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<document id="0A68B31C9E6D43442EF7FDA4E4A45F90" ID-CLB-Dataset="32278" ID-DOI="10.11646/zootaxa.4312.2.10" ID-GBIF-Dataset="fbc8059e-55b6-46b1-b18e-8854b042d3fd" ID-ISSN="1175-5326" ID-Zenodo-Dep="852742" ID-ZooBank="618B7E19-Af76-4Bcb-Bb95-2B31F1E4Bc9C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1503986340795" checkinUser="plazi" docAuthor="Zhang, Yalin" docDate="2017" docId="D06CCB3BFFE24F30D6C6FDEEE8A64E45" docLanguage="en" docName="zootaxa.4312.2.10.pdf" docOrigin="Zootaxa 4312 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kodaianellissus Zhang 2017, gen. nov." docType="treatment" docVersion="7" lastPageNumber="357" masterDocId="2C55B343FFE34F32D651FF8FEA574F05" masterDocTitle="New Oriental genera in the family Issidae (Hemiptera: Fulgoromorpha)" masterLastPageNumber="367" masterPageNumber="355" pageNumber="356" updateTime="1698467455336" updateUser="ExternalLinkService">
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<mods:title id="9A4E5254204C6299A1C3C935C1234535">New Oriental genera in the family Issidae (Hemiptera: Fulgoromorpha)</mods:title>
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<mods:namePart id="931564A3C2E55D6772413B0F204C2AEE">Zhang, Yalin</mods:namePart>
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<mods:date id="A9C7438A2FA72DE67532F3CE0990B203">2017</mods:date>
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<treatment id="D06CCB3BFFE24F30D6C6FDEEE8A64E45" ID-DOI="http://doi.org/10.5281/zenodo.6032764" ID-GBIF-Taxon="133010551" ID-Zenodo-Dep="6032764" LSID="urn:lsid:plazi:treatment:D06CCB3BFFE24F30D6C6FDEEE8A64E45" httpUri="http://treatment.plazi.org/id/D06CCB3BFFE24F30D6C6FDEEE8A64E45" lastPageId="2" lastPageNumber="357" pageId="1" pageNumber="356">
<subSubSection id="10DF29A6FFE24F33D6C6FDEEEB9D4D7E" box="[151,458,609,635]" pageId="1" pageNumber="356" type="nomenclature">
<paragraph id="587A7A2DFFE24F33D6C6FDEEEB9D4D7E" blockId="1.[151,458,609,635]" box="[151,458,609,635]" pageId="1" pageNumber="356">
<heading id="0332CD41FFE24F33D6C6FDEEEB9D4D7E" bold="true" box="[151,458,609,635]" fontSize="11" level="1" pageId="1" pageNumber="356" reason="1">
<emphasis id="6AB1A63FFFE24F33D6C6FDEEEB9D4D7E" bold="true" box="[151,458,609,635]" pageId="1" pageNumber="356">
<taxonomicName id="9FC501AEFFE24F33D6C6FDEEEB014D7E" ID-CoL="58N3" ID-ENA="2725186" authority="Zhang, 2017" authorityName="Zhang" authorityYear="2017" box="[151,342,609,635]" class="Insecta" family="Issidae" genus="Kodaianellissus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="356" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6AB1A63FFFE24F33D6C6FDEEEB014D7E" bold="true" box="[151,342,609,635]" italics="true" pageId="1" pageNumber="356">Kodaianellissus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71821B44FFE24F33D70DFDEEEB9D4D7E" box="[348,458,609,635]" pageId="1" pageNumber="356" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="10DF29A6FFE24F33D6C6FD26E9144DBB" box="[151,835,680,702]" pageId="1" pageNumber="356" type="materials_examined">
<paragraph id="587A7A2DFFE24F33D6C6FD26E9144DBB" blockId="1.[151,835,680,702]" box="[151,835,680,702]" pageId="1" pageNumber="356">
Type species:
<taxonomicName id="9FC501AEFFE24F33D777FD27E86D4DBB" authority="Zhang, 2017" authorityName="Zhang" authorityYear="2017" box="[294,570,680,702]" class="Insecta" family="Issidae" genus="Kodaianellissus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="356" phylum="Arthropoda" rank="species" species="intorqueus" status="sp. nov.">
<emphasis id="6AB1A63FFFE24F33D777FD27E86D4DBB" box="[294,570,680,702]" italics="true" pageId="1" pageNumber="356">Kodaianellissus intorqueus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71821B44FFE24F33D46EFD27E8C74DB8" box="[575,656,680,701]" pageId="1" pageNumber="356" rank="species">
<emphasis id="6AB1A63FFFE24F33D46EFD27E8C74DB8" bold="true" box="[575,656,680,701]" pageId="1" pageNumber="356">sp. nov.</emphasis>
</taxonomicNameLabel>
, here designated.
</paragraph>
</subSubSection>
<subSubSection id="10DF29A6FFE24F33D6C6FD65EFC54C9D" pageId="1" pageNumber="356" type="diagnosis">
<paragraph id="587A7A2DFFE24F33D6C6FD65EFC54C9D" blockId="1.[151,1437,746,2012]" pageId="1" pageNumber="356">
<emphasis id="6AB1A63FFFE24F33D6C6FD65EB464C06" bold="true" box="[151,273,746,771]" pageId="1" pageNumber="356">Diagnosis.</emphasis>
This genus is very similar to the genus
<taxonomicName id="9FC501AEFFE24F33D49DFD65E9A14C06" authority="Fennah, 1956" authorityName="Fennah" authorityYear="1956" box="[716,1014,746,771]" class="Insecta" family="Issidae" genus="Kodaianella" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="356" phylum="Arthropoda" rank="genus">
<emphasis id="6AB1A63FFFE24F33D49DFD65E9004C06" box="[716,855,746,771]" italics="true" pageId="1" pageNumber="356">Kodaianella</emphasis>
Fennah,
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in general appearance, but differs by: 1) ScP-R-MP-Cu lobe of hindwing around 1.5 times wider than Pcu-A1 lobe (
<figureCitation id="C0FE66A8FFE24F33D5ACFC81EE164C22" box="[1021,1089,782,807]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1369,1392]" captionTargetBox="[253,1333,384,1336]" captionTargetId="figure@2.[238,1348,375,1347]" captionTargetPageId="2" captionText="FIGURES 1 6. Kodaianellissus intorqueus sp. nov. 1. Adult, dorsal view; 2. Adult, lateral view; 3. Head and thorax; 4. Frons and clypeus; 5. Forewing; 6. Hindwing." httpUri="https://zenodo.org/record/852744/files/figure.png" pageId="1" pageNumber="356">Fig. 6</figureCitation>
), but the latter much narrower, 4 times wider than Pcu-A1 lobe (
<bibRefCitation id="3C5407DCFFE24F33D45EFCBAE94C4C4B" author="Gnezdilov" box="[527,795,821,846]" pageId="1" pageNumber="356" refString="Gnezdilov, V. M. (2013) On the genera Sivaloka Distant, 1906 and Kodaianella Fennah, 1956 (Hemiptera: Fulgoroidea: Issidae). Deutsche Entomologische Zeitschrift, 60 (1), 41 - 44." type="journal article" year="2013">Gnezdilov, 2013, fig. 6</bibRefCitation>
); 2) The forewing, in lateral view, broadest at basal 1/3 (
<figureCitation id="C0FE66A8FFE24F33D6CEFCD3EAB24C70" box="[159,229,860,885]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1369,1392]" captionTargetBox="[253,1333,384,1336]" captionTargetId="figure@2.[238,1348,375,1347]" captionTargetPageId="2" captionText="FIGURES 1 6. Kodaianellissus intorqueus sp. nov. 1. Adult, dorsal view; 2. Adult, lateral view; 3. Head and thorax; 4. Frons and clypeus; 5. Forewing; 6. Hindwing." httpUri="https://zenodo.org/record/852744/files/figure.png" pageId="1" pageNumber="356">Fig. 2</figureCitation>
), but broadest from basal 1/3 to apex in
<taxonomicName id="9FC501AEFFE24F33D4E7FCD3E9164C70" authorityName="Fennah" authorityYear="1956" box="[694,833,860,885]" class="Insecta" family="Issidae" genus="Kodaianella" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="356" phylum="Arthropoda" rank="genus">
<emphasis id="6AB1A63FFFE24F33D4E7FCD3E9164C70" box="[694,833,860,885]" italics="true" pageId="1" pageNumber="356">Kodaianella</emphasis>
</taxonomicName>
(
<bibRefCitation id="3C5407DCFFE24F33D500FCD3EE0D4C70" author="Gnezdilov" box="[849,1114,860,885]" pageId="1" pageNumber="356" refString="Gnezdilov, V. M. (2013) On the genera Sivaloka Distant, 1906 and Kodaianella Fennah, 1956 (Hemiptera: Fulgoroidea: Issidae). Deutsche Entomologische Zeitschrift, 60 (1), 41 - 44." type="journal article" year="2013">Gnezdilov, 2013, fig. 3</bibRefCitation>
); 3) Frons slightly narrower than in
<taxonomicName id="9FC501AEFFE24F33D6BAFC0FEB224C9C" authorityName="Fennah" authorityYear="1956" box="[235,373,896,921]" class="Insecta" family="Issidae" genus="Kodaianella" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="356" phylum="Arthropoda" rank="genus">
<emphasis id="6AB1A63FFFE24F33D6BAFC0FEB224C9C" box="[235,373,896,921]" italics="true" pageId="1" pageNumber="356">Kodaianella</emphasis>
</taxonomicName>
, frons just 1.3 times broader at widest part than long in midline, but 1.5 times in
<taxonomicName id="9FC501AEFFE24F33D351FC0FEFDC4C9C" authorityName="Fennah" authorityYear="1956" box="[1280,1419,896,921]" class="Insecta" family="Issidae" genus="Kodaianella" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="356" phylum="Arthropoda" rank="genus">
<emphasis id="6AB1A63FFFE24F33D351FC0FEFDC4C9C" box="[1280,1419,896,921]" italics="true" pageId="1" pageNumber="356">Kodaianella</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="10DF29A6FFE24F33D696FC2BEF2F48BD" pageId="1" pageNumber="356" type="description">
<paragraph id="587A7A2DFFE24F33D696FC2BE9314929" blockId="1.[151,1437,746,2012]" pageId="1" pageNumber="356">
<emphasis id="6AB1A63FFFE24F33D696FC2BEB0E4CB8" bold="true" box="[199,345,932,957]" pageId="1" pageNumber="356">Description.</emphasis>
Head with compound eyes almost as wide as thorax (
<figureCitation id="C0FE66A8FFE24F33D599FC2BEE614CB8" box="[968,1078,932,957]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1369,1392]" captionTargetBox="[253,1333,384,1336]" captionTargetId="figure@2.[238,1348,375,1347]" captionTargetPageId="2" captionText="FIGURES 1 6. Kodaianellissus intorqueus sp. nov. 1. Adult, dorsal view; 2. Adult, lateral view; 3. Head and thorax; 4. Frons and clypeus; 5. Forewing; 6. Hindwing." httpUri="https://zenodo.org/record/852744/files/figure.png" pageId="1" pageNumber="356">Figs. 1, 3</figureCitation>
). Vertex rectangular, obviously broader than long, median carina present; anterior margin slightly convex, posterior margin strongly angular concave medially (
<figureCitation id="C0FE66A8FFE24F33D720FC63EBB54B00" box="[369,482,1004,1029]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1369,1392]" captionTargetBox="[253,1333,384,1336]" captionTargetId="figure@2.[238,1348,375,1347]" captionTargetPageId="2" captionText="FIGURES 1 6. Kodaianellissus intorqueus sp. nov. 1. Adult, dorsal view; 2. Adult, lateral view; 3. Head and thorax; 4. Frons and clypeus; 5. Forewing; 6. Hindwing." httpUri="https://zenodo.org/record/852744/files/figure.png" pageId="1" pageNumber="356">Figs. 1, 3</figureCitation>
). Frons roundly widened below antenna, apical margin almost straight, apical and lateral margins carinate and elevated, median carina obviously elevated from apex extending to near base, but not reaching frontoclypeal sulcus (
<figureCitation id="C0FE66A8FFE24F33D7AFFBBBE81F4B48" box="[510,584,1076,1101]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1369,1392]" captionTargetBox="[253,1333,384,1336]" captionTargetId="figure@2.[238,1348,375,1347]" captionTargetPageId="2" captionText="FIGURES 1 6. Kodaianellissus intorqueus sp. nov. 1. Adult, dorsal view; 2. Adult, lateral view; 3. Head and thorax; 4. Frons and clypeus; 5. Forewing; 6. Hindwing." httpUri="https://zenodo.org/record/852744/files/figure.png" pageId="1" pageNumber="356">Fig. 4</figureCitation>
); frons with apical and lateral area distributed with numerous tubercles, tubercles change from small to large from apical to lateral part (
<figureCitation id="C0FE66A8FFE24F33D532FBD7E9FE4B75" box="[867,937,1112,1137]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1369,1392]" captionTargetBox="[253,1333,384,1336]" captionTargetId="figure@2.[238,1348,375,1347]" captionTargetPageId="2" captionText="FIGURES 1 6. Kodaianellissus intorqueus sp. nov. 1. Adult, dorsal view; 2. Adult, lateral view; 3. Head and thorax; 4. Frons and clypeus; 5. Forewing; 6. Hindwing." httpUri="https://zenodo.org/record/852744/files/figure.png" pageId="1" pageNumber="356">Fig. 4</figureCitation>
). Frontoclypeal suture strongly convex (
<figureCitation id="C0FE66A8FFE24F33D33EFBD7EAFF4B90" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1369,1392]" captionTargetBox="[253,1333,384,1336]" captionTargetId="figure@2.[238,1348,375,1347]" captionTargetPageId="2" captionText="FIGURES 1 6. Kodaianellissus intorqueus sp. nov. 1. Adult, dorsal view; 2. Adult, lateral view; 3. Head and thorax; 4. Frons and clypeus; 5. Forewing; 6. Hindwing." httpUri="https://zenodo.org/record/852744/files/figure.png" pageId="1" pageNumber="356">Fig. 4</figureCitation>
). Clypeus with several transverse thin stripes on lateral sides (
<figureCitation id="C0FE66A8FFE24F33D50CFBF3E9F54B90" box="[861,930,1148,1173]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1369,1392]" captionTargetBox="[253,1333,384,1336]" captionTargetId="figure@2.[238,1348,375,1347]" captionTargetPageId="2" captionText="FIGURES 1 6. Kodaianellissus intorqueus sp. nov. 1. Adult, dorsal view; 2. Adult, lateral view; 3. Head and thorax; 4. Frons and clypeus; 5. Forewing; 6. Hindwing." httpUri="https://zenodo.org/record/852744/files/figure.png" pageId="1" pageNumber="356">Fig. 4</figureCitation>
). Rostrum short, reaching midcoxae. Gena in lateral view flattened and oblique, with small protuberance near base (
<figureCitation id="C0FE66A8FFE24F33D5FDFB2FE9A64BBD" box="[940,1009,1184,1209]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1369,1392]" captionTargetBox="[253,1333,384,1336]" captionTargetId="figure@2.[238,1348,375,1347]" captionTargetPageId="2" captionText="FIGURES 1 6. Kodaianellissus intorqueus sp. nov. 1. Adult, dorsal view; 2. Adult, lateral view; 3. Head and thorax; 4. Frons and clypeus; 5. Forewing; 6. Hindwing." httpUri="https://zenodo.org/record/852744/files/figure.png" pageId="1" pageNumber="356">Fig. 2</figureCitation>
). Pronotum triangular, with two small incisions on disc, median carina present (
<figureCitation id="C0FE66A8FFE24F33D420FB4BE8B64BD8" box="[625,737,1220,1245]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1369,1392]" captionTargetBox="[253,1333,384,1336]" captionTargetId="figure@2.[238,1348,375,1347]" captionTargetPageId="2" captionText="FIGURES 1 6. Kodaianellissus intorqueus sp. nov. 1. Adult, dorsal view; 2. Adult, lateral view; 3. Head and thorax; 4. Frons and clypeus; 5. Forewing; 6. Hindwing." httpUri="https://zenodo.org/record/852744/files/figure.png" pageId="1" pageNumber="356">Figs. 1, 3</figureCitation>
). Mesonotum smooth, without carina (
<figureCitation id="C0FE66A8FFE24F33D2CEFB4BEF594BD8" box="[1183,1294,1220,1245]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1369,1392]" captionTargetBox="[253,1333,384,1336]" captionTargetId="figure@2.[238,1348,375,1347]" captionTargetPageId="2" captionText="FIGURES 1 6. Kodaianellissus intorqueus sp. nov. 1. Adult, dorsal view; 2. Adult, lateral view; 3. Head and thorax; 4. Frons and clypeus; 5. Forewing; 6. Hindwing." httpUri="https://zenodo.org/record/852744/files/figure.png" pageId="1" pageNumber="356">Figs. 1, 3</figureCitation>
). Forewings elongate, distinctly longer than wide, with very narrow hypocostal plate, longitudinal veins obvious and elevated, forewing slightly broadened near the basal 1/3 (
<figureCitation id="C0FE66A8FFE24F33D4FEFA83E9604A20" box="[687,823,1292,1317]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1369,1392]" captionTargetBox="[253,1333,384,1336]" captionTargetId="figure@2.[238,1348,375,1347]" captionTargetPageId="2" captionText="FIGURES 1 6. Kodaianellissus intorqueus sp. nov. 1. Adult, dorsal view; 2. Adult, lateral view; 3. Head and thorax; 4. Frons and clypeus; 5. Forewing; 6. Hindwing." httpUri="https://zenodo.org/record/852744/files/figure.png" pageId="1" pageNumber="356">Figs. 1, 2, 5</figureCitation>
). Vein ScP+R first separating near base, ScP+RA vein extremely long, surpassing the apical 1/3 of costal margin, the terminal of RP vein almost reaching the outer margin of forewing; MP vein first fork near the basal 1/4, MP1+2 straight, fork again at apical 1/9, MP3+4 sinuate, without fork; CuA vein first separate at apical 1/3; MP first fork before the first separation of CuA (
<figureCitation id="C0FE66A8FFE24F33D2A4FAF4EF6E4A91" box="[1269,1337,1403,1428]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1369,1392]" captionTargetBox="[253,1333,384,1336]" captionTargetId="figure@2.[238,1348,375,1347]" captionTargetPageId="2" captionText="FIGURES 1 6. Kodaianellissus intorqueus sp. nov. 1. Adult, dorsal view; 2. Adult, lateral view; 3. Head and thorax; 4. Frons and clypeus; 5. Forewing; 6. Hindwing." httpUri="https://zenodo.org/record/852744/files/figure.png" pageId="1" pageNumber="356">Fig. 5</figureCitation>
). Clavus obvious, veins Pcu and A1 fused at basal 1/2 of clavus (
<figureCitation id="C0FE66A8FFE24F33D550FA10E9124ABD" box="[769,837,1439,1464]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1369,1392]" captionTargetBox="[253,1333,384,1336]" captionTargetId="figure@2.[238,1348,375,1347]" captionTargetPageId="2" captionText="FIGURES 1 6. Kodaianellissus intorqueus sp. nov. 1. Adult, dorsal view; 2. Adult, lateral view; 3. Head and thorax; 4. Frons and clypeus; 5. Forewing; 6. Hindwing." httpUri="https://zenodo.org/record/852744/files/figure.png" pageId="1" pageNumber="356">Fig. 5</figureCitation>
). Forewings with numerous transverse veins (
<figureCitation id="C0FE66A8FFE24F33D319FA10EA934ADB" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1369,1392]" captionTargetBox="[253,1333,384,1336]" captionTargetId="figure@2.[238,1348,375,1347]" captionTargetPageId="2" captionText="FIGURES 1 6. Kodaianellissus intorqueus sp. nov. 1. Adult, dorsal view; 2. Adult, lateral view; 3. Head and thorax; 4. Frons and clypeus; 5. Forewing; 6. Hindwing." httpUri="https://zenodo.org/record/852744/files/figure.png" pageId="1" pageNumber="356">Figs. 1, 2, 5</figureCitation>
). Hindwing well developed, 3-lobed; Pcu-A1 lobe distinctly thinner than ScP-R-MP-Cu lobe; A2 lobe narrow with anterior and posterior margins subparallel and distinctly surpassing half length of Pcu-A1 lobe; A2 not branched (
<figureCitation id="C0FE66A8FFE24F33D75CF99CEB064929" box="[269,337,1555,1580]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1369,1392]" captionTargetBox="[253,1333,384,1336]" captionTargetId="figure@2.[238,1348,375,1347]" captionTargetPageId="2" captionText="FIGURES 1 6. Kodaianellissus intorqueus sp. nov. 1. Adult, dorsal view; 2. Adult, lateral view; 3. Head and thorax; 4. Frons and clypeus; 5. Forewing; 6. Hindwing." httpUri="https://zenodo.org/record/852744/files/figure.png" pageId="1" pageNumber="356">Fig. 6</figureCitation>
). Hind tibia with 2 lateral spines on apical half.
</paragraph>
<paragraph id="587A7A2DFFE24F33D696F9B8E91A4801" blockId="1.[151,1437,746,2012]" pageId="1" pageNumber="356">
<emphasis id="6AB1A63FFFE24F33D696F9B8EBD94955" bold="true" box="[199,398,1591,1616]" pageId="1" pageNumber="356">Male terminalia.</emphasis>
Anal tube in lateral view long and narrow (
<figureCitation id="C0FE66A8FFE24F33D527F9B8E9ED4955" box="[886,954,1591,1616]" captionStart="FIGURES 7 11" captionStartId="3.[151,264,1360,1383]" captionTargetBox="[230,1357,457,1322]" captionTargetId="figure@3.[218,1369,445,1339]" captionTargetPageId="3" captionText="FIGURES 7 11. Kodaianellissus intorqueus sp. nov. 7. Male terminalia, lateral view; 8. Male anal tube, dorsal view; 9. Gonostylus, lateral view; 10. Phallic complex, lateral view; 11. Phallic complex, ventral view." httpUri="https://zenodo.org/record/852746/files/figure.png" pageId="1" pageNumber="356">Fig. 7</figureCitation>
). Gonostyli triangular in profile, becoming broader from base to apex, caudo-ventral angle rounded, dorso-lateral margin with process; capitulum very long and narrow, with auricular process (
<figureCitation id="C0FE66A8FFE24F33D461F9F0E8C9499D" box="[560,670,1663,1688]" captionStart="FIGURES 7 11" captionStartId="3.[151,264,1360,1383]" captionTargetBox="[230,1357,457,1322]" captionTargetId="figure@3.[218,1369,445,1339]" captionTargetPageId="3" captionText="FIGURES 7 11. Kodaianellissus intorqueus sp. nov. 7. Male terminalia, lateral view; 8. Male anal tube, dorsal view; 9. Gonostylus, lateral view; 10. Phallic complex, lateral view; 11. Phallic complex, ventral view." httpUri="https://zenodo.org/record/852746/files/figure.png" pageId="1" pageNumber="356">Figs. 7, 9</figureCitation>
). Pygofer rectangular in lateral view, apparently longer than width, posterior margin not obviously caudally produced (
<figureCitation id="C0FE66A8FFE24F33D4B3F92CE97F49B9" box="[738,808,1699,1724]" captionStart="FIGURES 7 11" captionStartId="3.[151,264,1360,1383]" captionTargetBox="[230,1357,457,1322]" captionTargetId="figure@3.[218,1369,445,1339]" captionTargetPageId="3" captionText="FIGURES 7 11. Kodaianellissus intorqueus sp. nov. 7. Male terminalia, lateral view; 8. Male anal tube, dorsal view; 9. Gonostylus, lateral view; 10. Phallic complex, lateral view; 11. Phallic complex, ventral view." httpUri="https://zenodo.org/record/852746/files/figure.png" pageId="1" pageNumber="356">Fig. 7</figureCitation>
). Periandrium symmetrical, basally tubular, apical part divided into dorso-lateral lobe and ventral lobe (
<figureCitation id="C0FE66A8FFE24F33D49AF948E97549E5" box="[715,802,1735,1760]" captionStart="FIGURES 7 11" captionStartId="3.[151,264,1360,1383]" captionTargetBox="[230,1357,457,1322]" captionTargetId="figure@3.[218,1369,445,1339]" captionTargetPageId="3" captionText="FIGURES 7 11. Kodaianellissus intorqueus sp. nov. 7. Male terminalia, lateral view; 8. Male anal tube, dorsal view; 9. Gonostylus, lateral view; 10. Phallic complex, lateral view; 11. Phallic complex, ventral view." httpUri="https://zenodo.org/record/852746/files/figure.png" pageId="1" pageNumber="356">Fig. 10</figureCitation>
). Dorso-lateral lobe longer than ventral lobe (
<figureCitation id="C0FE66A8FFE24F33D368F948EFD949E5" box="[1337,1422,1735,1760]" captionStart="FIGURES 7 11" captionStartId="3.[151,264,1360,1383]" captionTargetBox="[230,1357,457,1322]" captionTargetId="figure@3.[218,1369,445,1339]" captionTargetPageId="3" captionText="FIGURES 7 11. Kodaianellissus intorqueus sp. nov. 7. Male terminalia, lateral view; 8. Male anal tube, dorsal view; 9. Gonostylus, lateral view; 10. Phallic complex, lateral view; 11. Phallic complex, ventral view." httpUri="https://zenodo.org/record/852746/files/figure.png" pageId="1" pageNumber="356">Fig. 11</figureCitation>
). Aedeagus with pair of ventral processes directed dorso-basad.
</paragraph>
<paragraph id="587A7A2DFFE24F33D696F880EF2F48BD" blockId="1.[151,1437,746,2012]" pageId="1" pageNumber="356">
<emphasis id="6AB1A63FFFE24F33D696F880EBFD482D" bold="true" box="[199,426,1807,1832]" pageId="1" pageNumber="356">Female terminalia.</emphasis>
Anal tube relatively long in lateral view, reaching level of posterior margin of gonoplac. Anterior connective lamina of gonapophysis VIII with teeth at apex and outer lateral margin, inner lateral margin without teeth (
<figureCitation id="C0FE66A8FFE24F33D76CF8D8EBC44875" box="[317,403,1879,1904]" captionStart="FIGURES 12 18" captionStartId="4.[151,264,1740,1763]" captionTargetBox="[246,1339,481,1698]" captionTargetId="figure@4.[226,1361,481,1719]" captionTargetPageId="4" captionText="FIGURES 12 18. Kodaianellissus intorqueus sp. nov. 12. Female anal tube, dorsal view; 13. Gonoplac, dorsal view; 14. Gonapophysis IX and gonaspiculum bridge, ventral view; 15. Gonoplac, lateral view; 16. Gonapophysis IX and gonaspiculum bridge, lateral view; 17. Sternite VII; 18. Gonocoxa VIII and gonapophysis VIII, lateral view." httpUri="https://zenodo.org/record/852748/files/figure.png" pageId="1" pageNumber="356">Fig. 18</figureCitation>
). Gonocoxa VIII subquadrangular, connected to gonapophysis VIII with rectangular angle (
<figureCitation id="C0FE66A8FFE24F33D6CEF8F4EAA24891" box="[159,245,1915,1940]" captionStart="FIGURES 12 18" captionStartId="4.[151,264,1740,1763]" captionTargetBox="[246,1339,481,1698]" captionTargetId="figure@4.[226,1361,481,1719]" captionTargetPageId="4" captionText="FIGURES 12 18. Kodaianellissus intorqueus sp. nov. 12. Female anal tube, dorsal view; 13. Gonoplac, dorsal view; 14. Gonapophysis IX and gonaspiculum bridge, ventral view; 15. Gonoplac, lateral view; 16. Gonapophysis IX and gonaspiculum bridge, lateral view; 17. Sternite VII; 18. Gonocoxa VIII and gonapophysis VIII, lateral view." httpUri="https://zenodo.org/record/852748/files/figure.png" pageId="1" pageNumber="356">Fig. 18</figureCitation>
). Gonapophysis IX in lateral view boat-shaped (
<figureCitation id="C0FE66A8FFE24F33D577F8F4E92C4891" box="[806,891,1915,1940]" captionStart="FIGURES 12 18" captionStartId="4.[151,264,1740,1763]" captionTargetBox="[246,1339,481,1698]" captionTargetId="figure@4.[226,1361,481,1719]" captionTargetPageId="4" captionText="FIGURES 12 18. Kodaianellissus intorqueus sp. nov. 12. Female anal tube, dorsal view; 13. Gonoplac, dorsal view; 14. Gonapophysis IX and gonaspiculum bridge, ventral view; 15. Gonoplac, lateral view; 16. Gonapophysis IX and gonaspiculum bridge, lateral view; 17. Sternite VII; 18. Gonocoxa VIII and gonapophysis VIII, lateral view." httpUri="https://zenodo.org/record/852748/files/figure.png" pageId="1" pageNumber="356">Fig. 16</figureCitation>
). Gonoplac subrectangular in lateral view (
<figureCitation id="C0FE66A8FFE24F33D33EF8F4EAE248BD" captionStart="FIGURES 12 18" captionStartId="4.[151,264,1740,1763]" captionTargetBox="[246,1339,481,1698]" captionTargetId="figure@4.[226,1361,481,1719]" captionTargetPageId="4" captionText="FIGURES 12 18. Kodaianellissus intorqueus sp. nov. 12. Female anal tube, dorsal view; 13. Gonoplac, dorsal view; 14. Gonapophysis IX and gonaspiculum bridge, ventral view; 15. Gonoplac, lateral view; 16. Gonapophysis IX and gonaspiculum bridge, lateral view; 17. Sternite VII; 18. Gonocoxa VIII and gonapophysis VIII, lateral view." httpUri="https://zenodo.org/record/852748/files/figure.png" pageId="1" pageNumber="356">Fig. 15</figureCitation>
), fused at middle near base, widest at base (
<figureCitation id="C0FE66A8FFE24F33D4F0F810E8A348BD" box="[673,756,1951,1976]" captionStart="FIGURES 12 18" captionStartId="4.[151,264,1740,1763]" captionTargetBox="[246,1339,481,1698]" captionTargetId="figure@4.[226,1361,481,1719]" captionTargetPageId="4" captionText="FIGURES 12 18. Kodaianellissus intorqueus sp. nov. 12. Female anal tube, dorsal view; 13. Gonoplac, dorsal view; 14. Gonapophysis IX and gonaspiculum bridge, ventral view; 15. Gonoplac, lateral view; 16. Gonapophysis IX and gonaspiculum bridge, lateral view; 17. Sternite VII; 18. Gonocoxa VIII and gonapophysis VIII, lateral view." httpUri="https://zenodo.org/record/852748/files/figure.png" pageId="1" pageNumber="356">Fig. 13</figureCitation>
). Hind margin of sternite VII concave medially (
<figureCitation id="C0FE66A8FFE24F33D346F810EF3C48BD" box="[1303,1387,1951,1976]" captionStart="FIGURES 12 18" captionStartId="4.[151,264,1740,1763]" captionTargetBox="[246,1339,481,1698]" captionTargetId="figure@4.[226,1361,481,1719]" captionTargetPageId="4" captionText="FIGURES 12 18. Kodaianellissus intorqueus sp. nov. 12. Female anal tube, dorsal view; 13. Gonoplac, dorsal view; 14. Gonapophysis IX and gonaspiculum bridge, ventral view; 15. Gonoplac, lateral view; 16. Gonapophysis IX and gonaspiculum bridge, lateral view; 17. Sternite VII; 18. Gonocoxa VIII and gonapophysis VIII, lateral view." httpUri="https://zenodo.org/record/852748/files/figure.png" pageId="1" pageNumber="356">Fig. 17</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="10DF29A6FFE24F30D696F84CE8A64E45" lastPageId="2" lastPageNumber="357" pageId="1" pageNumber="356" type="etymology">
<paragraph id="587A7A2DFFE24F30D696F84CEB474FD0" blockId="1.[151,1437,746,2012]" lastBlockId="2.[151,1436,151,320]" lastPageId="2" lastPageNumber="357" pageId="1" pageNumber="356">
<emphasis id="6AB1A63FFFE24F33D696F84CEB1948D9" bold="true" box="[199,334,1987,2012]" pageId="1" pageNumber="356">Etymology.</emphasis>
The name of this new genus is an arbitrary combination between two generic names of issids:
<taxonomicName id="9FC501AEFFE14F30D6C6FF18EB754FB5" authorityName="Fennah" authorityYear="1956" box="[151,290,151,176]" class="Insecta" family="Issidae" genus="Kodaianella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="357" phylum="Arthropoda" rank="genus">
<emphasis id="6AB1A63FFFE14F30D6C6FF18EB754FB5" box="[151,290,151,176]" italics="true" pageId="2" pageNumber="357">Kodaianella</emphasis>
</taxonomicName>
and
<taxonomicName id="9FC501AEFFE14F30D70CFF16EBC24FB5" authorityName="Fabricius" authorityYear="1803" box="[349,405,153,176]" class="Insecta" family="Issidae" genus="Issus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="357" phylum="Arthropoda" rank="genus">
<emphasis id="6AB1A63FFFE14F30D70CFF16EBC24FB5" box="[349,405,153,176]" italics="true" pageId="2" pageNumber="357">Issus</emphasis>
</taxonomicName>
, indicating this genus has a close relationship with the genus
<taxonomicName id="9FC501AEFFE14F30D232FF18EEBA4FB5" authorityName="Fennah" authorityYear="1956" box="[1123,1261,151,176]" class="Insecta" family="Issidae" genus="Kodaianella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="357" phylum="Arthropoda" rank="genus">
<emphasis id="6AB1A63FFFE14F30D232FF18EEBA4FB5" box="[1123,1261,151,176]" italics="true" pageId="2" pageNumber="357">Kodaianella</emphasis>
</taxonomicName>
. The gender is masculine.
</paragraph>
<paragraph id="587A7A2DFFE14F30D696FF50E8A64E45" blockId="2.[151,1436,151,320]" pageId="2" pageNumber="357">
<emphasis id="6AB1A63FFFE14F30D696FF50EB514FFD" bold="true" box="[199,262,223,248]" pageId="2" pageNumber="357">Note.</emphasis>
This genus refers to the taxon “
<emphasis id="6AB1A63FFFE14F30D43EFF50E88E4FFD" bold="true" box="[623,729,223,248]" pageId="2" pageNumber="357">gen. nov.</emphasis>
<emphasis id="6AB1A63FFFE14F30D4B1FF50E9FC4FFD" box="[736,939,223,248]" italics="true" pageId="2" pageNumber="357">
apud
<taxonomicName id="9FC501AEFFE14F30D571FF50E9FC4FFD" authorityName="Fennah" authorityYear="1956" box="[800,939,223,248]" class="Insecta" family="Issidae" genus="Kodaianella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="357" phylum="Arthropoda" rank="genus">Kodaianella</taxonomicName>
</emphasis>
” on the molecular phylogeny trees of
<bibRefCitation id="3C5407DCFFE14F30D30AFF6FEB7C4E18" pageId="2" pageNumber="357" refString="Wang, M. L., Zhang, Y. L. &amp; Bourgoin, T. (2016) Planthopper family Issidae (Insecta: Hemiptera: Fulgoromorpha): linking molecular phylogeny with classification. Molecular Phylogenetics and Evolution, 105, 224 - 234. https: // doi. org / 10.1016 / j. ympev. 2016.08.012" type="journal article">
Wang
<emphasis id="6AB1A63FFFE14F30D6C6FE8AEA844E18" box="[151,211,260,285]" italics="true" pageId="2" pageNumber="357">et al.</emphasis>
(2016)
</bibRefCitation>
. The position of this genus within Kodaianellini is supported by both the molecular phylogenetic results and the morphological characters listed above.
</paragraph>
</subSubSection>
</treatment>
</document>