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<document id="D09D3F6EC5EE3A78EA8B1BE29ED9297B" ID-CLB-Dataset="270787" ID-DOI="10.11646/zootaxa.5350.1.1" ID-GBIF-Dataset="503ff6a9-9260-469d-af97-456e103cbfae" ID-ISSN="1175-5334" ID-ZooBank="23B4C9BF-A1A4-4D27-8C30-E4A23DAD01AB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695987895977" checkinUser="plazi" docAuthor="Saad, Adib, Çiçek, Erdoğan, Esmaeili, Hamid Reza, Fricke, Ronald, Sungur, Sevil &amp; Eagderi, Soheil" docDate="2023" docId="039B4C3C2E04FF90FF42F88DE0FEFDAF" docLanguage="en" docName="zootaxa.5350.1.1.pdf" docOrigin="Zootaxa 5350 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5350.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Carasobarbus chantrei" docType="treatment" docVersion="5" lastPageNumber="17" masterDocId="FFA234442E0BFF80FFD5FFE2E357FFCE" masterDocTitle="Freshwater fishes of Syria: a revised and updated annotated checklist- 2023" masterLastPageNumber="62" masterPageNumber="1" pageNumber="16" updateTime="1699692675315" updateUser="plazi">
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<mods:title id="28ABFF67A3E00A0F2A73AFFC6DC8C282">Freshwater fishes of Syria: a revised and updated annotated checklist- 2023</mods:title>
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<mods:namePart id="47F5FF094797131123D2295294751A8F">Saad, Adib</mods:namePart>
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<mods:affiliation id="525C12B2D22A3F02D9DBE0FF212504C0">Al Andalus University, Kadmus, Tartus, Syria / Syrian Society for Aquatic Environment Protection, Lattakia, Syria. https: // orcid. org / 0000 - 0001 - 6393 - 2836</mods:affiliation>
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<mods:namePart id="5BC408C753D225A3E6AAB5C5FABA4D04">Çiçek, Erdoğan</mods:namePart>
<mods:affiliation id="8113EA065F76311EECDF8A18E460D149">Department of Biology, Faculty of Art and Sciences, Nevþehir Hacý BektaþVeli University, Nevþehir, T ¸ rkiye.</mods:affiliation>
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<mods:namePart id="6C8CA61B670E40F8929A8FFD16565DB8">Esmaeili, Hamid Reza</mods:namePart>
<mods:nameIdentifier id="898C6DBC98E20D18AC66491FAEF6BCA1" type="ORCID">0000-0002-9736-397X</mods:nameIdentifier>
<mods:affiliation id="A49E7BED98921970503951EA8F420F48">Ichthyology and Molecular Systematics Research Laboratory, Zoology Section, Department of Biology, School of Science, Shiraz University, Shiraz, Iran. https: // orcid. org / 0000 - 0002 - 9736 - 397 X</mods:affiliation>
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<mods:namePart id="90DE69F23BF0434A4C900F53BD766414">Fricke, Ronald</mods:namePart>
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<mods:affiliation id="C039D8EF9347E7C4CCC623C4365F9BF4">Staatliches Museum f ¸ r Naturkunde in Stuttgart, Rosenstein 1, 70191 Stuttgart, Germany. https: // orcid. org / 0000 - 0003 - 1476 - 6990</mods:affiliation>
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<mods:namePart id="CF796688C0B3A95DF66A4A07CFEAE55F">Sungur, Sevil</mods:namePart>
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<mods:affiliation id="8E25D0B4FF5187082541A79B4965ACF2">Health Services Vocational School, Nevþehir Hacý BektaþVeli University, Nevþehir, T ¸ rkiye. https: // orcid. org / 0000 - 0003 - 4018 - 6375</mods:affiliation>
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<mods:namePart id="0CA798C56C8C59FC68B1188FD96AC396">Eagderi, Soheil</mods:namePart>
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<mods:affiliation id="E2B3B0389D2D6155D171DB8F1845BA8E">Al Andalus University, Kadmus, Tartus, Syria / Syrian Society for Aquatic Environment Protection, Lattakia, Syria. https: // orcid. org / 0000 - 0001 - 6393 - 2836 &amp; Department of Fisheries, Faculty of Natural Resources, University of Tehran, Karaj, Iran. https: // orcid. org / 0000 - 0002 - 1134 - 0356 * Corresponding author: erdogancicek 50 @ gmail. com; https: // orcid. org / 0000 - 0002 - 5334 - 5737 &amp; Al Andalus University, Kadmus, Tartus, Syria / Syrian Society for Aquatic Environment Protection, Lattakia, Syria. https: // orcid. org / 0000 - 0001 - 6393 - 2836</mods:affiliation>
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<treatment id="039B4C3C2E04FF90FF42F88DE0FEFDAF" ID-GBIF-Taxon="213683870" LSID="urn:lsid:plazi:treatment:039B4C3C2E04FF90FF42F88DE0FEFDAF" httpUri="http://treatment.plazi.org/id/039B4C3C2E04FF90FF42F88DE0FEFDAF" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<subSubSection id="C328AEA12E04FF8FFF42F88DE0CAF844" box="[151,925,1903,1930]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8B8DFD2A2E04FF8FFF42F88DE0CAF844" blockId="15.[151,1437,1903,2039]" box="[151,925,1903,1930]" pageId="15" pageNumber="16">
<heading id="D0C54A462E04FF8FFF42F88DE0CAF844" box="[151,925,1903,1930]" fontSize="11" level="2" pageId="15" pageNumber="16" reason="5">
<taxonomicName id="4C3286A92E04FF8FFF42F88DE133F844" ID-CoL="R2ZT" ID-ENA="137083" authority="(Sauvage, 1882)" baseAuthorityName="Sauvage" baseAuthorityYear="1882" box="[151,612,1903,1930]" family="Cyprinidae" genus="Carasobarbus" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="chantrei">
<emphasis id="B94621382E04FF8FFF42F88DE133F844" bold="true" box="[151,612,1903,1930]" pageId="15" pageNumber="16">
<emphasis id="B94621382E04FF8FFF42F88DE2F7F844" bold="true" box="[151,416,1903,1930]" italics="true" pageId="15" pageNumber="16">Carasobarbus chantrei</emphasis>
(
<bibRefCitation id="EFA380DB2E04FF8FFE7AF892E10BF844" author="Sauvage, H. - E." box="[431,604,1904,1930]" pageId="15" pageNumber="16" pagination="163 - 168" refId="ref58872" refString="Sauvage, H. - E. (1882) Catalogue des poissons recueillis par M. E. Chantre pendant son voyage en Syrie, Haute-Mesopotamie, Kurdistan et Caucase. Bulletin de la Societe philomathique de Paris, Serie 7, 6, 163 - 168. https: // doi. org / 10.11646 / zootaxa. 4885.2.2" type="journal article" year="1882">Sauvage, 1882</bibRefCitation>
)
</emphasis>
</taxonomicName>
[N]—Orontes himri; Bonni
</heading>
</paragraph>
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<subSubSection id="C328AEA12E04FF90FF42F876E709FF37" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="8B8DFD2A2E04FF90FF42F876E709FF37" blockId="15.[151,1437,1903,2039]" lastBlockId="16.[151,1437,151,609]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="B94621382E04FF8FFF42F876E24DF860" bold="true" box="[151,282,1940,1966]" pageId="15" pageNumber="16">Taxonomy.</emphasis>
Original description:
<taxonomicName id="4C3286A92E04FF8FFDF0F876E74AF861" ID-CoL="3RJ44" authority="Sauvage, 1882: 165" authorityName="Sauvage" authorityPageNumber="165" authorityYear="1882" box="[549,1053,1940,1967]" family="Cyprinidae" genus="Labeobarbus" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="chantrei">
<emphasis id="B94621382E04FF8FFDF0F876E074F860" box="[549,803,1940,1966]" italics="true" pageId="15" pageNumber="16">Labeobarbus chantrei</emphasis>
<bibRefCitation id="EFA380DB2E04FF8FFCE4F876E74AF861" author="Sauvage, H. - E." box="[817,1053,1940,1967]" pageId="15" pageNumber="16" pagination="163 - 168" refId="ref58872" refString="Sauvage, H. - E. (1882) Catalogue des poissons recueillis par M. E. Chantre pendant son voyage en Syrie, Haute-Mesopotamie, Kurdistan et Caucase. Bulletin de la Societe philomathique de Paris, Serie 7, 6, 163 - 168. https: // doi. org / 10.11646 / zootaxa. 4885.2.2" type="journal article" year="1882">Sauvage, 1882: 165</bibRefCitation>
</taxonomicName>
[Antakya Lake (= Amik Ĝlü) (
<geoCoordinate id="EE069BED2E04FF8FFF39F85AE222F81D" box="[236,373,1976,2003]" degrees="36" direction="north" minutes="12" orientation="latitude" pageId="15" pageNumber="16" precision="15" seconds="14" value="36.203888">36°12’14”N</geoCoordinate>
,
<geoCoordinate id="EE069BED2E04FF8FFE57F85AE15CF81C" box="[386,523,1976,2002]" degrees="36" direction="east" minutes="09" orientation="longitude" pageId="15" pageNumber="16" precision="15" seconds="26" value="36.157223">36°09’26”E</geoCoordinate>
),
<collectingRegion id="49F633C82E04FF8FFDCBF85BE19AF81C" box="[542,717,1976,2003]" country="Turkey" name="Hatay" pageId="15" pageNumber="16">Hatay Province</collectingRegion>
,
<collectingCountry id="F325BDBA2E04FF8FFD02F85AE07CF81C" box="[727,811,1976,2002]" name="Turkey" pageId="15" pageNumber="16">Turkey</collectingCountry>
;
<typeStatus id="548943882E04FF8FFCE0F85AE0F7F81C" box="[821,928,1976,2002]" pageId="15" pageNumber="16" type="lectotype">lectotype</typeStatus>
: MNHN A-3866;
<typeStatus id="548943882E04FF8FFBB3F85AE79AF81C" box="[1126,1229,1976,2002]" pageId="15" pageNumber="16" type="lectotype">lectotype</typeStatus>
selected by
<bibRefCitation id="EFA380DB2E04FF8FFA80F85BE208F839" author="Krupp, F." pageId="15" pageNumber="16" pagination="17 - 25" refId="ref56231" refString="Krupp, F. (1985 b) Barbus chantrei (Sauvage 1882), a valid species of cyprinid fish from the northern Levant. Senckenbergiana Biologica, 66, 17 - 25." type="journal article" year="1985">Krupp (1985b:19)</bibRefCitation>
and again later (same specimen) by
<bibRefCitation id="EFA380DB2E04FF8FFCCAF83EE7C9F838" author="Ekmekci, F. G. &amp; Banarescu P. M." box="[799,1182,2012,2038]" pageId="15" pageNumber="16" pagination="87 - 96" refId="ref52820" refString="Ekmekci, F. G. &amp; Banarescu P. M. (1998) A revision of the generic position of Barynotus (Systomus) verhoeffi, and the validity of the genera Carasobarbus, Kosswigobarbus and Mesopotamichthys (Pisces, Cyprinidae). Folia Zoologica: International Journal of Vertebrate Zoology, 47 (Supplement 1), 87 - 96." type="journal article" year="1998">Ekmekçi &amp; Bănărescu (1998:92)</bibRefCitation>
].—Syrian synonyms:
<taxonomicName id="4C3286A92E1BFF90FF36FF75E133FF7F" ID-CoL="KN54" authority="(Sauvage, 1882)" baseAuthorityName="Sauvage" baseAuthorityYear="1882" box="[227,612,151,177]" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="chantrei">
<emphasis id="B94621382E1BFF90FF36FF75E2CCFF7F" box="[227,411,151,177]" italics="true" pageId="16" pageNumber="17">Barbus chantrei</emphasis>
(
<bibRefCitation id="EFA380DB2E1BFF90FE7BFF75E10BFF7F" author="Sauvage, H. - E." box="[430,604,151,177]" pageId="16" pageNumber="17" pagination="163 - 168" refId="ref58872" refString="Sauvage, H. - E. (1882) Catalogue des poissons recueillis par M. E. Chantre pendant son voyage en Syrie, Haute-Mesopotamie, Kurdistan et Caucase. Bulletin de la Societe philomathique de Paris, Serie 7, 6, 163 - 168. https: // doi. org / 10.11646 / zootaxa. 4885.2.2" type="journal article" year="1882">Sauvage, 1882</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C3286A92E1BFF90FDA1FF75E05CFF7E" ID-CoL="KN45" authorityName="Valenciennes" authorityYear="1842" box="[628,779,151,177]" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="canis">
<emphasis id="B94621382E1BFF90FDA1FF75E05CFF7E" box="[628,779,151,177]" italics="true" pageId="16" pageNumber="17">Barbus canis</emphasis>
</taxonomicName>
(non-Valenciennes, 1842);
<taxonomicName id="4C3286A92E1BFF90FB98FF75E7EEFF7E" ID-CoL="7CHC6" baseAuthorityName="Valenciennes" baseAuthorityYear="1842" box="[1101,1209,151,176]" family="Cyprinidae" genus="Tor" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="canis">
<emphasis id="B94621382E1BFF90FB98FF75E7EEFF7E" box="[1101,1209,151,176]" italics="true" pageId="16" pageNumber="17">Tor canis</emphasis>
</taxonomicName>
(non Valenciennes, 1842).—Revisions:
<bibRefCitation id="EFA380DB2E1BFF90FE1CFF59E002FF1B" author="Ekmekci, F. G. &amp; Banarescu P. M." box="[457,853,187,213]" pageId="16" pageNumber="17" pagination="87 - 96" refId="ref52820" refString="Ekmekci, F. G. &amp; Banarescu P. M. (1998) A revision of the generic position of Barynotus (Systomus) verhoeffi, and the validity of the genera Carasobarbus, Kosswigobarbus and Mesopotamichthys (Pisces, Cyprinidae). Folia Zoologica: International Journal of Vertebrate Zoology, 47 (Supplement 1), 87 - 96." type="journal article" year="1998">Ekmekçi &amp; Bănărescu (1998: 92)</bibRefCitation>
;
<bibRefCitation id="EFA380DB2E1BFF90FCB0FF59E781FF1B" author="Borkenhagen, K. &amp; Esmaeili, H. R. &amp; Mohsenzadeh, S. &amp; Shahryari, F. &amp; Gholamifard, A." box="[869,1238,187,213]" pageId="16" pageNumber="17" pagination="327 - 335" refId="ref51292" refString="Borkenhagen, K., Esmaeili, H. R. Mohsenzadeh, S., Shahryari, F. &amp; Gholamifard, A. (2011) The molecular systematics of the Carasobarbus species from Iran and adjacent areas, with comments on Carasobarbus albus (Heckel, 1843). Environmental Biology of Fishes, 91, 327 - 335. https: // doi. org / 10.1007 / s 10641 - 011 - 9787 - 1" type="journal article" year="2011">
Borkenhagen
<emphasis id="B94621382E1BFF90FBD0FF59E715FF1B" box="[1029,1090,187,213]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
(2011: 328)
</bibRefCitation>
;
<bibRefCitation id="EFA380DB2E1BFF90FB33FF59E2F4FF37" author="Borkenhagen, K. &amp; Krupp, F." pageId="16" pageNumber="17" pagination="1 - 53" refId="ref51098" refString="Borkenhagen, K. &amp; Krupp, F. (2013) Taxonomic revision of the genus Carasobarbus Karaman, 1971 (Actinopterygii, Cyprinidae). ZooKeys, 339, 1 - 53. https: // doi. org / 10.3897 / zookeys. 339.4903" type="journal article" year="2013">Borkenhagen &amp; Krupp (2013: 17)</bibRefCitation>
.—Illustration:
<bibRefCitation id="EFA380DB2E1BFF90FD86FF3DE09EFF37" author="Borkenhagen, K. &amp; Krupp, F." box="[595,969,223,249]" pageId="16" pageNumber="17" pagination="1 - 53" refId="ref51098" refString="Borkenhagen, K. &amp; Krupp, F. (2013) Taxonomic revision of the genus Carasobarbus Karaman, 1971 (Actinopterygii, Cyprinidae). ZooKeys, 339, 1 - 53. https: // doi. org / 10.3897 / zookeys. 339.4903" type="journal article" year="2013">Borkenhagen &amp; Krupp (2013: 19</bibRefCitation>
, figs. 10-11).
</paragraph>
</subSubSection>
<subSubSection id="C328AEA12E1BFF90FF42FEE0E665FE8F" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8B8DFD2A2E1BFF90FF42FEE0E665FE8F" blockId="16.[151,1437,151,609]" pageId="16" pageNumber="17">
<emphasis id="B94621382E1BFF90FF42FEE0E207FED2" bold="true" box="[151,336,258,284]" pageId="16" pageNumber="17">
Status in
<collectingCountry id="F325BDBA2E1BFF90FEDCFEE0E21CFED2" box="[265,331,258,284]" name="Syria" pageId="16" pageNumber="17">Syria</collectingCountry>
.
</emphasis>
First record from Syria by
<bibRefCitation id="EFA380DB2E1BFF90FD5CFEE1E03EFED3" author="Beckman, W. C." box="[649,873,259,285]" pageId="16" pageNumber="17" refId="ref50605" refString="Beckman, W. C. (1962) The Freshwater Fishes of Syria and their General Biology and Management. FAO, Rome, 297 pp." type="book" year="1962">Beckman (1962:98)</bibRefCitation>
as
<taxonomicName id="4C3286A92E1BFF90FC44FEE1E711FED3" baseAuthorityName="Sauvage" baseAuthorityYear="1882" box="[913,1094,259,285]" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="chantrei">
<emphasis id="B94621382E1BFF90FC44FEE1E711FED3" box="[913,1094,259,285]" italics="true" pageId="16" pageNumber="17">Barbus chantrei</emphasis>
</taxonomicName>
; confirmed by
<bibRefCitation id="EFA380DB2E1BFF90FB20FEE1E3AFFE8F" author="Krupp, F." pageId="16" pageNumber="17" pagination="17 - 25" refId="ref56231" refString="Krupp, F. (1985 b) Barbus chantrei (Sauvage 1882), a valid species of cyprinid fish from the northern Levant. Senckenbergiana Biologica, 66, 17 - 25." type="journal article" year="1985">Krupp (1985b: 18)</bibRefCitation>
;
<bibRefCitation id="EFA380DB2E1BFF90FED0FEC5E29BFE8F" author="Saad, A. &amp; Khalaf, G. &amp; Alzein, G. &amp; Abdellatif, A. &amp; Sabour, W. &amp; Alhawi, I. &amp; Hammoud, N." box="[261,460,295,321]" pageId="16" pageNumber="17" refId="ref58691" refString="Saad, A., Khalaf, G., Alzein, G., Abdellatif, A., Sabour, W., Alhawi, I. &amp; Hammoud, N. (2009) An updated check list of Fish in the Orontes River Basin within Syria and Lebanon. Final report of the joint Lebanese-Syrian scientific research project, Contract No. 119 of 2006. s. n., s. l., 62 pp. [in Arabic with an English summary]" type="book" year="2009">
Saad
<emphasis id="B94621382E1BFF90FE97FEC5E22CFE8F" box="[322,379,295,321]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
(2009)
</bibRefCitation>
as
<taxonomicName id="4C3286A92E1BFF90FE24FEC5E1F2FE8F" baseAuthorityName="Sauvage" baseAuthorityYear="1882" box="[497,677,295,321]" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="chantrei">
<emphasis id="B94621382E1BFF90FE24FEC5E1F2FE8F" box="[497,677,295,321]" italics="true" pageId="16" pageNumber="17">Barbus chantrei</emphasis>
</taxonomicName>
.—Syrian material: BMNH, CMN, MNHN, SMNS, MSL.
</paragraph>
</subSubSection>
<subSubSection id="C328AEA12E1BFF90FF42FEA8E753FE3B" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8B8DFD2A2E1BFF90FF42FEA8E1E6FE1F" blockId="16.[151,1437,151,609]" pageId="16" pageNumber="17">
<emphasis id="B94621382E1BFF90FF42FEA8E297FEAA" bold="true" box="[151,448,330,357]" pageId="16" pageNumber="17">Distribution and habitat.</emphasis>
Distribution in
<collectingCountry id="F325BDBA2E1BFF90FDA5FEA9E1F8FEAB" box="[624,687,331,357]" name="Syria" pageId="16" pageNumber="17">Syria</collectingCountry>
: Orontes River system.—Distribution in River Basin: 4-Orontes.— General distribution:
<collectingCountry id="F325BDBA2E1BFF90FE0FFE8DE108FE47" box="[474,607,367,393]" name="Turkey" pageId="16" pageNumber="17">Asia Minor</collectingCountry>
and Middle East:
<collectingCountry id="F325BDBA2E1BFF90FCEAFE8DE0D8FE47" box="[831,911,367,393]" name="Turkey" pageId="16" pageNumber="17">Turkey</collectingCountry>
and
<collectingCountry id="F325BDBA2E1BFF90FC1AFE8DE75DFE47" box="[975,1034,367,393]" name="Syria" pageId="16" pageNumber="17">Syria</collectingCountry>
.—Distribution in Ecoregion: 437- Orontes.—Habitat: This species inhabits lowland rivers, backwaters, lakes, reservoirs, springs, and ponds, rarely in fast-flowing waters. Freshwater.
</paragraph>
<paragraph id="8B8DFD2A2E1BFF90FF42FE38E753FE3B" blockId="16.[151,1437,151,609]" box="[151,1028,474,501]" pageId="16" pageNumber="17">
<emphasis id="B94621382E1BFF90FF42FE38E2F3FE3A" bold="true" box="[151,420,474,500]" pageId="16" pageNumber="17">Economic importance.</emphasis>
Locally consumed, but of no commercial importance.
</paragraph>
</subSubSection>
<subSubSection id="C328AEA12E1BFF90FF42FE1CE0FEFDAF" pageId="16" pageNumber="17" type="conservation">
<paragraph id="8B8DFD2A2E1BFF90FF42FE1CE0FEFDAF" blockId="16.[151,1437,151,609]" pageId="16" pageNumber="17">
<emphasis id="B94621382E1BFF90FF42FE1CE268FDD6" bold="true" box="[151,319,510,536]" pageId="16" pageNumber="17">Conservation.</emphasis>
Conservation status in
<collectingCountry id="F325BDBA2E1BFF90FD84FE1DE1C7FDD7" box="[593,656,511,537]" name="Syria" pageId="16" pageNumber="17">Syria</collectingCountry>
: Unknown.—IUCN: NT (
<bibRefCitation id="EFA380DB2E1BFF90FC69FE1DE711FDD7" author="IUCN" box="[956,1094,511,537]" pageId="16" pageNumber="17" refId="ref55612" refString="IUCN (2023) The IUCN Red List of Threatened Species. Version 2022 - 2. Available from: https: // www. iucnredlist. org / (accessed date 12 July 2023)" type="url" year="2023">IUCN 2023</bibRefCitation>
).—Threats: ABS, CLI, CON, EUT, HAB.—Moderate sensitivity to human activities.—Not considered as a keystone species.—Decline status: Decreasing.—Moderate priority for conservation action.
</paragraph>
</subSubSection>
</treatment>
</document>
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