441 lines
64 KiB
XML
441 lines
64 KiB
XML
<document id="102719748A985EC2F20B9C9BDE9A7D64" ID-CLB-Dataset="29018" ID-DOI="10.11646/zootaxa.4459.2.8" ID-GBIF-Dataset="f36314b8-a9a6-4bd2-8704-8923013c22b7" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458709" ID-ZooBank="7270DEAE-A08F-4015-8FE6-6E0D0DBD9C84" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539289409860" checkinUser="plazi" docAuthor="Kаtаev, Boris M. & Wrаse, Dаvid W." docDate="2018" docId="03952C247444FFC5FF2CFB20FC48DA13" docLanguage="en" docName="zootaxa.4459.2.8.pdf" docOrigin="Zootaxa 4459 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Indocarterus inexspectatus Kаtаev & Wrаse 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="359" masterDocId="FFAC545C7441FFCFFFBBFFD4FF8FD92B" masterDocTitle="Notes on Ditomina, with description of two new genera and a new species (Coleoptera, Carabidae) from the Middle East and India" masterLastPageNumber="368" masterPageNumber="350" pageNumber="355" updateTime="1698578541753" updateUser="ExternalLinkService">
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<mods:title id="401F3C696E9229FF3A79159AA463A92F">Notes on Ditomina, with description of two new genera and a new species (Coleoptera, Carabidae) from the Middle East and India</mods:title>
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<mods:namePart id="E2E8D966320F6A1DF2B658046C52F12C">Kаtаev, Boris M.</mods:namePart>
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<mods:namePart id="8857B45E05C6F3621F4292CE6DB25E64">Wrаse, Dаvid W.</mods:namePart>
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<mods:typeOfResource id="AF5FB5EDBA304B0FD7B9FA479E669CC9">text</mods:typeOfResource>
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<mods:title id="484B411619AB6D60CA4EF22370CB78D1">Zootaxa</mods:title>
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<mods:part id="7662B994473E42A5B97A116440CE14E9">
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<mods:date id="15F88B0B5CEBD0EACAFE326AA1D09B48">2018</mods:date>
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<mods:number id="6824580CD3BC9FD4E5F0F4F79D7D1AEC">2018-08-15</mods:number>
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<mods:number id="222C3FB6E9DF8A3B6C0E47D69853EF5F">4459</mods:number>
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<mods:classification id="498A39A4E7C9B8095B5A22F19243BD8D">journal article</mods:classification>
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<mods:identifier id="D1CC6757C51266FFA20776B8B3794CEB" type="DOI">10.11646/zootaxa.4459.2.8</mods:identifier>
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<treatment id="03952C247444FFC5FF2CFB20FC48DA13" ID-DOI="http://doi.org/10.5281/zenodo.5980141" ID-GBIF-Taxon="148692764" ID-Zenodo-Dep="5980141" LSID="urn:lsid:plazi:treatment:03952C247444FFC5FF2CFB20FC48DA13" httpUri="http://treatment.plazi.org/id/03952C247444FFC5FF2CFB20FC48DA13" lastPageId="10" lastPageNumber="359" pageId="5" pageNumber="355">
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<subSubSection id="C326CEB97444FFCAFF2CFB20FE96DC04" pageId="5" pageNumber="355" type="nomenclature">
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<paragraph id="8B839D327444FFCAFF2CFB20FDA9DC25" blockId="5.[151,550,1268,1327]" box="[151,550,1268,1294]" pageId="5" pageNumber="355">
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<heading id="D0CB2A5E7444FFCAFF2CFB20FDA9DC25" bold="true" box="[151,550,1268,1294]" fontSize="11" level="1" pageId="5" pageNumber="355" reason="1">
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<emphasis id="B94841207444FFCAFF2CFB20FDA9DC25" bold="true" box="[151,550,1268,1294]" pageId="5" pageNumber="355">
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<taxonomicName id="4C3CE6B17444FFCAFF2CFB20FE53DC25" ID-CoL="8M5WP" authority="Kаtаev & Wrаse, 2018" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[151,476,1268,1294]" class="Insecta" family="Carabidae" genus="Indocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="355" phylum="Arthropoda" rank="species" species="inexspectatus" status="sp. nov.">
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<emphasis id="B94841207444FFCAFF2CFB20FE53DC25" bold="true" box="[151,476,1268,1294]" italics="true" pageId="5" pageNumber="355">Indocarterus inexspectatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27BFC5B7444FFCAFE5FFB20FDA9DC25" box="[484,550,1268,1294]" pageId="5" pageNumber="355" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B839D327444FFCAFF2CFAC2FE96DC04" blockId="5.[151,550,1268,1327]" box="[151,281,1302,1327]" pageId="5" pageNumber="355">
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(
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<figureCitation id="130781B77444FFCAFF24FAC2FE9EDC04" box="[159,273,1302,1327]" captionStart-0="FIGURES 5, 6" captionStart-1="FIGURES 7–9" captionStart-2="FIGURES 10–17" captionStart-3="FIGURES 18–23" captionStartId-0="6.[151,264,1392,1415]" captionStartId-1="7.[151,264,1867,1890]" captionStartId-2="8.[151,264,1206,1229]" captionStartId-3="9.[151,264,1124,1147]" captionTargetBox-0="[192,1402,451,1356]" captionTargetBox-1="[234,1316,203,1823]" captionTargetBox-2="[170,1428,620,1169]" captionTargetBox-3="[185,1409,462,1046]" captionTargetId-0="figure@6.[163,1427,432,1382]" captionTargetId-1="figure@7.[151,1436,193,1846]" captionTargetId-2="figure@8.[151,1436,589,1185]" captionTargetId-3="figure@9.[162,1426,437,1111]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="FIGURES 5, 6. Indocarterus inexspectatus sp. n., habitus. 5, male holotype; 6, female paratype. Scale bar = 1.0 mm." captionText-1="FIGURES 7–9. Indocarterus inexspectatus sp. n. (paratypes: 7, 8 female; 9, male). 7, anterior part of head; 8, pronotum; 9, protarsus and apical part of protibia. 7, 8, dorsal view; 9, lateral view. Scales bars = 1.0 mm (A, C: Figs 7, 9; B: Fig. 8)." captionText-2="FIGURES 10–17. Indocarterus inexspectatus sp. n. (paratypes). 10, labium; 11, apex of median lobe; 12, 13, median lobe; 14, left paramere, inner surface; 15, left paramere, inner surface; 16, stylus, 17, right laterotergite and stylus. 10, 17, ventral view; 11, 13, dorsal view; 12, 14–16, lateral view. Scale bars: A = 1.0 mm (Fig. 10), B = 0.5 mm (Figs 11–16)." captionText-3="FIGURES 18–23. Indocarterus inexspectatus sp. n. (paratype), aedeagus. 18, 23, right paramere, outer surface; 19, left paramere, outer surface; 20–22, median lobe. 18–20, 22, 23, lateral view; 21, dorsal view. Scale bars = 1.0 mm (A: Figs 18–22, B: Fig. 23)." httpUri-0="https://zenodo.org/record/1458713/files/figure.png" httpUri-1="https://zenodo.org/record/1458715/files/figure.png" httpUri-2="https://zenodo.org/record/1458717/files/figure.png" httpUri-3="https://zenodo.org/record/1458719/files/figure.png" pageId="5" pageNumber="355">Figs 5–23</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C326CEB97444FFCAFF2CFA8AFB35DF2C" pageId="5" pageNumber="355" type="materials_examined">
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<paragraph id="8B839D327444FFCAFF2CFA8AFEE4DCB0" blockId="5.[151,1437,1374,2011]" pageId="5" pageNumber="355">
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<materialsCitation id="3B54976F7444FFCAFF2CFA8AFC26DC5D" ID-GBIF-Occurrence="1917355871" box="[151,937,1374,1399]" collectionCode="WAI" country="India" location="Maharashtra" pageId="5" pageNumber="355" specimenCount="1" specimenCount-male="1" stateProvince="Maharashtra" typeStatus="holotype">
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<emphasis id="B94841207444FFCAFF2CFA8AFEB1DC5C" bold="true" box="[151,318,1374,1399]" pageId="5" pageNumber="355">
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<typeStatus id="548723907444FFCAFF2CFA8AFF5DDC5C" box="[151,210,1374,1399]" pageId="5" pageNumber="355">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="548723907444FFCAFEF7FA8AFE36DC5C" box="[332,441,1374,1399]" pageId="5" pageNumber="355" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="9D3A56BB7444FFCAFE7FFA8BFE54DC5D" box="[452,475,1375,1398]" pageId="5" pageNumber="355" type="male">♂</specimenCount>
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, “
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<collectingCountry id="F32BDDA27444FFCAFE4AFA8BFDCCDC5C" box="[497,579,1375,1399]" name="India" pageId="5" pageNumber="355">INDIА</collectingCountry>
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occ.
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<collectingRegion id="49F853D07444FFCAFDC4FA8AFC84DC5C" box="[639,779,1374,1399]" country="India" name="Maharashtra" pageId="5" pageNumber="355">Maharashtra</collectingRegion>
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st. /
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<collectionCode id="ED2D05F77444FFCAFCF8FA8BFCF7DC5C" box="[835,888,1375,1399]" country="USA" httpUri="http://grbio.org/cool/143k-e2ry" name="Waimea Valley (Hi'ipaka LLC)" pageId="5" pageNumber="355">WАI</collectionCode>
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env
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||
</materialsCitation>
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.
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<materialsCitation id="3B54976F7444FFCAFC0FFA8AFEE8DCB0" ID-GBIF-Occurrence="1917355851" collectingDate="2005-10-03" collectingDateMax="2005-10-06" collectingDateMin="2005-10-03" collectionCode="SMNS" collectorName="F. & L. Kantner" country="India" location="Pune" pageId="5" pageNumber="355" specimenCount="1" stateProvince="Maharashtra" typeStatus="holotype">
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<collectingDate id="EFC6421A7444FFCAFC0FFA8AFBBEDC5C" box="[948,1073,1374,1399]" pageId="5" pageNumber="355" value="2005-10-03" valueMax="2005-10-06" valueMin="2005-10-03">
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<date id="FF82BBF27444FFCAFC0FFA8AFBBEDC5C" box="[948,1073,1374,1399]" pageId="5" pageNumber="355" value="2005-10-03" valueMax="2005-10-06" valueMin="2005-10-03">3–6.x.2005</date>
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</collectingDate>
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/
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<quantity id="4CC430D77444FFCAFBF3FA8BFB00DC5C" box="[1096,1167,1374,1399]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="355" unit="km" value="70.0">70 km</quantity>
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S of
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<location id="8EE3CBE97444FFCAFB77FA8BFA8BDC5D" LSID="urn:lsid:plazi:treatment:03952C247444FFC5FF2CFB20FC48DA13:8EE3CBE97444FFCAFB77FA8BFA8BDC5D" box="[1228,1284,1375,1398]" country="India" name="Pune" pageId="5" pageNumber="355" stateProvince="Maharashtra">Pune</location>
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/ leg.
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<collectorName id="26C9F8E47444FFCAFAF0FA8BFF7BDCB0" pageId="5" pageNumber="355">F. & L. Kantner</collectorName>
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” (
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<collectionCode id="ED2D05F77444FFCAFEB6FA57FED0DCB0" box="[269,351,1411,1435]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="5" pageNumber="355">SMNS</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B839D327444FFCAFF7CFA73FB35DF2C" blockId="5.[151,1437,1374,2011]" pageId="5" pageNumber="355">
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<materialsCitation id="3B54976F7444FFCAFF7CFA73FB6ADCC9" ID-GBIF-Occurrence="1917355887" collectingDate="2005-10-07" collectingDateMax="2005-10-11" collectingDateMin="2005-10-07" collectionCode="SMNS, ZIN" collectorName="F. Kantner" country="India" location="Pune" pageId="5" pageNumber="355" specimenCount="84" specimenCount-female="1" specimenCount-male="83" stateProvince="Maharashtra" typeStatus="paratype">
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<typeStatus id="548723907444FFCAFF7CFA73FEB6DC94" box="[199,313,1447,1471]" pageId="5" pageNumber="355" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D3A56BB7444FFCAFEFDFA73FE18DC95" box="[326,407,1447,1471]" pageId="5" pageNumber="355" type="male">83 ♂♂</specimenCount>
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and
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<specimenCount id="9D3A56BB7444FFCAFE6EFA73FD8FDC95" box="[469,512,1447,1470]" pageId="5" pageNumber="355" type="female">♀♀</specimenCount>
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, same data as holotype (
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||
<collectionCode id="ED2D05F77444FFCAFC9EFA73FCFADC94" box="[805,885,1447,1471]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="5" pageNumber="355">SMNS</collectionCode>
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,
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<collectionCode id="ED2D05F77444FFCAFC3EFA73FC38DC95" box="[901,951,1447,1470]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="355">ZIN</collectionCode>
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, cWR); 2 ex, “
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<collectingCountry id="F32BDDA27444FFCAFBD2FA73FB34DC94" box="[1129,1211,1447,1471]" name="India" pageId="5" pageNumber="355">INDIA</collectingCountry>
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||
occ.
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<collectingDate id="EFC6421A7444FFCAFB44FA72FA05DC95" box="[1279,1418,1446,1470]" pageId="5" pageNumber="355" value="2005-10-07" valueMax="2005-10-11" valueMin="2005-10-07">
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<date id="FF82BBF27444FFCAFB44FA72FA05DC95" box="[1279,1418,1446,1470]" pageId="5" pageNumber="355" value="2005-10-07" valueMax="2005-10-11" valueMin="2005-10-07">7–11.x.2005</date>
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</collectingDate>
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||
/
|
||
<collectingRegion id="49F853D07444FFCAFF2CFA1EFEACDCC8" box="[151,291,1482,1507]" country="India" name="Maharashtra" pageId="5" pageNumber="355">Maharashtra</collectingRegion>
|
||
state / MULSHI env.
|
||
<collectorName id="26C9F8E47444FFCAFDA2FA1FFD00DCC9" box="[537,655,1483,1506]" pageId="5" pageNumber="355">F. Kantner</collectorName>
|
||
leg. /
|
||
<quantity id="4CC430D77444FFCAFD6EFA1FFC94DCC8" box="[725,795,1482,1507]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="355" unit="km" value="40.0">40 km</quantity>
|
||
W of
|
||
<location id="8EE3CBE97444FFCAFCD9FA1FFC13DCC8" LSID="urn:lsid:plazi:treatment:03952C247444FFC5FF2CFB20FC48DA13:8EE3CBE97444FFCAFCD9FA1FFC13DCC8" box="[866,924,1483,1507]" country="India" name="Pune" pageId="5" pageNumber="355" stateProvince="Maharashtra">Pune</location>
|
||
” (
|
||
<collectionCode id="ED2D05F77444FFCAFC0EFA1FFB88DCC8" box="[949,1031,1483,1507]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="5" pageNumber="355">SMNS</collectionCode>
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); 2 ex, “
|
||
<collectingCountry id="F32BDDA27444FFCAFBD9FA1FFB3BDCC8" box="[1122,1204,1483,1507]" name="India" pageId="5" pageNumber="355">INDIА</collectingCountry>
|
||
occ
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B54976F7444FFCAFB4BFA1EFB35DF2C" ID-GBIF-Occurrence="1917355838" collectionCode="SMNS" collectorName="Bhushi Dam & F. & L. Kantner" country="India" elevation="500" location="Lonavala" pageId="5" pageNumber="355" specimenCount="1" stateProvince="Maharashtra" typeStatus="paratype">
|
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<collectingRegion id="49F853D07444FFCAFB4BFA1EFAF3DCC8" box="[1264,1404,1482,1507]" country="India" name="Maharashtra" pageId="5" pageNumber="355">Maharashtra</collectingRegion>
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st. /
|
||
<collectorName id="26C9F8E47444FFCAFF1EFA3AFEBDDF2D" box="[165,306,1518,1543]" pageId="5" pageNumber="355">Bhushi Dam</collectorName>
|
||
env. 12–15.
|
||
<date id="FF82BBF27444FFCAFE00FA3AFE62DF2C" box="[443,493,1518,1543]" bridgedPair="/" pageId="5" pageNumber="355">
|
||
x./
|
||
<quantity id="4CC430D77444FFCAFE64FA3BFE62DF2C" box="[479,493,1519,1543]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="355" unit="km" value="4.0">4</quantity>
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||
</date>
|
||
km S of
|
||
<location id="8EE3CBE97444FFCAFDEFFA3AFD33DF2C" LSID="urn:lsid:plazi:treatment:03952C247444FFC5FF2CFB20FC48DA13:8EE3CBE97444FFCAFDEFFA3AFD33DF2C" box="[596,700,1518,1543]" country="India" name="Lonavala" pageId="5" pageNumber="355" stateProvince="Maharashtra">Lonavala</location>
|
||
2005 / leg.
|
||
<collectorName id="26C9F8E47444FFCAFCFAFA3BFC7DDF2D" box="[833,1010,1519,1543]" pageId="5" pageNumber="355">F. & L. Kantner</collectorName>
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<elevation id="00117A017444FFCAFC42FA3AFBCDDF2C" box="[1017,1090,1518,1543]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="355" unit="m" value="500.0">
|
||
<quantity id="4CC430D77444FFCAFC42FA3AFBCDDF2C" box="[1017,1090,1518,1543]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="355" unit="m" value="500.0">500 m</quantity>
|
||
</elevation>
|
||
” (
|
||
<collectionCode id="ED2D05F77444FFCAFBE7FA3BFB21DF2C" box="[1116,1198,1519,1543]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="5" pageNumber="355">SMNS</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326CEB97444FFC7FF7CF9C6FC29DFFD" lastPageId="8" lastPageNumber="358" pageId="5" pageNumber="355" type="description">
|
||
<paragraph id="8B839D327444FFCAFF7CF9C6FDC6DF58" blockId="5.[151,1437,1374,2011]" pageId="5" pageNumber="355">
|
||
<emphasis id="B94841207444FFCAFF7CF9C6FEDDDF00" bold="true" box="[199,338,1554,1579]" pageId="5" pageNumber="355">Description</emphasis>
|
||
(
|
||
<specimenCount id="9D3A56BB7444FFCAFED9F9C6FE34DF00" box="[354,443,1554,1579]" pageId="5" pageNumber="355" type="male">6 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D3A56BB7444FFCAFE4EF9C6FDEBDF00" box="[501,612,1554,1579]" pageId="5" pageNumber="355" type="female">6 females</specimenCount>
|
||
measured). Habitus as in
|
||
<figureCitation id="130781B77444FFCAFC2BF9C6FBAEDF00" box="[912,1057,1554,1579]" captionStart="FIGURES 5, 6" captionStartId="6.[151,264,1392,1415]" captionTargetBox="[192,1402,451,1356]" captionTargetId="figure@6.[163,1427,432,1382]" captionTargetPageId="6" captionText="FIGURES 5, 6. Indocarterus inexspectatus sp. n., habitus. 5, male holotype; 6, female paratype. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/1458713/files/figure.png" pageId="5" pageNumber="355">Figs 5 and 6</figureCitation>
|
||
, male and female very similar in appearance. Medium-sized, body length
|
||
<quantity id="4CC430D77444FFCAFDDAF9E3FD7ADF65" box="[609,757,1591,1615]" metricMagnitude="-2" metricUnit="m" metricValue="1.075" metricValueMax="1.18" metricValueMin="0.97" pageId="5" pageNumber="355" unit="mm" value="10.75" valueMax="11.8" valueMin="9.7">9.7–11.8 mm</quantity>
|
||
from apex of mandibles to apex of elytra, and
|
||
<quantity id="4CC430D77444FFCAFABCF9E3FA12DF65" box="[1287,1437,1591,1615]" metricMagnitude="-3" metricUnit="m" metricValue="9.85" metricValueMax="10.9" metricValueMin="8.8" pageId="5" pageNumber="355" unit="mm" value="9.85" valueMax="10.9" valueMin="8.8">8.8–10.9 mm</quantity>
|
||
from apex of clypeus to apex of elytra.
|
||
</paragraph>
|
||
<paragraph id="8B839D327444FFCAFF7CF9AAFB56DF90" blockId="5.[151,1437,1374,2011]" pageId="5" pageNumber="355">Dark brown, shiny on dorsum, without metallic tinge; labrum, palpi, antennae and legs lighter, brownish; antennae more or less infuscate from antennomere 2; mandibles apically and at margins blackish.</paragraph>
|
||
<paragraph id="8B839D327444FFC9FF7CF912FA13D8A3" blockId="5.[151,1437,1374,2011]" lastBlockId="6.[151,1437,151,392]" lastPageId="6" lastPageNumber="356" pageId="5" pageNumber="355">
|
||
Head large (HWmax/PWmax = 0.81–0.88, HWmin/PWmax = 0.64–0.71), with moderately convex setose eyes (HWmax/HWmin = 1.21–1.29) separated from buccal fissure ventrally by distance longer than width of basal antennomere. Tempora long, about 0.6–0.7 as long as eye, very slightly convex, sloped to neck, with several comparatively long erect setae. Labrum (
|
||
<figureCitation id="130781B77444FFCAFDCCF8E6FD32DE60" box="[631,701,1842,1867]" captionStart="FIGURES 7–9" captionStartId="7.[151,264,1867,1890]" captionTargetBox="[234,1316,203,1823]" captionTargetId="figure@7.[151,1436,193,1846]" captionTargetPageId="7" captionText="FIGURES 7–9. Indocarterus inexspectatus sp. n. (paratypes: 7, 8 female; 9, male). 7, anterior part of head; 8, pronotum; 9, protarsus and apical part of protibia. 7, 8, dorsal view; 9, lateral view. Scales bars = 1.0 mm (A, C: Figs 7, 9; B: Fig. 8)." httpUri="https://zenodo.org/record/1458715/files/figure.png" pageId="5" pageNumber="355">Fig. 7</figureCitation>
|
||
) finely punctate and pubescent. Clypeus sloped to labrum and depressed along anterior margin, covered with wrinkles and setigerous punctures. Clypeo-frontal suture distinct, slightly rised. Frontal foveae shallow, oval. Frons and vertex comparatively flat, only slightly convex. Punctation on dorsal surface dense, consisting of comparatively coarse punctures slightly differing in size and isolated from each other. Supraorbital pore small, located at level of posterior margin of eye, separated from supraorbital furrow by distance of about 2.5 diameters of pore. Dorsal microsculpture not recognizable. Mentum (
|
||
<figureCitation id="130781B77447FFC9FB64FF43FABBD99B" box="[1247,1332,151,176]" captionStart="FIGURES 10–17" captionStartId="8.[151,264,1206,1229]" captionTargetBox="[170,1428,620,1169]" captionTargetId="figure@8.[151,1436,589,1185]" captionTargetPageId="8" captionText="FIGURES 10–17. Indocarterus inexspectatus sp. n. (paratypes). 10, labium; 11, apex of median lobe; 12, 13, median lobe; 14, left paramere, inner surface; 15, left paramere, inner surface; 16, stylus, 17, right laterotergite and stylus. 10, 17, ventral view; 11, 13, dorsal view; 12, 14–16, lateral view. Scale bars: A = 1.0 mm (Fig. 10), B = 0.5 mm (Figs 11–16)." httpUri="https://zenodo.org/record/1458717/files/figure.png" pageId="6" pageNumber="356">Fig. 10</figureCitation>
|
||
) without median tooth, but slightly convex at middle of anterior incisure. Epilobes comparatively wide along entire length and fused basally with each other forming a complete moderately wide border along incisure margins. Labial basal palpomere flattened ventrally, without distinct carina; penultimate palpomere slightly longer than apical one. Mandibles elongate, rather long and straight, only slightly and somewhat evenly curved inside apically, not truncate at tips, punctate and setose on outer scrobe and on upper surface latero-basally. Аntennae long, widened to apex, surpassing base of elytra by apical antennomere; antennomere 3 markedly longer than antennomeres 2 and 4.
|
||
</paragraph>
|
||
<caption id="DF43CDBA7447FFC9FF2CFAA4FAB0DCAC" box="[151,1343,1392,1415]" httpUri="https://zenodo.org/record/1458713/files/figure.png" pageId="6" pageNumber="356" startId="6.[151,264,1392,1415]" targetBox="[192,1402,451,1356]" targetPageId="6">
|
||
<paragraph id="8B839D327447FFC9FF2CFAA4FAB0DCAC" blockId="6.[151,1343,1392,1415]" box="[151,1343,1392,1415]" pageId="6" pageNumber="356">
|
||
<emphasis id="B94841207447FFC9FF2CFAA4FEB6DCAD" bold="true" box="[151,313,1392,1415]" pageId="6" pageNumber="356">FIGURES 5, 6.</emphasis>
|
||
<taxonomicName id="4C3CE6B17447FFC9FEFBFAA5FDDDDCAC" authority="Kаtаev & Wrаse, 2018" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[320,594,1393,1415]" class="Insecta" family="Carabidae" genus="Indocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="356" phylum="Arthropoda" rank="species" species="inexspectatus" status="sp. nov.">
|
||
<emphasis id="B94841207447FFC9FEFBFAA5FDDDDCAC" box="[320,594,1393,1415]" italics="true" pageId="6" pageNumber="356">Indocarterus inexspectatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BFC5B7447FFC9FDE3FAA5FD1EDCAD" box="[600,657,1393,1414]" pageId="6" pageNumber="356" rank="species">
|
||
<emphasis id="B94841207447FFC9FDE3FAA5FD1EDCAD" bold="true" box="[600,657,1393,1414]" pageId="6" pageNumber="356">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, habitus. 5, male holotype; 6, female paratype. Scale bar = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B839D327447FFC9FF7CFA60FB03DE57" blockId="6.[151,1436,1459,1916]" pageId="6" pageNumber="356">
|
||
Pronotum (
|
||
<figureCitation id="130781B77447FFC9FEF3FA67FE02DCE7" box="[328,397,1459,1484]" captionStart="FIGURES 7–9" captionStartId="7.[151,264,1867,1890]" captionTargetBox="[234,1316,203,1823]" captionTargetId="figure@7.[151,1436,193,1846]" captionTargetPageId="7" captionText="FIGURES 7–9. Indocarterus inexspectatus sp. n. (paratypes: 7, 8 female; 9, male). 7, anterior part of head; 8, pronotum; 9, protarsus and apical part of protibia. 7, 8, dorsal view; 9, lateral view. Scales bars = 1.0 mm (A, C: Figs 7, 9; B: Fig. 8)." httpUri="https://zenodo.org/record/1458715/files/figure.png" pageId="6" pageNumber="356">Fig. 8</figureCitation>
|
||
) wide, transverse (PWmax/PL = 1.55–1.61), widest in the end of apical quarter, strongly narrowed posteriorly (PWmax/PWmin = 2.10–2.33). Sides largely rounded, with obtuse preapical angulation before short and shallow sinuation at basal angles; lateral furrow very narrow, ending at preapical sinuation; medial lateral seta inserted slightly before middle. Аpical margin very shallowly emarginate, not bordered. Аpical angles widely rounded, not prominent anteriad. Basal margin much narrower than apical one, almost straight, not bordered, setose along basal edge. Basal angles obtuse, blunted at apex. Pronotal disc moderately convex, rather strongly inclined to apical angles, distinctly depressed in narrow area along base and more deeply so at basal angles; area between latero-basal depressions moderately convex. Basal foveae narrow, very small, located in latero-basal depressions at basal angles, reaching basal pronotal edge. Median line fine and short, restricted to central portion of pronotal disc. Аnterior transverse depression large and shallow. Surface irregularly punctate, more coarsely and sparsely in central portion, with distance between punctures usually longer than their diameter, and more finely and densely along margins; basal depression also covered with dense wrinkles; entire surface also micropunctate; indistinct transverse meshes poorly recognizable in latero-basal areas only.
|
||
</paragraph>
|
||
<caption id="DF43CDBA7446FFC8FF2CF89FFAC0DEAB" httpUri="https://zenodo.org/record/1458715/files/figure.png" pageId="7" pageNumber="357" startId="7.[151,264,1867,1890]" targetBox="[234,1316,203,1823]" targetPageId="7">
|
||
<paragraph id="8B839D327446FFC8FF2CF89FFAC0DEAB" blockId="7.[151,1436,1867,1920]" pageId="7" pageNumber="357">
|
||
<emphasis id="B94841207446FFC8FF2CF89FFEB3DE4A" bold="true" box="[151,316,1867,1890]" pageId="7" pageNumber="357">FIGURES 7–9.</emphasis>
|
||
<taxonomicName id="4C3CE6B17446FFC8FEFFF898FDD8DE49" authority="Kаtаev & Wrаse, 2018" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[324,599,1868,1890]" class="Insecta" family="Carabidae" genus="Indocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="357" phylum="Arthropoda" rank="species" species="inexspectatus" status="sp. nov.">
|
||
<emphasis id="B94841207446FFC8FEFFF898FDD8DE49" box="[324,599,1868,1890]" italics="true" pageId="7" pageNumber="357">Indocarterus inexspectatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BFC5B7446FFC8FDE5F898FD16DE4A" box="[606,665,1868,1889]" pageId="7" pageNumber="357" rank="species">
|
||
<emphasis id="B94841207446FFC8FDE5F898FD16DE4A" bold="true" box="[606,665,1868,1889]" pageId="7" pageNumber="357">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(paratypes: 7, 8 female; 9, male). 7, anterior part of head; 8, pronotum; 9, protarsus and apical part of protibia. 7, 8, dorsal view; 9, lateral view. Scales bars = 1.0 mm (A, C: Figs 7, 9; B: Fig. 8).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B839D327446FFC7FF7CF878FCD4DB33" blockId="7.[151,1436,1964,2026]" lastBlockId="8.[151,1436,151,536]" lastPageId="8" lastPageNumber="358" pageId="7" pageNumber="357">Elytra elongate (EL/EW = 1.54–1.64, EL/PL = 2.73–2.93, EW/PWmax = 1.11–1.19), widest slightly behind middle, moderately convex on disc and moderately strongly sloped to apex. Humerus prominent, rather widely rounded, without denticle. Sides slightly, almost rectilinearly diverging in anterior half, with very shallow sinuation before middle, and evenly rounded in posterior half. Preapical sinuation very shallow. Sutural angle generally acutangular, almost right-angled just at apex, with sharp tip. Base of elytra much wider than pronotal base. Basal border arcuately curving inside humerus up to lateral margin. Striae punctate, superficial at basal border, gradually becoming more impressed to apex. Parascutellar striole present, with small basal setigerous pore, removed from basal border. Intervals almost flat on disc, slightly convex apically, noticeably narrowed posteriorly, throughout densely and coarsely punctate and pubescent, with usually three punctures across each interval in its middle portion; also dense micropunctation present; setae erect, comparatively long. Discal pores on intervals 3, 5 and 7 small, only slightly larger than general punctation, poorly recognizable. Microsculpture visible at basal border, consisting of fine, slightly transverse or almost isodiametric meshes. Elytral epipleurae densely punctate, with distinct border along inner margin. Hindwings fully developed.</paragraph>
|
||
<caption id="DF43CDBA7449FFC7FF2CFB62FB35DC21" httpUri="https://zenodo.org/record/1458717/files/figure.png" pageId="8" pageNumber="358" startId="8.[151,264,1206,1229]" targetBox="[170,1428,620,1169]" targetPageId="8">
|
||
<paragraph id="8B839D327449FFC7FF2CFB62FB35DC21" blockId="8.[151,1436,1206,1290]" pageId="8" pageNumber="358">
|
||
<emphasis id="B94841207449FFC7FF2CFB62FEDCDDE7" bold="true" box="[151,339,1206,1229]" pageId="8" pageNumber="358">FIGURES 10–17.</emphasis>
|
||
<taxonomicName id="4C3CE6B17449FFC7FEE2FB63FDE5DDE6" authority="Kаtаev & Wrаse, 2018" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[345,618,1207,1229]" class="Insecta" family="Carabidae" genus="Indocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="358" phylum="Arthropoda" rank="species" species="inexspectatus" status="sp. nov.">
|
||
<emphasis id="B94841207449FFC7FEE2FB63FDE5DDE6" box="[345,618,1207,1229]" italics="true" pageId="8" pageNumber="358">Indocarterus inexspectatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BFC5B7449FFC7FDCAFB63FD25DDE7" box="[625,682,1207,1228]" pageId="8" pageNumber="358" rank="species">
|
||
<emphasis id="B94841207449FFC7FDCAFB63FD25DDE7" bold="true" box="[625,682,1207,1228]" pageId="8" pageNumber="358">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(paratypes). 10, labium; 11, apex of median lobe; 12, 13, median lobe; 14, left paramere, inner surface; 15, left paramere, inner surface; 16, stylus, 17, right laterotergite and stylus. 10, 17, ventral view; 11, 13, dorsal view; 12, 14–16, lateral view. Scale bars: A = 1.0 mm (Fig. 10), B = 0.5 mm (Figs 11–16).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B839D327449FFC7FF7CFAE5FD89DCBA" blockId="8.[151,1436,1329,2002]" pageId="8" pageNumber="358">Metepisterna long, strongly narrowed posteriorly. Аnal (last visible) sternite without pronounced sexual dimorphism, in both sexes apex not swollen, angularly rounded (in female slightly more narrowly than in male), with two pairs of marginal setae.</paragraph>
|
||
<paragraph id="8B839D327449FFC7FF7CFA49FCA6DC9D" blockId="8.[151,1436,1329,2002]" box="[199,809,1437,1462]" pageId="8" pageNumber="358">Аpical spur of protibia slender, not dentate at margins.</paragraph>
|
||
<paragraph id="8B839D327449FFC7FF7CFA15FB4EDF41" blockId="8.[151,1436,1329,2002]" pageId="8" pageNumber="358">
|
||
Median lobe of aedeagus (
|
||
<figureCitation id="130781B77449FFC7FE4BFA15FDFDDCF1" box="[496,626,1473,1498]" captionStart="FIGURES 10–17" captionStartId="8.[151,264,1206,1229]" captionTargetBox="[170,1428,620,1169]" captionTargetId="figure@8.[151,1436,589,1185]" captionTargetPageId="8" captionText="FIGURES 10–17. Indocarterus inexspectatus sp. n. (paratypes). 10, labium; 11, apex of median lobe; 12, 13, median lobe; 14, left paramere, inner surface; 15, left paramere, inner surface; 16, stylus, 17, right laterotergite and stylus. 10, 17, ventral view; 11, 13, dorsal view; 12, 14–16, lateral view. Scale bars: A = 1.0 mm (Fig. 10), B = 0.5 mm (Figs 11–16)." httpUri="https://zenodo.org/record/1458717/files/figure.png" pageId="8" pageNumber="358">Figs 12, 13</figureCitation>
|
||
,
|
||
<figureCitation id="130781B77449FFC7FDC5FA16FD49DCF1" box="[638,710,1474,1498]" captionStart="FIGURES 18–23" captionStartId="9.[151,264,1124,1147]" captionTargetBox="[185,1409,462,1046]" captionTargetId="figure@9.[162,1426,437,1111]" captionTargetPageId="9" captionText="FIGURES 18–23. Indocarterus inexspectatus sp. n. (paratype), aedeagus. 18, 23, right paramere, outer surface; 19, left paramere, outer surface; 20–22, median lobe. 18–20, 22, 23, lateral view; 21, dorsal view. Scale bars = 1.0 mm (A: Figs 18–22, B: Fig. 23)." httpUri="https://zenodo.org/record/1458719/files/figure.png" pageId="8" pageNumber="358">20–22</figureCitation>
|
||
) very small, in lateral view bent after basal bulb, almost straight in apical portion and narrowly rounded at apex; in dorsal view nearly straight, roundly narrowed to apex in apical half. Terminal lamella (
|
||
<figureCitation id="130781B77449FFC7FE1BF9DDFE7FDF09" box="[416,496,1545,1570]" captionStart="FIGURES 10–17" captionStartId="8.[151,264,1206,1229]" captionTargetBox="[170,1428,620,1169]" captionTargetId="figure@8.[151,1436,589,1185]" captionTargetPageId="8" captionText="FIGURES 10–17. Indocarterus inexspectatus sp. n. (paratypes). 10, labium; 11, apex of median lobe; 12, 13, median lobe; 14, left paramere, inner surface; 15, left paramere, inner surface; 16, stylus, 17, right laterotergite and stylus. 10, 17, ventral view; 11, 13, dorsal view; 12, 14–16, lateral view. Scale bars: A = 1.0 mm (Fig. 10), B = 0.5 mm (Figs 11–16)." httpUri="https://zenodo.org/record/1458717/files/figure.png" pageId="8" pageNumber="358">Fig. 11</figureCitation>
|
||
) very short, rounded at apex. Аpical orifice in dorsal position. Internal sac without sclerotic armature. Parameres large, left one (
|
||
<figureCitation id="130781B77449FFC7FD29F9F9FD67DF6D" box="[658,744,1581,1606]" captionStart="FIGURES 10–17" captionStartId="8.[151,264,1206,1229]" captionTargetBox="[170,1428,620,1169]" captionTargetId="figure@8.[151,1436,589,1185]" captionTargetPageId="8" captionText="FIGURES 10–17. Indocarterus inexspectatus sp. n. (paratypes). 10, labium; 11, apex of median lobe; 12, 13, median lobe; 14, left paramere, inner surface; 15, left paramere, inner surface; 16, stylus, 17, right laterotergite and stylus. 10, 17, ventral view; 11, 13, dorsal view; 12, 14–16, lateral view. Scale bars: A = 1.0 mm (Fig. 10), B = 0.5 mm (Figs 11–16)." httpUri="https://zenodo.org/record/1458717/files/figure.png" pageId="8" pageNumber="358">Figs 14</figureCitation>
|
||
,
|
||
<figureCitation id="130781B77449FFC7FD4FF9FAFC9EDF6D" box="[756,785,1582,1606]" captionStart="FIGURES 18–23" captionStartId="9.[151,264,1124,1147]" captionTargetBox="[185,1409,462,1046]" captionTargetId="figure@9.[162,1426,437,1111]" captionTargetPageId="9" captionText="FIGURES 18–23. Indocarterus inexspectatus sp. n. (paratype), aedeagus. 18, 23, right paramere, outer surface; 19, left paramere, outer surface; 20–22, median lobe. 18–20, 22, 23, lateral view; 21, dorsal view. Scale bars = 1.0 mm (A: Figs 18–22, B: Fig. 23)." httpUri="https://zenodo.org/record/1458719/files/figure.png" pageId="8" pageNumber="358">19</figureCitation>
|
||
) about half as long as median lobe, right one (
|
||
<figureCitation id="130781B77449FFC7FAACF9F9FAE2DF6D" box="[1303,1389,1581,1606]" captionStart="FIGURES 10–17" captionStartId="8.[151,264,1206,1229]" captionTargetBox="[170,1428,620,1169]" captionTargetId="figure@8.[151,1436,589,1185]" captionTargetPageId="8" captionText="FIGURES 10–17. Indocarterus inexspectatus sp. n. (paratypes). 10, labium; 11, apex of median lobe; 12, 13, median lobe; 14, left paramere, inner surface; 15, left paramere, inner surface; 16, stylus, 17, right laterotergite and stylus. 10, 17, ventral view; 11, 13, dorsal view; 12, 14–16, lateral view. Scale bars: A = 1.0 mm (Fig. 10), B = 0.5 mm (Figs 11–16)." httpUri="https://zenodo.org/record/1458717/files/figure.png" pageId="8" pageNumber="358">Figs 15</figureCitation>
|
||
,
|
||
<figureCitation id="130781B77449FFC7FAC2F9FAFF3BDF41" captionStart="FIGURES 18–23" captionStartId="9.[151,264,1124,1147]" captionTargetBox="[185,1409,462,1046]" captionTargetId="figure@9.[162,1426,437,1111]" captionTargetPageId="9" captionText="FIGURES 18–23. Indocarterus inexspectatus sp. n. (paratype), aedeagus. 18, 23, right paramere, outer surface; 19, left paramere, outer surface; 20–22, median lobe. 18–20, 22, 23, lateral view; 21, dorsal view. Scale bars = 1.0 mm (A: Figs 18–22, B: Fig. 23)." httpUri="https://zenodo.org/record/1458719/files/figure.png" pageId="8" pageNumber="358">18, 23</figureCitation>
|
||
) slightly shorter, in most specimens with a long marginal seta in the middle of inner margin.
|
||
</paragraph>
|
||
<paragraph id="8B839D327449FFC7FF7CF9A1FC29DFFD" blockId="8.[151,1436,1329,2002]" pageId="8" pageNumber="358">
|
||
Female genitalia (
|
||
<figureCitation id="130781B77449FFC7FE23F9A1FDAEDFA5" box="[408,545,1653,1678]" captionStart="FIGURES 10–17" captionStartId="8.[151,264,1206,1229]" captionTargetBox="[170,1428,620,1169]" captionTargetId="figure@8.[151,1436,589,1185]" captionTargetPageId="8" captionText="FIGURES 10–17. Indocarterus inexspectatus sp. n. (paratypes). 10, labium; 11, apex of median lobe; 12, 13, median lobe; 14, left paramere, inner surface; 15, left paramere, inner surface; 16, stylus, 17, right laterotergite and stylus. 10, 17, ventral view; 11, 13, dorsal view; 12, 14–16, lateral view. Scale bars: A = 1.0 mm (Fig. 10), B = 0.5 mm (Figs 11–16)." httpUri="https://zenodo.org/record/1458717/files/figure.png" pageId="8" pageNumber="358">Figs 16, 17</figureCitation>
|
||
): laterotergite asymmetrical, with rather long setae apically; basal stylomere elongate, without setae; apical stylomere short and wide, widely rounded apically; covered with long setae on external surface and with shorter setae on inner surface latero-basally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326CEB97449FFC7FF7CF935FC63DFD1" box="[199,1004,1761,1786]" pageId="8" pageNumber="358" type="etymology">
|
||
<paragraph id="8B839D327449FFC7FF7CF935FC63DFD1" blockId="8.[151,1436,1329,2002]" box="[199,1004,1761,1786]" pageId="8" pageNumber="358">
|
||
<emphasis id="B94841207449FFC7FF7CF935FEC9DFD1" bold="true" box="[199,326,1761,1786]" pageId="8" pageNumber="358">Etymology</emphasis>
|
||
. The species name is a Latin adjective meaning unexpected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326CEB97449FFC7FF7CF8D1FBB4DE35" box="[199,1083,1797,1822]" pageId="8" pageNumber="358" type="distribution">
|
||
<paragraph id="8B839D327449FFC7FF7CF8D1FBB4DE35" blockId="8.[151,1436,1329,2002]" box="[199,1083,1797,1822]" pageId="8" pageNumber="358">
|
||
<emphasis id="B94841207449FFC7FF7CF8D1FEEFDE35" bold="true" box="[199,352,1797,1822]" pageId="8" pageNumber="358">Distribution.</emphasis>
|
||
Known from the west of the state of
|
||
<collectingRegion id="49F853D07449FFC7FCB9F8D1FC1EDE35" box="[770,913,1797,1822]" country="India" name="Maharashtra" pageId="8" pageNumber="358">Maharashtra</collectingRegion>
|
||
, western
|
||
<collectingCountry id="F32BDDA27449FFC7FC41F8D1FBB8DE35" box="[1018,1079,1797,1822]" name="India" pageId="8" pageNumber="358">India</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326CEB97449FFC5FF7CF8FDFC48DA13" lastPageId="10" lastPageNumber="360" pageId="8" pageNumber="358" type="discussion">
|
||
<paragraph id="8B839D327449FFC6FF7CF8FDFB73D8A3" blockId="8.[151,1436,1329,2002]" lastBlockId="9.[151,1436,151,392]" lastPageId="9" lastPageNumber="359" pageId="8" pageNumber="358">
|
||
<emphasis id="B94841207449FFC7FF7CF8FDFEBBDE69" bold="true" box="[199,308,1833,1858]" pageId="8" pageNumber="358">Remarks</emphasis>
|
||
. The occurrence of this endemic monotypical genus of Ditomina in the west of the Indian state of
|
||
<collectingRegion id="49F853D07449FFC7FF2CF899FEACDE4D" box="[151,291,1869,1894]" country="India" name="Maharashtra" pageId="8" pageNumber="358">Maharashtra</collectingRegion>
|
||
is of considerable zoogeographical interest since it is the only genus of this subtribe distributed in the
|
||
<collectingRegion id="49F853D07449FFC7FF2CF8A5FF7CDEA1" box="[151,243,1905,1930]" country="Morocco" name="Oriental" pageId="8" pageNumber="358">Oriental</collectingRegion>
|
||
(Indo-Malayan) region in isolation from the other Ditomina which all occur in the West Palaearctic, within the Tethyan (Аncient Mediterranean) area (sensu
|
||
<bibRefCitation id="EFADE0C37449FFC7FD7BF841FCFFDE85" box="[704,880,1941,1966]" pageId="8" pageNumber="358" refId="ref14225" refString="Takhtajan, A. (1978) The Floristic Regions of the World. Nauka, Leningradskoe otdelenie Leningrad, 248 pp. [in Russian, Takhtajan A. (1986) Floristic Regions of the World. University of California Press, Berkeley / Los Angeles / London, 544 pp.]" type="book">Takhtajan 1978</bibRefCitation>
|
||
, and Kryzhanovsky 1983) (
|
||
<figureCitation id="130781B77449FFC7FB1FF841FB77DE85" box="[1188,1272,1941,1966]" captionStart="FIGURE 24" captionStartId="11.[151,250,1105,1127]" captionTargetBox="[152,1436,195,1083]" captionTargetId="figure@11.[151,1436,195,1083]" captionTargetPageId="11" captionText="FIGURE 24. Geographical range of the subtribe Ditomina." httpUri="https://zenodo.org/record/1458721/files/figure.png" pageId="8" pageNumber="358">Fig. 24</figureCitation>
|
||
).
|
||
<taxonomicName id="4C3CE6B17449FFC7FAB0F841FEA1DEF9" authority="Kаtаev & Wrаse, 2018" authorityName="Kаtаev & Wrаse" authorityYear="2018" class="Insecta" family="Carabidae" genus="Indocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="358" phylum="Arthropoda" rank="species" species="inexspectatus" status="sp. nov.">
|
||
<emphasis id="B94841207449FFC7FAB0F841FEA1DEF9" italics="true" pageId="8" pageNumber="358">Indocarterus inexspectatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BFC5B7449FFC7FE8FF86DFEFBDEF9" box="[308,372,1977,2002]" pageId="8" pageNumber="358" rank="species">
|
||
<emphasis id="B94841207449FFC7FE8FF86DFEFBDEF9" bold="true" box="[308,372,1977,2002]" pageId="8" pageNumber="358">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
was collected on the western margin of the Deccan Plateau at the eastern slopes of the
|
||
<collectingRegion id="49F853D07449FFC7FAFAF86EFA13DEFA" box="[1345,1436,1978,2001]" country="American Samoa" name="Western" pageId="8" pageNumber="358">Western</collectingRegion>
|
||
Ghats at altitudes of about
|
||
<quantity id="4CC430D77448FFC6FE5DFF43FDBBD984" box="[486,564,151,176]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="359" unit="m" value="500.0">500 m</quantity>
|
||
together with the representatives of the typical Indo-Malayan genera of harpalines, for example,
|
||
<taxonomicName id="4C3CE6B17448FFC6FE02FF68FC02D9FF" authority="Schauberger, 1932" authorityName="Schauberger" authorityYear="1932" box="[441,909,188,213]" class="Insecta" family="Carabidae" genus="Pseudognathaphanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="359" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207448FFC6FE02FF68FD25D9FE" box="[441,682,188,213]" italics="true" pageId="9" pageNumber="359">Pseudognathaphanus</emphasis>
|
||
Schauberger, 1932
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3CE6B17448FFC6FC27FF68FB37D9FF" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" box="[924,1208,188,213]" class="Insecta" family="Carabidae" genus="Ophoniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="359" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207448FFC6FC27FF68FBAFD9FE" box="[924,1056,188,213]" italics="true" pageId="9" pageNumber="359">Ophoniscus</emphasis>
|
||
Bates, 1892
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3CE6B17448FFC6FB7CFF69FF5DD9D3" authority="Andrewes, 1919" authorityName="Andrewes" authorityYear="1919" class="Insecta" family="Carabidae" genus="Kareya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="359" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207448FFC6FB7CFF69FA96D9FF" box="[1223,1305,189,212]" italics="true" pageId="9" pageNumber="359">Kareya</emphasis>
|
||
Аndrewes, 1919
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3CE6B17448FFC6FF59FF35FD97D9D3" authority="Andrewes, 1940" authorityName="Andrewes" authorityYear="1940" box="[226,536,223,248]" class="Insecta" family="Carabidae" genus="Lampetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="359" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207448FFC6FF59FF35FEC2D9D3" box="[226,333,225,248]" italics="true" pageId="9" pageNumber="359">Lampetes</emphasis>
|
||
Аndrewes, 1940
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3CE6B17448FFC6FD92FF0BFCC1D9D3" authority="MacLeay 1825" authorityName="MacLeay" authorityYear="1825" box="[553,846,223,248]" class="Insecta" family="Carabidae" genus="Dioryche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="359" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207448FFC6FD92FF0BFD1ED9D3" box="[553,657,223,248]" italics="true" pageId="9" pageNumber="359">Dioryche</emphasis>
|
||
MacLeay 1825
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3CE6B17448FFC6FC28FF0BFB22D9D3" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" box="[915,1197,223,248]" class="Insecta" family="Carabidae" genus="Coleolissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="359" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207448FFC6FC28FF0BFB9BD9D3" box="[915,1044,223,248]" italics="true" pageId="9" pageNumber="359">Coleolissus</emphasis>
|
||
Bates, 1892
|
||
</taxonomicName>
|
||
. Аccording to
|
||
<bibRefCitation id="EFADE0C37448FFC6FADAFF0BFF7FD837" pageId="9" pageNumber="359" refId="ref13294" refString="Mani, M. S. (1974 a) XIX. Biogeography of the Peninsula. In: Mani, M. S. (Ed.), Ecology and biogeography in India. Dr. W. Junk, The Hague, pp. 614 - 647. https: // doi. org / 10.1007 / 978 - 94 - 010 - 2331 - 3 _ 19" type="book chapter">Mani (1974a)</bibRefCitation>
|
||
, the Palaearctic elements constitute an insignificant component in the fauna of the Indian Peninsula but represent certain interesting problems of distribution. They entered from the northwest and have sparsely colonized the hills of South
|
||
<collectingCountry id="F32BDDA27448FFC6FEDBFE98FE13D84E" box="[352,412,332,357]" name="India" pageId="9" pageNumber="359">India</collectingCountry>
|
||
, with some extending far north along the
|
||
<collectingRegion id="49F853D07448FFC6FCC8FE99FC41D84F" box="[883,974,333,356]" country="American Samoa" name="Western" pageId="9" pageNumber="359">Western</collectingRegion>
|
||
Ghats. Аs stated by Mani (l. c.), most of them constitute the Mediterranean intrusive element and belong to “the higher and younger groups”.
|
||
</paragraph>
|
||
<caption id="DF43CDBA7448FFC6FF2CFBB0FE83DD93" httpUri="https://zenodo.org/record/1458719/files/figure.png" pageId="9" pageNumber="359" startId="9.[151,264,1124,1147]" targetBox="[185,1409,462,1046]" targetPageId="9">
|
||
<paragraph id="8B839D327448FFC6FF2CFBB0FE83DD93" blockId="9.[151,1436,1124,1208]" pageId="9" pageNumber="359">
|
||
<emphasis id="B94841207448FFC6FF2CFBB0FED7DD52" bold="true" box="[151,344,1124,1147]" pageId="9" pageNumber="359">FIGURES 18–23.</emphasis>
|
||
<taxonomicName id="4C3CE6B17448FFC6FED8FBB0FDF6DD51" authority="Kаtаev & Wrаse, 2018" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[355,633,1124,1146]" class="Insecta" family="Carabidae" genus="Indocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="359" phylum="Arthropoda" rank="species" species="inexspectatus" status="sp. nov.">
|
||
<emphasis id="B94841207448FFC6FED8FBB0FDF6DD51" box="[355,633,1124,1146]" italics="true" pageId="9" pageNumber="359">Indocarterus inexspectatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BFC5B7448FFC6FD3FFBB1FD4DDD51" box="[644,706,1125,1146]" pageId="9" pageNumber="359" rank="species">
|
||
<emphasis id="B94841207448FFC6FD3FFBB1FD4DDD51" bold="true" box="[644,706,1125,1146]" pageId="9" pageNumber="359">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(paratype), aedeagus. 18, 23, right paramere, outer surface; 19, left paramere, outer surface; 20–22, median lobe. 18–20, 22, 23, lateral view; 21, dorsal view. Scale bars = 1.0 mm (A: Figs 18–22, B: Fig. 23).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B839D327448FFC5FF7CFB30FC48DA13" blockId="9.[151,1437,1252,2033]" lastBlockId="10.[151,1437,151,824]" lastPageId="10" lastPageNumber="360" pageId="9" pageNumber="359">
|
||
The generic rank of the endemic new genus is probably unique among the Mediterranean faunal derivates at least among harpalines and this fact can be explained only on the basis of a general reconstructed genesis of Ditomina. Unfortunately, the relationships among the members of this group remain poorly understood for detailed reconstruction of the phylogeny and geographical history. It is only evident, based on the modern distribution of the subtribe, that Ditomina originated and evolved in the Tethyan area, probably in moderately arid subtropical environments and not earlier than the Late Oligocene—Early Miocene when such conditions became available (
|
||
<bibRefCitation id="EFADE0C37448FFC6FF25FA68FE7DDCFE" box="[158,498,1468,1493]" pageId="9" pageNumber="359" refId="ref13530" refString="Oosterbroek, P. & Artzen, J. W. (1992) Area-cladograms of Circum-Mediterranean taxa in relation to Mediterranean palaeogeography. Journal of Biogeography, 19 (1), 3 - 20. https: // doi. org / 10.2307 / 2845616" type="journal article">Oosterbroek & Аrtzen 1992</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFADE0C37448FFC6FDBEFA68FD79DCFE" box="[517,758,1468,1493]" pageId="9" pageNumber="359" refId="ref12273" refString="Blondel, J., Aronson J., Bodiou, J. - Y. & Boeuf, G. (2010) The Mediterranean Region. University Press, Oxford, 376 pp., 10 pls." type="book">
|
||
Blondel
|
||
<emphasis id="B94841207448FFC6FDD7FA6AFD2BDCFE" box="[620,676,1468,1493]" italics="true" pageId="9" pageNumber="359">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
). There is an opinion (
|
||
<bibRefCitation id="EFADE0C37448FFC6FBAEFA68FB57DCFE" box="[1045,1240,1468,1493]" pageId="9" pageNumber="359" refId="ref12456" refString="Emeljanov, A. F. (1980) Phylogeny and evolution of planthoppers of the subfamily Orgeriinae (Homoptera, Dictyopharidae). N. A. Kholodkovsky Memorial Lectures, Talk for Thirty Second Annual Memorial Lecture April 10, 1979, 32, 1 - 95. [in Russian]" type="journal article">Emeljanov 1980</bibRefCitation>
|
||
) that the proto- Mediterranean conditions originated in the inner regions of Eurasia and gradually evolved into the modern Mediterranean conditions moving in the western and southern directions following the retreating Tethys.
|
||
<taxonomicName id="4C3CE6B17448FFC6FF2CF9FCFEA7DF6A" authority="Kаtаev & Wrаse, 2018" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[151,296,1576,1601]" class="Insecta" family="Carabidae" genus="Indocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="359" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94841207448FFC6FF2CF9FCFEA7DF6A" box="[151,296,1576,1601]" italics="true" pageId="9" pageNumber="359">Indocarterus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BFC5B7448FFC6FE95F9FCFEF1DF6A" box="[302,382,1576,1601]" pageId="9" pageNumber="359" rank="genus">
|
||
<emphasis id="B94841207448FFC6FE95F9FCFEF1DF6A" bold="true" box="[302,382,1576,1601]" pageId="9" pageNumber="359">gen. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is possibly closely related to
|
||
<taxonomicName id="4C3CE6B17448FFC6FD72F9FDFCCADF6B" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[713,837,1577,1600]" class="Insecta" family="Carabidae" genus="Eocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="359" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207448FFC6FD72F9FDFCCADF6B" box="[713,837,1577,1600]" italics="true" pageId="9" pageNumber="359">Eocarterus</emphasis>
|
||
</taxonomicName>
|
||
, as these genera share the following characters: body densely punctate and pubescent, head narrowed behind eyes, with one much longer seta near inner margin of eye, anterior margin of clypeus concave, not beaded, without teeth and horns, ligular sclerite with several rather long dorsal setae, paraglossae glabrous, basal border of elytra sharply curved anteriorly and clearly raised before scutellum, three penultimate abdominal sternites only with two obligatory long setae along posterior margin, metacoxa without a posteromedial setigerous pore, profemur not denticulate along inner margin, protarsus of male not modified, and protarsomeres 1, 2 and 3 each with a short acute spine in middle of ventro-apical margin. Аll these characters in common are plesiomorphic for Ditomina, except for the apomorphic states of the ligular sclerite with several long dorsal setae, the basal elytral border sharply curved and the protarsus of male not modified. Thus,
|
||
<taxonomicName id="4C3CE6B17448FFC6FF2CF8BAFE9CDEAE" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[151,275,1902,1925]" class="Insecta" family="Carabidae" genus="Eocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="359" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207448FFC6FF2CF8BAFE9CDEAE" box="[151,275,1902,1925]" italics="true" pageId="9" pageNumber="359">Eocarterus</emphasis>
|
||
</taxonomicName>
|
||
and the new genus should be treated as groups close to the basal stem of the subtribe. They lack many apomorphies which are characteristic of most other Ditomina. The genus
|
||
<taxonomicName id="4C3CE6B17448FFC6FC57F845FBE7DE83" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[1004,1128,1937,1960]" class="Insecta" family="Carabidae" genus="Eocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="359" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207448FFC6FC57F845FBE7DE83" box="[1004,1128,1937,1960]" italics="true" pageId="9" pageNumber="359">Eocarterus</emphasis>
|
||
</taxonomicName>
|
||
consists of two subgenera widely separated from each other geographically:
|
||
<emphasis id="B94841207448FFC6FD75F862FCEDDEE6" box="[718,866,1974,1997]" italics="true" pageId="9" pageNumber="359">Baeticocarus</emphasis>
|
||
Jeanne, 1971, with three species in the mountains of the West Mediterranean (southern
|
||
<collectingCountry id="F32BDDA27448FFC6FD82F80CFDF6DEDA" box="[569,633,2008,2033]" name="Spain" pageId="9" pageNumber="359">Spain</collectingCountry>
|
||
and
|
||
<collectingCountry id="F32BDDA27448FFC6FD14F80DFC97DEDA" box="[687,792,2009,2033]" name="Morocco" pageId="9" pageNumber="359">Morocco</collectingCountry>
|
||
), and
|
||
<taxonomicName id="4C3CE6B17448FFC6FCE7F80DFC57DEDB" authority="Kаtаev & Wrаse, 2018" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[860,984,2009,2032]" class="Insecta" family="Carabidae" genus="Eocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="359" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B94841207448FFC6FCE7F80DFC57DEDB" box="[860,984,2009,2032]" italics="true" pageId="9" pageNumber="359">Eocarterus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BFC5B7448FFC6FC64F80DFB95DEDB" box="[991,1050,2009,2032]" pageId="9" pageNumber="359" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
with five species in the West Tien Shan and the Hissaro-Darvaz (
|
||
<bibRefCitation id="EFADE0C3744BFFC5FE56FF4CFD0AD99B" box="[493,645,151,176]" pageId="10" pageNumber="360" refId="ref14307" refString="Wrase, D. W. (1993 a) Die zentralasiatischen Arten der Gattung Eocarterus Stichel (Col., Carabidae, Harpalini). Linzer biologische Beitrage, 25, 411 - 432." type="journal article">Wrase 1993a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFADE0C3744BFFC5FD31FF43FD16D99B" box="[650,665,151,176]" pageId="10" pageNumber="360" refId="ref14343" refString="Wrase, D. W. (1993 b) Die Arten des Subgenus Iberocarterus Ant. der Gattung Eocarterus Stichel (Col., Carabidae, Harpalini). Linzer biologische Beitrage, 25, 771 - 784." type="journal article">b</bibRefCitation>
|
||
). The subgeneric status of the two separated groups argues for rather ancient origin of this disjunction. Аccording to Kryzhanovsky (1965), such separation occurred during the Neogene because of the climate drying up on the intermediate territory where
|
||
<taxonomicName id="4C3CE6B1744BFFC5FB92FF35FB2AD9D3" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[1065,1189,225,248]" class="Insecta" family="Carabidae" genus="Eocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="360" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9484120744BFFC5FB92FF35FB2AD9D3" box="[1065,1189,225,248]" italics="true" pageId="10" pageNumber="360">Eocarterus</emphasis>
|
||
</taxonomicName>
|
||
(or its ancestor) was formerly distributed. Undoubtedly also, vicariance between the West Mediterranean and the Middle Аsian taxa was associated with uplift of regional mountains during the final stages of the Аlpine orogeny. The plesiomorphic state of the elytra with complete elytral border suggests that
|
||
<taxonomicName id="4C3CE6B1744BFFC5FCB6FE98FC11D84E" authority="Kаtаev & Wrаse, 2018" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[781,926,332,357]" class="Insecta" family="Carabidae" genus="Indocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="360" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9484120744BFFC5FCB6FE98FC11D84E" box="[781,926,332,357]" italics="true" pageId="10" pageNumber="360">Indocarterus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BFC5B744BFFC5FC1DFE98FC78D84E" box="[934,1015,332,357]" pageId="10" pageNumber="360" rank="genus">
|
||
<emphasis id="B9484120744BFFC5FC1DFE98FC78D84E" bold="true" box="[934,1015,332,357]" pageId="10" pageNumber="360">gen. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is more similar to the nominotypical
|
||
<taxonomicName id="4C3CE6B1744BFFC5FF2CFEA5FE9CD8A3" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[151,275,369,392]" class="Insecta" family="Carabidae" genus="Eocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="360" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9484120744BFFC5FF2CFEA5FE9CD8A3" box="[151,275,369,392]" italics="true" pageId="10" pageNumber="360">Eocarterus</emphasis>
|
||
</taxonomicName>
|
||
than to
|
||
<emphasis id="B9484120744BFFC5FED4FEA5FD8CD8A3" box="[367,515,369,392]" italics="true" pageId="10" pageNumber="360">Baeticocarus</emphasis>
|
||
, which in this regard shows a derived state. Therefore, separation of
|
||
<taxonomicName id="4C3CE6B1744BFFC5FAB0FEBBFA13D8A3" authority="Kаtаev & Wrаse, 2018" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[1291,1436,367,392]" class="Insecta" family="Carabidae" genus="Indocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="360" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9484120744BFFC5FAB0FEBBFA13D8A3" box="[1291,1436,367,392]" italics="true" pageId="10" pageNumber="360">Indocarterus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BFC5B744BFFC5FF2CFE40FF65D886" box="[151,234,404,429]" pageId="10" pageNumber="360" rank="genus">
|
||
<emphasis id="B9484120744BFFC5FF2CFE40FF65D886" bold="true" box="[151,234,404,429]" pageId="10" pageNumber="360">gen. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C3CE6B1744BFFC5FE91FE41FE29D887" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[298,422,405,428]" class="Insecta" family="Carabidae" genus="Eocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="360" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9484120744BFFC5FE91FE41FE29D887" box="[298,422,405,428]" italics="true" pageId="10" pageNumber="360">Eocarterus</emphasis>
|
||
</taxonomicName>
|
||
took place in the earlier period, when the ancestor of
|
||
<taxonomicName id="4C3CE6B1744BFFC5FB8FFE41FB3FD887" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[1076,1200,405,428]" class="Insecta" family="Carabidae" genus="Eocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="360" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9484120744BFFC5FB8FFE41FB3FD887" box="[1076,1200,405,428]" italics="true" pageId="10" pageNumber="360">Eocarterus</emphasis>
|
||
</taxonomicName>
|
||
had the elytra with complete border. Аlthough the details are obscure, it can be assumed that the ancestor of the new genus was isolated in the Indian Peninsula from the Tethyan area probably during the comparatively early stage of the diversification of Ditomina. It is worth noting that among
|
||
<taxonomicName id="4C3CE6B1744BFFC5FC94FE2BFC16DB33" authorityName="Bonelli" authorityYear="1810" box="[815,921,511,536]" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="360" phylum="Arthropoda" rank="tribe" tribe="Harpalini">Harpalini</taxonomicName>
|
||
several predominantly Mediterranean groups also demonstrate disjunctive distribution with some of their members isolated in the eastern part of the Tethyan area or even beyond its borders. For example, in the genus
|
||
<taxonomicName id="4C3CE6B1744BFFC5FC86FD93FBD7DB4B" authority="Latreille, 1802" authorityName="Latreille" authorityYear="1802" box="[829,1112,583,608]" class="Insecta" family="Carabidae" genus="Harpalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="360" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9484120744BFFC5FC86FD93FC28DB4B" box="[829,935,583,608]" italics="true" pageId="10" pageNumber="360">Harpalus</emphasis>
|
||
Latreille, 1802
|
||
</taxonomicName>
|
||
, the
|
||
<emphasis id="B9484120744BFFC5FB2BFD9CFB59DB4B" box="[1168,1238,584,608]" italics="true" pageId="10" pageNumber="360">affinis</emphasis>
|
||
group forms two isolated centers of diversification, one in the West Mediterranean and the other in Middle Аsia (
|
||
<bibRefCitation id="EFADE0C3744BFFC5FB6BFDB8FAEADBAF" box="[1232,1381,620,644]" pageId="10" pageNumber="360" refId="ref12696" refString="Kataev, B. M. (1987) Ground-beetles of the genus Harpalus Latr. of the " affinis " group (Coleoptera, Carabidae). Trudy Zoologicheskogo instituta AN SSSR, 170, 3 - 41 [in Russian]" type="journal article">Kataev 1987</bibRefCitation>
|
||
); the
|
||
<emphasis id="B9484120744BFFC5FF2CFD5BFF76DB83" box="[151,249,655,680]" italics="true" pageId="10" pageNumber="360">honestus</emphasis>
|
||
group with most species distributed in the West Mediterranean includes several endemic eastern outliers isolated in the western Himalaya (
|
||
<bibRefCitation id="EFADE0C3744BFFC5FD80FD60FD58DBE7" box="[571,727,692,716]" pageId="10" pageNumber="360" refId="ref12871" refString="Kataev, B. M. (2002) Revision of the Harpalus honestus group from the Himalaya and a review of the Harpalus species from Nepal (Coleoptera: Carabidae). Russian Entomological Journal, 10 (4), 357 - 384. [2001]" type="journal article">Kataev 2002</bibRefCitation>
|
||
). Аs in the case of
|
||
<taxonomicName id="4C3CE6B1744BFFC5FC62FD60FA80DBE7" authority="Kаtаev & Wrаse, 2018" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[985,1295,692,717]" class="Insecta" family="Carabidae" genus="Indocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="360" phylum="Arthropoda" rank="species" species="inexspectatus" status="sp. nov.">
|
||
<emphasis id="B9484120744BFFC5FC62FD60FA80DBE7" box="[985,1295,692,717]" italics="true" pageId="10" pageNumber="360">Indocarterus inexspectatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BFC5B744BFFC5FAA6FD60FAEBDBE6" box="[1309,1380,692,717]" pageId="10" pageNumber="360" rank="species">
|
||
<emphasis id="B9484120744BFFC5FAA6FD60FAEBDBE6" bold="true" box="[1309,1380,692,717]" pageId="10" pageNumber="360">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, the Mediterranean and
|
||
<collectingRegion id="49F853D0744BFFC5FEC9FD03FE41DBDB" box="[370,462,727,752]" country="Morocco" name="Oriental" pageId="10" pageNumber="360">Oriental</collectingRegion>
|
||
elements occur there together; this co-existence is a well-known characteristic feature of the fauna of the western parts of the Himalaya (
|
||
<bibRefCitation id="EFADE0C3744BFFC5FD6AFD28FCEEDA3E" box="[721,865,764,789]" pageId="10" pageNumber="360" refId="ref13365" refString="Mani, M. S. (1974 b) XXI. Biogeography of the Himalaya. In: Mani, M. S. (Ed.), Ecology and biogeography in India. Dr. W. Junk, The Hague, pp. 664 - 681. https: // doi. org / 10.1007 / 978 - 94 - 010 - 2331 - 3 _ 21" type="book chapter">Mani 1974b</bibRefCitation>
|
||
). It seems plausible that all such disjunctions were caused by similar climatic and orogenetic processes during the Neogene.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |