257 lines
34 KiB
XML
257 lines
34 KiB
XML
<document id="A4B89DEB8CDE1FC95BA1488FBD73659A" ID-CLB-Dataset="29018" ID-DOI="10.11646/zootaxa.4459.2.8" ID-GBIF-Dataset="f36314b8-a9a6-4bd2-8704-8923013c22b7" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458709" ID-ZooBank="7270DEAE-A08F-4015-8FE6-6E0D0DBD9C84" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539289409860" checkinUser="plazi" docAuthor="Kаtаev, Boris M. & Wrаse, Dаvid W." docDate="2018" docId="03952C247443FFCCFF2CFC22FB0DDC26" docLanguage="en" docName="zootaxa.4459.2.8.pdf" docOrigin="Zootaxa 4459 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parapenthus Kаtаev & Wrаse 2018, gen. n." docType="treatment" docVersion="8" lastPageNumber="353" masterDocId="FFAC545C7441FFCFFFBBFFD4FF8FD92B" masterDocTitle="Notes on Ditomina, with description of two new genera and a new species (Coleoptera, Carabidae) from the Middle East and India" masterLastPageNumber="368" masterPageNumber="350" pageNumber="352" updateTime="1698578541753" updateUser="ExternalLinkService">
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<mods:title id="1EEEC728C0C3B94D822B71CC61AF66C2">Notes on Ditomina, with description of two new genera and a new species (Coleoptera, Carabidae) from the Middle East and India</mods:title>
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<mods:namePart id="CE9AE4B7040DE4B416F0F80A33DA2410">Kаtаev, Boris M.</mods:namePart>
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<mods:namePart id="0D5DA65530519894C9B4702DA4176FB7">Wrаse, Dаvid W.</mods:namePart>
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<mods:title id="AD02697AB68F1D236B6EFB487C050455">Zootaxa</mods:title>
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<mods:date id="C70B1B29FA48B22BC423AB6269E1E463">2018</mods:date>
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<mods:number id="EB18D8065074A2B2F648A2B3D915214F">2018-08-15</mods:number>
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<treatment id="03952C247443FFCCFF2CFC22FB0DDC26" ID-DOI="http://doi.org/10.5281/zenodo.5980133" ID-GBIF-Taxon="148692765" ID-Zenodo-Dep="5980133" LSID="urn:lsid:plazi:treatment:03952C247443FFCCFF2CFC22FB0DDC26" httpUri="http://treatment.plazi.org/id/03952C247443FFCCFF2CFC22FB0DDC26" lastPageId="3" lastPageNumber="353" pageId="2" pageNumber="352">
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<subSubSection id="C326CEB97443FFCDFF2CFC22FE04DD3B" box="[151,395,1014,1040]" pageId="2" pageNumber="352" type="nomenclature">
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<paragraph id="8B839D327443FFCDFF2CFC22FE04DD3B" blockId="2.[151,395,1014,1040]" box="[151,395,1014,1040]" pageId="2" pageNumber="352">
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<heading id="D0CB2A5E7443FFCDFF2CFC22FE04DD3B" bold="true" box="[151,395,1014,1040]" fontSize="11" level="1" pageId="2" pageNumber="352" reason="1">
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<emphasis id="B94841207443FFCDFF2CFC22FE04DD3B" bold="true" box="[151,395,1014,1040]" pageId="2" pageNumber="352">
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<taxonomicName id="4C3CE6B17443FFCDFF2CFC22FEBDDD3B" authority="Kаtаev & Wrаse, 2018" authorityName="Kаtаev & Wrаse" authorityYear="2018" box="[151,306,1014,1040]" class="Insecta" family="Carabidae" genus="Parapenthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="352" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B94841207443FFCDFF2CFC22FEBDDD3B" bold="true" box="[151,306,1014,1040]" italics="true" pageId="2" pageNumber="352">Parapenthus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27BFC5B7443FFCDFE83FC22FE04DD3B" box="[312,395,1014,1040]" pageId="2" pageNumber="352" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C326CEB97443FFCDFF2CFBEAFDE7DD78" box="[151,616,1085,1107]" pageId="2" pageNumber="352" type="materials_examined">
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<paragraph id="8B839D327443FFCDFF2CFBEAFDE7DD78" blockId="2.[151,616,1085,1107]" box="[151,616,1085,1107]" pageId="2" pageNumber="352">
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Type species
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<taxonomicName id="4C3CE6B17443FFCDFEA4FBEAFDE7DD78" authority="Peyron, 1858" authorityName="Peyron" authorityYear="1858" box="[287,616,1085,1107]" class="Insecta" family="Carabidae" genus="Ditomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="352" phylum="Arthropoda" rank="species" species="solitarius">
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<emphasis id="B94841207443FFCDFEA4FBEAFE54DD78" box="[287,475,1085,1107]" italics="true" pageId="2" pageNumber="352">Ditomus solitarius</emphasis>
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<bibRefCitation id="EFADE0C37443FFCDFE5AFBEAFDE7DD78" box="[481,616,1085,1107]" pageId="2" pageNumber="352" refId="ref13577" refString="Peyron, E. (1858) Catalogue des coleopteres des environs de Tarsous (Caramanie) avec la description des especes nouvelle. Annales de la Societe Entomologique de France, Series 3, 6, 353 - 434." type="journal article">Peyron, 1858</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C326CEB97443FFCDFF2CFBABFEAADF0E" pageId="2" pageNumber="352" type="diagnosis">
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<paragraph id="8B839D327443FFCDFF2CFBABFE8EDC03" blockId="2.[151,1437,1151,2005]" pageId="2" pageNumber="352">
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<emphasis id="B94841207443FFCDFF2CFBABFE85DDB3" bold="true" box="[151,266,1151,1176]" pageId="2" pageNumber="352">Diagnosis</emphasis>
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. Most similar in combination of characters to
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<taxonomicName id="4C3CE6B17443FFCDFCACFBABFCFEDDB3" authorityName="Chaudoir" authorityYear="1843" box="[791,881,1151,1176]" class="Insecta" family="Carabidae" genus="Penthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="352" phylum="Arthropoda" rank="genus">
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<emphasis id="B94841207443FFCDFCACFBABFCFEDDB3" box="[791,881,1151,1176]" italics="true" pageId="2" pageNumber="352">Penthus</emphasis>
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</taxonomicName>
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differing as follows: dorsum with violet metallic tinge; labrum rather deeply emarginate anteriorly; pronotum not bordered along apical and basal margins, its apical angles markedly protruding anteriad, very narrowly rounded at apex, basal angles almost rectangular, with sharp apex; and protibia with about 10–12 lateral spines situated in apical half of outer edge (versus 6–8 spines in
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<taxonomicName id="4C3CE6B17443FFCDFF2CFADBFF7EDC03" authorityName="Chaudoir" authorityYear="1843" box="[151,241,1295,1320]" class="Insecta" family="Carabidae" genus="Penthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="352" phylum="Arthropoda" rank="genus">
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<emphasis id="B94841207443FFCDFF2CFADBFF7EDC03" box="[151,241,1295,1320]" italics="true" pageId="2" pageNumber="352">Penthus</emphasis>
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</taxonomicName>
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).
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</paragraph>
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<paragraph id="8B839D327443FFCDFF7CFAE0FEAADF0E" blockId="2.[151,1437,1151,2005]" pageId="2" pageNumber="352">
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The new genus differs from
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<taxonomicName id="4C3CE6B17443FFCDFDB3FAE0FD27DC66" authorityName="Reitter" authorityYear="1900" box="[520,680,1332,1357]" class="Insecta" family="Carabidae" genus="Penthophonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="352" phylum="Arthropoda" rank="genus">
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<emphasis id="B94841207443FFCDFDB3FAE0FD27DC66" box="[520,680,1332,1357]" italics="true" pageId="2" pageNumber="352">Penthophonus</emphasis>
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</taxonomicName>
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in having the following distinctive features: head relatively larger; paraglossae narrow, separated from ligular sclerite by wide notch; apical angles of pronotum markedly protruding anteriad, very narrowly rounded at apex; three penultimate abdominal sternites, in addition to short general pubescence, also with several long setae along hind margin; metatrochanter large, more than a half as long as metafemur, slightly acute at apex; and protibia with about 10–12 lateral spines situated in apical half of outer edge (versus 3–4 spines in
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<taxonomicName id="4C3CE6B17443FFCDFE2FFA33FDBBDF2B" authorityName="Reitter" authorityYear="1900" box="[404,564,1511,1536]" class="Insecta" family="Carabidae" genus="Penthophonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="352" phylum="Arthropoda" rank="genus">
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<emphasis id="B94841207443FFCDFE2FFA33FDBBDF2B" box="[404,564,1511,1536]" italics="true" pageId="2" pageNumber="352">Penthophonus</emphasis>
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</taxonomicName>
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), with outer angle not dentiform and without additional short spur on inner apical angle.
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</paragraph>
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</subSubSection>
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<subSubSection id="C326CEB97443FFCCFF7CF9FBFB5EDB4B" lastPageId="3" lastPageNumber="353" pageId="2" pageNumber="352" type="description">
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<paragraph id="8B839D327443FFCDFF7CF9FBFE23DF47" blockId="2.[151,1437,1151,2005]" pageId="2" pageNumber="352">
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<emphasis id="B94841207443FFCDFF7CF9FBFEDEDF63" bold="true" box="[199,337,1583,1608]" pageId="2" pageNumber="352">Description</emphasis>
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. Body (
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<figureCitation id="130781B77443FFCDFE09F9FBFE79DF63" box="[434,502,1583,1608]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1592,1615]" captionTargetBox="[286,1307,608,1559]" captionTargetId="figure@4.[274,1313,569,1568]" captionTargetPageId="4" captionText="FIGURES 1–4. Parapenthus solitarius (Peyron, 1858), female (Ceyhan, Turkey). 1, habitus; 2, left protibia; 3, head; 4, labium. 1–3, dorsal view, 4, ventral view. Scale bars = 1.0 mm (A: Fig. 1; B: Figs 2, 3; C: Fig. 4)." httpUri="https://zenodo.org/record/1458711/files/figure.png" pageId="2" pageNumber="352">Fig. 1</figureCitation>
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) robust, dorsum with violet metallic tinge, rather densely and roughly punctate, covered with erect setae.
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</paragraph>
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<paragraph id="8B839D327443FFCDFF7CF9A3FAA5DE4C" blockId="2.[151,1437,1151,2005]" pageId="2" pageNumber="352">
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Labrum (
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<figureCitation id="130781B77443FFCDFE94F9A3FEFEDFBB" box="[303,369,1655,1680]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1592,1615]" captionTargetBox="[286,1307,608,1559]" captionTargetId="figure@4.[274,1313,569,1568]" captionTargetPageId="4" captionText="FIGURES 1–4. Parapenthus solitarius (Peyron, 1858), female (Ceyhan, Turkey). 1, habitus; 2, left protibia; 3, head; 4, labium. 1–3, dorsal view, 4, ventral view. Scale bars = 1.0 mm (A: Fig. 1; B: Figs 2, 3; C: Fig. 4)." httpUri="https://zenodo.org/record/1458711/files/figure.png" pageId="2" pageNumber="352">Fig. 3</figureCitation>
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) rather deeply emarginate anteriorly. Аpical margin of clypeus shallowly emarginate. Tempora setose. Genae wide. One very long supraorbital seta on each side. Lateral margin of frons above insertion of antennae simple, not widened. Mentum (
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<figureCitation id="130781B77443FFCDFDC1F96BFD4DDFF3" box="[634,706,1727,1752]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1592,1615]" captionTargetBox="[286,1307,608,1559]" captionTargetId="figure@4.[274,1313,569,1568]" captionTargetPageId="4" captionText="FIGURES 1–4. Parapenthus solitarius (Peyron, 1858), female (Ceyhan, Turkey). 1, habitus; 2, left protibia; 3, head; 4, labium. 1–3, dorsal view, 4, ventral view. Scale bars = 1.0 mm (A: Fig. 1; B: Figs 2, 3; C: Fig. 4)." httpUri="https://zenodo.org/record/1458711/files/figure.png" pageId="2" pageNumber="352">Fig. 4</figureCitation>
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) with wide and very short median tooth. Epilobes noticeably widened apically. Ligular sclerite very slightly narrowed before truncate apex, with two long ventro-apical setae and a few very short setae on dorsal side. Paraglossae glabrous, narrow, longer than ligular sclerite and separated from it by wide notch. Terminal maxillary and labial palpomeres more or less cylindrical, usually narrowed apically. Basal labial palpomere carinate. Penultimate labial palpomere markedly longer than apical one.
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</paragraph>
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<paragraph id="8B839D327443FFCDFF7CF8A1FD40DEFE" blockId="2.[151,1437,1151,2005]" pageId="2" pageNumber="352">Pronotum cordate, with two or several setigerous pores at sides before middle. Аpical angles markedly protruding anteriad, very narrowly rounded at apex. Basal angles almost rectangular. Аpical and basal margins not bordered, basal pronotal edge glabrous, not ciliate.</paragraph>
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<paragraph id="8B839D327442FFCCFF7CFF43FB28D99B" blockId="3.[151,1437,151,1293]" box="[199,1191,151,176]" pageId="3" pageNumber="353">Elytra not serrate at sides, basal border complete, not bent and not raised near scutellum.</paragraph>
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<paragraph id="8B839D327442FFCCFF7CFF68FBF0D9D3" blockId="3.[151,1437,151,1293]" pageId="3" pageNumber="353">Hindwings fully developed, folded at apex (obviously the species is capable of flight; one of the examined specimens was covered with Lepidopteran scales and might have been collected at light).</paragraph>
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<paragraph id="8B839D327442FFCCFF7CFED1FAE6D8A3" blockId="3.[151,1437,151,1293]" pageId="3" pageNumber="353">
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Prosternum with rather long setae medially. Metepisternum markedly longer than wide. Three penultimate abdominal sternites, in addition to short general pubescence, also with several long setae along hind margin (similar to those of
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<taxonomicName id="4C3CE6B17442FFCCFEC7FE98FE59D84E" authorityName="Chaudoir" authorityYear="1843" box="[380,470,332,357]" class="Insecta" family="Carabidae" genus="Penthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="genus">
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<emphasis id="B94841207442FFCCFEC7FE98FE59D84E" box="[380,470,332,357]" italics="true" pageId="3" pageNumber="353">Penthus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C3CE6B17442FFCCFE5DFE98FD73D84E" authority=", Chilotomus" authorityName="Chilotomus" box="[486,764,332,357]" class="Insecta" family="Carabidae" genus="Pachycarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="genus">
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<emphasis id="B94841207442FFCCFE5DFE98FDE5D84E" box="[486,618,332,357]" italics="true" pageId="3" pageNumber="353">Pachycarus</emphasis>
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,
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<emphasis id="B94841207442FFCCFDC0FE98FD73D84E" box="[635,764,332,357]" italics="true" pageId="3" pageNumber="353">Chilotomus</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C3CE6B17442FFCCFC85FE98FC4FD84E" authorityName="Semenov" authorityYear="1891" box="[830,960,332,357]" class="Insecta" family="Carabidae" genus="Bronislavia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="genus">
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<emphasis id="B94841207442FFCCFC85FE98FC4FD84E" box="[830,960,332,357]" italics="true" pageId="3" pageNumber="353">Bronislavia</emphasis>
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</taxonomicName>
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, but such setae absent in
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<taxonomicName id="4C3CE6B17442FFCCFB50FE98FA04D84E" authorityName="Reitter" authorityYear="1900" box="[1259,1419,332,357]" class="Insecta" family="Carabidae" genus="Penthophonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="genus">
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<emphasis id="B94841207442FFCCFB50FE98FA04D84E" box="[1259,1419,332,357]" italics="true" pageId="3" pageNumber="353">Penthophonus</emphasis>
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</taxonomicName>
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). Аnal (last visible) abdominal sternite in female with distal margin medially somewhat enlarged and plate-like.
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</paragraph>
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<paragraph id="8B839D327442FFCCFF7CFE40FB5EDB4B" blockId="3.[151,1437,151,1293]" pageId="3" pageNumber="353">
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Protibia (
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<figureCitation id="130781B77442FFCCFE94FE40FEFDD887" box="[303,370,404,429]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1592,1615]" captionTargetBox="[286,1307,608,1559]" captionTargetId="figure@4.[274,1313,569,1568]" captionTargetPageId="4" captionText="FIGURES 1–4. Parapenthus solitarius (Peyron, 1858), female (Ceyhan, Turkey). 1, habitus; 2, left protibia; 3, head; 4, labium. 1–3, dorsal view, 4, ventral view. Scale bars = 1.0 mm (A: Fig. 1; B: Figs 2, 3; C: Fig. 4)." httpUri="https://zenodo.org/record/1458711/files/figure.png" pageId="3" pageNumber="353">Fig. 2</figureCitation>
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) with outer angle not dentiform, without additional short spur on inner apical angle, with about 10–12 lateral spines situated in apical half of outer edge and with two ventroapical spines. Аpical spur of protibia slender. Profemur not denticulate along inner margin. Metafemur with many setigerous punctures along posterior margin. Metacoxa with a posteromedial setigerous pore and with numerous additional long setae. Metatrochanter large, more than a half as long as metafemur, slightly acute at apex. Tarsi setose dorsally. Metatarsomere 1 short, slightly longer than 2, noticeably shorter than 2+3. Protarsus and mesotarsus of male not dilated.
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</paragraph>
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</subSubSection>
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<subSubSection id="C326CEB97442FFCCFF7CFDB8FDCADBDB" pageId="3" pageNumber="353" type="etymology">
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<paragraph id="8B839D327442FFCCFF7CFDB8FECFDB8C" blockId="3.[151,1437,151,1293]" pageId="3" pageNumber="353">
|
||
<emphasis id="B94841207442FFCCFF7CFDB8FEC9DBAE" bold="true" box="[199,326,620,645]" pageId="3" pageNumber="353">Etymology</emphasis>
|
||
. The genus name is a combination of the Greek
|
||
<emphasis id="B94841207442FFCCFCC8FDB9FC27DBAF" box="[883,936,621,644]" italics="true" pageId="3" pageNumber="353">para</emphasis>
|
||
, meaning near, and the name of the carabid genus
|
||
<taxonomicName id="4C3CE6B17442FFCCFF64FD5BFEB6DB83" authorityName="Chaudoir" authorityYear="1843" box="[223,313,655,680]" class="Insecta" family="Carabidae" genus="Penthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207442FFCCFF64FD5BFEB6DB83" box="[223,313,655,680]" italics="true" pageId="3" pageNumber="353">Penthus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B839D327442FFCCFF7CFD60FDCADBDB" blockId="3.[151,1437,151,1293]" pageId="3" pageNumber="353">
|
||
<emphasis id="B94841207442FFCCFF7CFD60FDA4DBE6" bold="true" box="[199,555,692,717]" pageId="3" pageNumber="353">Composition and distribution</emphasis>
|
||
. The genus includes only one species,
|
||
<taxonomicName id="4C3CE6B17442FFCCFC50FD61FBFDDBE6" authorityName="Peyron" authorityYear="1858" box="[1003,1138,692,717]" class="Insecta" family="Carabidae" genus="Ditomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="species" species="solitarius">
|
||
<emphasis id="B94841207442FFCCFC50FD61FBFDDBE6" box="[1003,1138,692,717]" italics="true" pageId="3" pageNumber="353">P. solitarius</emphasis>
|
||
</taxonomicName>
|
||
, distributed in the Middle East (
|
||
<collectingCountry id="F32BDDA27442FFCCFF6DFD03FEA6DBDB" box="[214,297,727,752]" name="Turkey" pageId="3" pageNumber="353">Turkey</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32BDDA27442FFCCFE8FFD03FEFDDBDB" box="[308,370,727,752]" name="Syria" pageId="3" pageNumber="353">Syria</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32BDDA27442FFCCFEC7FD03FE35DBDB" box="[380,442,727,752]" name="Israel" pageId="3" pageNumber="353">Israel</collectingCountry>
|
||
and
|
||
<collectingCountry id="F32BDDA27442FFCCFE4AFD0CFDB6DBDB" box="[497,569,728,752]" name="Egypt" pageId="3" pageNumber="353">Egypt</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326CEB97442FFCCFF7CFD28FB0DDC26" pageId="3" pageNumber="353" type="discussion">
|
||
<paragraph id="8B839D327442FFCCFF7CFD28FB0DDC26" blockId="3.[151,1437,151,1293]" pageId="3" pageNumber="353">
|
||
<emphasis id="B94841207442FFCCFF7CFD28FEBBDA3E" bold="true" box="[199,308,764,789]" pageId="3" pageNumber="353">Remarks</emphasis>
|
||
. This new genus is erected for the single species
|
||
<taxonomicName id="4C3CE6B17442FFCCFCE6FD29FC6DDA3E" authorityName="Peyron" authorityYear="1858" box="[861,994,764,789]" class="Insecta" family="Carabidae" genus="Ditomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="species" species="solitarius">
|
||
<emphasis id="B94841207442FFCCFCE6FD29FC6DDA3E" box="[861,994,764,789]" italics="true" pageId="3" pageNumber="353">P. solitarius</emphasis>
|
||
</taxonomicName>
|
||
, which was originally described within
|
||
<taxonomicName id="4C3CE6B17442FFCCFF2CFCF5FE12DA13" authority="Bonelli, 1810" authorityName="Bonelli" authorityYear="1810" box="[151,413,799,824]" class="Insecta" family="Carabidae" genus="Ditomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207442FFCCFF2CFCF5FF79DA13" box="[151,246,801,824]" italics="true" pageId="3" pageNumber="353">Ditomus</emphasis>
|
||
Bonelli, 1810
|
||
</taxonomicName>
|
||
, but later was treated as a member of the genus
|
||
<taxonomicName id="4C3CE6B17442FFCCFC6AFCCBFB6BDA13" authority="Chaudoir, 1843" authorityName="Chaudoir" authorityYear="1843" box="[977,1252,799,824]" class="Insecta" family="Carabidae" genus="Penthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207442FFCCFC6AFCCBFBA3DA13" box="[977,1068,799,824]" italics="true" pageId="3" pageNumber="353">Penthus</emphasis>
|
||
Chaudoir, 1843
|
||
</taxonomicName>
|
||
(Piochard de la
|
||
<bibRefCitation id="EFADE0C37442FFCCFF2CFC90FEBBDA77" box="[151,308,836,861]" pageId="3" pageNumber="353" refId="ref13618" refString="Piochard de la Brulerie, C. J. (1873) Monographie des Ditomides, tribu des carabiques. L'Abeille, Journal d'Entomologie, 15, i - viii, 1 - 100." type="book chapter">Brûlerie 1873</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFADE0C37442FFCCFE85FC90FE43DA77" box="[318,460,836,861]" pageId="3" pageNumber="353" refId="ref13653" refString="Reitter, E. (1900) Bestimmungs-Tabelle der europaischen Coleopteren. XLI. Heft. Enthaltend: Carabidae. Abtheilung: Harpalini und Licinini. Verhandlungen des naturforschenden Vereines in Brunn, 38, 33 - 155. [1899]" type="journal article">Reitter 1900</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFADE0C37442FFCCFE6DFC90FD37DA77" box="[470,696,836,861]" pageId="3" pageNumber="353" refId="ref14280" refString="Tschitscherine, T. (1901) Genera des Harpalini des regions Palearctique et Paleanarctique. Horae Societatis Entomologicae Rossicae, 35, 217 - 251." type="journal article">Tschitschérine 1901</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFADE0C37442FFCCFD7FFC90FCE2DA77" box="[708,877,836,861]" pageId="3" pageNumber="353" refId="ref12605" refString="Jakobson, G. G. (1907) Fasc. 5. In: Zhuki Rossii i Zapadnoi Evropy. A. F. Devrien, Sankt-Petersburg, pp. 321 - 400." type="book chapter">Jakobson 1907</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFADE0C37442FFCCFCC3FC90FB89DA77" box="[888,1030,836,861]" pageId="3" pageNumber="353" refId="ref14177" refString="Stichel, W. (1923) Zur Phylogenesis eines geologisch jungen Formenkreises der Kafer, der Ditominen (Carab. Harpal.). Zeitschrift fur wissenschaftliche Insektenbiologie, 18, 41 - 50 + 81 - 100 + 145 - 162 + 209 - 242." type="journal article">Stichel 1923</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFADE0C37442FFCCFBAAFC90FB01DA77" box="[1041,1166,836,861]" pageId="3" pageNumber="353" refId="ref12345" refString="Csiki, E. (1932) Carabidae: Harpalinae. Vol. VI. Pars 121. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. Vol. III. Carabidae III. W. Junk, Berlin, pp. 1023 - 1278." type="book chapter">Csiki 1932</bibRefCitation>
|
||
). Based on the available descriptions of this species, without examination of the specimens,
|
||
<bibRefCitation id="EFADE0C37442FFCCFC11FCB3FBC3DAAB" box="[938,1100,871,896]" pageId="3" pageNumber="353" refId="ref13767" refString="Sciaky, R. (1987) Revisione delle specie paleartiche occidentali del genere Ophonus Dejean, 1821 (Coleoptera, Carabidae). (XVIII contributo alla conoscenza dei Coleoptera Carabidae). Memorie della Societa Entomologica Italiana, 65, 29 - 120. [1986]" type="journal article">Sciaky (1987)</bibRefCitation>
|
||
included it in
|
||
<taxonomicName id="4C3CE6B17442FFCCFB47FCB3FEA8DA8F" authority="Reitter, 1900" authorityName="Reitter" authorityYear="1900" class="Insecta" family="Carabidae" genus="Penthophonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207442FFCCFB47FCB3FA13DAAB" box="[1276,1436,871,896]" italics="true" pageId="3" pageNumber="353">Penthophonus</emphasis>
|
||
<bibRefCitation id="EFADE0C37442FFCCFF2CFC58FEA8DA8F" box="[151,295,908,933]" pageId="3" pageNumber="353" refId="ref13653" refString="Reitter, E. (1900) Bestimmungs-Tabelle der europaischen Coleopteren. XLI. Heft. Enthaltend: Carabidae. Abtheilung: Harpalini und Licinini. Verhandlungen des naturforschenden Vereines in Brunn, 38, 33 - 155. [1899]" type="journal article">Reitter, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
(a taxon very close to the genus
|
||
<taxonomicName id="4C3CE6B17442FFCCFD28FC58FC19DA8F" authority="Dejean, 1921" authorityName="Dejean" authorityYear="1921" box="[659,918,908,933]" class="Insecta" family="Carabidae" genus="Ophonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207442FFCCFD28FC58FD77DA8E" box="[659,760,908,933]" italics="true" pageId="3" pageNumber="353">Ophonus</emphasis>
|
||
Dejean, 1921
|
||
</taxonomicName>
|
||
), and this treatment was tacitly accepted in the subsequent literature (
|
||
<bibRefCitation id="EFADE0C37442FFCCFE2BFC64FDACDAE3" box="[400,547,944,968]" pageId="3" pageNumber="353" refId="ref13151" refString="Lorenz, W. (1998) Systematic List of Extant Ground Beetles of the World (Insecta Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 1 st Edition. W. Lorenz, Tutzing, 502 pp." type="book">Lorenz 1998</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFADE0C37442FFCCFD8BFC7BFDE4DAE3" box="[560,619,943,968]" pageId="3" pageNumber="353" refId="ref13199" refString="Lorenz, W. (2005) Systematic List of Extant Ground Beetles of the World (Insecta Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. W. Lorenz, Tutzing, 530 pp." type="book">2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFADE0C37442FFCCFDC3FC64FC8DDAE3" box="[632,770,944,968]" pageId="3" pageNumber="353" refId="ref14452" refString="Wrase, D. W. (2003) Subtribe Ditomina Bonelli, 1810. In: Lobl, L. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata-Myxophaga-Adephaga. Apollo Books, Stenstrup, pp. 363 - 367." type="book chapter">Wrase 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFADE0C37442FFCCFCB5FC64FB80DAE3" box="[782,1039,944,968]" pageId="3" pageNumber="353" refId="ref14504" refString="Wrase, D. W. & Kataev, B. M. (2017) Subtribe Ditomina Bonelli, 1810. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Revised and Updated Edition. Archostemata-Myxophaga-Adephaga. Brill, Leiden / Boston, pp. 509 - 514." type="book chapter">Wrase & Kataev 2017</bibRefCitation>
|
||
). Re-examination of the specimens of
|
||
<taxonomicName id="4C3CE6B17442FFCCFF0CFC01FEB0DAC6" authorityName="Peyron" authorityYear="1858" box="[183,319,980,1005]" class="Insecta" family="Carabidae" genus="Ditomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="species" species="solitarius">
|
||
<emphasis id="B94841207442FFCCFF0CFC01FEB0DAC6" box="[183,319,980,1005]" italics="true" pageId="3" pageNumber="353">P. solitarius</emphasis>
|
||
</taxonomicName>
|
||
revealed that this species is distinguished from the members of
|
||
<taxonomicName id="4C3CE6B17442FFCCFB98FC00FB4CDAC6" authorityName="Reitter" authorityYear="1900" box="[1059,1219,980,1005]" class="Insecta" family="Carabidae" genus="Penthophonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207442FFCCFB98FC00FB4CDAC6" box="[1059,1219,980,1005]" italics="true" pageId="3" pageNumber="353">Penthophonus</emphasis>
|
||
</taxonomicName>
|
||
in many sufficient characters (see diagnosis above) and actually most similar and apparently most related to
|
||
<taxonomicName id="4C3CE6B17442FFCCFB7AFC23FA94DD3B" authorityName="Chaudoir" authorityYear="1843" box="[1217,1307,1015,1040]" class="Insecta" family="Carabidae" genus="Penthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207442FFCCFB7AFC23FA94DD3B" box="[1217,1307,1015,1040]" italics="true" pageId="3" pageNumber="353">Penthus</emphasis>
|
||
</taxonomicName>
|
||
. The most important characters in common for
|
||
<taxonomicName id="4C3CE6B17442FFCCFDEDFBC8FCD1DD1E" baseAuthorityName="Peyron" baseAuthorityYear="1858" box="[598,862,1052,1077]" class="Insecta" family="Carabidae" genus="Parapenthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="species" species="solitarius">
|
||
<emphasis id="B94841207442FFCCFDEDFBC8FCD1DD1E" box="[598,862,1052,1077]" italics="true" pageId="3" pageNumber="353">Parapenthus solitarius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3CE6B17442FFCCFC18FBC8FC72DD1E" authorityName="Chaudoir" authorityYear="1843" box="[931,1021,1052,1077]" class="Insecta" family="Carabidae" genus="Penthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207442FFCCFC18FBC8FC72DD1E" box="[931,1021,1052,1077]" italics="true" pageId="3" pageNumber="353">Penthus</emphasis>
|
||
</taxonomicName>
|
||
, and absent in
|
||
<taxonomicName id="4C3CE6B17442FFCCFB78FBC8FAECDD1E" authorityName="Reitter" authorityYear="1900" box="[1219,1379,1052,1077]" class="Insecta" family="Carabidae" genus="Penthophonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94841207442FFCCFB78FBC8FAECDD1E" box="[1219,1379,1052,1077]" italics="true" pageId="3" pageNumber="353">Penthophonus</emphasis>
|
||
</taxonomicName>
|
||
are: paraglossae narrow, separated from ligular sclerite by wide notch; abdominal sternites with transverse row of long setae along posterior margin; protibia without additional short spur on inner apical angle, with more than one ventroapical and more than four outer preapical spines; metafemur with many setigerous punctures along posterior margin; and metatrochanter long, about half as long as metafemur, acute at apex.
|
||
<taxonomicName id="4C3CE6B17442FFCCFB88FB78FABBDDEE" baseAuthorityName="Peyron" baseAuthorityYear="1858" box="[1075,1332,1196,1221]" class="Insecta" family="Carabidae" genus="Parapenthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="species" species="solitarius">
|
||
<emphasis id="B94841207442FFCCFB88FB78FABBDDEE" box="[1075,1332,1196,1221]" italics="true" pageId="3" pageNumber="353">Parapenthus solitarius</emphasis>
|
||
</taxonomicName>
|
||
warrants separate generic status due to the peculiar combination of the sufficient characters distinguishing it from the single representative of the genus
|
||
<emphasis id="B94841207442FFCCFE70FB20FDCEDC27" box="[459,577,1268,1293]" italics="true" pageId="3" pageNumber="353">
|
||
<taxonomicName id="4C3CE6B17442FFCCFE70FB20FDAADC26" authorityName="Chaudoir" authorityYear="1843" box="[459,549,1268,1293]" class="Insecta" family="Carabidae" genus="Penthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="genus">Penthus</taxonomicName>
|
||
—
|
||
</emphasis>
|
||
<taxonomicName id="4C3CE6B17442FFCCFDFAFB21FC0DDC27" authority="(Waltl, 1838)" baseAuthorityName="Waltl" baseAuthorityYear="1838" box="[577,898,1268,1293]" class="Insecta" family="Carabidae" genus="Penthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="species" species="tenebrioides">
|
||
<emphasis id="B94841207442FFCCFDFAFB21FD69DC26" box="[577,742,1268,1293]" italics="true" pageId="3" pageNumber="353">P. tenebrioides</emphasis>
|
||
(Waltl, 1838)
|
||
</taxonomicName>
|
||
(see diagnosis above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |