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<document id="430952373FCAA8AEC90F0ED5020498F4" ID-DOI="10.11646/zootaxa.3838.1.1" ID-GBIF-Dataset="e5103854-af5b-4e33-958b-163592f0b388" ID-ISSN="1175-5326" ID-Zenodo-Dep="286727" ID-ZooBank="1534C775-D28D-470F-9AEC-8BABB3D8FA56" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460458262828" checkinUser="plazi" docAuthor="Albuquerque, Felícia Pereira De, Manfio, Daiara &amp; Ribeiro-Costa, Cibele Stramare" docDate="2014" docId="03FF87F5FFC8FFDF38ADFF75FD0E76D7" docLanguage="en" docName="zt03838p045.pdf" docOrigin="Zootaxa 3838 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ctenocolum triangulatus Albuquerque &amp; Ribeiro-Costa, sp. nov." docType="treatment" docVersion="8" lastPageNumber="34" masterDocId="FFC6FF8DFFE9FFFE383AFFE2FFB17008" masterDocTitle="A contribution to the knowledge of New World Bruchinae (Coleoptera, Chrysomelidae): taxonomic revision of Ctenocolum Kingsolver &amp; Whitehead, with description of five new species" masterLastPageNumber="45" masterPageNumber="1" pageNumber="34" updateTime="1698302949382" updateUser="plazi">
<mods:mods id="E360D6457E8F428A85C183D184E1C4F0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="897E3AA770399543D73F90C690C088D7">
<mods:title id="25176E63A27531260FC532893A395DFF">A contribution to the knowledge of New World Bruchinae (Coleoptera, Chrysomelidae): taxonomic revision of Ctenocolum Kingsolver &amp; Whitehead, with description of five new species</mods:title>
</mods:titleInfo>
<mods:name id="63B9BF8B5A7005C86474721315480019" type="personal">
<mods:role id="4136D4407D02A7FD799BFFBDD2D9BE17">
<mods:roleTerm id="1CCFFE414D9482EDC03A3B8849ACA435">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0F72F737C8B807E08CC60BFDD40BEAE5">Albuquerque, Felícia Pereira De</mods:namePart>
</mods:name>
<mods:name id="B0B2CC6D52A002A9145FE147D3360CA2" type="personal">
<mods:role id="D13E40B08CD452D32AFD8686B87062EC">
<mods:roleTerm id="F309337769D36BEEB240EF7D2B1630ED">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="706AAC9F0096299B25E9051EC22B908D">Manfio, Daiara</mods:namePart>
</mods:name>
<mods:name id="54D8044A3507B64898308F9B4F44CCF3" type="personal">
<mods:role id="E3193EC205F47DA8BD72C0EFE7BD53B7">
<mods:roleTerm id="A5FD1C593407665C5BF28F15B09B8C44">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B2038DB000CAD86691A40162F0979335">Ribeiro-Costa, Cibele Stramare</mods:namePart>
</mods:name>
<mods:typeOfResource id="6F7F34E8F7D16847E745D0A2131E0DA2">text</mods:typeOfResource>
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<mods:titleInfo id="C3B35598F1D188EBB38AD10055780FAF">
<mods:title id="A2577CDFE9E7C939CD4EC07CBFD2AA22">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A1394337C4100F157175284354AE57B5">
<mods:date id="CF3D7060EC326B69ECAFCF582AA78C5E">2014</mods:date>
<mods:detail id="45C483D83752358D49DB309C61092399" type="volume">
<mods:number id="BC2F8591117F60DD155DAB1818B2E8A9">3838</mods:number>
</mods:detail>
<mods:detail id="0DBCD1253C2D870D248D3BAFF92B37EA" type="issue">
<mods:number id="0C7F6658E47ECB5D4FB0E619D61DBD62">1</mods:number>
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<mods:start id="C4A052ACAA4D09CA829EE0882D23502B">1</mods:start>
<mods:end id="52B3346E3F4A029DB6C52355E51C4EF5">45</mods:end>
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<mods:classification id="6CBD7FF34CCAB126FAAA71631C9150AF">journal article</mods:classification>
<mods:identifier id="B64F809D2CE83BCD0FC2177E6CBD98E0" type="DOI">10.11646/zootaxa.3838.1.1</mods:identifier>
<mods:identifier id="3DA0562CC963230ACC6D057565F58B1A" type="GBIF-Dataset">e5103854-af5b-4e33-958b-163592f0b388</mods:identifier>
<mods:identifier id="F39E3E6FA9B13366B6409DCE1969AAC2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="94EB83DB2950E2791FA9BD5D48825B2E" type="Zenodo-Dep">286727</mods:identifier>
<mods:identifier id="C9FEC6121AED0F3496B9AEF5DF303CA7" type="ZooBank">1534C775-D28D-470F-9AEC-8BABB3D8FA56</mods:identifier>
</mods:mods>
<treatment id="03FF87F5FFC8FFDF38ADFF75FD0E76D7" ID-DOI="http://doi.org/10.5281/zenodo.6124249" ID-GBIF-Taxon="127668880" ID-Zenodo-Dep="6124249" LSID="urn:lsid:plazi:treatment:03FF87F5FFC8FFDF38ADFF75FD0E76D7" httpUri="http://treatment.plazi.org/id/03FF87F5FFC8FFDF38ADFF75FD0E76D7" lastPageNumber="34" pageId="33" pageNumber="34">
<subSubSection id="C34C6568FFC8FFDF38ADFF75FE4E70DB" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="8BE936E3FFC8FFDF38ADFF75FC1F70BA" blockId="33.[151,942,151,211]" box="[151,942,151,178]" pageId="33" pageNumber="34">
<heading id="D0A1818FFFC8FFDF38ADFF75FC1F70BA" bold="true" box="[151,942,151,178]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
<emphasis id="B922EAF1FFC8FFDF38ADFF75FC1F70BA" bold="true" box="[151,942,151,178]" pageId="33" pageNumber="34">
<taxonomicName id="4C564D60FFC8FFDF38ADFF75FCF970BA" authority="Albuquerque &amp; Ribeiro-Costa" authorityName="Albuquerque &amp; Ribeiro-Costa" box="[151,840,151,178]" class="Insecta" family="Chrysomelidae" genus="Ctenocolum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="triangulatus" status="sp. nov.">
<emphasis id="B922EAF1FFC8FFDF38ADFF75FE7770B9" bold="true" box="[151,454,151,177]" italics="true" pageId="33" pageNumber="34">Ctenocolum triangulatus</emphasis>
Albuquerque &amp; Ribeiro-Costa
</taxonomicName>
<taxonomicNameLabel id="A211578AFFC8FFDF3B6AFF7AFC1F70BA" box="[848,942,152,178]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE936E3FFC8FFDF38ADFF58FE4E70DB" blockId="33.[151,942,151,211]" box="[151,511,186,211]" pageId="33" pageNumber="34">
(
<figureCitation id="136D2A66FFC8FFDF38A5FF58FF4D70DB" box="[159,252,186,211]" captionStart="FIGURES 17 – 21" captionStartId="5.[151,262,1469,1492]" captionTargetBox="[184,1433,256,1431]" captionTargetId="figure@5.[151,1436,193,1448]" captionTargetPageId="5" captionText="FIGURES 17 – 21. Dorsal habitus: 17 Ctenocolum triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 18 C. acapulcensis; 19 C. janzeni; 20 C. salvini (lectotype); 21 C. tuberculatum. Scale: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3698643" httpUri="https://zenodo.org/record/3698643/files/figure.png" pageId="33" pageNumber="34">Figs. 17</figureCitation>
,
<figureCitation id="136D2A66FFC8FFDF3933FF59FE9670DB" box="[265,295,187,211]" captionStart="FIGURES 22 – 30" captionStartId="6.[151,263,1916,1939]" captionTargetBox="[214,1372,193,1894]" captionTargetId="figure@6.[214,1372,193,1896]" captionTargetPageId="6" captionText="FIGURES 22 – 30. Frontal habitus: 22 Ctenocolum aquilus Albuquerque &amp; Ribeiro-Costa sp. nov. (paratype); 23 C. biolleyi (paratype); 24 C. colburni; 25 C. martiale; 26 C. milelo Albuquerque &amp; Ribeiro-Costa sp. nov. (paratype); 27 C. podagricus; 28 C. punctinotatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 29 C. pygospilotos Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 30 C. triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype). Scale: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3698645" httpUri="https://zenodo.org/record/3698645/files/figure.png" pageId="33" pageNumber="34">30</figureCitation>
,
<figureCitation id="136D2A66FFC8FFDF3909FF59FEE070DB" box="[307,337,187,211]" captionStart="FIGURES 35 – 43" captionStartId="8.[151,263,1916,1939]" captionTargetBox="[214,1372,193,1894]" captionTargetId="figure@8.[214,1372,193,1896]" captionTargetPageId="8" captionText="FIGURES 35 – 43. Pygidium of male: 35 Ctenocolum aquilus Albuquerque &amp; Ribeiro-Costa sp. nov. (paratype); 36 C. colburni; 37 C. martiale; 38 C. milelo Albuquerque &amp; Ribeiro-Costa sp. nov. (paratype); 39 C. podagricus; 40 C. punctinotatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 41 C. pygospilotos Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 42 C. triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 43 C. acapulcensis. Scale: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3698649" httpUri="https://zenodo.org/record/3698649/files/figure.png" pageId="33" pageNumber="34">42</figureCitation>
,
<figureCitation id="136D2A66FFC8FFDF3967FF58FECA70DB" box="[349,379,186,211]" captionStart="FIGURES 61 – 65" captionStartId="12.[151,263,1650,1673]" captionTargetBox="[156,1426,261,1604]" captionTargetId="figure@12.[151,1436,193,1629]" captionTargetPageId="12" captionText="FIGURES 61 – 65. Lateral habitus: 61 Ctenocolum triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 62 C. acapulcensis; 63 C. janzeni; 64 C. salvini (lectotype); 65 C. tuberculatum. Scale: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3698657" httpUri="https://zenodo.org/record/3698657/files/figure.png" pageId="33" pageNumber="34">61</figureCitation>
,
<figureCitation id="136D2A66FFC8FFDF39BCFF59FE1570DB" box="[390,420,187,211]" captionStart="FIGURES 72 – 76" captionStartId="14.[151,263,1952,1975]" captionTargetBox="[216,1383,211,1917]" captionTargetId="figure@14.[200,1387,193,1931]" captionTargetPageId="14" captionText="FIGURES 72 – 76. Posterior leg, external view: 72 Ctenocolum punctinotatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 73 C. pygospilotos Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 74 C. triangulatus Albuquerque &amp; Ribeiro- Costa sp. nov. (holotype); 75 C. acapulcensis; 76 C. janzeni. Scale: 0.50 mm." httpUri="https://zenodo.org/record/286739/files/figure.png" pageId="33" pageNumber="34">74</figureCitation>
,
<figureCitation id="136D2A66FFC8FFDF398AFF58FE7F70DB" box="[432,462,186,211]" captionStart="FIGURES 85 – 90" captionStartId="18.[151,263,1953,1976]" captionTargetBox="[202,1384,193,1930]" captionTargetId="figure@18.[202,1385,193,1932]" captionTargetPageId="18" captionText="FIGURES 85 – 90. Male genitalia, median lobe: 85 Ctenocolum pygospilotos Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 86 C. triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 87 C. acapulcensis; 88 C. janzeni; 89 C. salvini (paralectotype); 90 C. tuberculatum. Scale: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.3698661" httpUri="https://zenodo.org/record/3698661/files/figure.png" pageId="33" pageNumber="34">86</figureCitation>
,
<figureCitation id="136D2A66FFC8FFDF39E0FF59FE4670DB" box="[474,503,187,211]" captionStart="FIGURES 91 – 99" captionStartId="19.[151,263,1916,1939]" captionTargetBox="[214,1372,193,1894]" captionTargetId="figure@19.[214,1372,193,1896]" captionTargetPageId="19" captionText="FIGURES 91 – 99. Male genitalia, tegmen, lateral lobe: 91 C. aquilus (paratype); 92 C. colburni; 93 C. martiale (holotype); 94 C. milelo Albuquerque &amp; Ribeiro-Costa sp. nov. (paratype); 95 C. podagricus; 96 C. punctinotatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 97 C. pygospilotos Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 98 C. triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 99 C. acapulcensis. Scale: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.3698663" httpUri="https://zenodo.org/record/3698663/files/figure.png" pageId="33" pageNumber="34">98</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34C6568FFC8FFDF38ADFEE0FADA71A3" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="8BE936E3FFC8FFDF38ADFEE0FADA71A3" blockId="33.[151,1437,258,1759]" pageId="33" pageNumber="34">
<emphasis id="B922EAF1FFC8FFDF38ADFEE0FEF57113" bold="true" box="[151,324,258,283]" pageId="33" pageNumber="34">
<typeStatus id="54ED8841FFC8FFDF38ADFEE0FF637113" box="[151,210,258,283]" pageId="33" pageNumber="34">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54ED8841FFC8FFDF3968FEE0FE0B7113" box="[338,442,258,283]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
deposited in
<collectionCode id="ED47AE26FFC8FFDF3A6AFEE1FD157112" box="[592,676,259,282]" pageId="33" pageNumber="34">ZMHB</collectionCode>
, male: “53758” [white label printed in black]; “misotur/ m.” [white label handwritten in black]; “Hist. -Coll. (
<taxonomicName id="4C564D60FFC8FFDF3AAAFEC4FCBE7137" box="[656,783,294,319]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
)/ Nr. 53758/
<taxonomicName id="4C564D60FFC8FFDF3B8CFEC4FBA37137" box="[950,1042,294,319]" class="Insecta" family="Chrysomelidae" genus="Bruchus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="43" phylum="Arthropoda" rank="genus">Bruchus</taxonomicName>
/ aequinoctialis Erichs./ Columb., Moritz./ Zool. Mus. Berlin” [green label with black margin, printed in black]; “
<taxonomicName id="4C564D60FFC8FFDF3C4BFEA8FB4D716B" box="[1137,1276,330,355]" class="Insecta" family="Chrysomelidae" genus="Ctenocolum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="44" phylum="Arthropoda" rank="genus">Ctenocolum</taxonomicName>
/ SP/ det./ J. M.Kingsolver” [white label handwritten, printed in black]; “♂” [white label printed in black];]; “
<typeStatus id="54ED8841FFC8FFDF3CC7FE8DFA25718F" box="[1277,1428,367,391]" pageId="33" pageNumber="34" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C564D60FFC8FFDF38ADFE71FE1E71A3" box="[151,431,403,427]" class="Insecta" family="Chrysomelidae" genus="Ctenocolum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="triangulatus">
<emphasis id="B922EAF1FFC8FFDF38ADFE71FE1E71A3" box="[151,431,403,427]" italics="true" pageId="33" pageNumber="34">Ctenocolum triangulatus</emphasis>
</taxonomicName>
/ Det. Albuquerque &amp; Ribeiro-Costa” [white label with red margin, printed in black].
</paragraph>
</subSubSection>
<subSubSection id="C34C6568FFC8FFDF38FDFE54FA237233" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="8BE936E3FFC8FFDF38FDFE54FA237233" blockId="33.[151,1437,258,1759]" pageId="33" pageNumber="34">
<emphasis id="B922EAF1FFC8FFDF38FDFE54FEF071C7" bold="true" box="[199,321,438,463]" pageId="33" pageNumber="34">Diagnosis.</emphasis>
<taxonomicName id="4C564D60FFC8FFDF3971FE54FC6071C7" authority="Albuquerque &amp; Ribeiro-Costa" authorityName="Albuquerque &amp; Ribeiro-Costa" box="[331,977,438,463]" class="Insecta" family="Chrysomelidae" genus="Ctenocolum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="triangulatus" status="sp. nov.">
<emphasis id="B922EAF1FFC8FFDF3971FE54FDD971C6" box="[331,616,438,462]" italics="true" pageId="33" pageNumber="34">Ctenocolum triangulatus</emphasis>
Albuquerque &amp; Ribeiro-Costa
</taxonomicName>
<emphasis id="B922EAF1FFC8FFDF3BE6FE54FB8B71C7" bold="true" box="[988,1082,438,463]" pageId="33" pageNumber="34">
<taxonomicNameLabel id="A211578AFFC8FFDF3BE6FE54FB8B71C7" box="[988,1082,438,463]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is different from
<taxonomicName id="4C564D60FFC8FFDF3D1CFE55FE5F71FB" authority="Albuquerque &amp; Ribeiro-Costa" authorityName="Albuquerque &amp; Ribeiro-Costa" class="Insecta" family="Chrysomelidae" genus="Ctenocolum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="milelo" status="sp. nov.">
<emphasis id="B922EAF1FFC8FFDF3D1CFE55FA2D71C6" box="[1318,1436,438,462]" italics="true" pageId="33" pageNumber="34">C. milelo</emphasis>
Albuquerque &amp; Ribeiro-Costa
</taxonomicName>
<emphasis id="B922EAF1FFC8FFDF39C2FE38FDE271FB" bold="true" box="[504,595,474,499]" pageId="33" pageNumber="34">
<taxonomicNameLabel id="A211578AFFC8FFDF39C2FE38FDE271FB" box="[504,595,474,499]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C564D60FFC8FFDF3AB6FE39FC9C71FB" box="[652,813,475,499]" class="Insecta" family="Chrysomelidae" genus="Ctenocolum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="podagricus">
<emphasis id="B922EAF1FFC8FFDF3AB6FE39FC9C71FB" box="[652,813,475,499]" italics="true" pageId="33" pageNumber="34">C. podagricus</emphasis>
</taxonomicName>
by having ocular index 4.5 and pygidium oval in both sex; differs from
<taxonomicName id="4C564D60FFC8FFDF3966FE1DFE6C721E" box="[348,477,510,534]" class="Insecta" family="Chrysomelidae" genus="Ctenocolum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="martiale">
<emphasis id="B922EAF1FFC8FFDF3966FE1DFE6C721E" box="[348,477,510,534]" italics="true" pageId="33" pageNumber="34">C. martiale</emphasis>
</taxonomicName>
by elytral striae 3 and 4 with less conspicuous teeth at base. This species differs from all others by the male genitalia at submedian region with squamous subtriangular-shape sclerite (
<figureCitation id="136D2A66FFC8FFDF3D0BFDC0FA347233" box="[1329,1413,546,571]" captionStart="FIGURES 85 – 90" captionStartId="18.[151,263,1953,1976]" captionTargetBox="[202,1384,193,1930]" captionTargetId="figure@18.[202,1385,193,1932]" captionTargetPageId="18" captionText="FIGURES 85 – 90. Male genitalia, median lobe: 85 Ctenocolum pygospilotos Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 86 C. triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 87 C. acapulcensis; 88 C. janzeni; 89 C. salvini (paralectotype); 90 C. tuberculatum. Scale: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.3698661" httpUri="https://zenodo.org/record/3698661/files/figure.png" pageId="33" pageNumber="34">Fig. 86</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34C6568FFC8FFDF38FDFDA4FDAB767B" pageId="33" pageNumber="34" type="description">
<paragraph id="8BE936E3FFC8FFDF38FDFDA4FD3C7256" blockId="33.[151,1437,258,1759]" box="[199,653,582,607]" pageId="33" pageNumber="34">
<emphasis id="B922EAF1FFC8FFDF38FDFDA4FEE97257" bold="true" box="[199,344,582,607]" pageId="33" pageNumber="34">Description.</emphasis>
BL:
<quantity id="4CAE9B06FFC8FFDF39ABFDA5FE5B7256" box="[401,490,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="33" pageNumber="34" unit="mm" value="2.4">2.4 mm</quantity>
; BW:
<quantity id="4CAE9B06FFC8FFDF3A0AFDA4FD387256" box="[560,649,582,607]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="33" pageNumber="34" unit="mm" value="1.5">1.5 mm</quantity>
.
</paragraph>
<paragraph id="8BE936E3FFC8FFDF38FDFD89FCFD72C3" blockId="33.[151,1437,258,1759]" pageId="33" pageNumber="34">
Integument. Dorsum reddish brown and dark brown. Antenna dark brown, first 3 antennomeres paler (
<figureCitation id="136D2A66FFC8FFDF3D55FD88FF0772AF" captionStart="FIGURES 61 – 65" captionStartId="12.[151,263,1650,1673]" captionTargetBox="[156,1426,261,1604]" captionTargetId="figure@12.[151,1436,193,1629]" captionTargetPageId="12" captionText="FIGURES 61 – 65. Lateral habitus: 61 Ctenocolum triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 62 C. acapulcensis; 63 C. janzeni; 64 C. salvini (lectotype); 65 C. tuberculatum. Scale: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3698657" httpUri="https://zenodo.org/record/3698657/files/figure.png" pageId="33" pageNumber="34">Fig. 61</figureCitation>
). Pygidium reddish brown and dark brown.Ventral region reddish brown and black. Front and middle femur and tibia brown; hind femur brown, reddish brown to dark brown.
</paragraph>
<paragraph id="8BE936E3FFC8FFDF38FDFD34FC047377" blockId="33.[151,1437,258,1759]" pageId="33" pageNumber="34">
Pubescence. Pronotum reddish brown and white; sparse setae exposing the integument forming an oval, wide area from anterior to posterior region and on each lateral region one small area (
<figureCitation id="136D2A66FFC8FFDF3C45FD18FB6B731B" box="[1151,1242,762,787]" captionStart="FIGURES 17 – 21" captionStartId="5.[151,262,1469,1492]" captionTargetBox="[184,1433,256,1431]" captionTargetId="figure@5.[151,1436,193,1448]" captionTargetPageId="5" captionText="FIGURES 17 – 21. Dorsal habitus: 17 Ctenocolum triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 18 C. acapulcensis; 19 C. janzeni; 20 C. salvini (lectotype); 21 C. tuberculatum. Scale: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3698643" httpUri="https://zenodo.org/record/3698643/files/figure.png" pageId="33" pageNumber="34">Fig. 17</figureCitation>
). Elytra slightly variegated, reddish brown and black setae; interstria 3 without white setae at base and at submedian region with short sparse strip of white setae (
<figureCitation id="136D2A66FFC8FFDF3A3FFCA0FDE67353" box="[517,599,834,859]" captionStart="FIGURES 17 – 21" captionStartId="5.[151,262,1469,1492]" captionTargetBox="[184,1433,256,1431]" captionTargetId="figure@5.[151,1436,193,1448]" captionTargetPageId="5" captionText="FIGURES 17 – 21. Dorsal habitus: 17 Ctenocolum triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 18 C. acapulcensis; 19 C. janzeni; 20 C. salvini (lectotype); 21 C. tuberculatum. Scale: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3698643" httpUri="https://zenodo.org/record/3698643/files/figure.png" pageId="33" pageNumber="34">Fig. 17</figureCitation>
). Pygidium dense, white and yellowish gray in a spotted pattern (
<figureCitation id="136D2A66FFC8FFDF3D02FCA0FA217353" box="[1336,1424,834,859]" captionStart="FIGURES 35 – 43" captionStartId="8.[151,263,1916,1939]" captionTargetBox="[214,1372,193,1894]" captionTargetId="figure@8.[214,1372,193,1896]" captionTargetPageId="8" captionText="FIGURES 35 – 43. Pygidium of male: 35 Ctenocolum aquilus Albuquerque &amp; Ribeiro-Costa sp. nov. (paratype); 36 C. colburni; 37 C. martiale; 38 C. milelo Albuquerque &amp; Ribeiro-Costa sp. nov. (paratype); 39 C. podagricus; 40 C. punctinotatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 41 C. pygospilotos Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 42 C. triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 43 C. acapulcensis. Scale: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3698649" httpUri="https://zenodo.org/record/3698649/files/figure.png" pageId="33" pageNumber="34">Fig. 42</figureCitation>
). Ventral region with setae yellowish gray, brown and white (
<figureCitation id="136D2A66FFC8FFDF3B6EFC84FC197377" box="[852,936,870,895]" captionStart="FIGURES 61 – 65" captionStartId="12.[151,263,1650,1673]" captionTargetBox="[156,1426,261,1604]" captionTargetId="figure@12.[151,1436,193,1629]" captionTargetPageId="12" captionText="FIGURES 61 – 65. Lateral habitus: 61 Ctenocolum triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 62 C. acapulcensis; 63 C. janzeni; 64 C. salvini (lectotype); 65 C. tuberculatum. Scale: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3698657" httpUri="https://zenodo.org/record/3698657/files/figure.png" pageId="33" pageNumber="34">Fig. 61</figureCitation>
).
</paragraph>
<paragraph id="8BE936E3FFC8FFDF38FDFC68FBC773CF" blockId="33.[151,1437,258,1759]" pageId="33" pageNumber="34">
Head. Ocular sinus
<quantity id="4CAE9B06FFC8FFDF399EFC69FE4D73AA" box="[420,508,907,931]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="33" pageNumber="34" unit="mm" value="0.2">0.2 mm</quantity>
; ocular index 4.5; length of eyes in frontal view behind sinus
<quantity id="4CAE9B06FFC8FFDF3C80FC69FA9373AA" box="[1210,1314,907,931]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="34" unit="mm" value="0.09">0.09 mm</quantity>
(
<figureCitation id="136D2A66FFC8FFDF3D0CFC68FA2173AB" box="[1334,1424,906,931]" captionStart="FIGURES 22 – 30" captionStartId="6.[151,263,1916,1939]" captionTargetBox="[214,1372,193,1894]" captionTargetId="figure@6.[214,1372,193,1896]" captionTargetPageId="6" captionText="FIGURES 22 – 30. Frontal habitus: 22 Ctenocolum aquilus Albuquerque &amp; Ribeiro-Costa sp. nov. (paratype); 23 C. biolleyi (paratype); 24 C. colburni; 25 C. martiale; 26 C. milelo Albuquerque &amp; Ribeiro-Costa sp. nov. (paratype); 27 C. podagricus; 28 C. punctinotatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 29 C. pygospilotos Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 30 C. triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype). Scale: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3698645" httpUri="https://zenodo.org/record/3698645/files/figure.png" pageId="33" pageNumber="34">Fig. 30</figureCitation>
). Antenna serrate from antennomere 4-10 (
<figureCitation id="136D2A66FFC8FFDF3AB8FC4CFD6A73CF" box="[642,731,942,967]" captionStart="FIGURES 61 – 65" captionStartId="12.[151,263,1650,1673]" captionTargetBox="[156,1426,261,1604]" captionTargetId="figure@12.[151,1436,193,1629]" captionTargetPageId="12" captionText="FIGURES 61 – 65. Lateral habitus: 61 Ctenocolum triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 62 C. acapulcensis; 63 C. janzeni; 64 C. salvini (lectotype); 65 C. tuberculatum. Scale: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3698657" httpUri="https://zenodo.org/record/3698657/files/figure.png" pageId="33" pageNumber="34">Fig. 61</figureCitation>
). Frons with frontal carina (
<figureCitation id="136D2A66FFC8FFDF3C2EFC4CFBD973CF" box="[1044,1128,942,967]" captionStart="FIGURES 22 – 30" captionStartId="6.[151,263,1916,1939]" captionTargetBox="[214,1372,193,1894]" captionTargetId="figure@6.[214,1372,193,1896]" captionTargetPageId="6" captionText="FIGURES 22 – 30. Frontal habitus: 22 Ctenocolum aquilus Albuquerque &amp; Ribeiro-Costa sp. nov. (paratype); 23 C. biolleyi (paratype); 24 C. colburni; 25 C. martiale; 26 C. milelo Albuquerque &amp; Ribeiro-Costa sp. nov. (paratype); 27 C. podagricus; 28 C. punctinotatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 29 C. pygospilotos Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 30 C. triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype). Scale: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3698645" httpUri="https://zenodo.org/record/3698645/files/figure.png" pageId="33" pageNumber="34">Fig. 30</figureCitation>
).
</paragraph>
<paragraph id="8BE936E3FFC8FFDF38FDFC30FF72743B" blockId="33.[151,1437,258,1759]" pageId="33" pageNumber="34">
Prothorax. Pronotum with median gibbosity slightly elevated, not divided by longitudinal and transversal sulcus (
<figureCitation id="136D2A66FFC8FFDF38CEFC14FEFE7407" box="[244,335,1014,1039]" captionStart="FIGURES 61 – 65" captionStartId="12.[151,263,1650,1673]" captionTargetBox="[156,1426,261,1604]" captionTargetId="figure@12.[151,1436,193,1629]" captionTargetPageId="12" captionText="FIGURES 61 – 65. Lateral habitus: 61 Ctenocolum triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 62 C. acapulcensis; 63 C. janzeni; 64 C. salvini (lectotype); 65 C. tuberculatum. Scale: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3698657" httpUri="https://zenodo.org/record/3698657/files/figure.png" pageId="33" pageNumber="34">Fig. 61</figureCitation>
); lateral gibbosity slightly elevated; basal lobe with depression and slightly emarginated (
<figureCitation id="136D2A66FFC8FFDF3D55FC14FF07743B" captionStart="FIGURES 17 – 21" captionStartId="5.[151,262,1469,1492]" captionTargetBox="[184,1433,256,1431]" captionTargetId="figure@5.[151,1436,193,1448]" captionTargetPageId="5" captionText="FIGURES 17 – 21. Dorsal habitus: 17 Ctenocolum triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 18 C. acapulcensis; 19 C. janzeni; 20 C. salvini (lectotype); 21 C. tuberculatum. Scale: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3698643" httpUri="https://zenodo.org/record/3698643/files/figure.png" pageId="33" pageNumber="34">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="8BE936E3FFC8FFDF38FDFBDCFDB374EE" blockId="33.[151,1437,258,1759]" pageId="33" pageNumber="34">
Mesothorax and metathorax. Elytra, striae with punctures moderately impressed; less conspicuous teeth at base of striae 3 and 4; tooth of stria 4 closer to base of tooth of stria 3 than to anterior margin of elytra; stria 6 conspicuously impressed.
<collectingCountry id="F3417673FFC8FFDF39FBFB64FE497497" box="[449,504,1158,1183]" name="India" pageId="33" pageNumber="34">Hind</collectingCountry>
femur (
<figureCitation id="136D2A66FFC8FFDF3A6FFB64FD197497" box="[597,680,1158,1183]" captionStart="FIGURES 72 – 76" captionStartId="14.[151,263,1952,1975]" captionTargetBox="[216,1383,211,1917]" captionTargetId="figure@14.[200,1387,193,1931]" captionTargetPageId="14" captionText="FIGURES 72 – 76. Posterior leg, external view: 72 Ctenocolum punctinotatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 73 C. pygospilotos Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 74 C. triangulatus Albuquerque &amp; Ribeiro- Costa sp. nov. (holotype); 75 C. acapulcensis; 76 C. janzeni. Scale: 0.50 mm." httpUri="https://zenodo.org/record/286739/files/figure.png" pageId="33" pageNumber="34">Fig. 74</figureCitation>
) on external ventral margin with toothed carina; without denticles above of external ventral margin; pecten with 8 teeth.
<collectingCountry id="F3417673FFC8FFDF3B21FB48FCE374CB" box="[795,850,1194,1219]" name="India" pageId="33" pageNumber="34">Hind</collectingCountry>
tibia (
<figureCitation id="136D2A66FFC8FFDF3B9EFB48FC4874CB" box="[932,1017,1194,1219]" captionStart="FIGURES 72 – 76" captionStartId="14.[151,263,1952,1975]" captionTargetBox="[216,1383,211,1917]" captionTargetId="figure@14.[200,1387,193,1931]" captionTargetPageId="14" captionText="FIGURES 72 – 76. Posterior leg, external view: 72 Ctenocolum punctinotatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 73 C. pygospilotos Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 74 C. triangulatus Albuquerque &amp; Ribeiro- Costa sp. nov. (holotype); 75 C. acapulcensis; 76 C. janzeni. Scale: 0.50 mm." httpUri="https://zenodo.org/record/286739/files/figure.png" pageId="33" pageNumber="34">Fig. 74</figureCitation>
) strongly emarginated beside mucro; lateral coronal denticles present.
</paragraph>
<paragraph id="8BE936E3FFC8FFDF38FDFB10FEB17527" blockId="33.[151,1437,258,1759]" pageId="33" pageNumber="34">
Abdomen. Pygidium longer than wide, oval, at median basal region with moderately impressed punctures (
<figureCitation id="136D2A66FFC8FFDF38A5FAF4FF427527" box="[159,243,1302,1327]" captionStart="FIGURES 35 – 43" captionStartId="8.[151,263,1916,1939]" captionTargetBox="[214,1372,193,1894]" captionTargetId="figure@8.[214,1372,193,1896]" captionTargetPageId="8" captionText="FIGURES 35 – 43. Pygidium of male: 35 Ctenocolum aquilus Albuquerque &amp; Ribeiro-Costa sp. nov. (paratype); 36 C. colburni; 37 C. martiale; 38 C. milelo Albuquerque &amp; Ribeiro-Costa sp. nov. (paratype); 39 C. podagricus; 40 C. punctinotatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 41 C. pygospilotos Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 42 C. triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 43 C. acapulcensis. Scale: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3698649" httpUri="https://zenodo.org/record/3698649/files/figure.png" pageId="33" pageNumber="34">Fig. 42</figureCitation>
).
</paragraph>
<paragraph id="8BE936E3FFC8FFDF38FDFAD8FDEC7646" blockId="33.[151,1437,258,1759]" pageId="33" pageNumber="34">
Male genitalia. Median lobe, ventral valve as long as wide, lateral margin concave on subapical region (
<figureCitation id="136D2A66FFC8FFDF3D55FAD8FF07757F" captionStart="FIGURES 85 – 90" captionStartId="18.[151,263,1953,1976]" captionTargetBox="[202,1384,193,1930]" captionTargetId="figure@18.[202,1385,193,1932]" captionTargetPageId="18" captionText="FIGURES 85 – 90. Male genitalia, median lobe: 85 Ctenocolum pygospilotos Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 86 C. triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 87 C. acapulcensis; 88 C. janzeni; 89 C. salvini (paralectotype); 90 C. tuberculatum. Scale: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.3698661" httpUri="https://zenodo.org/record/3698661/files/figure.png" pageId="33" pageNumber="34">Fig. 86</figureCitation>
), basal margin not emarginated. Internal sac, lateral apex with short tuft of setae, hinge sclerite with curved apex, long, extending through subapical region; subapical region with homogeneously distributed spicules, dense medially and in lateral forming elongated strips; median region with homogeneously distributed spicules; submedian region with squamous subtriangular- shape sclerite (
<figureCitation id="136D2A66FFC8FFDF3B5CFA28FC0A75EB" box="[870,955,1482,1507]" captionStart="FIGURES 85 – 90" captionStartId="18.[151,263,1953,1976]" captionTargetBox="[202,1384,193,1930]" captionTargetId="figure@18.[202,1385,193,1932]" captionTargetPageId="18" captionText="FIGURES 85 – 90. Male genitalia, median lobe: 85 Ctenocolum pygospilotos Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 86 C. triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 87 C. acapulcensis; 88 C. janzeni; 89 C. salvini (paralectotype); 90 C. tuberculatum. Scale: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.3698661" httpUri="https://zenodo.org/record/3698661/files/figure.png" pageId="33" pageNumber="34">Fig. 86</figureCitation>
); basal region with spicules. Tegmen (
<figureCitation id="136D2A66FFC8FFDF3D55FA28FF07760F" captionStart="FIGURES 91 – 99" captionStartId="19.[151,263,1916,1939]" captionTargetBox="[214,1372,193,1894]" captionTargetId="figure@19.[214,1372,193,1896]" captionTargetPageId="19" captionText="FIGURES 91 – 99. Male genitalia, tegmen, lateral lobe: 91 C. aquilus (paratype); 92 C. colburni; 93 C. martiale (holotype); 94 C. milelo Albuquerque &amp; Ribeiro-Costa sp. nov. (paratype); 95 C. podagricus; 96 C. punctinotatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 97 C. pygospilotos Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 98 C. triangulatus Albuquerque &amp; Ribeiro-Costa sp. nov. (holotype); 99 C. acapulcensis. Scale: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.3698663" httpUri="https://zenodo.org/record/3698663/files/figure.png" pageId="33" pageNumber="34">Fig. 98</figureCitation>
), lateral lobes separated by emargination about 0.85 times the length of lateral lobes; internal margin near end of emargination straight, forming a “V”; expanded at apex, about 2.5 times the smallest width on median region; without membranous projection at apex.
</paragraph>
<paragraph id="8BE936E3FFC8FFDF38FDF9B9FDAB767B" blockId="33.[151,1437,258,1759]" box="[199,538,1626,1651]" pageId="33" pageNumber="34">Note. The female is unknown.</paragraph>
</subSubSection>
<subSubSection id="C34C6568FFC8FFDF38FDF99CFEB176B3" pageId="33" pageNumber="34" type="etymology">
<paragraph id="8BE936E3FFC8FFDF38FDF99CFEB176B3" blockId="33.[151,1437,258,1759]" pageId="33" pageNumber="34">
<emphasis id="B922EAF1FFC8FFDF38FDF99CFEFF769F" bold="true" box="[199,334,1662,1687]" pageId="33" pageNumber="34">Etymology.</emphasis>
The specific name “
<emphasis id="B922EAF1FFC8FFDF3A79F99CFD6D769E" box="[579,732,1662,1686]" italics="true" pageId="33" pageNumber="34">
<taxonomicName id="4C564D60FFC8FFDF3A79F99CFD65769E" box="[579,724,1662,1686]" class="Insecta" family="Chrysomelidae" genus="Ctenocolum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="triangulatus">triangulatus</taxonomicName>
”
</emphasis>
refers to the squamous subtriangular-shaped sclerite of male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C34C6568FFC8FFDF38FDF924FD0E76D7" box="[199,703,1734,1759]" pageId="33" pageNumber="34" type="distribution">
<paragraph id="8BE936E3FFC8FFDF38FDF924FD0E76D7" blockId="33.[151,1437,258,1759]" box="[199,703,1734,1759]" pageId="33" pageNumber="34">
<emphasis id="B922EAF1FFC8FFDF38FDF924FED176D7" bold="true" box="[199,352,1734,1759]" pageId="33" pageNumber="34">Distribution.</emphasis>
Neotropical region:
<collectingCountry id="F3417673FFC8FFDF3A72F924FD0B76D7" box="[584,698,1734,1759]" name="Colombia" pageId="33" pageNumber="34">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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