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<document id="DEE185AB2A55F0BDDFAA748A4243711C" ID-CLB-Dataset="29018" ID-DOI="10.11646/zootaxa.4459.2.8" ID-GBIF-Dataset="f36314b8-a9a6-4bd2-8704-8923013c22b7" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458709" ID-ZooBank="7270DEAE-A08F-4015-8FE6-6E0D0DBD9C84" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539289409860" checkinUser="plazi" docAuthor="Kаtаev, Boris M. &amp; Wrаse, Dаvid W." docDate="2018" docId="03952C24744BFFC5FF2CFC52FCADDEC2" docLanguage="en" docName="zootaxa.4459.2.8.pdf" docOrigin="Zootaxa 4459 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Proditomus Schauberger 1934" docType="treatment" docVersion="7" lastPageNumber="360" masterDocId="FFAC545C7441FFCFFFBBFFD4FF8FD92B" masterDocTitle="Notes on Ditomina, with description of two new genera and a new species (Coleoptera, Carabidae) from the Middle East and India" masterLastPageNumber="368" masterPageNumber="350" pageNumber="360" updateTime="1698578541753" updateUser="ExternalLinkService">
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<mods:title id="B45F3B3657380ABFBCEE4830C3224865">Notes on Ditomina, with description of two new genera and a new species (Coleoptera, Carabidae) from the Middle East and India</mods:title>
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<mods:namePart id="532A51FA46E6479E4DBB08A6DFDA135C">Kаtаev, Boris M.</mods:namePart>
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<mods:namePart id="58FBC98D740BCA2195B2DC5499733BBE">Wrаse, Dаvid W.</mods:namePart>
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<treatment id="03952C24744BFFC5FF2CFC52FCADDEC2" ID-DOI="http://doi.org/10.5281/zenodo.5980143" ID-GBIF-Taxon="148692759" ID-Zenodo-Dep="5980143" LSID="urn:lsid:plazi:treatment:03952C24744BFFC5FF2CFC52FCADDEC2" httpUri="http://treatment.plazi.org/id/03952C24744BFFC5FF2CFC52FCADDEC2" lastPageNumber="360" pageId="10" pageNumber="360">
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<paragraph id="8B839D32744BFFC5FF2CFC52FD9EDA8B" blockId="10.[151,529,902,928]" box="[151,529,902,928]" pageId="10" pageNumber="360">
<heading id="D0CB2A5E744BFFC5FF2CFC52FD9EDA8B" bold="true" box="[151,529,902,928]" fontSize="11" level="1" pageId="10" pageNumber="360" reason="1">
<taxonomicName id="4C3CE6B1744BFFC5FF2CFC52FD9EDA8B" ID-CoL="8KYJV" authority="Schauberger, 1934" authorityName="Schauberger" authorityYear="1934" box="[151,529,902,928]" class="Insecta" family="Carabidae" genus="Proditomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="360" phylum="Arthropoda" rank="genus">
<emphasis id="B9484120744BFFC5FF2CFC52FD9EDA8B" bold="true" box="[151,529,902,928]" pageId="10" pageNumber="360">
<emphasis id="B9484120744BFFC5FF2CFC52FEACDA8B" bold="true" box="[151,291,902,928]" italics="true" pageId="10" pageNumber="360">Proditomus</emphasis>
Schauberger, 1934
</emphasis>
</taxonomicName>
</heading>
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<subSubSection id="C326CEB9744BFFC5FF2CFC19FD80DAC8" box="[151,527,973,995]" pageId="10" pageNumber="360" type="reference_group">
<paragraph id="8B839D32744BFFC5FF2CFC19FD80DAC8" blockId="10.[151,527,973,995]" box="[151,527,973,995]" pageId="10" pageNumber="360">
<treatmentCitationGroup id="AB2CBA1C744BFFC5FF2CFC19FD80DAC8" box="[151,527,973,995]" pageId="10" pageNumber="360">
<taxonomicName id="4C3CE6B1744BFFC5FF2CFC19FD80DAC8" ID-CoL="8KYJV" authority="Schauberger, 1934 b: 232" authorityName="Schauberger" authorityPageNumber="232" authorityYear="1934" box="[151,527,973,995]" class="Insecta" family="Carabidae" genus="Proditomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="360" phylum="Arthropoda" rank="genus">
<emphasis id="B9484120744BFFC5FF2CFC19FE83DAC8" box="[151,268,973,995]" italics="true" pageId="10" pageNumber="360">Proditomus</emphasis>
<treatmentCitation id="0A9DBB23744BFFC5FEA9FC19FD80DAC8" author="" box="[274,527,973,995]" page="232" pageId="10" pageNumber="360">
<bibRefCitation id="EFADE0C3744BFFC5FEA9FC19FE50DAC8" box="[274,479,973,995]" pageId="10" pageNumber="360" refId="ref13722" refString="Schauberger, E. (1934 b) Zur Kenntnis der palaarktischen Harpalinen (Col.). Stylops, 3, 232 - 240. https: // doi. org / 10.1111 / j. 1365 - 3113.1934. tb 01528. x" type="journal article">Schauberger, 1934b</bibRefCitation>
: 232
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</paragraph>
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<subSubSection id="C326CEB9744BFFC5FF2CFBDFFC1BDD0B" box="[151,916,1034,1056]" pageId="10" pageNumber="360" type="materials_examined">
<paragraph id="8B839D32744BFFC5FF2CFBDFFC1BDD0B" blockId="10.[151,916,1034,1056]" box="[151,916,1034,1056]" pageId="10" pageNumber="360">
Type species
<taxonomicName id="4C3CE6B1744BFFC5FEA4FBDEFD16DD0B" authority="Schauberger, 1934" authorityName="Schauberger" authorityYear="1934" box="[287,665,1034,1056]" class="Insecta" family="Carabidae" genus="Proditomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="360" phylum="Arthropoda" rank="species" species="mirus">
<emphasis id="B9484120744BFFC5FEA4FBDEFE5BDD0B" box="[287,468,1034,1056]" italics="true" pageId="10" pageNumber="360">Proditomus mirus</emphasis>
Schauberger, 1934
</taxonomicName>
, by original designation.
</paragraph>
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<subSubSection id="C326CEB9744BFFC5FF2CFB99FCD2DD85" pageId="10" pageNumber="360" type="diagnosis">
<paragraph id="8B839D32744BFFC5FF2CFB99FCD2DD85" blockId="10.[151,1437,1101,2026]" pageId="10" pageNumber="360">
<emphasis id="B9484120744BFFC5FF2CFB99FE9EDD4D" bold="true" box="[151,273,1101,1126]" pageId="10" pageNumber="360">Diagnosis.</emphasis>
<taxonomicName id="4C3CE6B1744BFFC5FEA3FB99FE15DD4D" authorityName="Schauberger" authorityYear="1934" box="[280,410,1101,1126]" class="Insecta" family="Carabidae" genus="Proditomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="360" phylum="Arthropoda" rank="genus">
<emphasis id="B9484120744BFFC5FEA3FB99FE15DD4D" box="[280,410,1101,1126]" italics="true" pageId="10" pageNumber="360">Proditomus</emphasis>
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is similar to
<taxonomicName id="4C3CE6B1744BFFC5FD91FB9AFC9CDD4D" authority="Billberg, 1820" authorityName="Billberg" authorityYear="1820" box="[554,787,1101,1126]" class="Insecta" family="Carabidae" genus="Dixus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="360" phylum="Arthropoda" rank="genus">
<emphasis id="B9484120744BFFC5FD91FB9AFDE4DD4E" box="[554,619,1102,1125]" italics="true" pageId="10" pageNumber="360">Dixus</emphasis>
Billberg,
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in head wide, without neck constriction and mentum with long median tooth, but differs in having anterior margin of clypeus deeply concave, labrum with deep emargination anteriorly, bilobed, and sides of pronotum not bordered basally.
</paragraph>
</subSubSection>
<subSubSection id="C326CEB9744BFFC5FF7CFB6DFA89DE89" pageId="10" pageNumber="360" type="description">
<paragraph id="8B839D32744BFFC5FF7CFB6DFD23DE55" blockId="10.[151,1437,1101,2026]" pageId="10" pageNumber="360">
<emphasis id="B9484120744BFFC5FF7CFB6DFED6DDF9" bold="true" box="[199,345,1209,1234]" pageId="10" pageNumber="360">Description.</emphasis>
Body (
<figureCitation id="130781B7744BFFC5FE14FB6DFD8CDDF9" box="[431,515,1209,1234]" captionStart="FIGURES 2529" captionStartId="12.[151,264,1946,1969]" captionTargetBox="[189,1404,537,1897]" captionTargetId="figure@12.[151,1436,517,1925]" captionTargetPageId="12" captionText="FIGURES 2529. Proditomus mirus Schauberger, 1934 (holotype). 25, labium; 26, right laterotergite and stylus; 27, stylus; 28, habitus; 29, left metepisternum. 25, 26, 29, ventral view; 27, lateral view; 28, dorsal view. Scale bars = 1.0 mm (A: Fig. 25, 29; C: Fig. 28), 0.5 mm (B: Figs 26, 27)." httpUri="https://zenodo.org/record/1458723/files/figure.png" pageId="10" pageNumber="360">Fig. 28</figureCitation>
) punctate, superior surface very convex, covered with very fine, scattered setae, obviously most of them rubbed off (as a very old specimen), and hence difficult to observe. Punctation on head and pronotum coarse and rather dense, on elytra fine and sparse. Head large, as wide as pronotum, without neck constriction and with only one long seta (supraorbital setigerous pore) near inner margin of eyes on each side. Lateral margin of frons above insertion of antennae somewhat roundly widened forming an angulate projection on each side before eye. Eyes small, noticeably removed from buccal fissure ventrally. Tempora long, slightly convex, evenly curved to neck. Labrum with deep emargination anteriorly, bilobed. Аnterior margin of clypeus deeply concave at middle. Mentum (
<figureCitation id="130781B7744BFFC5FE66FA61FDA2DCE5" box="[477,557,1461,1486]" captionStart="FIGURES 2529" captionStartId="12.[151,264,1946,1969]" captionTargetBox="[189,1404,537,1897]" captionTargetId="figure@12.[151,1436,517,1925]" captionTargetPageId="12" captionText="FIGURES 2529. Proditomus mirus Schauberger, 1934 (holotype). 25, labium; 26, right laterotergite and stylus; 27, stylus; 28, habitus; 29, left metepisternum. 25, 26, 29, ventral view; 27, lateral view; 28, dorsal view. Scale bars = 1.0 mm (A: Fig. 25, 29; C: Fig. 28), 0.5 mm (B: Figs 26, 27)." httpUri="https://zenodo.org/record/1458723/files/figure.png" pageId="10" pageNumber="360">Fig. 25</figureCitation>
) with large median tooth, without setae. Epilobes wide, prolonged medially up to tooth apex where they connect with each other. Ligular sclerite very wide, long, its sides roundly converging anteriorly and apex shortly sharpened, apicad distinctly deflexed; dorsal side with numerous rather long setae, ventrally unsetose. Paraglossae very small and narrow, much shorter than ligular sclerite, glabrous. Basal palpomere not carinate. Mandibles long, evenly curved, with acute apices only slightly blunted at tips. Аntennae slender, with antennomeres narrow, much longer than wide. Pronotum cordate, strongly narrowed posteriorly, almost parallel-sided in apical third and before almost right basal angles, with apical angles acute, projecting anteriad. Pronotal apical and basal margins not bordered, sides very narrowly bordered in apical half only. Mesothorax and extreme base of elytra narrowed forming a short neck-like constriction with scutellum above. Elytral base not bordered, sides border beginning from base of stria 7. Metacoxa without a posteromedial setigerous pore, punctate. Profemur not denticulate along inner margin. Protibia with prominent outer angle and with a transverse row of strong ventroapical setae. Metatrochanter short, about 0.3 times as long as metafemur. Metatarsomere 1 approximately as long as 2+3.
</paragraph>
<paragraph id="8B839D32744BFFC5FF7CF85DFA89DE89" blockId="10.[151,1437,1101,2026]" box="[199,1286,1929,1954]" pageId="10" pageNumber="360">
<emphasis id="B9484120744BFFC5FF7CF85DFDA8DE89" bold="true" box="[199,551,1929,1954]" pageId="10" pageNumber="360">Composition and distribution</emphasis>
. Includes only one species,
<taxonomicName id="4C3CE6B1744BFFC5FCD8F85DFBA3DE8A" authorityName="Schauberger" authorityYear="1934" box="[867,1068,1929,1954]" class="Insecta" family="Carabidae" genus="Proditomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="360" phylum="Arthropoda" rank="species" species="mirus">
<emphasis id="B9484120744BFFC5FCD8F85DFBA3DE8A" box="[867,1068,1929,1954]" italics="true" pageId="10" pageNumber="360">Proditomus mirus</emphasis>
</taxonomicName>
, from “Turkestan”.
</paragraph>
</subSubSection>
<subSubSection id="C326CEB9744BFFC5FF7CF879FCADDEC2" pageId="10" pageNumber="360" type="discussion">
<paragraph id="8B839D32744BFFC5FF7CF879FCADDEC2" blockId="10.[151,1437,1101,2026]" pageId="10" pageNumber="360">
<emphasis id="B9484120744BFFC5FF7CF879FEB4DEED" bold="true" box="[199,315,1965,1990]" pageId="10" pageNumber="360">Remarks.</emphasis>
Re-examination of the
<typeStatus id="54872390744BFFC5FDFAF879FD2CDEED" box="[577,675,1965,1990]" pageId="10" pageNumber="360" type="holotype">holotype</typeStatus>
confirmed Schaubergers opinion (1934) that this taxon is closest to
<taxonomicName id="4C3CE6B1744BFFC5FF2CF806FE0FDEC1" authority="Billberg, 1820" authorityName="Billberg" authorityYear="1820" box="[151,384,2001,2026]" class="Insecta" family="Carabidae" genus="Dixus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="360" phylum="Arthropoda" rank="genus">
<emphasis id="B9484120744BFFC5FF2CF806FF57DEC2" box="[151,216,2002,2025]" italics="true" pageId="10" pageNumber="360">Dixus</emphasis>
Billberg, 1820
</taxonomicName>
and warrants separate generic status.
</paragraph>
</subSubSection>
</treatment>
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