treatments-xml/data/03/C7/87/03C787B9FFC0EC37FF7DD457FBEFFED9.xml

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<mods:title id="A494559D57C7D6D977B26FB155785DC8">Tanyproctus (Tanyproctus) arher (Coleoptera: Scarabaeidae: Melolonthinae: Tanyproctini), a new species from the Socotra Island, Yemen</mods:title>
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<mods:namePart id="2080A6F021BB495140CFBC48E7EFA0CC">Bezděk, Aleš</mods:namePart>
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<treatment id="03C787B9FFC0EC37FF7DD457FBEFFED9" ID-DOI="http://doi.org/10.5281/zenodo.5698259" ID-GBIF-Taxon="119582865" ID-Zenodo-Dep="5698259" LSID="urn:lsid:plazi:treatment:03C787B9FFC0EC37FF7DD457FBEFFED9" httpUri="http://treatment.plazi.org/id/03C787B9FFC0EC37FF7DD457FBEFFED9" lastPageId="5" lastPageNumber="196" pageId="1" pageNumber="192">
<subSubSection id="C3746524FFC0EC33FF7DD457FEA3FD19" pageId="1" pageNumber="192" type="nomenclature">
<paragraph id="8BD136AFFFC0EC33FF7DD457FC17FD3E" blockId="1.[151,1003,617,677]" box="[151,1003,617,643]" pageId="1" pageNumber="192">
<heading id="D09981C3FFC0EC33FF7DD457FC17FD3E" bold="true" box="[151,1003,617,643]" fontSize="11" level="1" pageId="1" pageNumber="192" reason="1">
<emphasis id="B91AEABDFFC0EC33FF7DD457FC17FD3E" bold="true" box="[151,1003,617,643]" pageId="1" pageNumber="192">
<taxonomicName id="4C6E4D2CFFC0EC33FF7DD457FCB2FD3E" authority="Bezdek, Sehnal &amp; Kral" authorityName="Bezdek, Sehnal &amp; Kral" box="[151,846,617,643]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="192" phylum="Arthropoda" rank="species" species="arher" status="sp. nov." subGenus="Tanyproctus">
<emphasis id="B91AEABDFFC0EC33FF7DD457FED7FD3E" bold="true" box="[151,299,617,643]" italics="true" pageId="1" pageNumber="192">Tanyproctus</emphasis>
(
<emphasis id="B91AEABDFFC0EC33FED6D457FE2CFD3E" bold="true" box="[316,464,617,643]" italics="true" pageId="1" pageNumber="192">Tanyproctus</emphasis>
) 
<emphasis id="B91AEABDFFC0EC33FE08D457FDD9FD3E" bold="true" box="[482,549,617,643]" italics="true" pageId="1" pageNumber="192">arher</emphasis>
Bezděk, Sehnal &amp; Král
</taxonomicName>
, 
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</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136AFFFC0EC33FF7DD4B2FEA3FD19" blockId="1.[151,1003,617,677]" box="[151,351,652,677]" pageId="1" pageNumber="192">
(
<figureCitation id="13552A2AFFC0EC33FF75D4B2FEFFFD19" box="[159,259,652,677]" captionStart="FIGURES 1 – 4. 1 – 2" captionStartId="2.[151,269,1850,1873]" captionTargetBox="[166,1430,189,1817]" captionTargetId="figure@2.[151,1436,176,1828]" captionTargetPageId="2" captionText="FIGURES 1 – 4. 1 – 2: Tanyproctus (Tanyproctus) arher Bezděk, Sehnal &amp; Král, new species, holotype, 8.5 mm: 1. habitus; 2. head; 3 – 4: T. (T.) wraniki Král, Sehnal &amp; Bezděk, 2012, holotype (Socotra, Diksam plateau, IBUR), 6.8 mm: 3. habitus; 4. head. Not in scale." httpUri="https://zenodo.org/record/224231/files/figure.png" pageId="1" pageNumber="192">Figs 1, 2</figureCitation>
, 
<figureCitation id="13552A2AFFC0EC33FEE5D4B2FEC7FD18" box="[271,315,652,677]" captionStart="FIGURES 5 – 8. 5 – 6" captionStartId="3.[151,269,1404,1427]" captionTargetBox="[189,1400,209,1378]" captionTargetId="figure@3.[174,1412,194,1382]" captionTargetPageId="3" captionText="FIGURES 5 – 8. 5 – 6: Tanyproctus (Tanyproctus) arher Bezděk, Sehnal &amp; Král, new species, holotype: 5. aedeagus in lateral aspect; 6. parameres in dorsal aspect; 7 – 8: T. (T.) wraniki Král, Sehnal &amp; Bezděk, 2012, holotype (Socotra, Diksam plateau, IBUR): 7. aedeagus in lateral aspect; 8. parameres in dorsal aspect. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/224232/files/figure.png" pageId="1" pageNumber="192">5, 6</figureCitation>
, 
<figureCitation id="13552A2AFFC0EC33FEADD4B2FEAAFD19" box="[327,342,652,676]" captionStart="FIGURE 9" captionStartId="4.[151,255,1061,1084]" captionTargetBox="[154,1418,197,1060]" captionTargetId="figure@4.[151,1436,194,1124]" captionTargetPageId="4" captionText="FIGURE 9. Type locality of Tanyproctus (Tanyproctus) arher Bezděk, Sehnal &amp; Král, new species, Socotra, Haalla area," httpUri="https://zenodo.org/record/224233/files/figure.png" pageId="1" pageNumber="192">9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3746524FFC0EC33FF7DD4EAFB37FC1D" pageId="1" pageNumber="192" type="materials_examined">
<paragraph id="8BD136AFFFC0EC33FF7DD4EAFE85FCAD" blockId="1.[151,1436,724,2009]" pageId="1" pageNumber="192">
<emphasis id="B91AEABDFFC0EC33FF7DD4EAFEC4FD50" bold="true" box="[151,312,724,749]" pageId="1" pageNumber="192">
<typeStatus id="54D5880DFFC0EC33FF7DD4EAFF2EFD50" box="[151,210,724,749]" pageId="1" pageNumber="192">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F379763FFFC0EC33FEABD4EBFE6EFD51" box="[321,402,725,748]" name="Yemen" pageId="1" pageNumber="192">Yemen</collectingCountry>
, 
<collectingCountry id="F379763FFFC0EC33FE75D4EAFE08FD51" box="[415,500,724,748]" name="Yemen" pageId="1" pageNumber="192">Socotra</collectingCountry>
Island, Hala [= Haalla] area, 
<taxonomicName id="4C6E4D2CFFC0EC33FCB2D4EAFC67FD50" box="[856,923,724,749]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="195" phylum="Arthropoda" rank="species" species="arher">Arher</taxonomicName>
, 
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, 
<geoCoordinate id="EE5A5068FFC0EC33FBC1D4EAFB61FD50" box="[1067,1181,724,749]" direction="east" orientation="longitude" pageId="1" pageNumber="192" precision="92" value="54.46">54°27.6′E</geoCoordinate>
, 
<quantity id="4C969B4AFFC0EC33FB41D4EAFB24FD51" box="[1195,1240,724,749]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="192" unit="m" value="5.0">5 m</quantity>
a.s.l., freshwater spring in sand dune.
</paragraph>
<paragraph id="8BD136AFFFC0EC33FF2DD522FB37FC1D" blockId="1.[151,1436,724,2009]" pageId="1" pageNumber="192">
<emphasis id="B91AEABDFFC0EC33FF2DD522FD81FC88" bold="true" box="[199,637,796,821]" pageId="1" pageNumber="192">
<typeStatus id="54D5880DFFC0EC33FF2DD522FEFEFC88" box="[199,258,796,821]" pageId="1" pageNumber="192">Type</typeStatus>
material. 
<collectingCountry id="F379763FFFC0EC33FE94D522FE33FC88" box="[382,463,796,821]" name="Yemen" pageId="1" pageNumber="192">Yemen</collectingCountry>
(
<collectingCountry id="F379763FFFC0EC33FE35D522FDC1FC88" box="[479,573,796,821]" name="Yemen" pageId="1" pageNumber="192">Socotra</collectingCountry>
Isl.):
</emphasis>
<typeStatus id="54D5880DFFC0EC33FD6FD522FD11FC88" box="[645,749,796,821]" pageId="1" pageNumber="192" type="holotype">Holotype</typeStatus>
♂ (NMPC): “ 
<collectingCountry id="F379763FFFC0EC33FC65D522FC1EFC89" box="[911,994,796,820]" name="Yemen" pageId="1" pageNumber="192">Yemen</collectingCountry>
, 
<collectingCountry id="F379763FFFC0EC33FC05D522FBA7FC89" box="[1007,1115,796,820]" name="Yemen" pageId="1" pageNumber="192">SOCOTRA</collectingCountry>
<collectingCountry id="F379763FFFC0EC33FB89D522FB54FC88" box="[1123,1192,796,821]" name="Iceland" pageId="1" pageNumber="192">Island</collectingCountry>
/ Hala area, 
<taxonomicName id="4C6E4D2CFFC0EC33FAD6D522FA70FC89" box="[1340,1420,796,820]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="195" phylum="Arthropoda" rank="species" species="arher">ARHER</taxonomicName>
/ freshwater spring in sand dune / 9–10.+ 
<date id="FFD0106FFFC0EC33FDBED501FD2FFCE5" box="[596,723,831,856]" pageId="1" pageNumber="192" value="2012-06-15">15-vi-2012</date>
/ 
<geoCoordinate id="EE5A5068FFC0EC33FD00D501FCA3FCE5" box="[746,863,831,856]" direction="north" orientation="latitude" pageId="1" pageNumber="192" precision="92" value="12.55">12°33.0′N</geoCoordinate>
, 
<geoCoordinate id="EE5A5068FFC0EC33FC81D501FC21FCE5" box="[875,989,831,856]" direction="east" orientation="longitude" pageId="1" pageNumber="192" precision="92" value="54.46">54°27.6′E</geoCoordinate>
, 
<quantity id="4C969B4AFFC0EC33FC03D501FBE8FCEA" box="[1001,1044,831,856]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="192" unit="m" value="5.0">5 m</quantity>
// 
<collectingCountry id="F379763FFFC0EC33FBD9D57EFB4AFCE5" box="[1075,1206,832,856]" name="Yemen" pageId="1" pageNumber="192">SOCOTRA</collectingCountry>
expedition 2012 / J. Bezděk, J. Hájek, V. Hula, / P. Kment, I. Malenovský, / J. Niedobová &amp; L. Purchart leg. [printed label]”. 
<typeStatus id="54D5880DFFC0EC33FAD0D55BFA60FCC0" box="[1338,1436,869,893]" pageId="1" pageNumber="192" type="paratype">Paratype</typeStatus>
♂ [pinned specimen, left middle leg preserved in pure alcohol] (ZFMK): same data as 
<typeStatus id="54D5880DFFC0EC33FB8BD5B9FB3BFC1D" box="[1121,1223,903,928]" pageId="1" pageNumber="192" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3746524FFC0EC36FF2DD592FE68F994" lastPageId="4" lastPageNumber="195" pageId="1" pageNumber="192" type="description">
<paragraph id="8BD136AFFFC0EC33FF2DD592FB63FC55" blockId="1.[151,1436,724,2009]" pageId="1" pageNumber="192">
<emphasis id="B91AEABDFFC0EC33FF2DD592FE18FC78" bold="true" box="[199,484,940,965]" pageId="1" pageNumber="192">
Description of 
<typeStatus id="54D5880DFFC0EC33FE97D592FE18FC78" box="[381,484,940,965]" pageId="1" pageNumber="192" type="holotype">holotype</typeStatus>
</emphasis>
(♂). Body length 
<quantity id="4C969B4AFFC0EC33FD55D592FCE7FC79" box="[703,795,940,965]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="192" unit="mm" value="8.5">8.5 mm</quantity>
. Body elongate; surface dark brown to black coloured, appendages and sutural elytral interval somewhat lighter; shiny; macrosetation pale (
<figureCitation id="13552A2AFFC0EC33FBA6D5F1FB71FC55" box="[1100,1165,975,1000]" captionStart="FIGURES 1 – 4. 1 – 2" captionStartId="2.[151,269,1850,1873]" captionTargetBox="[166,1430,189,1817]" captionTargetId="figure@2.[151,1436,176,1828]" captionTargetPageId="2" captionText="FIGURES 1 – 4. 1 – 2: Tanyproctus (Tanyproctus) arher Bezděk, Sehnal &amp; Král, new species, holotype, 8.5 mm: 1. habitus; 2. head; 3 – 4: T. (T.) wraniki Král, Sehnal &amp; Bezděk, 2012, holotype (Socotra, Diksam plateau, IBUR), 6.8 mm: 3. habitus; 4. head. Not in scale." httpUri="https://zenodo.org/record/224231/files/figure.png" pageId="1" pageNumber="192">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BD136AFFFC0EC33FF7DD5CAFE12FA00" blockId="1.[151,1436,724,2009]" pageId="1" pageNumber="192">
<emphasis id="B91AEABDFFC0EC33FF7DD5CAFF28FBB0" box="[151,212,1012,1037]" italics="true" pageId="1" pageNumber="192">Head</emphasis>
. Labrum small, bilobed; lobes round, coarsely, irregularly punctate. Outline of clypeus almost trapezoidal, with distinctly upturned margin; clypeus remarkably depressed along margin and excavated centrally; anterior margin shallowly emarginate, anterior angles round, sides broadly arcuate (
<figureCitation id="13552A2AFFC0EC33FBC4D202FB80FBE9" box="[1070,1148,1084,1109]" captionStart="FIGURES 1 – 4. 1 – 2" captionStartId="2.[151,269,1850,1873]" captionTargetBox="[166,1430,189,1817]" captionTargetId="figure@2.[151,1436,176,1828]" captionTargetPageId="2" captionText="FIGURES 1 – 4. 1 – 2: Tanyproctus (Tanyproctus) arher Bezděk, Sehnal &amp; Král, new species, holotype, 8.5 mm: 1. habitus; 2. head; 3 – 4: T. (T.) wraniki Král, Sehnal &amp; Bezděk, 2012, holotype (Socotra, Diksam plateau, IBUR), 6.8 mm: 3. habitus; 4. head. Not in scale." httpUri="https://zenodo.org/record/224231/files/figure.png" pageId="1" pageNumber="192">Fig. 2</figureCitation>
). Genae narrow, round. Frontoclypeal suture feebly arcuate, considerably impressed. Eyes relatively large, distinctly exceeding genae externally in dorsal aspect (
<figureCitation id="13552A2AFFC0EC33FE25D2BAFDE9FB21" box="[463,533,1156,1181]" captionStart="FIGURES 1 – 4. 1 – 2" captionStartId="2.[151,269,1850,1873]" captionTargetBox="[166,1430,189,1817]" captionTargetId="figure@2.[151,1436,176,1828]" captionTargetPageId="2" captionText="FIGURES 1 – 4. 1 – 2: Tanyproctus (Tanyproctus) arher Bezděk, Sehnal &amp; Král, new species, holotype, 8.5 mm: 1. habitus; 2. head; 3 – 4: T. (T.) wraniki Král, Sehnal &amp; Bezděk, 2012, holotype (Socotra, Diksam plateau, IBUR), 6.8 mm: 3. habitus; 4. head. Not in scale." httpUri="https://zenodo.org/record/224231/files/figure.png" pageId="1" pageNumber="192">Fig. 2</figureCitation>
); distance between eyes in ventral aspect extended remarkably diameter of eye. Punctation of clypeus coarse and dense, almost evenly distributed but impunctate centrally; punctures separated by approximately their diameter, each puncture bearing short, semierect macroseta. Vertex rather rugopunctate; punctures separated by less than their diameter to confluent, each puncture bearing very short, erect macroseta; macrosetation becoming shorter posteriad. Antennae with 10 antennomeres; antennomere 2 short, approximately as long as wide; antennomeres 3–5 elongate. Club pentamerous, straight, shorter than antennal shaft (antennomeres 1–5 combined). Antennomeres 1–5 with sparse, long macrosetae, club with sparse, short macrosetae. Terminal maxillar palpomeres elongate, rounded apically, absent from depression, approximately of same length as palpomeres 2 and 3 combined.
</paragraph>
<paragraph id="8BD136AFFFC0EC33FF2DD3F7FDF8F8BD" blockId="1.[151,1436,724,2009]" pageId="1" pageNumber="192">
<emphasis id="B91AEABDFFC0EC33FF2DD3F7FECAFA5D" box="[199,310,1481,1504]" italics="true" pageId="1" pageNumber="192">Pronotum</emphasis>
moderately convex, transversal, broadest just anterior to middle except broad basal interruption all around border; anterior bead flat, narrow, distinctly widened medially. Punctures irregular; each puncture bearing long, erect macroseta; macrosetae becoming regularly shorter mediad. Lateral margin considerably coarsely crenate, with row of long macrosetae; basal margin with row of finely and irregularly distributed punctures bearing long, recumbent macrosetae in posterior angles and short macrosetae medially. Anterior angles prominent, projecting over anterior margin, acute-angled, with round apex; sides in approximately anterior half almost straight, divergent posteriad to very broadly obtuse posterior angles; posterior margin broadly round. Surface glabrous and finely, coarsely, slightly irregularly punctate, punctures separated by 2–4 their diameters, area along medial longitudinal line smooth.
</paragraph>
<paragraph id="8BD136AFFFC0EC33FF2DD132FE0EF8F5" blockId="1.[151,1436,724,2009]" pageId="1" pageNumber="192">
<taxonomicName id="4C6E4D2CFFC0EC33FF2DD132FEC9F898" box="[199,309,1804,1829]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="1" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEABDFFC0EC33FF2DD132FEC9F898" box="[199,309,1804,1829]" italics="true" pageId="1" pageNumber="192">Scutellum</emphasis>
</taxonomicName>
approximately as wide as long, triangulate, sides broadly arcuate, apex acute, surface with several punctures basally and laterally.
</paragraph>
<paragraph id="8BD136AFFFC0EC33FF2DD16AFB0CF865" blockId="1.[151,1436,724,2009]" pageId="1" pageNumber="192">
<emphasis id="B91AEABDFFC0EC33FF2DD16AFEF1F8D0" box="[199,269,1876,1901]" italics="true" pageId="1" pageNumber="192">Elytra</emphasis>
convex, slightly dilated posteriad, sutural angle rounded; striae excepting sutural stria missing or only very feebly indicated; distinctly microsculptured with disc feebly transversally wrinkled; punctation coarse, dense, almost regular; punctures separated by 1–2 their diameters. Sutural interval slightly convex; sutural stria with row of irregularly distributed punctures; lateral margin distinctly bordered with row of long, erect setae.
</paragraph>
<caption id="DF116627FFC3EC30FF7DD104FE11F82C" httpUri="https://zenodo.org/record/224231/files/figure.png" pageId="2" pageNumber="193" targetBox="[166,1430,189,1817]" targetPageId="2">
<paragraph id="8BD136AFFFC3EC30FF7DD104FE11F82C" blockId="2.[151,1437,1850,1937]" pageId="2" pageNumber="193">
<emphasis id="B91AEABDFFC3EC30FF7DD104FEBBF8EC" bold="true" box="[151,327,1850,1873]" pageId="2" pageNumber="193">FIGURES 1–4.</emphasis>
1–2: 
<taxonomicName id="4C6E4D2CFFC3EC30FE64D104FBF5F8EC" authority="Bezdek, Sehnal &amp; Kral" authorityName="Bezdek, Sehnal &amp; Kral" box="[398,1033,1850,1873]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="193" phylum="Arthropoda" rank="species" species="arher" status="sp. nov." subGenus="Tanyproctus">
<emphasis id="B91AEABDFFC3EC30FE64D104FDECF8EC" box="[398,528,1850,1873]" italics="true" pageId="2" pageNumber="193">Tanyproctus</emphasis>
(
<emphasis id="B91AEABDFFC3EC30FDCED104FD5AF8EC" box="[548,678,1850,1873]" italics="true" pageId="2" pageNumber="193">Tanyproctus</emphasis>
) 
<emphasis id="B91AEABDFFC3EC30FD50D104FD08F8EC" box="[698,756,1850,1873]" italics="true" pageId="2" pageNumber="193">arher</emphasis>
Bezděk, Sehnal &amp; Král
</taxonomicName>
, 
<taxonomicNameLabel id="A22957C6FFC3EC30FBF2D104FB5CF8EC" box="[1048,1184,1850,1873]" pageId="2" pageNumber="193" rank="species">new species</taxonomicNameLabel>
, holotype, 8.5 mm: 1. habitus; 2. head; 3–4: 
<emphasis id="B91AEABDFFC3EC30FE6ED164FE6DF8CC" box="[388,401,1882,1905]" italics="true" pageId="2" pageNumber="193">T</emphasis>
. (
<emphasis id="B91AEABDFFC3EC30FE4DD164FE48F8CC" box="[423,436,1882,1905]" italics="true" pageId="2" pageNumber="193">T</emphasis>
.) 
<taxonomicName id="4C6E4D2CFFC3EC30FE21D164FC9DF8CC" authority="Kral, Sehnal &amp; Bezdek, 2012" authorityName="Kral, Sehnal &amp; Bezdek" authorityYear="2012" box="[459,865,1882,1905]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="193" phylum="Arthropoda" rank="species" species="wraniki">
<emphasis id="B91AEABDFFC3EC30FE21D164FDE0F8CC" box="[459,540,1882,1905]" italics="true" pageId="2" pageNumber="193">wraniki</emphasis>
Král, Sehnal &amp; Bezděk, 2012
</taxonomicName>
, holotype (Socotra, Diksam plateau, IBUR), 6.8 mm: 3. habitus; 4. head. Not in scale.
</paragraph>
</caption>
<caption id="DF116627FFC2EC31FF7DD342FB0FFA6E" httpUri="https://zenodo.org/record/224232/files/figure.png" pageId="3" pageNumber="194" targetBox="[189,1400,209,1378]" targetPageId="3">
<paragraph id="8BD136AFFFC2EC31FF7DD342FB0FFA6E" blockId="3.[151,1436,1404,1491]" pageId="3" pageNumber="194">
<emphasis id="B91AEABDFFC2EC31FF7DD342FEB9FA2E" bold="true" box="[151,325,1404,1427]" pageId="3" pageNumber="194">FIGURES 5–8.</emphasis>
5–6: 
<taxonomicName id="4C6E4D2CFFC2EC31FE6CD342FC0FFA2E" authority="Bezdek, Sehnal &amp; Kral" authorityName="Bezdek, Sehnal &amp; Kral" box="[390,1011,1404,1427]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="194" phylum="Arthropoda" rank="species" species="arher" status="sp. nov." subGenus="Tanyproctus">
<emphasis id="B91AEABDFFC2EC31FE6CD342FDFBFA2E" box="[390,519,1404,1427]" italics="true" pageId="3" pageNumber="194">Tanyproctus</emphasis>
(
<emphasis id="B91AEABDFFC2EC31FDF0D342FD60FA2E" box="[538,668,1404,1427]" italics="true" pageId="3" pageNumber="194">Tanyproctus</emphasis>
) 
<emphasis id="B91AEABDFFC2EC31FD44D342FD15FA2E" box="[686,745,1404,1427]" italics="true" pageId="3" pageNumber="194">arher</emphasis>
Bezděk, Sehnal &amp; Král
</taxonomicName>
, 
<taxonomicNameLabel id="A22957C6FFC2EC31FBEAD342FB79FA2E" box="[1024,1157,1404,1427]" pageId="3" pageNumber="194" rank="species">new species</taxonomicNameLabel>
, holotype: 5. aedeagus in lateral aspect; 6. parameres in dorsal aspect; 7–8: 
<emphasis id="B91AEABDFFC2EC31FD21D3A2FD24FA0E" box="[715,728,1436,1459]" italics="true" pageId="3" pageNumber="194">T</emphasis>
. (
<emphasis id="B91AEABDFFC2EC31FD19D3A2FCFCFA0E" box="[755,768,1436,1459]" italics="true" pageId="3" pageNumber="194">T</emphasis>
.) 
<taxonomicName id="4C6E4D2CFFC2EC31FCF0D3A2FB38FA0E" authority="Kral, Sehnal &amp; Bezdek, 2012" authorityName="Kral, Sehnal &amp; Bezdek" authorityYear="2012" box="[794,1220,1436,1459]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="194" phylum="Arthropoda" rank="species" species="wraniki">
<emphasis id="B91AEABDFFC2EC31FCF0D3A2FC90FA0E" box="[794,876,1436,1459]" italics="true" pageId="3" pageNumber="194">wraniki</emphasis>
Král, Sehnal &amp; Bezděk, 2012
</taxonomicName>
, holotype (Socotra, Diksam plateau, IBUR): 7. aedeagus in lateral aspect; 8. parameres in dorsal aspect. Scale bar = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BD136AFFFC2EC31FF2DD3C3FE96F9A9" blockId="3.[151,1437,1533,2025]" box="[199,362,1533,1556]" pageId="3" pageNumber="194">
<emphasis id="B91AEABDFFC2EC31FF2DD3C3FE9FF9A9" box="[199,355,1533,1556]" italics="true" pageId="3" pageNumber="194">Macropterous</emphasis>
.
</paragraph>
<paragraph id="8BD136AFFFC2EC31FF2DD01FFE6CF8E4" blockId="3.[151,1437,1533,2025]" pageId="3" pageNumber="194">
<emphasis id="B91AEABDFFC2EC31FF2DD01FFF00F985" box="[199,252,1569,1592]" italics="true" pageId="3" pageNumber="194">Legs</emphasis>
. Femora moderately shiny; metafemora with entire surface sparsely, almost regularly punctate; punctures bearing long, recumbent macrosetae. Protibia tridentate; basal tooth considerably weak; terminal calcar long, sharp, slightly curved externally, acute apically, inserted against emargination between basal and medial teeth. Mesotibia and metatibia slightly expanded apicad, with two setiferous transversal carinae. Mesotibial terminal calcars equal in length, flattened, acute apically. Metatibial terminal calcars equal in length, considerably flattened, acute apically. Protarsomeres 2–4 considerably dilated, protarsomere 2 with length-width ratio: 1:2. Mesotarsomeres 2–4 remarkably more slightly dilated that those of protibiae; protarsomeres and mesotarsomeres 1-4 with shortly and densely macrosetaceous pads ventrally; metatarsomeres covered with long sparse macrosetae ventrally. Claws bifid.
</paragraph>
<paragraph id="8BD136AFFFC2EC31FF2DD15AFC31F8C0" blockId="3.[151,1437,1533,2025]" box="[199,973,1892,1917]" pageId="3" pageNumber="194">
<emphasis id="B91AEABDFFC2EC31FF2DD15AFE8EF8C0" box="[199,370,1892,1917]" italics="true" pageId="3" pageNumber="194">Ventral surface</emphasis>
of thorax with dense, long, recumbent macrosetation.
</paragraph>
<paragraph id="8BD136AFFFC2EC31FF2DD1B6FCD0F879" blockId="3.[151,1437,1533,2025]" pageId="3" pageNumber="194">
<emphasis id="B91AEABDFFC2EC31FF2DD1B6FEA4F81C" box="[199,344,1928,1953]" italics="true" pageId="3" pageNumber="194">Propygidium</emphasis>
microsculptured, coarsely, sparsely, irregularly punctate; pygidium finely microsculptured; border complete; surface coarsely and irregularly punctate.
</paragraph>
<paragraph id="8BD136AFFFC2EC31FF2DD1EFFC41F855" blockId="3.[151,1437,1533,2025]" box="[199,957,2000,2025]" pageId="3" pageNumber="194">
<emphasis id="B91AEABDFFC2EC31FF2DD1EFFEDAF855" box="[199,294,2001,2024]" italics="true" pageId="3" pageNumber="194">Ventrit es</emphasis>
almost bare, remarkably coarsely and irregularly punctate.
</paragraph>
<caption id="DF116627FFC5EC36FF7DD21BFA60FB81" box="[151,1436,1061,1084]" httpUri="https://zenodo.org/record/224233/files/figure.png" pageId="4" pageNumber="195" targetBox="[154,1418,197,1060]" targetPageId="4">
<paragraph id="8BD136AFFFC5EC36FF7DD21BFA60FB81" blockId="4.[151,1436,1061,1084]" box="[151,1436,1061,1084]" pageId="4" pageNumber="195">
<emphasis id="B91AEABDFFC5EC36FF7DD21BFEE4FB81" bold="true" box="[151,280,1061,1084]" pageId="4" pageNumber="195">FIGURE 9.</emphasis>
Type locality of 
<taxonomicName id="4C6E4D2CFFC5EC36FE26D21BFBD4FB81" authority="Bezdek, Sehnal &amp; Kral" authorityName="Bezdek, Sehnal &amp; Kral" box="[460,1064,1061,1084]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="arher" status="sp. nov." subGenus="Tanyproctus">
<emphasis id="B91AEABDFFC5EC36FE26D21BFDB2FB81" box="[460,590,1061,1084]" italics="true" pageId="4" pageNumber="195">Tanyproctus</emphasis>
(
<emphasis id="B91AEABDFFC5EC36FDB7D21BFD23FB81" box="[605,735,1061,1084]" italics="true" pageId="4" pageNumber="195">Tanyproctus</emphasis>
) 
<emphasis id="B91AEABDFFC5EC36FD04D21BFCD5FB81" box="[750,809,1061,1084]" italics="true" pageId="4" pageNumber="195">arher</emphasis>
Bezděk, Sehnal &amp; Král
</taxonomicName>
, 
<taxonomicNameLabel id="A22957C6FFC5EC36FBD8D21BFB49FB81" box="[1074,1205,1061,1084]" pageId="4" pageNumber="195" rank="species">new species</taxonomicNameLabel>
, Socotra, Haalla area,
</paragraph>
</caption>
<paragraph id="8BD136AFFFC5EC36FF2DD2B7FD3AFB7B" blockId="4.[151,1436,1161,1474]" pageId="4" pageNumber="195">
<emphasis id="B91AEABDFFC5EC36FF2DD2B7FE97FB1F" box="[199,363,1161,1186]" italics="true" pageId="4" pageNumber="195">Male genitalia</emphasis>
(
<figureCitation id="13552A2AFFC5EC36FE92D2B7FE23FB1F" box="[376,479,1161,1186]" captionStart="FIGURES 5 – 8. 5 – 6" captionStartId="3.[151,269,1404,1427]" captionTargetBox="[189,1400,209,1378]" captionTargetId="figure@3.[174,1412,194,1382]" captionTargetPageId="3" captionText="FIGURES 5 – 8. 5 – 6: Tanyproctus (Tanyproctus) arher Bezděk, Sehnal &amp; Král, new species, holotype: 5. aedeagus in lateral aspect; 6. parameres in dorsal aspect; 7 – 8: T. (T.) wraniki Král, Sehnal &amp; Bezděk, 2012, holotype (Socotra, Diksam plateau, IBUR): 7. aedeagus in lateral aspect; 8. parameres in dorsal aspect. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/224232/files/figure.png" pageId="4" pageNumber="195">Figs 5–6</figureCitation>
). Aedeagus symmetrical; parameres slender, relatively long, only slightly shorter than phallobase, distal part not dilated in dorsal aspect.
</paragraph>
<paragraph id="8BD136AFFFC5EC36FF2DD2EFFE5EFB57" blockId="4.[151,1436,1161,1474]" box="[199,418,1233,1258]" pageId="4" pageNumber="195">
<emphasis id="B91AEABDFFC5EC36FF2DD2EFFED9FB57" bold="true" box="[199,293,1233,1258]" pageId="4" pageNumber="195">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BD136AFFFC5EC36FF2DD2CBFEBBFA8F" blockId="4.[151,1436,1161,1474]" pageId="4" pageNumber="195">
<emphasis id="B91AEABDFFC5EC36FF2DD2CBFEB9FAB3" bold="true" box="[199,325,1269,1294]" pageId="4" pageNumber="195">Varability.</emphasis>
The only known 
<typeStatus id="54D5880DFFC5EC36FDF1D2C9FD87FAB2" box="[539,635,1271,1295]" pageId="4" pageNumber="195" type="paratype">paratype</typeStatus>
slightly differs in size (total body length: 
<quantity id="4C969B4AFFC5EC36FB9ED2C8FB2CFAB3" box="[1140,1232,1270,1294]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="4" pageNumber="195" unit="mm" value="8.4">8.4 mm</quantity>
) and with lighter coloured elytra.
</paragraph>
<paragraph id="8BD136AFFFC5EC36FF2DD303FE7AFA7F" blockId="4.[151,1436,1161,1474]" pageId="4" pageNumber="195">
<emphasis id="B91AEABDFFC5EC36FF2DD303FE33FAEB" bold="true" box="[199,463,1341,1366]" pageId="4" pageNumber="195">Differential diagnosis.</emphasis>
<taxonomicName id="4C6E4D2CFFC5EC36FE3CD301FDA2FAEB" box="[470,606,1343,1366]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEABDFFC5EC36FE3CD301FDA2FAEB" box="[470,606,1343,1366]" italics="true" pageId="4" pageNumber="195">Tanyproctus</emphasis>
</taxonomicName>
(
<emphasis id="B91AEABDFFC5EC36FD9AD301FD81FAEB" box="[624,637,1343,1366]" italics="true" pageId="4" pageNumber="195">T</emphasis>
.) 
<taxonomicName id="4C6E4D2CFFC5EC36FD7CD303FD28FAEB" box="[662,724,1341,1366]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="arher">
<emphasis id="B91AEABDFFC5EC36FD7CD303FD28FAEB" box="[662,724,1341,1366]" italics="true" pageId="4" pageNumber="195">arher</emphasis>
</taxonomicName>
is morphologically very similar to the other endemic Socotran 
<taxonomicName id="4C6E4D2CFFC5EC36FF7DD35DFEE3FAC7" box="[151,287,1379,1402]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEABDFFC5EC36FF7DD35DFEE3FAC7" box="[151,287,1379,1402]" italics="true" pageId="4" pageNumber="195">Tanyproctus</emphasis>
</taxonomicName>
species (Král 
<emphasis id="B91AEABDFFC5EC36FE2BD35DFE05FAC7" box="[449,505,1377,1402]" italics="true" pageId="4" pageNumber="195">et al.</emphasis>
2012). In the key to males of Socotran 
<taxonomicName id="4C6E4D2CFFC5EC36FC52D35FFBBBFAC7" box="[952,1095,1377,1402]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="tribe" tribe="Tanyproctini">Tanyproctini</taxonomicName>
(Král 
<emphasis id="B91AEABDFFC5EC36FB7AD35DFB35FAC7" box="[1168,1225,1377,1402]" italics="true" pageId="4" pageNumber="195">et al.</emphasis>
2012: 180), 
<emphasis id="B91AEABDFFC5EC36FAB3D35DFA9AFAC7" box="[1369,1382,1379,1402]" italics="true" pageId="4" pageNumber="195">T</emphasis>
. (
<emphasis id="B91AEABDFFC5EC36FA94D35DFA77FAC7" box="[1406,1419,1379,1402]" italics="true" pageId="4" pageNumber="195">T</emphasis>
.) 
<taxonomicName id="4C6E4D2CFFC5EC36FF7DD3BBFF29FA23" box="[151,213,1413,1438]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="arher">
<emphasis id="B91AEABDFFC5EC36FF7DD3BBFF29FA23" box="[151,213,1413,1438]" italics="true" pageId="4" pageNumber="195">arher</emphasis>
</taxonomicName>
will key to the couplet with 
<emphasis id="B91AEABDFFC5EC36FDFBD3B9FDE2FA23" box="[529,542,1415,1438]" italics="true" pageId="4" pageNumber="195">T</emphasis>
. (
<emphasis id="B91AEABDFFC5EC36FDDFD3B9FDBFFA23" box="[565,579,1415,1438]" italics="true" pageId="4" pageNumber="195">T</emphasis>
.) 
<taxonomicName id="4C6E4D2CFFC5EC36FDB3D3BBFD53FA23" box="[601,687,1413,1438]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="wraniki">
<emphasis id="B91AEABDFFC5EC36FDB3D3BBFD53FA23" box="[601,687,1413,1438]" italics="true" pageId="4" pageNumber="195">wraniki</emphasis>
</taxonomicName>
. For differentiation from this species see the complex of diagnostic characters in 
<tableCitation id="C6EC0314FFC5EC36FEC6D397FE7CFA7F" box="[300,384,1449,1474]" captionStart="TABLE 1" captionStartId="4.[151,239,1522,1545]" captionTargetBox="[151,1436,1602,2003]" captionTargetPageId="4" captionText="TABLE 1. Differential characters of Tanyproctus (T.) arher Bezděk, Sehnal &amp; Král, new species and T. (T.) wraniki Král," httpUri="http://table.plazi.org/id/DF116627FFC5EC36FF7DD3CCFA60F9B4" pageId="4" pageNumber="195" tableUuid="DF116627FFC5EC36FF7DD3CCFA60F9B4">Table 1</tableCitation>
.
</paragraph>
<caption id="DF116627FFC5EC36FF7DD3CCFA60F9B4" ID-Table-UUID="DF116627FFC5EC36FF7DD3CCFA60F9B4" box="[151,1436,1522,1545]" httpUri="http://table.plazi.org/id/DF116627FFC5EC36FF7DD3CCFA60F9B4" pageId="4" pageNumber="195" targetBox="[151,1436,1602,2003]" targetIsTable="true" targetPageId="4">
<paragraph id="8BD136AFFFC5EC36FF7DD3CCFA60F9B4" blockId="4.[151,1436,1522,1577]" box="[151,1436,1522,1545]" pageId="4" pageNumber="195">
<emphasis id="B91AEABDFFC5EC36FF7DD3CCFEF5F9B4" bold="true" box="[151,265,1522,1545]" pageId="4" pageNumber="195">TABLE 1.</emphasis>
Differential characters of 
<taxonomicName id="4C6E4D2CFFC5EC36FDCAD3CCFD5EF9B4" box="[544,674,1522,1545]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEABDFFC5EC36FDCAD3CCFD5EF9B4" box="[544,674,1522,1545]" italics="true" pageId="4" pageNumber="195">Tanyproctus</emphasis>
</taxonomicName>
(
<emphasis id="B91AEABDFFC5EC36FD5BD3CCFD42F9B4" box="[689,702,1522,1545]" italics="true" pageId="4" pageNumber="195">T</emphasis>
.) 
<taxonomicName id="4C6E4D2CFFC5EC36FD39D3CCFBF1F9B4" authority="Bezdek, Sehnal &amp; Kral" authorityName="Bezdek, Sehnal &amp; Kral" box="[723,1037,1522,1545]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="arher" status="sp. nov.">
<emphasis id="B91AEABDFFC5EC36FD39D3CCFCF1F9B4" box="[723,781,1522,1545]" italics="true" pageId="4" pageNumber="195">arher</emphasis>
Bezděk, Sehnal &amp; Král
</taxonomicName>
, 
<taxonomicNameLabel id="A22957C6FFC5EC36FBFDD3CCFB6AF9B4" box="[1047,1174,1522,1545]" pageId="4" pageNumber="195" rank="species">new species</taxonomicNameLabel>
and 
<emphasis id="B91AEABDFFC5EC36FB23D3CCFB20F9B4" box="[1225,1244,1522,1545]" italics="true" pageId="4" pageNumber="195">T.</emphasis>
(
<emphasis id="B91AEABDFFC5EC36FB06D3CCFB05F9B4" box="[1260,1273,1522,1545]" italics="true" pageId="4" pageNumber="195">T</emphasis>
.) 
<taxonomicName id="4C6E4D2CFFC5EC36FAE4D3CCFA64F9B4" authority="Kral" authorityName="Kral" box="[1294,1432,1522,1545]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="wraniki">
<emphasis id="B91AEABDFFC5EC36FAE4D3CCFAA3F9B4" box="[1294,1375,1522,1545]" italics="true" pageId="4" pageNumber="195">wraniki</emphasis>
Král
</taxonomicName>
,
</paragraph>
</caption>
<paragraph id="8BD136AFFFC5EC36FF7DD02CFE68F994" blockId="4.[151,1436,1522,1577]" box="[151,404,1554,1577]" pageId="4" pageNumber="195">Sehnal &amp; Bezděk, 2012</paragraph>
</subSubSection>
<paragraph id="8BD136AFFFC5EC36FF76D07CFBA3F86E" pageId="4" pageNumber="195">
<table id="F96EC40FFFC513CDFF7DD07CFA60F86E" box="[151,1436,1602,2003]" gridcols="3" gridrows="5" pageId="4" pageNumber="195">
<tr id="355E34EDFFC513CDFF7DD07CFA60F9C6" box="[151,1436,1602,1659]" gridrow="0" pageId="4" pageNumber="195">
<th id="768F5D91FFC513CDFF7DD07CFE94F9C6" box="[151,360,1602,1659]" gridcol="0" gridrow="0" pageId="4" pageNumber="195">species / character shape of clypeus</th>
<th id="768F5D91FFC513CDFE62D07CFCC8F9C6" box="[392,820,1602,1659]" gridcol="1" gridrow="0" pageId="4" pageNumber="195">
<taxonomicName id="4C6E4D2CFFC5EC36FE62D07DFDBEF9E5" box="[392,578,1602,1624]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="arher">
<emphasis id="B91AEABDFFC5EC36FE62D07DFDBEF9E5" box="[392,578,1602,1624]" italics="true" pageId="4" pageNumber="195">Tanyproctus arher</emphasis>
</taxonomicName>
trapezoidal, longer (Fig. 2)
</th>
<th id="768F5D91FFC513CDFCAFD07CFA60F9C6" box="[837,1436,1602,1659]" gridcol="2" gridrow="0" pageId="4" pageNumber="195">
<taxonomicName id="4C6E4D2CFFC5EC36FCAFD07DFBE6F9E5" box="[837,1050,1602,1624]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="wraniki">
<emphasis id="B91AEABDFFC5EC36FCAFD07DFBE6F9E5" box="[837,1050,1602,1624]" italics="true" pageId="4" pageNumber="195">Tanyproctus wraniki</emphasis>
</taxonomicName>
trapezoidal, shorter (Fig. 4)
</th>
</tr>
<tr id="355E34EDFFC513CDFF7DD0B4FA60F95F" box="[151,1436,1674,1762]" gridrow="1" pageId="4" pageNumber="195">
<th id="768F5D91FFC513CDFF7DD0B4FE94F95F" box="[151,360,1674,1762]" gridcol="0" gridrow="1" pageId="4" pageNumber="195">sculpture of clypeus size of eyes</th>
<td id="768F5D91FFC513CDFE62D0B4FCC8F95F" box="[392,820,1674,1762]" gridcol="1" gridrow="1" pageId="4" pageNumber="195">excavate and impunctate centrally (Fig. 2) relatively large, considerably prominent laterad in dorsal aspect (Fig. 2)</td>
<td id="768F5D91FFC513CDFCAFD0B4FA60F95F" box="[837,1436,1674,1762]" gridcol="2" gridrow="1" pageId="4" pageNumber="195">depressed and punctate centrally (Fig. 4) relatively small, only weakly prominent externally in dorsal aspect (Fig. 4)</td>
</tr>
<tr id="355E34EDFFC513CDFF7DD0CEFA60F898" box="[151,1436,1776,1829]" gridrow="2" pageId="4" pageNumber="195">
<th id="768F5D91FFC513CDFF7DD0CEFE94F898" box="[151,360,1776,1829]" gridcol="0" gridrow="2" pageId="4" pageNumber="195">metafemora</th>
<td id="768F5D91FFC513CDFE62D0CEFCC8F898" box="[392,820,1776,1829]" gridcol="1" gridrow="2" pageId="4" pageNumber="195">entire surface sparsely, almost regularly punctate</td>
<td id="768F5D91FFC513CDFCAFD0CEFA60F898" box="[837,1436,1776,1829]" gridcol="2" gridrow="2" pageId="4" pageNumber="195">punctation coarser, somewhat irregular, restricted approximately to marginal areas, impunctate centrally</td>
</tr>
<tr id="355E34EDFFC513CDFF7DD10DFA60F836" box="[151,1436,1843,1931]" gridrow="3" pageId="4" pageNumber="195">
<th id="768F5D91FFC513CDFF7DD10DFE94F836" box="[151,360,1843,1931]" gridcol="0" gridrow="3" pageId="4" pageNumber="195">shape of protarsomeres 2–4 shape of parameres</th>
<td id="768F5D91FFC513CDFE62D10DFCC8F836" box="[392,820,1843,1931]" gridcol="1" gridrow="3" pageId="4" pageNumber="195">considerably dilated, protarsomere 2 with length-width ratio: 1:2 as in Figs 5–6</td>
<td id="768F5D91FFC513CDFCAFD10DFA60F836" box="[837,1436,1843,1931]" gridcol="2" gridrow="3" pageId="4" pageNumber="195">dilated, protarsomere 2 with length–width ratio: 1.0:3.7 as in Figs 7–8</td>
</tr>
<tr id="355E34EDFFC513CDFF7DD1A4FA60F86E" box="[151,1436,1946,2003]" gridrow="4" pageId="4" pageNumber="195">
<th id="768F5D91FFC513CDFF7DD1A4FE94F86E" box="[151,360,1946,2003]" gridcol="0" gridrow="4" pageId="4" pageNumber="195">body size distribution</th>
<td id="768F5D91FFC513CDFE62D1A4FCC8F86E" box="[392,820,1946,2003]" gridcol="1" gridrow="4" pageId="4" pageNumber="195">8.4–8.5 mm E Socotra (Haalla coastal area)</td>
<td id="768F5D91FFC513CDFCAFD1A4FA60F86E" box="[837,1436,1946,2003]" gridcol="2" gridrow="4" pageId="4" pageNumber="195">6.8 mm (holotype) C Socotra (Diksam plateau)</td>
</tr>
</table>
</paragraph>
<subSubSection id="C3746524FFC4EC37FF2DD6A9FEAAFEA0" pageId="5" pageNumber="196" type="etymology">
<paragraph id="8BD136AFFFC4EC37FF2DD6A9FE9BFF68" blockId="5.[151,1436,151,357]" pageId="5" pageNumber="196">
<emphasis id="B91AEABDFFC4EC37FF2DD6A9FEBAFF0D" bold="true" box="[199,326,151,176]" pageId="5" pageNumber="196">Etymology</emphasis>
. Derived from area of origin of the new species, the 
<taxonomicName id="4C6E4D2CFFC4EC37FC59D6A9FC08FF0D" box="[947,1012,151,176]" class="Insecta" family="Melolonthidae" genus="Tanyproctus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="195" phylum="Arthropoda" rank="species" species="arher">Arher</taxonomicName>
freshwater spring, 
<collectingCountry id="F379763FFFC4EC37FB3ED6A6FAD5FF0D" box="[1236,1321,152,176]" name="Yemen" pageId="5" pageNumber="196">Socotra</collectingCountry>
(
<collectingCountry id="F379763FFFC4EC37FAD1D6A6FA6CFF0D" box="[1339,1424,152,176]" name="Yemen" pageId="5" pageNumber="196">Yemen</collectingCountry>
); noun in appositon.
</paragraph>
<paragraph id="8BD136AFFFC4EC37FF2DD6E1FEAAFEA0" blockId="5.[151,1436,151,357]" pageId="5" pageNumber="196">
<emphasis id="B91AEABDFFC4EC37FF2DD6E1FE03FF45" bold="true" box="[199,511,223,248]" pageId="5" pageNumber="196">Collecting circumstances.</emphasis>
Both specimens were captured at light at about 
<specimenCount id="9D68FD26FFC4EC37FBB4D6E1FB73FF4A" box="[1118,1167,223,248]" pageId="5" pageNumber="196" type="generic">6 p.</specimenCount>
m. (J. Hájek, personal communication).
</paragraph>
</subSubSection>
<subSubSection id="C3746524FFC4EC37FF2DD719FBEFFED9" pageId="5" pageNumber="196" type="description">
<paragraph id="8BD136AFFFC4EC37FF2DD719FBEFFED9" blockId="5.[151,1436,151,357]" pageId="5" pageNumber="196">
<emphasis id="B91AEABDFFC4EC37FF2DD719FE03FEFD" bold="true" box="[199,511,295,320]" pageId="5" pageNumber="196">Geographical distribution</emphasis>
. Endemic species to the 
<collectingCountry id="F379763FFFC4EC37FCCBD716FC8BFEFD" box="[801,887,296,320]" name="Yemen" pageId="5" pageNumber="196">Socotra</collectingCountry>
Island; both so far known specimens originates from the coastal area of Haalla (E 
<collectingCountry id="F379763FFFC4EC37FDF2D772FD8EFED9" box="[536,626,332,356]" name="Yemen" pageId="5" pageNumber="196">Socotra</collectingCountry>
), for details see Bezděk 
<emphasis id="B91AEABDFFC4EC37FC68D773FC46FED8" box="[898,954,332,357]" italics="true" pageId="5" pageNumber="196">et al.</emphasis>
(2012).
</paragraph>
</subSubSection>
</treatment>
</document>