ID-CoL="7CQPT" authority="Endang, Thornton &amp; New" authorityName="Endang, Thornton &amp; New" box="[151,846,1764,1790]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="28" pageNumber="497" phylum="Arthropoda" rank="species" species="krucilense">
<emphasis id="B942375BFFE3FFE6FF20F97BFCC738E0" bold="true" box="[151,846,1764,1790]" pageId="28" pageNumber="497">
<emphasis id="B942375BFFE3FFE6FF20F97BFD8938E0" bold="true" box="[151,512,1764,1790]" italics="true" pageId="28" pageNumber="497">Trichadenotecnum krucilense</emphasis>
Endang, Thornton &amp; New
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(
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, 
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)
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<paragraph id="8B89EB49FFE3FFE6FF20F8D0FBDE397B" blockId="28.[151,1111,1871,1893]" box="[151,1111,1871,1893]" pageId="28" pageNumber="497">
<treatmentCitationGroup id="AB26CC67FFE3FFE6FF20F8D0FBDE397B" box="[151,1111,1871,1893]" pageId="28" pageNumber="497">
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<emphasis id="B942375BFFE3FFE6FF20F8D0FE4B397B" box="[151,450,1871,1893]" italics="true" pageId="28" pageNumber="497">Trichadenotecnum krucilense</emphasis>
<bibRefCitation id="EFA796B8FFE3FFE6FE7FF8D0FCCD397B" author="Endang, S. K. &amp; Thornton, I. W. B. &amp; New, T. R." box="[456,836,1871,1893]" pageId="28" pageNumber="497" pagination="107 - 176" refId="ref15093" refString="Endang, S. K., Thornton, I. W. B. &amp; New, T. R. (2002) The Psocidae (Insecta: Psocoptera) of Java and the eastern islands of Indonesia. Invertebrate Systematics, 16, 107 - 176." type="journal article" year="2002">Endang, Thornton &amp; New, 2002: 165</bibRefCitation>
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; 
<treatmentCitation id="0A97CD58FFE3FFE6FCF9F8D0FBDA397B" author="Endang, S. K. &amp; New, T. R." box="[846,1107,1871,1893]" page="23" pageId="28" pageNumber="497" year="2005">
<bibRefCitation id="EFA796B8FFE3FFE6FCF9F8D0FBDA397B" author="Endang, S. K. &amp; New, T. R." box="[846,1107,1871,1893]" pageId="28" pageNumber="497" pagination="15 - 38" refId="ref14985" refString="Endang, S. K. &amp; New, T. R. (2005) The genus Trichadenotecnum (Insecta: Psocoptera: Psocidae) in Sumatra, Indonesia, with description of thirteen new species. Records of the Australian Museum, 57, 15 - 38. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.57.2005.1437" type="journal article" year="2005">Endang &amp; New, 2005: 23</bibRefCitation>
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.
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</paragraph>
<paragraph id="8B89EB49FFE3FFE6FF20F80DFE7639D0" blockId="28.[151,1437,1938,1998]" pageId="28" pageNumber="497">
<emphasis id="B942375BFFE3FFE6FF20F80DFE0639B5" bold="true" box="[151,399,1938,1963]" inLineHeading="true" pageId="28" pageNumber="497" reason="1">Specimen examined.</emphasis>
[
<collectingCountry id="F321ABD9FFE3FFE6FE17F80DFD8539B5" box="[416,524,1938,1963]" name="Malaysia" pageId="28" pageNumber="497">Malaysia</collectingCountry>
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<specimenCount id="9D3020C0FFE3FFE6FDA9F80DFDF939B5" box="[542,624,1938,1963]" pageId="28" pageNumber="497" type="male">1 male</specimenCount>
, 
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, 
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3, 
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, 
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, 
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, 
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.etal (canopy fogging, St. 6).
</paragraph>
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<paragraph id="8B89EB49FFE2FFE7FF20FB4AFD443B14" blockId="29.[151,1436,1237,1290]" pageId="29" pageNumber="498">
<emphasis id="B942375BFFE2FFE7FF20FB4AFEAD3AF4" bold="true" box="[151,292,1237,1259]" pageId="29" pageNumber="498">FIGURE 29.</emphasis>
Male terminalia of 
<taxonomicName id="4C3690CAFFE2FFE7FDB5FB4AFCB83AF5" box="[514,817,1237,1259]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="29" pageNumber="498" phylum="Arthropoda" rank="species" species="krucilense">
<emphasis id="B942375BFFE2FFE7FDB5FB4AFCB83AF5" box="[514,817,1237,1259]" italics="true" pageId="29" pageNumber="498">Trichadenotecnum krucilense</emphasis>
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. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view.
</paragraph>
</caption>
<subSubSection id="C32CB8C2FFE2FFE7FF70FAA9FED03975" pageId="29" pageNumber="498" type="discussion">
<paragraph id="8B89EB49FFE2FFE7FF70FAA9FB433839" blockId="29.[151,1437,1334,1899]" pageId="29" pageNumber="498">
<emphasis id="B942375BFFE2FFE7FF70FAA9FEB23B50" box="[199,315,1334,1359]" inLineHeading="true" pageId="29" pageNumber="498" reason="1">
<emphasis id="B942375BFFE2FFE7FF70FAA9FEBD3B51" bold="true" box="[199,308,1334,1359]" pageId="29" pageNumber="498">Remarks</emphasis>
.
</emphasis>
The species was originally described from 
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, 
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, later also recorded from 
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, 
<collectingCountry id="F321ABD9FFE2FFE7FF20FAC5FE8E3B6D" box="[151,263,1370,1395]" name="Indonesia" pageId="29" pageNumber="498">Indonesia</collectingCountry>
, and is here recorded from Peninsula 
<collectingCountry id="F321ABD9FFE2FFE7FD60FAC5FCB43B6D" box="[727,829,1370,1395]" name="Malaysia" pageId="29" pageNumber="498">Malaysia</collectingCountry>
for the first time. The variation of the hypandrial structure, three (Javanese) or four (Sumatran) hypandrial spikes, has been pointed out by 
<bibRefCitation id="EFA796B8FFE2FFE7FB26FAE1FA113B89" author="Endang, S. K. &amp; New, T. R." box="[1169,1432,1406,1431]" pageId="29" pageNumber="498" pagination="15 - 38" refId="ref14985" refString="Endang, S. K. &amp; New, T. R. (2005) The genus Trichadenotecnum (Insecta: Psocoptera: Psocidae) in Sumatra, Indonesia, with description of thirteen new species. Records of the Australian Museum, 57, 15 - 38. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.57.2005.1437" type="journal article" year="2005">Endang &amp; New (2005)</bibRefCitation>
. The presently examined specimen has four hypandrial spikes. Judging from the original hypandrial illustration published by 
<bibRefCitation id="EFA796B8FFE2FFE7FE83FA59FD973BC1" author="Endang, S. K. &amp; Thornton, I. W. B. &amp; New, T. R." box="[308,542,1478,1503]" pageId="29" pageNumber="498" pagination="107 - 176" refId="ref15093" refString="Endang, S. K., Thornton, I. W. B. &amp; New, T. R. (2002) The Psocidae (Insecta: Psocoptera) of Java and the eastern islands of Indonesia. Invertebrate Systematics, 16, 107 - 176." type="journal article" year="2002">
Endang 
<emphasis id="B942375BFFE2FFE7FE23FA58FE463BC1" box="[404,463,1478,1503]" italics="true" pageId="29" pageNumber="498">et al.</emphasis>
(2002
</bibRefCitation>
: fig. 333), missing of some hypandrial parts on the left side of the specimen seems evident; on that side only one spike is figured whereas two on the other side. It is likely that the variation pointed out by 
<bibRefCitation id="EFA796B8FFE2FFE7FEF7F991FDB63839" author="Endang, S. K. &amp; New, T. R." box="[320,575,1550,1575]" pageId="29" pageNumber="498" pagination="15 - 38" refId="ref14985" refString="Endang, S. K. &amp; New, T. R. (2005) The genus Trichadenotecnum (Insecta: Psocoptera: Psocidae) in Sumatra, Indonesia, with description of thirteen new species. Records of the Australian Museum, 57, 15 - 38. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.57.2005.1437" type="journal article" year="2005">Endang &amp; New (2005)</bibRefCitation>
is actually an artifact caused by dissection or preparation.
</paragraph>
<paragraph id="8B89EB49FFE2FFE7FF70F9ADFED03975" blockId="29.[151,1437,1334,1899]" pageId="29" pageNumber="498">
The hypandrium of this species (
<figureCitation id="130DF7CCFFE2FFE7FDF1F9ADFD273855" box="[582,686,1586,1611]" captionStart="FIGURE 29" captionStartId="29.[151,250,1237,1259]" captionTargetBox="[153,1423,205,1211]" captionTargetId="figure-318@29.[151,1435,193,1216]" captionTargetPageId="29" captionText="FIGURE 29. Male terminalia of Trichadenotecnum krucilense. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921690" httpUri="https://zenodo.org/record/4921690/files/figure.png" pageId="29" pageNumber="498">Fig. 29C</figureCitation>
) is unique, very different from that of any other known species. Absence of the hypandrial median tongue may indicate its close affinity with 
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<emphasis id="B942375BFFE2FFE7FBB4F9C8FBF53870" box="[1027,1148,1623,1646]" italics="true" pageId="29" pageNumber="498">T. apertum</emphasis>
</taxonomicName>
and its close relatives. In the original description of this species, the phallosome is illustrated in highly deformed condition, but the present examination clearly shows that this species has the pseudo-parameres on the phallosome (
<figureCitation id="130DF7CCFFE2FFE7FB3CF901FB7D38A9" box="[1163,1268,1694,1719]" captionStart="FIGURE 29" captionStartId="29.[151,250,1237,1259]" captionTargetBox="[153,1423,205,1211]" captionTargetId="figure-318@29.[151,1435,193,1216]" captionTargetPageId="29" captionText="FIGURE 29. Male terminalia of Trichadenotecnum krucilense. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921690" httpUri="https://zenodo.org/record/4921690/files/figure.png" pageId="29" pageNumber="498">Fig. 29D</figureCitation>
). The presence of this structure is a synapomorphy of the 
<emphasis id="B942375BFFE2FFE7FDDBF95CFC8F38C4" box="[620,774,1731,1754]" italics="true" pageId="29" pageNumber="498">sexpunctatum</emphasis>
, 
<taxonomicName id="4C3690CAFFE2FFE7FCA3F95DFCE438C5" authorityName="Thornton" authorityYear="1961" box="[788,877,1730,1755]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="29" pageNumber="498" phylum="Arthropoda" rank="species" species="medium">
<emphasis id="B942375BFFE2FFE7FCA3F95DFCE438C5" box="[788,877,1730,1755]" italics="true" pageId="29" pageNumber="498">medium</emphasis>
</taxonomicName>
and New World species groups (see 
<bibRefCitation id="EFA796B8FFE2FFE7FAB0F95DFF7738E1" author="Yoshizawa, K. &amp; Garcia Aldrete, A. N. &amp; Mockford, E. L." pageId="29" pageNumber="498" pagination="651 - 723" refId="ref15976" refString="Yoshizawa, K., Garcia Aldrete, A. N. &amp; Mockford, E. L. (2008) Systematics and biogeography of the New World species of Trichadenotecnum Enderlein (Insecta: Psocodea: ' Psocoptera': Psocidae). Zoological Journal of the Linnean Society, 153, 651 - 723. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2008.00398. x" type="journal article" year="2008">
Yoshizawa 
<emphasis id="B942375BFFE2FFE7FA3FF95CFF2438E1" italics="true" pageId="29" pageNumber="498">et al</emphasis>
., 2008
</bibRefCitation>
) so that assignment of 
<taxonomicName id="4C3690CAFFE2FFE7FDBCF978FD1538E1" box="[523,668,1766,1791]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="29" pageNumber="498" phylum="Arthropoda" rank="species" species="krucilense">
<emphasis id="B942375BFFE2FFE7FDBCF978FD1538E1" box="[523,668,1766,1791]" italics="true" pageId="29" pageNumber="498">T. krucilense</emphasis>
</taxonomicName>
into the 
<taxonomicName id="4C3690CAFFE2FFE7FCB2F978FCC138E0" baseAuthorityName="Kolbe" baseAuthorityYear="1880" box="[773,840,1767,1790]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="29" pageNumber="498" phylum="Arthropoda" rank="species" species="majus">
<emphasis id="B942375BFFE2FFE7FCB2F978FCC138E0" box="[773,840,1767,1790]" italics="true" pageId="29" pageNumber="498">majus</emphasis>
</taxonomicName>
group is excluded. However, this species cannot be assigned to the 
<emphasis id="B942375BFFE2FFE7FEF5F894FE55393C" box="[322,476,1803,1826]" italics="true" pageId="29" pageNumber="498">sexpunctatum</emphasis>
, 
<taxonomicName id="4C3690CAFFE2FFE7FE5DF895FDCA393D" authorityName="Thornton" authorityYear="1961" box="[490,579,1802,1827]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="29" pageNumber="498" phylum="Arthropoda" rank="species" species="medium">
<emphasis id="B942375BFFE2FFE7FE5DF895FDCA393D" box="[490,579,1802,1827]" italics="true" pageId="29" pageNumber="498">medium</emphasis>
</taxonomicName>
or New World species groups because of lack of apomorphies characterizing any of these species group so that a new species group for its own is proposed here. Females of this species are unknown to date.
</paragraph>
</subSubSection>
</treatment>
</document>