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<document id="83F9C3726D3E0525572EA03E3B3FA5DC" ID-DOI="10.1080/00222930310001617742" ID-ISSN="1464-5262" ID-Zenodo-Dep="4676131" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617981004741" checkinUser="carolina" docAuthor="Wiles, P. R." docDate="2004" docId="03FA577FFFBAB11CFE71F992FD2BFDDF" docLanguage="en" docName="JNATHIST.38.2153-2165.pdf" docOrigin="Journal of Natural History 38 (17)" docSource="http://dx.doi.org/10.1080/00222930310001617742" docStyle="DocumentStyle:B3570C21952C5CF44265488F0CE432D6.2:JNatHist.2003-2004.journal_article.2cover" docStyleId="B3570C21952C5CF44265488F0CE432D6" docStyleName="JNatHist.2003-2004.journal_article.2cover" docStyleVersion="2" docTitle="Harpagopalpus indicus &quot; Cook 1967" docType="treatment" docVersion="3" lastPageNumber="2164" masterDocId="FFC32F07FFB1B111FFB5FF81FFBFFFA7" masterDocTitle="Water mites (Acari: Hydrachnidia) from South-East Asia, Thailand and Sulawesi Tenggara (Indonesia): descriptions of new species and new records" masterLastPageNumber="2165" masterPageNumber="2153" pageNumber="2162" updateTime="1698903822784" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="0C0E3E547600BEAD4B9C240D573E722B">Water mites (Acari: Hydrachnidia) from South-East Asia, Thailand and Sulawesi Tenggara (Indonesia): descriptions of new species and new records</mods:title>
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<mods:namePart id="7EDC9E5CAF88D71CFCD5B69D64448BF2">Wiles, P. R.</mods:namePart>
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<mods:date id="DA11A01E6D3937B9D8EA6B62B87A154F">2004</mods:date>
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<subSubSection id="C349B5E2FFBAB11AFE71F992FCEBF98C" box="[452,852,1555,1579]" pageId="11" pageNumber="2162" type="nomenclature">
<paragraph id="8BECE669FFBAB11AFE71F992FCEBF98C" blockId="11.[452,852,1555,1579]" box="[452,852,1555,1579]" pageId="11" pageNumber="2162">
<heading id="D0A45105FFBAB11AFE71F992FCEBF98C" box="[452,852,1555,1579]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="2162" reason="2">
<taxonomicName id="4C539DEAFFBAB11AFE71F992FCEBF98C" authority="Cook, 1967" authorityName="&quot; Cook" authorityYear="1967" box="[452,852,1555,1579]" class="Arachnida" family="Harpagopalpidae" genus="Harpagopalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="11" pageNumber="2162" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B9273A7BFFBAB11AFE71F992FD7BF98C" bold="true" box="[452,708,1555,1579]" italics="true" pageId="11" pageNumber="2162">Harpagopalpus indicus</emphasis>
Cook, 1967
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C349B5E2FFBAB11CFDEFF9B2FD2BFDDF" lastPageId="13" lastPageNumber="2164" pageId="11" pageNumber="2162" type="description">
<paragraph id="8BECE669FFBAB11AFDEFF9B2FD01F9EC" blockId="11.[602,702,1587,1611]" box="[602,702,1587,1611]" pageId="11" pageNumber="2162">
(
<figureCitation id="1368FAECFFBAB11AFDD4F9B2FD09F9EC" box="[609,694,1587,1611]" captionStart="FIG" captionStartId="12.[159,175,1195,1217]" captionTargetBox="[163,1115,170,1160]" captionTargetId="figure-161@12.[159,1117,170,1162]" captionTargetPageId="12" captionText="FIG. 7. Harpagopalpus indicus „ Cook, 1967. (a) Venter; (b) dorsal shield; (c) pedipalp; (d) IV-Leg; (e) I-Leg; (f) lateral view. Scale bar: 170 Mm (a, b, f); 23 Mm (c); 60 Mm (d, e)." figureDoi="http://doi.org/10.5281/zenodo.4676146" httpUri="https://zenodo.org/record/4676146/files/figure.png" pageId="11" pageNumber="2162">figure 7</figureCitation>
)
</paragraph>
<paragraph id="8BECE669FFBAB11AFF0AF9EAFF51F963" blockId="11.[154,1110,1642,1732]" pageId="11" pageNumber="2162">
<emphasis id="B9273A7BFFBAB11AFF0AF9EAFE1DF924" box="[191,418,1643,1667]" italics="true" pageId="11" pageNumber="2162">Material examined.</emphasis>
S376„,
<date id="FFEDC0A9FFBAB11AFDA4F9EBFD0EF925" box="[529,689,1642,1666]" pageId="11" pageNumber="2162" value="2001-07-31">31 July 2001</date>
, S. Lamatano, nr Labundobundo, Buton,
<collectingRegion id="4997288BFFBAB11AFF47F90BFE74F905" box="[242,459,1674,1698]" country="Indonesia" name="Sulawesi Tenggara" pageId="11" pageNumber="2162">Sulawesi Tenggara</collectingRegion>
,
<collectingCountry id="F344A6F9FFBAB11AFE6FF90BFDF4F905" box="[474,587,1674,1698]" name="Indonesia" pageId="11" pageNumber="2162">Indonesia</collectingCountry>
. M474”,
<materialsCitation id="3B3BEC34FFBAB11AFD09F90BFF55F963" collectingDate="1992-09-15" country="Brunei" location="Temburong" pageId="11" pageNumber="2162" specimenCount="1" stateProvince="Temburong">
<date id="FFEDC0A9FFBAB11AFD09F90BFC24F905" box="[700,923,1674,1698]" pageId="11" pageNumber="2162" value="1992-09-15">
<collectingDate id="EFA93941FFBAB11AFD09F90BFC24F905" box="[700,923,1674,1698]" pageId="11" pageNumber="2162" value="1992-09-15">15 September 1992</collectingDate>
</date>
, S.
<collectingRegion id="4997288BFFBAB11AFC79F90BFBEDF905" box="[972,1106,1674,1698]" country="Brunei" name="Temburong" pageId="11" pageNumber="2162">Temburong</collectingRegion>
,
<collectingCountry id="F344A6F9FFBAB11AFF2FF92DFF55F963" box="[154,234,1708,1732]" name="Brunei" pageId="11" pageNumber="2162">Brunei</collectingCountry>
</materialsCitation>
.
</paragraph>
<caption id="DF2CB6E1FFBDB11DFF2AFB2AFBFDFB7C" ID-DOI="http://doi.org/10.5281/zenodo.4676146" ID-Zenodo-Dep="4676146" httpUri="https://zenodo.org/record/4676146/files/figure.png" pageId="12" pageNumber="2163" startId="12.[159,175,1195,1217]" targetBox="[163,1115,170,1160]" targetPageId="12">
<paragraph id="8BECE669FFBDB11DFF2AFB2AFBFDFB7C" blockId="12.[159,1116,1195,1243]" pageId="12" pageNumber="2163">
FIG. 7.
<taxonomicName id="4C539DEAFFBDB11DFF4CFB2AFDC5FB66" authority="&quot; Cook, 1967" authorityName="&quot; Cook" authorityYear="1967" box="[249,634,1195,1217]" class="Arachnida" family="Harpagopalpidae" genus="Harpagopalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="12" pageNumber="2163" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B9273A7BFFBDB11DFF4CFB2AFE61FB67" box="[249,478,1195,1216]" italics="true" pageId="12" pageNumber="2163">Harpagopalpus indicus</emphasis>
„ Cook, 1967
</taxonomicName>
. (a) Venter; (b) dorsal shield; (c) pedipalp; (d) IV-Leg; (e) I-Leg; (f) lateral view. Scale bar: 170
<emphasis id="B9273A7BFFBDB11DFD75FB47FD72FB7C" box="[704,717,1222,1243]" italics="true" pageId="12" pageNumber="2163">M</emphasis>
m (a, b, f); 23
<emphasis id="B9273A7BFFBDB11DFCDDFB47FCCAFB7C" box="[872,885,1222,1243]" italics="true" pageId="12" pageNumber="2163">M</emphasis>
m (c); 60
<emphasis id="B9273A7BFFBDB11DFC6CFB47FC59FB7C" box="[985,998,1222,1243]" italics="true" pageId="12" pageNumber="2163">M</emphasis>
m (d, e).
</paragraph>
</caption>
<paragraph id="8BECE669FFBDB11DFF70FA94FBE4FA14" blockId="12.[159,1116,1301,1459]" pageId="12" pageNumber="2163">
<emphasis id="B9273A7BFFBDB11DFF70FA94FEF3FA8A" box="[197,332,1301,1325]" italics="true" pageId="12" pageNumber="2163">Description.</emphasis>
Male: colour: blue/green/brown. Body broadly rounded anteriorly and tapered to a truncated posterior end; in lateral view, dorsal surface behind EpIV curves ventrally, length 815, width 650, depth 484. R2, and three dorsoglandularia located on large dorsal plate; length 767, width 550. Dorsal furrow complete. R1, A1, A2, one pair of dorsoglandularia and L14 on ventral. L2
</paragraph>
<paragraph id="8BECE669FFBDB11CFF2AFA3FFDC8FF40" blockId="12.[159,1116,1470,1732]" lastBlockId="13.[154,1111,173,632]" lastPageId="13" lastPageNumber="2164" pageId="12" pageNumber="2163">located close to IV-Leg socket and L3L4 anteriorly displaced. EpI fused medially and not projecting beyond the anterior idiosoma. Ventral shield pilose with medial suture lines of epimera indistinct. Genital field with fourzfive asymmetrically distributed acetabula in genital pore and many acetabula on idiosoma located around the posterior epimera; genital opening oval, length 100, width 61. Pedipalp with PIV broad distally and PV narrow and claw-like, length PIPV 26, 68, 28, 81, 53. Anus posterior to genital opening. Legs with swimming setae III-Leg-4 10, III-Leg-5 13, IV-Leg-4 6z9, IV-Leg-4 9z14, IV-Leg-5 10. Length I-Leg-36 82, 103, 126, 113; II-Leg-36 87, 119, 139, 119; III-Leg-36 126, 142, 194, 148; IV-Leg-36 103, 123, 129, 135.</paragraph>
<paragraph id="8BECE669FFBCB11CFF0AFF6FFD2BFDDF" blockId="13.[154,1111,173,632]" pageId="13" pageNumber="2164">
<emphasis id="B9273A7BFFBCB11CFF0AFF6FFE95FEA1" box="[191,298,238,262]" italics="true" pageId="13" pageNumber="2164">Remarks.</emphasis>
The species is described from a single female from
<collectingCountry id="F344A6F9FFBCB11CFC2FFF6FFC65FEA1" box="[922,986,238,262]" name="India" pageId="13" pageNumber="2164">India</collectingCountry>
. A second female was collected in
<collectingCountry id="F344A6F9FFBCB11CFE0EFE8CFDB8FE82" box="[443,519,269,293]" name="Brunei" pageId="13" pageNumber="2164">Brunei</collectingCountry>
(
<bibRefCitation id="EFC29B98FFBCB11CFDAEFE8CFD16FE82" author="WILES, P. R." box="[539,681,269,293]" pageId="13" pageNumber="2164" pagination="409 - 439" refId="ref5623" refString="WILES, P. R., 1999, The water mites (Acari: Hydrachnidia) of Brunei and Thailand, Raffles Bulletin of Zoology, 47, 409 - 439." type="journal article" year="1999">Wiles, 1999</bibRefCitation>
). This is the first record of a male. The
<collectingCountry id="F344A6F9FFBCB11CFF61FEAEFE9FFEE0" box="[212,288,303,327]" name="Brunei" pageId="13" pageNumber="2164">Brunei</collectingCountry>
specimen differs from the
<typeStatus id="54E858CBFFBCB11CFDC4FEAEFD6AFEE0" box="[625,725,303,327]" pageId="13" pageNumber="2164" type="holotype">holotype</typeStatus>
in that it has an asymmetrical distribution of acetabula either side of the genital pore, five and six, respectively, compared with four on the
<typeStatus id="54E858CBFFBCB11CFE42FEF2FDE0FE2C" box="[503,607,371,395]" pageId="13" pageNumber="2164" type="holotype">holotype</typeStatus>
. However, it is almost identical in other respects and is considered to be
<taxonomicName id="4C539DEAFFBCB11CFDABFE14FD2DFE0A" authorityName="&quot; Cook" authorityYear="1967" box="[542,658,405,429]" class="Arachnida" family="Harpagopalpidae" genus="Harpagopalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="13" pageNumber="2164" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B9273A7BFFBCB11CFDABFE14FD2DFE0A" box="[542,658,405,429]" italics="true" pageId="13" pageNumber="2164">H. indicus</emphasis>
</taxonomicName>
. The designation of the female of the only other species,
<taxonomicName id="4C539DEAFFBCB11CFE36FE36FD19FE69" authority="Viets, 1925" authorityName="Viets" authorityYear="1925" box="[387,678,438,463]" class="Arachnida" family="Harpagopalpidae" genus="Harpagopalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="13" pageNumber="2164" phylum="Arthropoda" rank="species" species="octoporus">
<emphasis id="B9273A7BFFBCB11CFE36FE36FDA7FE68" box="[387,536,439,463]" italics="true" pageId="13" pageNumber="2164">H. octoporus</emphasis>
<bibRefCitation id="EFC29B98FFBCB11CFD91FE37FD19FE69" author="VIETS, K. H." box="[548,678,438,462]" pageId="13" pageNumber="2164" pagination="197 - 242" refId="ref5384" refString="VIETS, K. H., 1925, Nachtrage zur Hydracarinen - Fauna von Kamerun, Sammlungen aus Kamerun, Span. Guinea und Fernando Poo, Archiv fur Hydrobiologie, 16, 197 - 242." type="journal article" year="1925">Viets, 1925</bibRefCitation>
</taxonomicName>
from Africa, is dubious (see Cook, 1967: 210). Furthermore, Cooks (1967) newly hatched male
<taxonomicName id="4C539DEAFFBCB11CFCDFFE58FC40FE56" authorityName="Viets" authorityYear="1925" box="[874,1023,473,497]" class="Arachnida" family="Harpagopalpidae" genus="Harpagopalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="13" pageNumber="2164" phylum="Arthropoda" rank="species" species="octoporus">
<emphasis id="B9273A7BFFBCB11CFCDFFE58FC40FE56" box="[874,1023,473,497]" italics="true" pageId="13" pageNumber="2164">H. octoporus</emphasis>
</taxonomicName>
is very different from
<taxonomicName id="4C539DEAFFBCB11CFEFDFE7BFE03FDB5" authorityName="&quot; Cook" authorityYear="1967" box="[328,444,506,530]" class="Arachnida" family="Harpagopalpidae" genus="Harpagopalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="13" pageNumber="2164" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B9273A7BFFBCB11CFEFDFE7BFE03FDB5" box="[328,444,506,530]" italics="true" pageId="13" pageNumber="2164">H. indicus</emphasis>
</taxonomicName>
described above, being broader than long. The male
<taxonomicName id="4C539DEAFFBCB11CFB8CFE7BFF59FD93" authorityName="&quot; Cook" authorityYear="1967" class="Arachnida" family="Harpagopalpidae" genus="Harpagopalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="13" pageNumber="2164" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B9273A7BFFBCB11CFB8CFE7BFF59FD93" italics="true" pageId="13" pageNumber="2164">H. indicus</emphasis>
</taxonomicName>
is like
<taxonomicName id="4C539DEAFFBCB11CFEF7FD9DFE69FD93" authorityName="Viets" authorityYear="1925" box="[322,470,540,564]" class="Arachnida" family="Harpagopalpidae" genus="Harpagopalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="13" pageNumber="2164" phylum="Arthropoda" rank="species" species="octoporus">
<emphasis id="B9273A7BFFBCB11CFEF7FD9DFE69FD93" box="[322,470,540,564]" italics="true" pageId="13" pageNumber="2164">H. octoporus</emphasis>
</taxonomicName>
in that it has acetabula in the genital pore and widespread on the ventral shield. Better specimens of
<taxonomicName id="4C539DEAFFBCB11CFD50FDBFFCC6FDF1" authorityName="Viets" authorityYear="1925" box="[741,889,574,598]" class="Arachnida" family="Harpagopalpidae" genus="Harpagopalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="13" pageNumber="2164" phylum="Arthropoda" rank="species" species="octoporus">
<emphasis id="B9273A7BFFBCB11CFD50FDBFFCC6FDF1" box="[741,889,574,598]" italics="true" pageId="13" pageNumber="2164">H. octoporus</emphasis>
</taxonomicName>
are required for a comprehensive comparison with
<taxonomicName id="4C539DEAFFBCB11CFDACFDE1FD32FDDF" authorityName="&quot; Cook" authorityYear="1967" box="[537,653,608,632]" class="Arachnida" family="Harpagopalpidae" genus="Harpagopalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="13" pageNumber="2164" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B9273A7BFFBCB11CFDACFDE1FD32FDDF" box="[537,653,608,632]" italics="true" pageId="13" pageNumber="2164">H. indicus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>