346 lines
48 KiB
XML
346 lines
48 KiB
XML
<document id="E5EB47C49E52CE6B0CC7D1160F592E85" ID-DOI="10.11646/zootaxa.3647.3.4" ID-GBIF-Dataset="6160bdbc-2d0a-4407-8d89-fe24a10cbdfc" ID-ISSN="1175-5326" ID-Zenodo-Dep="216407" ID-ZooBank="E4DD8898-554F-42A9-B6C8-568144378732" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460606498401" checkinUser="plazi" docAuthor="Davie, Peter J. F." docDate="2013" docId="CC61A21E112AFFD1FF31C8FEFB7245E6" docLanguage="en" docName="zt03647p478.pdf" docOrigin="Zootaxa 3647 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chasmocarcinidae Serène 1964" docType="treatment" docVersion="8" lastPageNumber="473" masterDocId="3058DA66112BFFD2FFA6C821FFD14242" masterDocTitle="A new species of Megaesthesius Rathbun, 1909 (Crustacea: Brachyura: Chasmocarcinidae) from deep water off Western Australia" masterLastPageNumber="478" masterPageNumber="470" pageNumber="471" updateTime="1698327929202" updateUser="plazi">
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<mods:title id="1EB4F7BED7E0A2E62521E9511E7A72FE">A new species of Megaesthesius Rathbun, 1909 (Crustacea: Brachyura: Chasmocarcinidae) from deep water off Western Australia</mods:title>
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<mods:namePart id="3FD94E369A89818E774671E3E724FC5E">Davie, Peter J. F.</mods:namePart>
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<mods:title id="3B8522A1C9E2B813DC78016B727331B5">Zootaxa</mods:title>
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<mods:date id="85145153ACACD5B8D6EE4A18F1C1357E">2013</mods:date>
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<mods:number id="618EEE2B2DF9E4D5DB4506017E3516D2">3647</mods:number>
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<treatment id="CC61A21E112AFFD1FF31C8FEFB7245E6" ID-DOI="http://doi.org/10.5281/zenodo.5659896" ID-GBIF-Taxon="119567914" ID-Zenodo-Dep="5659896" LSID="urn:lsid:plazi:treatment:CC61A21E112AFFD1FF31C8FEFB7245E6" httpUri="http://treatment.plazi.org/id/CC61A21E112AFFD1FF31C8FEFB7245E6" lastPageId="3" lastPageNumber="473" pageId="1" pageNumber="471">
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<subSubSection id="0CD24083112AFFD3FF31C8FEFDB1437D" pageId="1" pageNumber="471" type="nomenclature">
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<paragraph id="44771308112AFFD3FF31C8FEFDAE42BB" blockId="1.[151,639,223,249]" box="[151,639,223,249]" pageId="1" pageNumber="471">
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<heading id="1F3FA464112AFFD3FF31C8FEFDAE42BB" bold="true" box="[151,639,223,249]" fontSize="11" level="1" pageId="1" pageNumber="471" reason="1">
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<taxonomicName id="83C8688B112AFFD3FF31C8FEFDAE42BB" ID-CoL="7NHWW" box="[151,639,223,249]" class="Malacostraca" family="Chasmocarcinidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="470" phylum="Arthropoda" rank="family">
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<emphasis id="76BCCF1A112AFFD3FF31C8FEFDAE42BB" bold="true" box="[151,639,223,249]" pageId="1" pageNumber="471">CHASMOCARCINIDAE Serène, 1964</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="44771308112AFFD3FF31C904FDB1437D" blockId="1.[151,608,292,319]" box="[151,608,292,319]" pageId="1" pageNumber="471">
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<emphasis id="76BCCF1A112AFFD3FF31C904FE27437D" bold="true" box="[151,502,293,319]" pageId="1" pageNumber="471">
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<taxonomicName id="83C8688B112AFFD3FF31C904FE68437D" box="[151,441,293,319]" class="Malacostraca" family="Chasmocarcinidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="471" phylum="Arthropoda" rank="subFamily" subFamily="Megaesthesiinae">MEGAESTHESIINAE</taxonomicName>
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Štev
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</emphasis>
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č
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<emphasis id="76BCCF1A112AFFD3FDA5C904FDDA437D" bold="true" box="[515,523,293,319]" pageId="1" pageNumber="471">i</emphasis>
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ć,
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<emphasis id="76BCCF1A112AFFD3FD80C904FDB1437D" bold="true" box="[550,608,293,319]" pageId="1" pageNumber="471">2005</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="0CD24083112AFFD3FF31C94BFBD84044" pageId="1" pageNumber="471" type="description">
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<paragraph id="44771308112AFFD3FF31C94BFE2643C1" blockId="1.[151,503,362,387]" box="[151,503,362,387]" pageId="1" pageNumber="471">
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<taxonomicName id="83C8688B112AFFD3FF31C94BFE2643C1" authority="Rathbun, 1909" authorityName="Rathbun" authorityYear="1909" box="[151,503,362,387]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="471" phylum="Arthropoda" rank="genus">
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<emphasis id="76BCCF1A112AFFD3FF31C94BFE2643C1" bold="true" box="[151,503,362,387]" pageId="1" pageNumber="471">
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<emphasis id="76BCCF1A112AFFD3FF31C94BFE9043C1" bold="true" box="[151,321,362,387]" italics="true" pageId="1" pageNumber="471">Megaesthesius</emphasis>
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Rathbun, 1909
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</emphasis>
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</taxonomicName>
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</paragraph>
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<paragraph id="44771308112AFFD3FF31C993FC0A43A5" blockId="1.[151,1436,434,518]" pageId="1" pageNumber="471">
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<taxonomicName id="83C8688B112AFFD3FF31C993FE2E438A" authority="Rathbun, 1909: 112" authorityName="Rathbun" authorityPageNumber="112" authorityYear="1909" box="[151,511,434,456]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="471" phylum="Arthropoda" rank="genus">
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<emphasis id="76BCCF1A112AFFD3FF31C993FEFD438A" box="[151,300,434,456]" italics="true" pageId="1" pageNumber="471">Megaesthesius</emphasis>
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Rathbun, 1909: 112
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</taxonomicName>
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; 1910: 344.—Tesch, 1918: 202 (key).—Sakai, 1939: 577; 1976: 552.—Serène, 1964: 175; 1968: 92 (list).—Takeda & Miyake, 1969: 460; 1972: 86.—Takeda, 1973: 54.
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</paragraph>
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<paragraph id="44771308112AFFD3FF31C9D0FBD84044" blockId="1.[151,1436,434,518]" box="[151,1033,496,518]" pageId="1" pageNumber="471">
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<typeStatus id="9B73ADAA112AFFD3FF31C9D0FF194044" box="[151,200,497,518]" pageId="1" pageNumber="471">Type</typeStatus>
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species:
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<taxonomicName id="83C8688B112AFFD3FE80C9D1FD634047" authority="Rathbun, 1909" authorityName="Rathbun" authorityYear="1909" box="[294,690,496,518]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="471" phylum="Arthropoda" rank="species" species="sagedae">
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<emphasis id="76BCCF1A112AFFD3FE80C9D1FDC54044" box="[294,532,496,518]" italics="true" pageId="1" pageNumber="471">Megaesthesius sagedae</emphasis>
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Rathbun, 1909
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</taxonomicName>
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, by monotypy; gender masculine.
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</paragraph>
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</subSubSection>
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<subSubSection id="0CD24083112AFFD3FF31CA13FE8B41B8" pageId="1" pageNumber="471" type="diagnosis">
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<paragraph id="44771308112AFFD3FF31CA13FEB84104" blockId="1.[151,1436,562,875]" pageId="1" pageNumber="471">
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||
<emphasis id="76BCCF1A112AFFD3FF31CA13FEDB4009" bold="true" box="[151,266,562,587]" pageId="1" pageNumber="471">Diagnosis</emphasis>
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. Carapace quadrangular, rounded anteriorly; lateral margins almost parallel; anterolateral margins forming regular curve with anterior margin. Front narrow, bilobed. Orbits long, transverse, occupying whole anterior border. Eyestalks very long, embedded in orbits; swollen proximally, tapering distally; cornea elongated, distally bent posteriorly, slightly protruding laterally, pigment reduced to speck. Antennules thick, cannot be folded because swollen basal segment fills fossa; palp with long thick plumose setae. Buccal cavern anteriorly narrowed, outer borders arcuate. Third maxillipeds with merus subtriangular; palp short, articulated apically. Chelipeds subequal, shorter than ambulatory legs. Ambulatory legs slender, with spinules on borders. Male abdomen with somites 3–5 fused.
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</paragraph>
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<paragraph id="44771308112AFFD3FF61CB73FA874129" blockId="1.[151,1436,562,875]" box="[199,1366,850,875]" pageId="1" pageNumber="471">
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Included species:
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<taxonomicName id="83C8688B112AFFD3FE29CB72FC974128" authority="Rathbun, 1909" authorityName="Rathbun" authorityYear="1909" box="[399,838,850,875]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="471" phylum="Arthropoda" rank="species" species="sagedae">
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<emphasis id="76BCCF1A112AFFD3FE29CB72FD464129" box="[399,663,851,875]" italics="true" pageId="1" pageNumber="471">Megaesthesius sagedae</emphasis>
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Rathbun, 1909
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</taxonomicName>
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;
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<taxonomicName id="83C8688B112AFFD3FCF4CB75FB8D4128" authority="Sakai, 1939" authorityName="Sakai" authorityYear="1939" box="[850,1116,850,875]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="471" phylum="Arthropoda" rank="species" species="yokoyai">
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<emphasis id="76BCCF1A112AFFD3FCF4CB75FC1F4129" box="[850,974,851,875]" italics="true" pageId="1" pageNumber="471">M. yokoyai</emphasis>
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Sakai, 1939
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</taxonomicName>
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;
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<taxonomicName id="83C8688B112AFFD3FBCECB75FB244129" box="[1128,1269,851,875]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="471" phylum="Arthropoda" rank="species" species="westralia" status="sp. nov.">
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<emphasis id="76BCCF1A112AFFD3FBCECB75FB244129" box="[1128,1269,851,875]" italics="true" pageId="1" pageNumber="471">M. westralia</emphasis>
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</taxonomicName>
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<emphasis id="76BCCF1A112AFFD3FB5ACB72FA874129" bold="true" box="[1276,1366,851,875]" pageId="1" pageNumber="471">
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||
<taxonomicNameLabel id="6D8F7261112AFFD3FB5ACB72FA874129" box="[1276,1366,851,875]" pageId="1" pageNumber="471" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</paragraph>
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||
<paragraph id="44771308112AFFD3FF31CB9FFE8B41B8" blockId="1.[151,528,958,1018]" pageId="1" pageNumber="471">
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||
<emphasis id="76BCCF1A112AFFD3FF31CB9FFDC14195" bold="true" box="[151,528,958,983]" pageId="1" pageNumber="471">
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||
<taxonomicName id="83C8688B112AFFD3FF31CB9FFE7E4195" box="[151,431,958,983]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="471" phylum="Arthropoda" rank="species" species="westralia" status="sp. nov.">
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<emphasis id="76BCCF1A112AFFD3FF31CB9FFE7E4195" bold="true" box="[151,431,958,983]" italics="true" pageId="1" pageNumber="471">Megaesthesius westralia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6D8F7261112AFFD3FE10CB9EFDC14195" box="[438,528,959,983]" pageId="1" pageNumber="471" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
(
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<figureCitation id="DCF30F8D112AFFD3FF39CBC3FF3441B8" box="[159,229,994,1018]" captionStart="FIGURE 1" captionStartId="2.[151,250,1436,1459]" captionTargetBox="[158,1436,310,1375]" captionTargetId="figure@2.[158,1444,301,1407]" captionTargetPageId="2" captionText="FIGURE 1. Megaesthesius westralia sp. nov., holotype male. A, carapace in dorsal view; B, fronto-ventral view showing third maxilliped and buccal frame; C, latero-ventral view; D, frontal view." httpUri="https://zenodo.org/record/216408/files/figure.png" pageId="1" pageNumber="471">Figs 1</figureCitation>
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,
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<figureCitation id="DCF30F8D112AFFD3FF55CBC3FED241B8" box="[243,259,994,1018]" captionStart="FIGURE 2" captionStartId="3.[151,250,1119,1142]" captionTargetBox="[151,1409,412,1079]" captionTargetId="figure@3.[151,1436,409,1098]" captionTargetPageId="3" captionText="FIGURE 2. Megaesthesius westralia sp. nov., holotype male. A, anterodorsal view of frontal and orbital regions; B, ventral view of infraorbital margin showing eyestalk; C, third maxilliped; D, fourth pereiopod; E, fifth pereiopod." httpUri="https://zenodo.org/record/216409/files/figure.png" pageId="1" pageNumber="471">2</figureCitation>
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,
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<figureCitation id="DCF30F8D112AFFD3FEA9CBC3FECA41B8" box="[271,283,994,1018]" captionStart="FIGURE 3" captionStartId="4.[151,250,1854,1877]" captionTargetBox="[168,1434,199,1819]" captionTargetId="figure@4.[168,1453,193,1821]" captionTargetPageId="4" captionText="FIGURE 3. Male gonopods and abdomens for the three known species of Megaesthesius. A – E, M. westralia sp. nov.: A, left G 1 in sternal view; B, left G 1 in abdominal view; C, left G 1, detail of tip; D, left G 2; E, male abdomen with lateral view of sternites 7 – 8 and penis. F – I, M. yokoyai Sakai, 1939: F, left G 1 in abdominal view; G, H. distal part of G 1 in abdominal and sternal view; I, male abdomen (somites 1 – 2 not shown) (After Takeda & Miyake, 1972). J – N, M. sagedae Rathbun, 1909: J, left G 1, abdominal face; K, detail of tip; L, left G 1 sternal face; M, left G 2 abdominal face; N, male abdomen (after Crosnier 1975)." httpUri="https://zenodo.org/record/216410/files/figure.png" pageId="1" pageNumber="471">3</figureCitation>
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A–E)
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0CD24083112AFFD3FF31CC0BFDAE4602" box="[151,639,1066,1088]" pageId="1" pageNumber="471" type="description">
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<paragraph id="44771308112AFFD3FF31CC0BFDAE4602" blockId="1.[151,639,1066,1088]" box="[151,639,1066,1088]" pageId="1" pageNumber="471">
|
||
<taxonomicName id="83C8688B112AFFD3FF31CC0BFE544602" box="[151,389,1066,1088]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="471" phylum="Arthropoda" rank="species" species="sagedae">
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||
<emphasis id="76BCCF1A112AFFD3FF31CC0BFE544602" box="[151,389,1066,1088]" italics="true" pageId="1" pageNumber="471">Megaesthesius sagedae</emphasis>
|
||
</taxonomicName>
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||
—Poore
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||
<emphasis id="76BCCF1A112AFFD3FE46CC0AFDC24602" box="[480,531,1066,1088]" italics="true" pageId="1" pageNumber="471">et al.</emphasis>
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||
, 2008: 46.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0CD24083112AFFD3FF31CC4CFAF446EB" pageId="1" pageNumber="471" type="materials_examined">
|
||
<paragraph id="44771308112AFFD3FF31CC4CFAF446EB" blockId="1.[151,1437,1133,2021]" pageId="1" pageNumber="471">
|
||
<emphasis id="76BCCF1A112AFFD3FF31CC4CFE5346C4" bold="true" box="[151,386,1133,1158]" pageId="1" pageNumber="471">Material Examined</emphasis>
|
||
.
|
||
<typeStatus id="9B73ADAA112AFFD3FE32CC4CFDFD46C7" box="[404,556,1133,1157]" pageId="1" pageNumber="471" type="holotype">HOLOTYPE</typeStatus>
|
||
: MV-
|
||
<accessionNumber id="5B9B8EEB112AFFD3FDD1CC4CFD1A46C7" box="[631,715,1133,1157]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J54597" pageId="1" pageNumber="471" type="EnaNcbi">J54597</accessionNumber>
|
||
, male (3.8×
|
||
<quantity id="8330BEED112AFFD3FCF2CC4CFC6346C7" box="[852,946,1133,1157]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="471" unit="mm" value="3.4">3.4 mm</quantity>
|
||
),
|
||
<geoCoordinate id="21FC75CF112AFFD3FC6ECC4CFB9E46C7" box="[968,1103,1133,1157]" direction="south" orientation="latitude" pageId="1" pageNumber="471" precision="9" value="-22.066668">22°04.00' S</geoCoordinate>
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,
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<geoCoordinate id="21FC75CF112AFFD3FBF8CC4CFB2546C7" box="[1118,1268,1133,1157]" direction="east" orientation="longitude" pageId="1" pageNumber="471" precision="9" value="113.80666">113°48.40' E</geoCoordinate>
|
||
, off Ningaloo, Western
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||
<collectingCountry id="3CDF5398112AFFD3FF5FCCB0FEB346EB" box="[249,354,1169,1193]" name="Australia" pageId="1" pageNumber="471">Australia</collectingCountry>
|
||
, depth
|
||
<quantity id="8330BEED112AFFD3FE15CCB0FDE246EB" box="[435,563,1169,1193]" metricMagnitude="2" metricUnit="m" metricValue="1.035" metricValueMax="1.06" metricValueMin="1.01" pageId="1" pageNumber="471" unit="m" value="103.5" valueMax="106.0" valueMin="101.0">101–106 m</quantity>
|
||
,
|
||
<emphasis id="76BCCF1A112AFFD3FD99CCB0FCC146EB" box="[575,784,1169,1193]" italics="true" pageId="1" pageNumber="471">Southern Surveyor</emphasis>
|
||
voyage SS10-2005, Sherman sled,
|
||
<date id="307635C8112AFFD3FB39CCB0FAF046EB" box="[1183,1313,1169,1193]" pageId="1" pageNumber="471" value="2005-10-12">10.12.2005</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0CD24083112AFFD0FF61CC94FCAC4582" lastPageId="2" lastPageNumber="472" pageId="1" pageNumber="471" type="description">
|
||
<paragraph id="44771308112AFFD3FF61CC94FCFC443F" blockId="1.[151,1437,1133,2021]" pageId="1" pageNumber="471">
|
||
<emphasis id="76BCCF1A112AFFD3FF61CC94FE0D468F" bold="true" box="[199,476,1205,1230]" pageId="1" pageNumber="471">
|
||
Description of
|
||
<typeStatus id="9B73ADAA112AFFD3FED3CC94FE0D468F" box="[373,476,1205,1229]" pageId="1" pageNumber="471" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. Carapace subquadrate, 1.1 times wider than long, strongly convex antero-posteriorly, flat from side to side; regions indistinct except for moderately well-defined urogastic-cardiac regions separated by broad shallow grooves; surface moderately rugose; conspicuously but sparsely granular anteriorly, laterally near margin, posterolaterally. Frontal margin strongly pronounced; bilobed; frontal region swollen either side of median sulcus. Supraorbital border moderately deflexed but margin clearly visible in dorsal view; weakly sinuous medially, broadly rounded laterally; sloping slightly below first anterolateral tooth; mostly smooth but granular near anterolateral margin. Lateral carapace margins almost parallel; bearing 4 teeth separated by clear notches. First lateral tooth commencing above lateral outer orbital rim; forming broad flat lobe with weakly granular rim; second tooth of similar form, shorter, separated from first by shallow notch; third tooth the largest, almost as long as first 2 teeth combined, separated from second tooth by deeper u-shaped sulcus, rim minutely granular only; fourth tooth in form of large granular tubercle, relatively more widely separated from third tooth, single large granule between teeth. Posterolateral border behind last tooth marked by sparse strong moderately sharp granules. Posterolateral walls of carapace diverging; visible dorsally.
|
||
</paragraph>
|
||
<paragraph id="44771308112AFFD3FF61CEA8FA4D45A7" blockId="1.[151,1437,1133,2021]" pageId="1" pageNumber="471">
|
||
Infraorbital border (
|
||
<figureCitation id="DCF30F8D112AFFD3FE13CEA8FDD144E3" box="[437,512,1673,1697]" captionStart="FIGURE 2" captionStartId="3.[151,250,1119,1142]" captionTargetBox="[151,1409,412,1079]" captionTargetId="figure@3.[151,1436,409,1098]" captionTargetPageId="3" captionText="FIGURE 2. Megaesthesius westralia sp. nov., holotype male. A, anterodorsal view of frontal and orbital regions; B, ventral view of infraorbital margin showing eyestalk; C, third maxilliped; D, fourth pereiopod; E, fifth pereiopod." httpUri="https://zenodo.org/record/216409/files/figure.png" pageId="1" pageNumber="471">Fig. 2</figureCitation>
|
||
B) smooth; straight over most of length, laterally curved; lacking obviously developed tooth at inner end. Eyestalk long; frontal edge armed with row of low, well-spaced, blunt to subacute tubercles; embedded in orbital hiatus, immovable; distal quarter of length projecting free of orbit, curved to lie closely abutting lateral carapace wall (
|
||
<figureCitation id="DCF30F8D112AFFD3FDFBCED4FD74454F" box="[605,677,1781,1805]" captionStart="FIGURE 2" captionStartId="3.[151,250,1119,1142]" captionTargetBox="[151,1409,412,1079]" captionTargetId="figure@3.[151,1436,409,1098]" captionTargetPageId="3" captionText="FIGURE 2. Megaesthesius westralia sp. nov., holotype male. A, anterodorsal view of frontal and orbital regions; B, ventral view of infraorbital margin showing eyestalk; C, third maxilliped; D, fourth pereiopod; E, fifth pereiopod." httpUri="https://zenodo.org/record/216409/files/figure.png" pageId="1" pageNumber="471">Fig. 2</figureCitation>
|
||
B), just visible dorsally as lateral protrusion below antero-lateral margin; cornea poorly defined, rounded, with only hint of pigment. Antennule greatly enlarged, projecting, lacking fossa, flagellum fringed with long fine setae; wholly visible dorsally except for basal segment. Antennal basal segment similar but slightly stouter than second; flagellum long, but not quite reaching level of antero-lateral tooth of carapace, with 2 long fine setae apically. Pterygostomial region coarsely granular distolaterally otherwise microscopically granular; strongly swollen, separated from subhepatic region by broad sulcus; strong clear suture line. Buccal cavity with raised lateral rim; margins broadly curved, narrowing anteriorly. Third maxillipeds (
|
||
<figureCitation id="DCF30F8D112AFFD3FAC8CFECFA4D45A7" box="[1390,1436,1997,2021]" captionStart="FIGURE 2" captionStartId="3.[151,250,1119,1142]" captionTargetBox="[151,1409,412,1079]" captionTargetId="figure@3.[151,1436,409,1098]" captionTargetPageId="3" captionText="FIGURE 2. Megaesthesius westralia sp. nov., holotype male. A, anterodorsal view of frontal and orbital regions; B, ventral view of infraorbital margin showing eyestalk; C, third maxilliped; D, fourth pereiopod; E, fifth pereiopod." httpUri="https://zenodo.org/record/216409/files/figure.png" pageId="1" pageNumber="471">Fig.</figureCitation>
|
||
</paragraph>
|
||
<paragraph id="447713081129FFD0FF31C8B6FD6242B5" blockId="2.[151,1436,151,247]" pageId="2" pageNumber="472">2C) wide, only narrow median gape; ischium longer than merus, subquadrate, armed with 4 or 5 spinules on outer lateral margin; merus subtriangular, uneven row of minute spinules near outer lateral margin; palp short, reaching only about half length of inner margin of merus.</paragraph>
|
||
<caption id="10B743801129FFD0FF31CDBDFC814793" httpUri="https://zenodo.org/record/216408/files/figure.png" pageId="2" pageNumber="472" targetBox="[158,1436,310,1375]" targetPageId="2">
|
||
<paragraph id="447713081129FFD0FF31CDBDFC814793" blockId="2.[151,1436,1436,1489]" pageId="2" pageNumber="472">
|
||
<emphasis id="76BCCF1A1129FFD0FF31CDBDFEC347F3" bold="true" box="[151,274,1436,1459]" pageId="2" pageNumber="472">FIGURE 1.</emphasis>
|
||
<taxonomicName id="83C8688B1129FFD0FEBECDBDFDC047F0" box="[280,529,1436,1458]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="472" phylum="Arthropoda" rank="species" species="westralia" status="sp. nov.">
|
||
<emphasis id="76BCCF1A1129FFD0FEBECDBDFDC047F0" box="[280,529,1436,1458]" italics="true" pageId="2" pageNumber="472">Megaesthesius westralia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="76BCCF1A1129FFD0FDB1CDBCFDB947F0" bold="true" box="[535,616,1437,1458]" pageId="2" pageNumber="472">
|
||
<taxonomicNameLabel id="6D8F72611129FFD0FDB1CDBCFDB947F0" box="[535,616,1437,1458]" pageId="2" pageNumber="472" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male. A, carapace in dorsal view; B, fronto-ventral view showing third maxilliped and buccal frame; C, latero-ventral view; D, frontal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="447713081129FFD0FF61CDD9FACA443E" blockId="2.[151,1437,1528,2021]" pageId="2" pageNumber="472">
|
||
Chelipeds missing. Walking legs (
|
||
<figureCitation id="DCF30F8D1129FFD0FDF0CDD9FD4F4452" box="[598,670,1528,1552]" captionStart="FIGURE 2" captionStartId="3.[151,250,1119,1142]" captionTargetBox="[151,1409,412,1079]" captionTargetId="figure@3.[151,1436,409,1098]" captionTargetPageId="3" captionText="FIGURE 2. Megaesthesius westralia sp. nov., holotype male. A, anterodorsal view of frontal and orbital regions; B, ventral view of infraorbital margin showing eyestalk; C, third maxilliped; D, fourth pereiopod; E, fifth pereiopod." httpUri="https://zenodo.org/record/216409/files/figure.png" pageId="2" pageNumber="472">Fig. 2</figureCitation>
|
||
D, E) long, slender; sparsely fringed with plumose setae; last leg shorter than preceding. P 4 merus c. 4.9 times longer than wide; anterior posterior borders with sparse row of small spinules. Carpus unarmed. Propodus/dactylus each with some large stout, movable spines along posterior border. P5 merus c. 4.4 times longer than wide; stout, movable spines absent on propodus, smaller on dactylus.
|
||
</paragraph>
|
||
<paragraph id="447713081129FFD0FF61CEA9FCAC4582" blockId="2.[151,1437,1528,2021]" pageId="2" pageNumber="472">
|
||
Thoracic sternum broad; sternite 3 separated from fused sternites 1/2 by clear suture line; sternite 3 fused to sternite 4, with only remnants of suture present near articulation of maxilliped 3; sternites 3–4 transversely sunken medially. Surface of sternites 4–8 coarsely granular. Penis with outer (ventral) surface calcified, forming a movable, plate-like structure between anterior supplementary plate and sternite 8. Male abdomen with third to fifth somites fused, though sutures partially visible; telson broadly rounded, c. 1.6 times wider than long, slightly shorter than sixth somite; sixth somite twice as wide as long. G1 (
|
||
<figureCitation id="DCF30F8D1129FFD0FCE4CF1DFC544516" box="[834,901,1852,1876]" captionStart="FIGURE 3" captionStartId="4.[151,250,1854,1877]" captionTargetBox="[168,1434,199,1819]" captionTargetId="figure@4.[168,1453,193,1821]" captionTargetPageId="4" captionText="FIGURE 3. Male gonopods and abdomens for the three known species of Megaesthesius. A – E, M. westralia sp. nov.: A, left G 1 in sternal view; B, left G 1 in abdominal view; C, left G 1, detail of tip; D, left G 2; E, male abdomen with lateral view of sternites 7 – 8 and penis. F – I, M. yokoyai Sakai, 1939: F, left G 1 in abdominal view; G, H. distal part of G 1 in abdominal and sternal view; I, male abdomen (somites 1 – 2 not shown) (After Takeda & Miyake, 1972). J – N, M. sagedae Rathbun, 1909: J, left G 1, abdominal face; K, detail of tip; L, left G 1 sternal face; M, left G 2 abdominal face; N, male abdomen (after Crosnier 1975)." httpUri="https://zenodo.org/record/216410/files/figure.png" pageId="2" pageNumber="472">Fig. 3</figureCitation>
|
||
A–C) stout, tapering to about middle, margins subparallel until distal end; apex markedly tapering towards mid-line; apical opening broad, elongated, triangular; row of short stout moveable spinules along long edge above opening. G2 (
|
||
<figureCitation id="DCF30F8D1129FFD0FC44CFA5FBF445DE" box="[994,1061,1924,1948]" captionStart="FIGURE 3" captionStartId="4.[151,250,1854,1877]" captionTargetBox="[168,1434,199,1819]" captionTargetId="figure@4.[168,1453,193,1821]" captionTargetPageId="4" captionText="FIGURE 3. Male gonopods and abdomens for the three known species of Megaesthesius. A – E, M. westralia sp. nov.: A, left G 1 in sternal view; B, left G 1 in abdominal view; C, left G 1, detail of tip; D, left G 2; E, male abdomen with lateral view of sternites 7 – 8 and penis. F – I, M. yokoyai Sakai, 1939: F, left G 1 in abdominal view; G, H. distal part of G 1 in abdominal and sternal view; I, male abdomen (somites 1 – 2 not shown) (After Takeda & Miyake, 1972). J – N, M. sagedae Rathbun, 1909: J, left G 1, abdominal face; K, detail of tip; L, left G 1 sternal face; M, left G 2 abdominal face; N, male abdomen (after Crosnier 1975)." httpUri="https://zenodo.org/record/216410/files/figure.png" pageId="2" pageNumber="472">Fig. 3</figureCitation>
|
||
D) sinuous, subequal in length to G1; markedly constricted medially to form long curved flagellum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0CD240831129FFD0FF61CFECFD7545A6" box="[199,676,1996,2021]" pageId="2" pageNumber="472" type="etymology">
|
||
<paragraph id="447713081129FFD0FF61CFECFD7545A6" blockId="2.[151,1437,1528,2021]" box="[199,676,1996,2021]" pageId="2" pageNumber="472">
|
||
<emphasis id="76BCCF1A1129FFD0FF61CFECFE9745A7" bold="true" box="[199,326,1997,2021]" pageId="2" pageNumber="472">Etymology</emphasis>
|
||
. Named for Western
|
||
<collectingCountry id="3CDF53981129FFD0FD93CFEDFD7145A6" box="[565,672,1996,2020]" name="Australia" pageId="2" pageNumber="472">Australia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0CD240831128FFD1FF61C8B6FA4D42ED" box="[199,1436,151,176]" pageId="3" pageNumber="473" type="distribution">
|
||
<paragraph id="447713081128FFD1FF61C8B6FA4D42ED" blockId="3.[151,1436,151,356]" box="[199,1436,151,176]" pageId="3" pageNumber="473">
|
||
<emphasis id="76BCCF1A1128FFD1FF61C8B6FE8842F2" bold="true" box="[199,345,151,176]" pageId="3" pageNumber="473">Distribution</emphasis>
|
||
. Only known from the
|
||
<typeStatus id="9B73ADAA1128FFD1FDF9C8B9FD5E42F2" box="[607,655,152,176]" pageId="3" pageNumber="473">type</typeStatus>
|
||
locality off Exmouth, Western
|
||
<collectingCountry id="3CDF53981128FFD1FC48C8B6FB8942ED" box="[1006,1112,151,175]" name="Australia" pageId="3" pageNumber="473">Australia</collectingCountry>
|
||
; from a depth of
|
||
<quantity id="8330BEED1128FFD1FAB0C8B6FA4942ED" box="[1302,1432,151,175]" metricMagnitude="2" metricUnit="m" metricValue="1.035" metricValueMax="1.06" metricValueMin="1.01" pageId="3" pageNumber="473" unit="m" value="103.5" valueMax="106.0" valueMin="101.0">
|
||
101–
|
||
<specimenCount id="52CED8811128FFD1FAE8C8B6FA4942ED" box="[1358,1432,151,175]" pageId="3" pageNumber="473" type="generic">106 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0CD240831128FFD1FF61C89DFB7245E6" pageId="3" pageNumber="473" type="discussion">
|
||
<paragraph id="447713081128FFD1FF61C89DFE474326" blockId="3.[151,1436,151,356]" pageId="3" pageNumber="473">
|
||
<emphasis id="76BCCF1A1128FFD1FF61C89DFEE54296" bold="true" box="[199,308,188,212]" pageId="3" pageNumber="473">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="83C8688B1128FFD1FEE2C89DFD8B4296" box="[324,602,188,212]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="473" phylum="Arthropoda" rank="species" species="westralia" status="sp. nov.">
|
||
<emphasis id="76BCCF1A1128FFD1FEE2C89DFD8B4296" box="[324,602,188,212]" italics="true" pageId="3" pageNumber="473">Megaesthesius westralia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6D8F72611128FFD1FDC5C89DFD6F4296" box="[611,702,188,212]" pageId="3" pageNumber="473" rank="species">
|
||
<emphasis id="76BCCF1A1128FFD1FDC5C89DFD6F4296" bold="true" box="[611,702,188,212]" pageId="3" pageNumber="473">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be easily separated from the
|
||
<typeStatus id="9B73ADAA1128FFD1FBE5C89CFBA24297" box="[1091,1139,189,213]" pageId="3" pageNumber="473">type</typeStatus>
|
||
of the genus,
|
||
<taxonomicName id="83C8688B1128FFD1FABFC89CFEEE42B5" authority="Rathbun, 1909" authorityName="Rathbun" authorityYear="1909" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="473" phylum="Arthropoda" rank="species" species="sagedae">
|
||
<emphasis id="76BCCF1A1128FFD1FABFC89CFA4D4296" box="[1305,1436,188,212]" italics="true" pageId="3" pageNumber="473">M. sagedae</emphasis>
|
||
Rathbun, 1909
|
||
</taxonomicName>
|
||
, by the lack of spination on the lateral and frontal margins of the carapace, the presence of truncate lobes on the lateral margins, the shape of the male abdomen, particularly a narrower sixth somite (compare
|
||
<figureCitation id="DCF30F8D1128FFD1FAECC925FA58435E" box="[1354,1417,260,284]" captionStart="FIGURE 3" captionStartId="4.[151,250,1854,1877]" captionTargetBox="[168,1434,199,1819]" captionTargetId="figure@4.[168,1453,193,1821]" captionTargetPageId="4" captionText="FIGURE 3. Male gonopods and abdomens for the three known species of Megaesthesius. A – E, M. westralia sp. nov.: A, left G 1 in sternal view; B, left G 1 in abdominal view; C, left G 1, detail of tip; D, left G 2; E, male abdomen with lateral view of sternites 7 – 8 and penis. F – I, M. yokoyai Sakai, 1939: F, left G 1 in abdominal view; G, H. distal part of G 1 in abdominal and sternal view; I, male abdomen (somites 1 – 2 not shown) (After Takeda & Miyake, 1972). J – N, M. sagedae Rathbun, 1909: J, left G 1, abdominal face; K, detail of tip; L, left G 1 sternal face; M, left G 2 abdominal face; N, male abdomen (after Crosnier 1975)." httpUri="https://zenodo.org/record/216410/files/figure.png" pageId="3" pageNumber="473">Fig. 3</figureCitation>
|
||
E and 3N), a more slender G1 (compare
|
||
<figureCitation id="DCF30F8D1128FFD1FDE8C906FD5E437D" box="[590,655,295,319]" captionStart="FIGURE 3" captionStartId="4.[151,250,1854,1877]" captionTargetBox="[168,1434,199,1819]" captionTargetId="figure@4.[168,1453,193,1821]" captionTargetPageId="4" captionText="FIGURE 3. Male gonopods and abdomens for the three known species of Megaesthesius. A – E, M. westralia sp. nov.: A, left G 1 in sternal view; B, left G 1 in abdominal view; C, left G 1, detail of tip; D, left G 2; E, male abdomen with lateral view of sternites 7 – 8 and penis. F – I, M. yokoyai Sakai, 1939: F, left G 1 in abdominal view; G, H. distal part of G 1 in abdominal and sternal view; I, male abdomen (somites 1 – 2 not shown) (After Takeda & Miyake, 1972). J – N, M. sagedae Rathbun, 1909: J, left G 1, abdominal face; K, detail of tip; L, left G 1 sternal face; M, left G 2 abdominal face; N, male abdomen (after Crosnier 1975)." httpUri="https://zenodo.org/record/216410/files/figure.png" pageId="3" pageNumber="473">Fig. 3</figureCitation>
|
||
A–C and 3J–L), and also a more slender and much longer G2 with a long curved flagellum.
|
||
</paragraph>
|
||
<caption id="10B743801128FFD1FF31CC7EFB1B46D6" httpUri="https://zenodo.org/record/216409/files/figure.png" pageId="3" pageNumber="473" targetBox="[151,1409,412,1079]" targetPageId="3">
|
||
<paragraph id="447713081128FFD1FF31CC7EFB1B46D6" blockId="3.[151,1436,1119,1172]" pageId="3" pageNumber="473">
|
||
<emphasis id="76BCCF1A1128FFD1FF31CC7EFEC54637" bold="true" box="[151,276,1119,1142]" pageId="3" pageNumber="473">FIGURE 2.</emphasis>
|
||
<taxonomicName id="83C8688B1128FFD1FEBACC41FDC64634" box="[284,535,1120,1142]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="473" phylum="Arthropoda" rank="species" species="westralia" status="sp. nov.">
|
||
<emphasis id="76BCCF1A1128FFD1FEBACC41FDC64634" box="[284,535,1120,1142]" italics="true" pageId="3" pageNumber="473">Megaesthesius westralia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="76BCCF1A1128FFD1FDB9CC41FDA04637" bold="true" box="[543,625,1120,1141]" pageId="3" pageNumber="473">
|
||
<taxonomicNameLabel id="6D8F72611128FFD1FDB9CC41FDA04637" box="[543,625,1120,1141]" pageId="3" pageNumber="473" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male. A, anterodorsal view of frontal and orbital regions; B, ventral view of infraorbital margin showing eyestalk; C, third maxilliped; D, fourth pereiopod; E, fifth pereiopod.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="447713081128FFD1FF61CC9DFDA14455" blockId="3.[151,1437,1212,1956]" pageId="3" pageNumber="473">
|
||
<taxonomicName id="83C8688B1128FFD1FF61CC9DFE0B4696" box="[199,474,1212,1236]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="473" phylum="Arthropoda" rank="species" species="westralia" status="sp. nov.">
|
||
<emphasis id="76BCCF1A1128FFD1FF61CC9DFE0B4696" box="[199,474,1212,1236]" italics="true" pageId="3" pageNumber="473">Megaesthesius westralia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="76BCCF1A1128FFD1FE47CC9DFDEB4696" bold="true" box="[481,570,1212,1236]" pageId="3" pageNumber="473">
|
||
<taxonomicNameLabel id="6D8F72611128FFD1FE47CC9DFDEB4696" box="[481,570,1212,1236]" pageId="3" pageNumber="473" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is more closely related to
|
||
<taxonomicName id="83C8688B1128FFD1FCC6CC9CFBB84696" authority="Sakai, 1939" authorityName="Sakai" authorityYear="1939" box="[864,1129,1212,1236]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="473" phylum="Arthropoda" rank="species" species="yokoyai">
|
||
<emphasis id="76BCCF1A1128FFD1FCC6CC9CFC0D4696" box="[864,988,1212,1236]" italics="true" pageId="3" pageNumber="473">M. yokoyai</emphasis>
|
||
Sakai, 1939
|
||
</taxonomicName>
|
||
, in that both share a similar carapace form, with bluntly truncate, unarmed lobes/teeth on the lateral margins, although the shape and length of these appear to differ between the two species. In
|
||
<taxonomicName id="83C8688B1128FFD1FD67CD24FC9C475E" box="[705,845,1284,1308]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="473" phylum="Arthropoda" rank="species" species="westralia" status="sp. nov.">
|
||
<emphasis id="76BCCF1A1128FFD1FD67CD24FC9C475E" box="[705,845,1284,1308]" italics="true" pageId="3" pageNumber="473">M. westralia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="76BCCF1A1128FFD1FCF2CD25FC7C475E" bold="true" box="[852,941,1284,1308]" pageId="3" pageNumber="473">
|
||
<taxonomicNameLabel id="6D8F72611128FFD1FCF2CD25FC7C475E" box="[852,941,1284,1308]" pageId="3" pageNumber="473" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
the third lobe is particularly long, extending further posteriorly, and is clearly demarcated from the last broad tubercular tooth. Also the last tooth in
|
||
<taxonomicName id="83C8688B1128FFD1FAB9CD08FA4D4702" box="[1311,1436,1320,1344]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="473" phylum="Arthropoda" rank="species" species="yokoyai">
|
||
<emphasis id="76BCCF1A1128FFD1FAB9CD08FA4D4702" box="[1311,1436,1320,1344]" italics="true" pageId="3" pageNumber="473">M. yokoyai</emphasis>
|
||
</taxonomicName>
|
||
seems to be lower, less pronounced, and more spinous than in the new species.
|
||
<taxonomicName id="83C8688B1128FFD1FB9BCD6CFB1C4726" box="[1085,1229,1356,1380]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="473" phylum="Arthropoda" rank="species" species="westralia" status="sp. nov.">
|
||
<emphasis id="76BCCF1A1128FFD1FB9BCD6CFB1C4726" box="[1085,1229,1356,1380]" italics="true" pageId="3" pageNumber="473">M. westralia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="76BCCF1A1128FFD1FB7ECD6DFAE44726" bold="true" box="[1240,1333,1356,1380]" pageId="3" pageNumber="473">
|
||
<taxonomicNameLabel id="6D8F72611128FFD1FB7ECD6DFAE44726" box="[1240,1333,1356,1380]" pageId="3" pageNumber="473" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
also has longer, more slender walking legs, and in particular the propodus and carpus of P3 are noticeably longer (compare
|
||
<figureCitation id="DCF30F8D1128FFD1FF31CDB5FF0847EE" box="[151,217,1428,1452]" captionStart="FIGURE 2" captionStartId="3.[151,250,1119,1142]" captionTargetBox="[151,1409,412,1079]" captionTargetId="figure@3.[151,1436,409,1098]" captionTargetPageId="3" captionText="FIGURE 2. Megaesthesius westralia sp. nov., holotype male. A, anterodorsal view of frontal and orbital regions; B, ventral view of infraorbital margin showing eyestalk; C, third maxilliped; D, fourth pereiopod; E, fifth pereiopod." httpUri="https://zenodo.org/record/216409/files/figure.png" pageId="3" pageNumber="473">Fig. 2</figureCitation>
|
||
D and 5B). The male abdomens also differ between the two species, with
|
||
<taxonomicName id="83C8688B1128FFD1FB86CDB4FB4C47EE" box="[1056,1181,1428,1452]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="473" phylum="Arthropoda" rank="species" species="yokoyai">
|
||
<emphasis id="76BCCF1A1128FFD1FB86CDB4FB4C47EE" box="[1056,1181,1428,1452]" italics="true" pageId="3" pageNumber="473">M. yokoyai</emphasis>
|
||
</taxonomicName>
|
||
having a much longer telson, and a relatively wider, narrower somite 6 (compare
|
||
<figureCitation id="DCF30F8D1128FFD1FC95CD96FCA2478D" box="[819,883,1463,1487]" captionStart="FIGURE 3" captionStartId="4.[151,250,1854,1877]" captionTargetBox="[168,1434,199,1819]" captionTargetId="figure@4.[168,1453,193,1821]" captionTargetPageId="4" captionText="FIGURE 3. Male gonopods and abdomens for the three known species of Megaesthesius. A – E, M. westralia sp. nov.: A, left G 1 in sternal view; B, left G 1 in abdominal view; C, left G 1, detail of tip; D, left G 2; E, male abdomen with lateral view of sternites 7 – 8 and penis. F – I, M. yokoyai Sakai, 1939: F, left G 1 in abdominal view; G, H. distal part of G 1 in abdominal and sternal view; I, male abdomen (somites 1 – 2 not shown) (After Takeda & Miyake, 1972). J – N, M. sagedae Rathbun, 1909: J, left G 1, abdominal face; K, detail of tip; L, left G 1 sternal face; M, left G 2 abdominal face; N, male abdomen (after Crosnier 1975)." httpUri="https://zenodo.org/record/216410/files/figure.png" pageId="3" pageNumber="473">Fig. 3</figureCitation>
|
||
E and
|
||
<date id="307635C81128FFD1FC18CD96FC09478D" box="[958,984,1463,1487]" pageId="3" pageNumber="473">3I</date>
|
||
). The G1 structure of the two species is also different (compare
|
||
<figureCitation id="DCF30F8D1128FFD1FE0BCDFDFE2147B6" box="[429,496,1500,1524]" captionStart="FIGURE 3" captionStartId="4.[151,250,1854,1877]" captionTargetBox="[168,1434,199,1819]" captionTargetId="figure@4.[168,1453,193,1821]" captionTargetPageId="4" captionText="FIGURE 3. Male gonopods and abdomens for the three known species of Megaesthesius. A – E, M. westralia sp. nov.: A, left G 1 in sternal view; B, left G 1 in abdominal view; C, left G 1, detail of tip; D, left G 2; E, male abdomen with lateral view of sternites 7 – 8 and penis. F – I, M. yokoyai Sakai, 1939: F, left G 1 in abdominal view; G, H. distal part of G 1 in abdominal and sternal view; I, male abdomen (somites 1 – 2 not shown) (After Takeda & Miyake, 1972). J – N, M. sagedae Rathbun, 1909: J, left G 1, abdominal face; K, detail of tip; L, left G 1 sternal face; M, left G 2 abdominal face; N, male abdomen (after Crosnier 1975)." httpUri="https://zenodo.org/record/216410/files/figure.png" pageId="3" pageNumber="473">Fig. 3</figureCitation>
|
||
A–C & 3F–H), with
|
||
<taxonomicName id="83C8688B1128FFD1FD41CDFCFCA947B6" box="[743,888,1500,1524]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="473" phylum="Arthropoda" rank="species" species="westralia">
|
||
<emphasis id="76BCCF1A1128FFD1FD41CDFCFCA947B6" box="[743,888,1500,1524]" italics="true" pageId="3" pageNumber="473">M. westralia</emphasis>
|
||
</taxonomicName>
|
||
being constricted more proximally, and the tip being more acutely and obliquely pointed.
|
||
</paragraph>
|
||
<paragraph id="447713081128FFD1FF61CE05FB7245E6" blockId="3.[151,1437,1212,1956]" pageId="3" pageNumber="473">
|
||
The shape of the male G2 is particularly noteworthy. In
|
||
<taxonomicName id="83C8688B1128FFD1FC9ECE05FB9D447E" box="[824,1100,1572,1596]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="473" phylum="Arthropoda" rank="species" species="westralia" status="sp. nov.">
|
||
<emphasis id="76BCCF1A1128FFD1FC9ECE05FB9D447E" box="[824,1100,1572,1596]" italics="true" pageId="3" pageNumber="473">Megaesthesius westralia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="76BCCF1A1128FFD1FBF5CE05FB7D447E" bold="true" box="[1107,1196,1572,1596]" pageId="3" pageNumber="473">
|
||
<taxonomicNameLabel id="6D8F72611128FFD1FBF5CE05FB7D447E" box="[1107,1196,1572,1596]" pageId="3" pageNumber="473" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
it is of similar length to the G1, and medially constricted with a long curved flagellum; however in the only other figure or description of the G2 (for the specimen of
|
||
<taxonomicName id="83C8688B1128FFD1FE76CE4CFD8144C6" box="[464,592,1644,1668]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="473" phylum="Arthropoda" rank="species" species="sagedae">
|
||
<emphasis id="76BCCF1A1128FFD1FE76CE4CFD8144C6" box="[464,592,1644,1668]" italics="true" pageId="3" pageNumber="473">M. sagedae</emphasis>
|
||
</taxonomicName>
|
||
from off
|
||
<collectingCountry id="3CDF53981128FFD1FD1DCE4DFC9344C6" box="[699,834,1644,1668]" name="Madagascar" pageId="3" pageNumber="473">Madagascar</collectingCountry>
|
||
by Crosnier1975), it is apparently shorter (about twothirds the length of the G1), stouter, and while tapering to a slender tip, the flagellum is markedly broadened at its base, and not obviously constricted at this point. Unfortunately, Serène’s (1964) refiguring of
|
||
<taxonomicName id="83C8688B1128FFD1FB12CE94FAE4448E" box="[1204,1333,1716,1740]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="473" phylum="Arthropoda" rank="species" species="sagedae">
|
||
<emphasis id="76BCCF1A1128FFD1FB12CE94FAE4448E" box="[1204,1333,1716,1740]" italics="true" pageId="3" pageNumber="473">M. sagedae</emphasis>
|
||
</taxonomicName>
|
||
from the Gulf of
|
||
<collectingCountry id="3CDF53981128FFD1FF48CEF9FE8044B2" box="[238,337,1752,1776]" name="Thailand" pageId="3" pageNumber="473">Thailand</collectingCountry>
|
||
was of a female specimen, so there is currently no information on the gonopod morphology of this species from either
|
||
<collectingCountry id="3CDF53981128FFD1FED3CEDDFE384556" box="[373,489,1788,1812]" name="Singapore" pageId="3" pageNumber="473">Singapore</collectingCountry>
|
||
, its
|
||
<typeStatus id="9B73ADAA1128FFD1FDB1CEDCFD964557" box="[535,583,1789,1813]" pageId="3" pageNumber="473">type</typeStatus>
|
||
locality, or elsewhere in Southeast Asia. It is highly unusual to have such a marked difference in G2 morphology within a genus, and this bears closer examination as part of a larger revisionary study. Unfortunately, although Takeda & Miyake (1969) gave excellent figures of the G1 of
|
||
<taxonomicName id="83C8688B1128FFD1FA86CF64FA4D451E" box="[1312,1436,1860,1884]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="473" phylum="Arthropoda" rank="species" species="yokoyai">
|
||
<emphasis id="76BCCF1A1128FFD1FA86CF64FA4D451E" box="[1312,1436,1860,1884]" italics="true" pageId="3" pageNumber="473">M. yokoyai</emphasis>
|
||
</taxonomicName>
|
||
they failed to either describe or figure the G2. Because of the close morphological similarities of
|
||
<taxonomicName id="83C8688B1128FFD1FB70CF48FAB245C2" box="[1238,1379,1896,1920]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="473" phylum="Arthropoda" rank="species" species="westralia">
|
||
<emphasis id="76BCCF1A1128FFD1FB70CF48FAB245C2" box="[1238,1379,1896,1920]" italics="true" pageId="3" pageNumber="473">M. westralia</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="83C8688B1128FFD1FF31CFACFEC245E6" box="[151,275,1932,1956]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="473" phylum="Arthropoda" rank="species" species="yokoyai">
|
||
<emphasis id="76BCCF1A1128FFD1FF31CFACFEC245E6" box="[151,275,1932,1956]" italics="true" pageId="3" pageNumber="473">M. yokoyai</emphasis>
|
||
</taxonomicName>
|
||
it seems likely that these two species will prove to share similar G2 morphology.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |