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<document ID-DOI="10.11646/palaeoentomology.5.5.10" ID-ISSN="2624-2834" ID-Zenodo-Dep="7333603" ID-ZooBank="3DE47527-54FD-405A-89C0-D71222CFF1B5" approvalRequired="1" approvalRequired_for_textStreams="1" checkinTime="1668766033449" checkinUser="plazi" docAuthor="ENGEL, MICHAEL S. &amp; WANG, BO" docDate="2022" docId="55471F0E6204FFBEFF363959E6D4940E" docLanguage="en" docName="palaeoentomology.5.5.10.pdf" docOrigin="Palaeoentomology 5 (5)" docSource="http://dx.doi.org/10.11646/palaeoentomology.5.5.10" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.2:Palaeontomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeontomology.2018-.journal_article" docStyleVersion="2" docTitle="Belaphotroctes grimaldii ENGEL &amp; WANG 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="490" masterDocId="A97E67766205FFBDFFBE3F23E5179C06" masterDocTitle="A new species of embidopsocine barklouse in Langhian amber from Zhangpu, China (Psocoptera: Liposcelididae)" masterLastPageNumber="492" masterPageNumber="487" pageNumber="488" updateTime="1668766413165" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of embidopsocine barklouse in Langhian amber from Zhangpu, China (Psocoptera: Liposcelididae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>ENGEL, MICHAEL S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>WANG, BO</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Palaeoentomology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-09-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>5</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>487</mods:start>
<mods:end>492</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.11646/palaeoentomology.5.5.10</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/palaeoentomology.5.5.10</mods:identifier>
<mods:identifier type="ISSN">2624-2834</mods:identifier>
<mods:identifier type="Zenodo-Dep">7333603</mods:identifier>
<mods:identifier type="ZooBank">3DE47527-54FD-405A-89C0-D71222CFF1B5</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:55471F0E6204FFBEFF363959E6D4940E" httpUri="http://treatment.plazi.org/id/55471F0E6204FFBEFF363959E6D4940E" lastPageId="3" lastPageNumber="490" pageId="1" pageNumber="488">
<subSubSection box="[136,518,1658,1685]" pageId="1" pageNumber="488" type="nomenclature">
<paragraph blockId="1.[136,518,1658,1722]" box="[136,518,1658,1685]" pageId="1" pageNumber="488">
<heading bold="true" box="[136,518,1658,1685]" fontSize="10" level="2" pageId="1" pageNumber="488" reason="3">
<emphasis bold="true" box="[136,518,1658,1685]" pageId="1" pageNumber="488">
<taxonomicName authority="ENGEL &amp; WANG, 2022" authorityName="ENGEL &amp; WANG" authorityYear="2022" box="[136,422,1658,1685]" class="Insecta" family="Liposcelididae" genus="Belaphotroctes" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="488" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
<emphasis bold="true" box="[136,422,1658,1685]" italics="true" pageId="1" pageNumber="488">Belaphotroctes grimaldii</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[429,518,1659,1685]" pageId="1" pageNumber="488" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[136,252,1696,1722]" pageId="1" pageNumber="488" type="description">
<paragraph blockId="1.[136,518,1658,1722]" box="[136,252,1696,1722]" pageId="1" pageNumber="488">
(
<figureCitation box="[144,244,1696,1722]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="2.[134,233,1584,1608]" captionStartId-1="3.[136,235,1509,1533]" captionStartId-2="4.[136,231,757,781]" captionTargetBox-0="[277,1305,189,1560]" captionTargetBox-1="[242,1346,193,1485]" captionTargetBox-2="[348,1204,231,710]" captionTargetId-0="figure-187@2.[277,1305,189,1561]" captionTargetId-1="figure-199@3.[242,1346,193,1485]" captionTargetId-2="figure-519@4.[339,1249,217,734]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 2. Habitus photographs of holotype female of Belaphotroctes grimaldii sp. nov. (NIGP200508); scale bar (for both images) = 1 mm. A, Dorsal habitus. B, Ventral habitus." captionText-1="FIGURE 3. Photographic details of holotype female of Belaphotroctes grimaldii sp. nov. (NIGP200508); scale bars = 0.25 mm. A, Head and pronotum, dorsal view. B, Head and pronotum, ventral view (P4 = maxillary palpomere IV). C, Apical abdominal sterna." captionText-2="FIGURE 4. Wing venation of Belaphotroctes grimaldii sp. nov. (NIGP200508), forewing above, hind wing below. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7333607" figureDoi-1="http://doi.org/10.5281/zenodo.7333609" figureDoi-2="http://doi.org/10.5281/zenodo.7333611" httpUri-0="https://zenodo.org/record/7333607/files/figure.png" httpUri-1="https://zenodo.org/record/7333609/files/figure.png" httpUri-2="https://zenodo.org/record/7333611/files/figure.png" pageId="1" pageNumber="488">Figs 24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="488" type="materials_examined">
<paragraph blockId="1.[136,778,1770,2093]" pageId="1" pageNumber="488">
<materialsCitation collectionCode="NIGP" country="China" county="Nanjing" location="Nanjing Institute of Geology" municipality="Chinese Academy of Sciences" pageId="1" pageNumber="488" specimenCode="NIGP200508" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis bold="true" box="[136,252,1770,1796]" pageId="1" pageNumber="488">
<typeStatus box="[136,247,1770,1796]" pageId="1" pageNumber="488">Holotype</typeStatus>
.
</emphasis>
<specimenCount box="[266,287,1771,1797]" pageId="1" pageNumber="488" type="female"></specimenCount>
,
<specimenCode box="[309,458,1770,1797]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="1" pageNumber="488">NIGP200508</specimenCode>
(
<figureCitation box="[481,557,1770,1796]" captionStart="FIGURE 2" captionStartId="2.[134,233,1584,1608]" captionTargetBox="[277,1305,189,1560]" captionTargetId="figure-187@2.[277,1305,189,1561]" captionTargetPageId="2" captionText="FIGURE 2. Habitus photographs of holotype female of Belaphotroctes grimaldii sp. nov. (NIGP200508); scale bar (for both images) = 1 mm. A, Dorsal habitus. B, Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.7333607" httpUri="https://zenodo.org/record/7333607/files/figure.png" pageId="1" pageNumber="488">Fig. 2</figureCitation>
), deposited in the
<location LSID="urn:lsid:plazi:treatment:55471F0E6204FFBEFF363959E6D4940E:D831F8C36204FFBCFF36382CE4DC9B2F" box="[136,459,1807,1833]" country="China" county="Nanjing" municipality="Chinese Academy of Sciences" name="Nanjing Institute of Geology" pageId="1" pageNumber="488">Nanjing Institute of Geology</location>
and
<location LSID="urn:lsid:plazi:treatment:55471F0E6204FFBEFF363959E6D4940E:D831F8C36204FFBCFDBD382CE7B39B2F" box="[515,676,1807,1833]" country="China" county="Nanjing" municipality="Chinese Academy of Sciences" name="Palaeontology" pageId="1" pageNumber="488">Palaeontology</location>
,
<collectingMunicipality pageId="1" pageNumber="488">Chinese Academy of Sciences</collectingMunicipality>
,
<collectingCounty box="[393,486,1844,1870]" pageId="1" pageNumber="488">Nanjing</collectingCounty>
,
<collectingCountry box="[497,568,1844,1870]" name="China" pageId="1" pageNumber="488">China</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="488" type="etymology">
<paragraph blockId="1.[136,778,1770,2093]" pageId="1" pageNumber="488">
<emphasis bold="true" box="[189,324,1881,1907]" pageId="1" pageNumber="488">Etymology.</emphasis>
The specific epithet honors David A. Grimaldi, worlds authority on insects, amber, evolution, and all things palaeoentomological, as well as a cherished friend.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="488" type="diagnosis">
<paragraph blockId="1.[136,778,1770,2093]" lastBlockId="1.[810,1452,820,2092]" pageId="1" pageNumber="488">
<emphasis bold="true" box="[189,312,2029,2055]" pageId="1" pageNumber="488">Diagnosis.</emphasis>
The new species belongs to that group of
<taxonomicName authorityName="Roesler" authorityYear="1943" box="[136,304,2066,2092]" class="Insecta" family="Liposcelididae" genus="Belaphotroctes" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis box="[136,304,2066,2092]" italics="true" pageId="1" pageNumber="488">Belaphotroctes</emphasis>
</taxonomicName>
lacking vesicles or pulvulli on the pretarsi. It can be further distinguished by the combination of a line of setae on R in the forewing, the presence of a second abscissa of R orthogonal to the longitudinal stem, the absence of Cu and A in the forewing, and the faint trace of C running along the anterior with margin and anterior to the short trace of Sc near the wing base (
<figureCitation box="[1265,1334,1003,1029]" captionStart="FIGURE 4" captionStartId="4.[136,231,757,781]" captionTargetBox="[348,1204,231,710]" captionTargetId="figure-519@4.[339,1249,217,734]" captionTargetPageId="4" captionText="FIGURE 4. Wing venation of Belaphotroctes grimaldii sp. nov. (NIGP200508), forewing above, hind wing below. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7333611" httpUri="https://zenodo.org/record/7333611/files/figure.png" pageId="1" pageNumber="488">Fig. 4</figureCitation>
).
</paragraph>
<paragraph blockId="1.[810,1452,820,2092]" pageId="1" pageNumber="488">
<emphasis bold="true" box="[863,1140,1039,1066]" pageId="1" pageNumber="488">Locality and horizon.</emphasis>
Zhangpu,
<collectingRegion box="[1281,1356,1040,1066]" country="China" name="Fujian" pageId="1" pageNumber="488">Fujian</collectingRegion>
,
<collectingCountry box="[1377,1447,1040,1066]" name="China" pageId="1" pageNumber="488">China</collectingCountry>
, middle Miocene Fotan Group (14.7 Ma).
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="490" pageId="1" pageNumber="488" type="description">
<paragraph blockId="1.[810,1452,820,2092]" pageId="1" pageNumber="488">
<emphasis bold="true" box="[863,1009,1113,1139]" pageId="1" pageNumber="488">Description.</emphasis>
<specimenCount box="[1030,1053,1114,1140]" pageId="1" pageNumber="488" type="female"></specimenCount>
: Total body length
<quantity box="[1329,1447,1113,1140]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" pageId="1" pageNumber="488" unit="mm" value="1.14">1.14 mm</quantity>
; integument finely imbricate to coriaceous; light brown except slightly darker on head and thoracic nota. Head slightly wider than long, length
<quantity box="[1186,1292,1223,1250]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="1" pageNumber="488" unit="mm" value="0.33">0.33 mm</quantity>
, width across compound eyes
<quantity box="[995,1099,1260,1286]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="488" unit="mm" value="0.37">0.37 mm</quantity>
, interocular distance
<quantity box="[1342,1447,1260,1286]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="488" unit="mm" value="0.24">0.24 mm</quantity>
; compound eye well-developed, diameter
<quantity box="[1283,1387,1296,1323]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="488" unit="mm" value="0.11">0.11 mm</quantity>
, with numerous ommatidia (over 30); ocelli arranged in tight triangle on slight swelling; epicranial cleavage scar absent; surface of head with sparse, minute, fine setae; antenna with scape and pedicel thicker than flagellomeres; flagellum with 12 flagellomeres; flagellomeres without annulations, each flagellomere with 24 long apical setae, scape longer than flagellomere I, length
<quantity box="[1342,1447,1553,1580]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="1" pageNumber="488" unit="mm" value="0.07">0.07 mm</quantity>
; pedicel short, 0.5× length of flagellomere I, flagellomeres IX lengths each
<quantity box="[1014,1119,1626,1653]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="488" unit="mm" value="0.05">0.05 mm</quantity>
, flagellomere XI length
<quantity metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="1" pageNumber="488" unit="mm" value="0.07">0.07 mm</quantity>
, flagellomere XII length
<quantity box="[1144,1248,1663,1690]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="488" unit="mm" value="0.08">0.08 mm</quantity>
. Maxillary palpus with four palpomeres; palpomere lengths (P
<subScript attach="left" box="[1314,1322,1718,1732]" fontSize="6" pageId="1" pageNumber="488">1</subScript>
, P
<subScript attach="left" box="[1354,1362,1718,1732]" fontSize="6" pageId="1" pageNumber="488">2</subScript>
, P
<subScript attach="left" box="[1394,1402,1718,1732]" fontSize="6" pageId="1" pageNumber="488">3</subScript>
, P
<subScript attach="left" box="[1434,1442,1718,1732]" fontSize="6" pageId="1" pageNumber="488">4</subScript>
) 0.05, 0.04, 0.02,
<quantity box="[997,1099,1736,1763]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="488" unit="mm" value="0.12">0.12 mm</quantity>
, respectively; apical palpomere (P
<subScript attach="left" box="[835,843,1791,1805]" fontSize="6" pageId="1" pageNumber="488">4</subScript>
) distinctly enlarged relative to preceding palpomeres (
<figureCitation box="[818,944,1810,1836]" captionStart="FIGURE 3" captionStartId="3.[136,235,1509,1533]" captionTargetBox="[242,1346,193,1485]" captionTargetId="figure-199@3.[242,1346,193,1485]" captionTargetPageId="3" captionText="FIGURE 3. Photographic details of holotype female of Belaphotroctes grimaldii sp. nov. (NIGP200508); scale bars = 0.25 mm. A, Head and pronotum, dorsal view. B, Head and pronotum, ventral view (P4 = maxillary palpomere IV). C, Apical abdominal sterna." figureDoi="http://doi.org/10.5281/zenodo.7333609" httpUri="https://zenodo.org/record/7333609/files/figure.png" pageId="1" pageNumber="488">Fig. 3A, B</figureCitation>
), width nearly 2× width of P
<subScript attach="left" box="[1278,1286,1828,1842]" fontSize="6" pageId="1" pageNumber="488">3</subScript>
, without stout conical sensilla but with some minute, fine sensilla apically and longer setae laterally, such setae not longer than palpomere breadth. Labial palpus with two short palpomeres, apicalmost palpomere densely covered with fine minute setae.
</paragraph>
<paragraph blockId="1.[810,1452,820,2092]" lastBlockId="2.[810,1452,1698,2084]" lastPageId="2" lastPageNumber="489" pageId="1" pageNumber="488">
Pronotum divided into three distinct lobes; medial lobe with distinct mediolongitudinal sulcus; lateral lobes narrower than medial lobe, lateral lobe with pronounced, proclined seta (
<figureCitation box="[302,387,1734,1760]" captionStart="FIGURE 3" captionStartId="3.[136,235,1509,1533]" captionTargetBox="[242,1346,193,1485]" captionTargetId="figure-199@3.[242,1346,193,1485]" captionTargetPageId="3" captionText="FIGURE 3. Photographic details of holotype female of Belaphotroctes grimaldii sp. nov. (NIGP200508); scale bars = 0.25 mm. A, Head and pronotum, dorsal view. B, Head and pronotum, ventral view (P4 = maxillary palpomere IV). C, Apical abdominal sterna." figureDoi="http://doi.org/10.5281/zenodo.7333609" httpUri="https://zenodo.org/record/7333609/files/figure.png" pageId="2" pageNumber="489">Fig. 3A</figureCitation>
) [much of finer details of pronotum obscured by fracture:
<figureCitation box="[383,453,1770,1796]" captionStart="FIGURE 2" captionStartId="2.[134,233,1584,1608]" captionTargetBox="[277,1305,189,1560]" captionTargetId="figure-187@2.[277,1305,189,1561]" captionTargetPageId="2" captionText="FIGURE 2. Habitus photographs of holotype female of Belaphotroctes grimaldii sp. nov. (NIGP200508); scale bar (for both images) = 1 mm. A, Dorsal habitus. B, Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.7333607" httpUri="https://zenodo.org/record/7333607/files/figure.png" pageId="2" pageNumber="489">Fig. 2</figureCitation>
]. Thoracic sterna broad, flat; mesosternum without sclerotized bands. Coxae broadly separated by sterna; femora slightly swollen except metafemur greatly swollen; tibiae slender, tibial spur formula apparently 1-1-1; all tarsi trimerous; pretarsal claw apparently simple, pulvillus lacking (pretarsal vesicle absent). Profemur without prominent setae ventrally, with scattered minute, fine setae; protibia with numerous fine, suberect setae along length; mesofemur length
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, mesotibia length
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, setation as on profemur and protibia; metafemur length
<quantity box="[1340,1447,1734,1760]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="489" unit="mm" value="0.31">0.31 mm</quantity>
, lacking tubercle, setation as on profemur; metatibia length
<quantity box="[810,912,1806,1832]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="489" unit="mm" value="0.27">0.27 mm</quantity>
, maximum width
<quantity box="[1115,1231,1806,1832]" metricMagnitude="-5" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="489" unit="mm" value="0.024">0.024 mm</quantity>
, with setation as on protibia except intermixed with fine, elongate, erect setae dorsally.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7333607" ID-Zenodo-Dep="7333607" httpUri="https://zenodo.org/record/7333607/files/figure.png" pageId="2" pageNumber="489" startId="2.[134,233,1584,1608]" targetBox="[277,1305,189,1560]" targetPageId="2">
<paragraph blockId="2.[134,1449,1584,1644]" pageId="2" pageNumber="489">
<emphasis bold="true" box="[134,260,1584,1608]" pageId="2" pageNumber="489">FIGURE 2.</emphasis>
Habitus photographs of holotype female of
<taxonomicName authority="ENGEL &amp; WANG, 2022" authorityName="ENGEL &amp; WANG" authorityYear="2022" box="[729,984,1584,1608]" class="Insecta" family="Liposcelididae" genus="Belaphotroctes" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="489" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
<emphasis box="[729,984,1584,1608]" italics="true" pageId="2" pageNumber="489">Belaphotroctes grimaldii</emphasis>
</taxonomicName>
<emphasis bold="true" box="[994,1077,1584,1608]" pageId="2" pageNumber="489">
<taxonomicNameLabel box="[994,1077,1584,1608]" pageId="2" pageNumber="489" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(NIGP200508); scale bar (for both images) = 1 mm.
<emphasis bold="true" box="[312,330,1620,1644]" pageId="2" pageNumber="489">A</emphasis>
, Dorsal habitus.
<emphasis bold="true" box="[502,519,1620,1644]" pageId="2" pageNumber="489">B</emphasis>
, Ventral habitus.
</paragraph>
</caption>
<paragraph blockId="2.[810,1452,1698,2084]" lastBlockId="3.[810,1452,1665,2056]" lastPageId="3" lastPageNumber="490" pageId="2" pageNumber="489">
Wings hyaline, diaphanous, membrane finely and irregularly wrinkled (
<figureCitation box="[1060,1134,1950,1976]" captionStart="FIGURE 2" captionStartId="2.[134,233,1584,1608]" captionTargetBox="[277,1305,189,1560]" captionTargetId="figure-187@2.[277,1305,189,1561]" captionTargetPageId="2" captionText="FIGURE 2. Habitus photographs of holotype female of Belaphotroctes grimaldii sp. nov. (NIGP200508); scale bar (for both images) = 1 mm. A, Dorsal habitus. B, Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.7333607" httpUri="https://zenodo.org/record/7333607/files/figure.png" pageId="2" pageNumber="489">Fig. 2</figureCitation>
), broadly rounded apically (
<figureCitation box="[818,891,1986,2012]" captionStart="FIGURE 2" captionStartId="2.[134,233,1584,1608]" captionTargetBox="[277,1305,189,1560]" captionTargetId="figure-187@2.[277,1305,189,1561]" captionTargetPageId="2" captionText="FIGURE 2. Habitus photographs of holotype female of Belaphotroctes grimaldii sp. nov. (NIGP200508); scale bar (for both images) = 1 mm. A, Dorsal habitus. B, Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.7333607" httpUri="https://zenodo.org/record/7333607/files/figure.png" pageId="2" pageNumber="489">Figs 2</figureCitation>
,
<figureCitation box="[903,919,1986,2012]" captionStart="FIGURE 4" captionStartId="4.[136,231,757,781]" captionTargetBox="[348,1204,231,710]" captionTargetId="figure-519@4.[339,1249,217,734]" captionTargetPageId="4" captionText="FIGURE 4. Wing venation of Belaphotroctes grimaldii sp. nov. (NIGP200508), forewing above, hind wing below. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7333611" httpUri="https://zenodo.org/record/7333611/files/figure.png" pageId="2" pageNumber="489">4</figureCitation>
), with veins indicated as light brown, nebulous traces (
<figureCitation box="[907,992,2022,2048]" captionStart="FIGURE 2" captionStartId="2.[134,233,1584,1608]" captionTargetBox="[277,1305,189,1560]" captionTargetId="figure-187@2.[277,1305,189,1561]" captionTargetPageId="2" captionText="FIGURE 2. Habitus photographs of holotype female of Belaphotroctes grimaldii sp. nov. (NIGP200508); scale bar (for both images) = 1 mm. A, Dorsal habitus. B, Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.7333607" httpUri="https://zenodo.org/record/7333607/files/figure.png" pageId="2" pageNumber="489">Fig. 2</figureCitation>
); forewing long, extending beyond abdominal apex, length
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, maximum width
<quantity lastPageId="3" lastPageNumber="490" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="489" unit="mm" value="0.43">0.43 mm</quantity>
; hind wing length
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, maximum width
<quantity metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="490" unit="mm" value="0.23">0.23 mm</quantity>
. Forewing (
<figureCitation box="[313,379,1702,1728]" captionStart="FIGURE 4" captionStartId="4.[136,231,757,781]" captionTargetBox="[348,1204,231,710]" captionTargetId="figure-519@4.[339,1249,217,734]" captionTargetPageId="4" captionText="FIGURE 4. Wing venation of Belaphotroctes grimaldii sp. nov. (NIGP200508), forewing above, hind wing below. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7333611" httpUri="https://zenodo.org/record/7333611/files/figure.png" pageId="3" pageNumber="490">Fig. 4</figureCitation>
) with trace of Sc scarcely evident in basal quarter of wing length; R extending nearly to wing midlength, with distinct setae along length, R1 evident diverging orthogonal to stem of R, with a single seta at juncture with faint trace of C, then extending apically for short distance before merging into anterior wing margin (
<figureCitation box="[144,221,1923,1949]" captionStart="FIGURE 4" captionStartId="4.[136,231,757,781]" captionTargetBox="[348,1204,231,710]" captionTargetId="figure-519@4.[339,1249,217,734]" captionTargetPageId="4" captionText="FIGURE 4. Wing venation of Belaphotroctes grimaldii sp. nov. (NIGP200508), forewing above, hind wing below. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7333611" httpUri="https://zenodo.org/record/7333611/files/figure.png" pageId="3" pageNumber="490">Fig. 4</figureCitation>
); Rs originating just prior to wing midlength, extending nearly to wing apex, weakly curved apically; M present and largely straight, extending nearly to wing apex; traces of Cu and A absent. Hind wing (
<figureCitation box="[644,712,2033,2060]" captionStart="FIGURE 4" captionStartId="4.[136,231,757,781]" captionTargetBox="[348,1204,231,710]" captionTargetId="figure-519@4.[339,1249,217,734]" captionTargetPageId="4" captionText="FIGURE 4. Wing venation of Belaphotroctes grimaldii sp. nov. (NIGP200508), forewing above, hind wing below. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7333611" httpUri="https://zenodo.org/record/7333611/files/figure.png" pageId="3" pageNumber="490">Fig. 4</figureCitation>
) with only trace R; R nearly extending to wing apex, slightly curved, paralleling curve in anterior wing margin.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7333609" ID-Zenodo-Dep="7333609" httpUri="https://zenodo.org/record/7333609/files/figure.png" pageId="3" pageNumber="490" startId="3.[136,235,1509,1533]" targetBox="[242,1346,193,1485]" targetPageId="3">
<paragraph blockId="3.[136,1452,1509,1605]" box="[136,1452,1509,1533]" pageId="3" pageNumber="490">
<emphasis bold="true" box="[136,260,1509,1533]" pageId="3" pageNumber="490">FIGURE 3.</emphasis>
Photographic details of holotype female of
<taxonomicName authority="ENGEL &amp; WANG, 2022" authorityName="ENGEL &amp; WANG" authorityYear="2022" box="[711,964,1509,1533]" class="Insecta" family="Liposcelididae" genus="Belaphotroctes" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="490" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
<emphasis box="[711,964,1509,1533]" italics="true" pageId="3" pageNumber="490">Belaphotroctes grimaldii</emphasis>
</taxonomicName>
<emphasis bold="true" box="[971,1053,1509,1533]" pageId="3" pageNumber="490">
<taxonomicNameLabel box="[971,1053,1509,1533]" pageId="3" pageNumber="490" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(NIGP200508); scale bars = 0.25 mm.
</paragraph>
<paragraph blockId="3.[136,1452,1509,1605]" pageId="3" pageNumber="490">
<emphasis bold="true" box="[136,154,1545,1569]" pageId="3" pageNumber="490">A</emphasis>
, Head and pronotum, dorsal view.
<emphasis bold="true" box="[520,537,1545,1569]" pageId="3" pageNumber="490">B</emphasis>
, Head and pronotum, ventral view (P
<subScript attach="left" box="[929,936,1561,1575]" fontSize="6" pageId="3" pageNumber="490">4</subScript>
= maxillary palpomere IV).
<emphasis bold="true" box="[1237,1255,1545,1569]" pageId="3" pageNumber="490">C</emphasis>
, Apical abdominal sterna.
</paragraph>
</caption>
<paragraph blockId="3.[810,1452,1665,2056]" pageId="3" pageNumber="490">
Abdomen length
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, maximum width
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; terga and sterna well-developed; preclunial terga without sclerotized bands; terga IVI apparently without setation except minute, suberect, fine, setae laterally; remaining terga with similar laterally setae as well as such setae sparsely scattered on dorsum, intermixed with a few long setae laterally and apically, longest marginal seta
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.
</paragraph>
<paragraph blockId="3.[810,1452,1665,2056]" box="[863,963,2030,2056]" pageId="3" pageNumber="490">
<specimenCount box="[863,887,2030,2056]" pageId="3" pageNumber="490" type="male"></specimenCount>
:
<emphasis box="[899,957,2030,2055]" italics="true" pageId="3" pageNumber="490">Latet</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>