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<mods:title id="5A28063A953538DD822E42C5184E3F4C">Notes on two species of the cavernicolous subgenus Neobisium (Blothrus) Schiödte, 1847 (Arachnida: Pseudoscorpiones) from Transylvania (Romania), with a key to the species of the Carpathian Mountains</mods:title>
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<treatment id="03C787B5B945FF842694FF1CFE90F80E" ID-DOI="http://doi.org/10.5281/zenodo.6141588" ID-GBIF-Taxon="119427244" ID-Zenodo-Dep="6141588" LSID="urn:lsid:plazi:treatment:03C787B5B945FF842694FF1CFE90F80E" httpUri="http://treatment.plazi.org/id/03C787B5B945FF842694FF1CFE90F80E" lastPageNumber="398" pageId="4" pageNumber="398">
<subSubSection id="C3746528B945FF842694FF1CFE8DFF4E" pageId="4" pageNumber="398" type="nomenclature">
<paragraph id="8BD136A3B945FF842694FF1CFD64FF2F" blockId="4.[151,767,151,211]" box="[151,767,151,178]" pageId="4" pageNumber="398">
<heading id="D09981CFB945FF842694FF1CFD64FF2F" bold="true" box="[151,767,151,178]" fontSize="11" level="1" pageId="4" pageNumber="398" reason="1">
<taxonomicName id="4C6E4D20B945FF842694FF1CFD64FF2F" ID-CoL="7VYWH" authority="Frivaldszky, 1865" authorityName="Frivaldszky" authorityYear="1865" box="[151,767,151,178]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="398" phylum="Arthropoda" rank="species" species="brevipes" subGenus="Blothrus">
<emphasis id="B91AEAB1B945FF842694FF1CFD64FF2F" bold="true" box="[151,767,151,178]" pageId="4" pageNumber="398">
<emphasis id="B91AEAB1B945FF842694FF1CFD93FF2C" bold="true" box="[151,520,151,177]" italics="true" pageId="4" pageNumber="398">Neobisium (Blothrus) brevipes</emphasis>
(
<bibRefCitation id="EFFF4B52B945FF842414FF13FD6CFF2F" author="Frivaldszky" box="[535,759,152,178]" pageId="4" pageNumber="398" refString="Frivaldszky, J. (1865) Adatok a magyarhoni barlangok faunajahoz. Mathematikai es Termeszettudomanyi Kozlemenyek, Vonatkozolag a Hazai Viszonyokra, 3, 17 - 53." type="journal article" year="1865">Frivaldszky, 1865</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD136A3B945FF842694FF31FE8DFF4E" blockId="4.[151,767,151,211]" box="[151,278,186,211]" pageId="4" pageNumber="398">
(
<figureCitation id="13552A26B945FF84269CFF31FF7BFF4E" box="[159,224,186,211]" captionStart="FIGURE 1" captionStartId="2.[151,250,1843,1865]" captionTargetBox="[163,1402,334,1823]" captionTargetId="figure@2.[151,1436,317,1829]" captionTargetPageId="2" captionText="FIGURE 1. Neobisium (Blothrus) minutum and N. (B.) brevipes. A: Carapace of N. (B.) minutum (male, HNHM Pseud- 1389); B: Left chelicera of N. (B.) minutum with seven setae (male, HNHM Pseud- 1388); C: Left chelicera of N. (B.) minutum with six setae (male, HNHM Pseud- 1390); D: Right pedipalp of N. (B.) minutum (male, HNHM Pseud- 1389); E: Right chela of N. (B.) minutum (male, HNHM Pseud- 1388); F: Carapace of N. (B.) brevipes (male, HNHM Pseud- 1392); G: Left chelicera of N. (B.) brevipes (female, HNHM Pseud- 1391); H: Left pedipalp of N. (B.) brevipes (male, HNHM Pseud- 1392); I: Right chela of N. (B.) brevipes (male, HNHM Pseud- 1392)." httpUri="https://zenodo.org/record/231660/files/figure.png" pageId="4" pageNumber="398">Figs 1</figureCitation>
FI)
</paragraph>
</subSubSection>
<subSubSection id="C3746528B945FF842694FE89FF4DFEFE" pageId="4" pageNumber="398" type="materials_examined">
<paragraph id="8BD136A3B945FF842694FE89FF4DFEFE" blockId="4.[151,1437,258,1939]" pageId="4" pageNumber="398">
<emphasis id="B91AEAB1B945FF842694FE89FE1FFE86" bold="true" box="[151,388,258,283]" pageId="4" pageNumber="398">Material examined.</emphasis>
<collectionCode id="ED7FAE66B945FF84278CFE88FE7FFE87" LSID="urn:lsid:biocol.org:col:34991" box="[399,484,259,282]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="4" pageNumber="398">HNHM</collectionCode>
Pseud-1391 (female): 2339/1931 Tamsay Cave (Transylvania,
<collectingCountry id="F3797633B945FF8422C6FE89FAABFE86" box="[1221,1328,258,283]" name="Romania" pageId="4" pageNumber="398">Romania</collectingCountry>
);
<collectionCode id="ED7FAE66B945FF842344FE88FA07FE87" LSID="urn:lsid:biocol.org:col:34991" box="[1351,1436,259,282]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="4" pageNumber="398">HNHM</collectionCode>
Pseud-1392 (male): 1282/1812 Rézbányai Cave, Bihar County (Transylvania,
<collectingCountry id="F3797633B945FF84221FFEADFB1DFEA2" box="[1052,1158,294,319]" name="Romania" pageId="4" pageNumber="398">Romania</collectingCountry>
), leg. Dr Elemér Bokor, 1912.
</paragraph>
</subSubSection>
<subSubSection id="C3746528B945FF8426C4FEE5FD74FACF" pageId="4" pageNumber="398" type="description">
<paragraph id="8BD136A3B945FF8426C4FEE5FAFDFE36" blockId="4.[151,1437,258,1939]" pageId="4" pageNumber="398">
<emphasis id="B91AEAB1B945FF8426C4FEE5FECAFE1A" bold="true" box="[199,337,366,391]" pageId="4" pageNumber="398">Description</emphasis>
. Medium-sized pseudoscorpions with troglomorphic habitus. Carapace, chelicerae and pedipalps reddish brown, opisthosoma and legs yellowish. Measurements and ratios of the new specimens as in
<tableCitation id="C6EC0318B945FF84230FFE19FAFBFE36" box="[1292,1376,402,427]" captionStart="TABLE 1" captionStartId="3.[151,239,152,175]" captionText="TABLE 1. Measurements and ratios of Neobisium (Blothrus) minutum (HNHM Pseud- 0051, Pseud- 1388 1390) and Neobisium (Blothrus) brevipes (HNHM Pseud- 1391 1392). Legs I and IV of the type of N. (B.) minutum were not measured, due to the state of the specimen." pageId="4" pageNumber="398">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BD136A3B945FF8426C4FE3CFF56FD8A" blockId="4.[151,1437,258,1939]" pageId="4" pageNumber="398">
Carapace (
<figureCitation id="13552A26B945FF84273CFE3DFEE5FE52" box="[319,382,438,463]" captionStart="FIGURE 1" captionStartId="2.[151,250,1843,1865]" captionTargetBox="[163,1402,334,1823]" captionTargetId="figure@2.[151,1436,317,1829]" captionTargetPageId="2" captionText="FIGURE 1. Neobisium (Blothrus) minutum and N. (B.) brevipes. A: Carapace of N. (B.) minutum (male, HNHM Pseud- 1389); B: Left chelicera of N. (B.) minutum with seven setae (male, HNHM Pseud- 1388); C: Left chelicera of N. (B.) minutum with six setae (male, HNHM Pseud- 1390); D: Right pedipalp of N. (B.) minutum (male, HNHM Pseud- 1389); E: Right chela of N. (B.) minutum (male, HNHM Pseud- 1388); F: Carapace of N. (B.) brevipes (male, HNHM Pseud- 1392); G: Left chelicera of N. (B.) brevipes (female, HNHM Pseud- 1391); H: Left pedipalp of N. (B.) brevipes (male, HNHM Pseud- 1392); I: Right chela of N. (B.) brevipes (male, HNHM Pseud- 1392)." httpUri="https://zenodo.org/record/231660/files/figure.png" pageId="4" pageNumber="398">Fig. 1</figureCitation>
F) somewhat longer than broad. Epistome triangular and pointed. Eyes or eyespots absent. A pair of ocular protuberances present, each bearing one seta. Preocular setae absent. Chaetotaxy of carapace: 4:6:6:6 (22).
</paragraph>
<paragraph id="8BD136A3B945FF8426C4FDA9FD8CFD1E" blockId="4.[151,1437,258,1939]" pageId="4" pageNumber="398">Coxal area. Manducatory process with 35 setae; rest of pedipalpal coxa with 911 setae; pedal coxa I with 58 setae; II: 78; III: 67; IV: 79. Anterolateral process of coxa I long, triangular and apically pointed; medial process prominent, with denticles.</paragraph>
<paragraph id="8BD136A3B945FF8426C4FD05FDFAFCAA" blockId="4.[151,1437,258,1939]" pageId="4" pageNumber="398">Opisthosoma with granulostriate pleural membrane. Chaetotaxy of tergites IX: 6:6:7:8:8:8:9:8:8:8. Segment XI with 12 setae, at least 2 of them tactile. Anal cone with 2 ventral and 2 dorsal setae. Male genital region: sternite II with 11 setae, sternite III with 17 small setae, 8 of them along posterior margin. Female genital region: sternite II with 6 small setae, sternite III with 12 small setae in a row. Chaetotaxy of sternites IVX: 1315:812:11:1014:813:1113:812.</paragraph>
<paragraph id="8BD136A3B945FF8426C4FCC9FD0AFB92" blockId="4.[151,1437,258,1939]" pageId="4" pageNumber="398">
Chelicera (
<figureCitation id="13552A26B945FF842745FCC9FE13FCC6" box="[326,392,834,859]" captionStart="FIGURE 1" captionStartId="2.[151,250,1843,1865]" captionTargetBox="[163,1402,334,1823]" captionTargetId="figure@2.[151,1436,317,1829]" captionTargetPageId="2" captionText="FIGURE 1. Neobisium (Blothrus) minutum and N. (B.) brevipes. A: Carapace of N. (B.) minutum (male, HNHM Pseud- 1389); B: Left chelicera of N. (B.) minutum with seven setae (male, HNHM Pseud- 1388); C: Left chelicera of N. (B.) minutum with six setae (male, HNHM Pseud- 1390); D: Right pedipalp of N. (B.) minutum (male, HNHM Pseud- 1389); E: Right chela of N. (B.) minutum (male, HNHM Pseud- 1388); F: Carapace of N. (B.) brevipes (male, HNHM Pseud- 1392); G: Left chelicera of N. (B.) brevipes (female, HNHM Pseud- 1391); H: Left pedipalp of N. (B.) brevipes (male, HNHM Pseud- 1392); I: Right chela of N. (B.) brevipes (male, HNHM Pseud- 1392)." httpUri="https://zenodo.org/record/231660/files/figure.png" pageId="4" pageNumber="398">Fig. 1</figureCitation>
G) with 6 setae on hand, subgaleal seta 0.65× from base of movable finger. Spinneret a flattened hyaline tubercle with 46 silk ducts. Fixed finger with 89 small and medium sized teeth, the apical denticles could not be investigated due to the poor condition of the material. Movable finger with 5 apical denticles and one large tooth in middle, other teeth not clearly visible. Rallum with 8 blades, two distalmost blades unilaterally pinnate on anterior face and placed on dilated base, separate from the others. Serrula exterior with 2225 blades, serrula interior with 23 blades.
</paragraph>
<paragraph id="8BD136A3B945FF8426C4FB91FB50FB7B" blockId="4.[151,1437,258,1939]" pageId="4" pageNumber="398">
Pedipalp (
<figureCitation id="13552A26B945FF84273BFB91FEECFBAE" box="[312,375,1050,1075]" captionStart="FIGURE 1" captionStartId="2.[151,250,1843,1865]" captionTargetBox="[163,1402,334,1823]" captionTargetId="figure@2.[151,1436,317,1829]" captionTargetPageId="2" captionText="FIGURE 1. Neobisium (Blothrus) minutum and N. (B.) brevipes. A: Carapace of N. (B.) minutum (male, HNHM Pseud- 1389); B: Left chelicera of N. (B.) minutum with seven setae (male, HNHM Pseud- 1388); C: Left chelicera of N. (B.) minutum with six setae (male, HNHM Pseud- 1390); D: Right pedipalp of N. (B.) minutum (male, HNHM Pseud- 1389); E: Right chela of N. (B.) minutum (male, HNHM Pseud- 1388); F: Carapace of N. (B.) brevipes (male, HNHM Pseud- 1392); G: Left chelicera of N. (B.) brevipes (female, HNHM Pseud- 1391); H: Left pedipalp of N. (B.) brevipes (male, HNHM Pseud- 1392); I: Right chela of N. (B.) brevipes (male, HNHM Pseud- 1392)." httpUri="https://zenodo.org/record/231660/files/figure.png" pageId="4" pageNumber="398">Fig. 1</figureCitation>
H). Surfaces of articles smooth, but femur bears some small tubercles. Trochanter with a small tubercle near distal end. Femur and patella elongated. Chelal fingers 1.611.75 times longer than hand with pedicel. Fixed finger with 145150 close set, equally long teeth. Movable finger 130140 teeth, those in distal third similar to those of fixed finger, teeth in proximal two thirds flattened. Dental line on movable finger ending somewhat distal to trichobothrium
<emphasis id="B91AEAB1B945FF842422FB20FDB4FB5E" box="[545,559,1195,1219]" italics="true" pageId="4" pageNumber="398">b</emphasis>
. Trichobothria on chelal fingers distributed as in
<figureCitation id="13552A26B945FF84225DFB21FB20FB5E" box="[1118,1211,1194,1219]" captionStart="FIGURE 1" captionStartId="2.[151,250,1843,1865]" captionTargetBox="[163,1402,334,1823]" captionTargetId="figure@2.[151,1436,317,1829]" captionTargetPageId="2" captionText="FIGURE 1. Neobisium (Blothrus) minutum and N. (B.) brevipes. A: Carapace of N. (B.) minutum (male, HNHM Pseud- 1389); B: Left chelicera of N. (B.) minutum with seven setae (male, HNHM Pseud- 1388); C: Left chelicera of N. (B.) minutum with six setae (male, HNHM Pseud- 1390); D: Right pedipalp of N. (B.) minutum (male, HNHM Pseud- 1389); E: Right chela of N. (B.) minutum (male, HNHM Pseud- 1388); F: Carapace of N. (B.) brevipes (male, HNHM Pseud- 1392); G: Left chelicera of N. (B.) brevipes (female, HNHM Pseud- 1391); H: Left pedipalp of N. (B.) brevipes (male, HNHM Pseud- 1392); I: Right chela of N. (B.) brevipes (male, HNHM Pseud- 1392)." httpUri="https://zenodo.org/record/231660/files/figure.png" pageId="4" pageNumber="398">
Figure
<date id="FFD01063B945FF8422ACFB20FB20FB5E" box="[1199,1211,1195,1219]" pageId="4" pageNumber="398">1</date>
</figureCitation>
I. Distance between trichobothria
<emphasis id="B91AEAB1B945FF84272DFB45FEDFFB7B" box="[302,324,1230,1254]" italics="true" pageId="4" pageNumber="398">ib</emphasis>
and
<emphasis id="B91AEAB1B945FF842778FB44FE0DFB7B" box="[379,406,1231,1254]" italics="true" pageId="4" pageNumber="398">ist</emphasis>
more than two and a half times longer (2.67) than that between
<emphasis id="B91AEAB1B945FF842267FB44FBE5FB7B" box="[1124,1150,1231,1254]" italics="true" pageId="4" pageNumber="398">ist</emphasis>
and
<emphasis id="B91AEAB1B945FF8422B7FB44FB58FB7B" box="[1204,1219,1231,1254]" italics="true" pageId="4" pageNumber="398">it</emphasis>
.
</paragraph>
<paragraph id="8BD136A3B945FF8426C4FB78FD74FACF" blockId="4.[151,1437,258,1939]" pageId="4" pageNumber="398">Legs distinctly elongated, with smooth surfaces. Claws smooth and simple, arolia shorter than claws on all legs. Subterminal seta of telotarsus IV with three rami, the two longer ones serrate, the shortest smooth. Basitarsus IV with one, telotarsus IV with two long tactile setae.</paragraph>
</subSubSection>
<subSubSection id="C3746528B945FF8426C4FAD5FE90F80E" pageId="4" pageNumber="398" type="discussion">
<paragraph id="8BD136A3B945FF8426C4FAD5FB31F942" blockId="4.[151,1437,258,1939]" pageId="4" pageNumber="398">
<emphasis id="B91AEAB1B945FF8426C4FAD5FEA0FAEA" bold="true" box="[199,315,1374,1399]" pageId="4" pageNumber="398">Remarks.</emphasis>
<bibRefCitation id="EFFF4B52B945FF842742FAD5FD8DFAEA" author="Frivaldszky" box="[321,534,1374,1399]" pageId="4" pageNumber="398" refString="Frivaldszky, J. (1865) Adatok a magyarhoni barlangok faunajahoz. Mathematikai es Termeszettudomanyi Kozlemenyek, Vonatkozolag a Hazai Viszonyokra, 3, 17 - 53." type="journal article" year="1865">Frivaldszky (1865)</bibRefCitation>
mentioned Oncsásza Cave (now Pesterea Zemeilor din Onceasa, near the village of Beius,
<collectingCountry id="F3797633B945FF842708FA09FEE9FA06" box="[267,370,1410,1435]" name="Romania" pageId="4" pageNumber="398">Romania</collectingCountry>
) as the
<typeStatus id="54D58801B945FF8427D0FA08FD98FA06" box="[467,515,1411,1435]" pageId="4" pageNumber="398">type</typeStatus>
locality, but gave no data about where the
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material was deposited. Later,
<bibRefCitation id="EFFF4B52B945FF842694FA2CFEF6FA22" author="Tomosvary" box="[151,365,1447,1471]" pageId="4" pageNumber="398" refString="Tomosvary, O. (1882) A Magyar fauna alskorpioi. Magyar Tudomanyos Akademia Matematikai es Termeszettudomanyi Kozlemenyek, 18, 135 - 256." type="journal article" year="1882">Tömösváry (1882)</bibRefCitation>
reported further localities (Caves at Fericse, Pestere, Mehadia and Szelistye villages) and referred to the HNHM as the place of deposition. In addition, Beier reported the species from the Tatarczy Cave at Herkulesbad (
<bibRefCitation id="EFFF4B52B945FF842730FA65FE2CF99A" author="Beier" box="[307,439,1518,1543]" pageId="4" pageNumber="398" refString="Beier, M. (1928) Die Pseudoskorpione des Wiener Naturhistorischen Museums. I. Hemictenodactyli. Annalen des Naturhistorischen Museums in Wien, 42, 285 - 314." type="journal article" year="1928">Beier, 1928</bibRefCitation>
) and from caves near the village of Vascau (
<bibRefCitation id="EFFF4B52B945FF8425A9FA65FBAAF99A" author="Beier" box="[938,1073,1518,1543]" pageId="4" pageNumber="398" refString="Beier, M. (1939) Pseudoscorpionidea de Roumanie. Bulletin du Musee Royal d'Histoire Naturelle de Belgique, 15 (39), 1 - 21." type="journal article" year="1939">Beier, 1939</bibRefCitation>
). Unfortunately, neither the
<typeStatus id="54D58801B945FF84236FFA64FA07F99A" box="[1388,1436,1519,1543]" pageId="4" pageNumber="398">type</typeStatus>
material nor any other identified specimens of this species could be found in the collection of the HNHM, having probably been lost or destroyed when the Department of Zoology burnt down in 1956. Examination of the two
<taxonomicName id="4C6E4D20B945FF842381F9BCFEB7F9EE" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="398" phylum="Arthropoda" rank="species" species="brevipes" subGenus="Blothrus">
<emphasis id="B91AEAB1B945FF842381F9BCFEB7F9EE" italics="true" pageId="4" pageNumber="398">N. (B.) brevipes</emphasis>
</taxonomicName>
specimens studied here revealed two important characteristics not previously mentioned in the literature. As in
<taxonomicName id="4C6E4D20B945FF84274DF9F4FD9FF90B" box="[334,516,1663,1686]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="398" phylum="Arthropoda" rank="species" species="minutum" subGenus="Blothrus">
<emphasis id="B91AEAB1B945FF84274DF9F4FD9FF90B" box="[334,516,1663,1686]" italics="true" pageId="4" pageNumber="398">N. (B.) minutum</emphasis>
</taxonomicName>
, the carapace bears two ocular protuberances. Furthermore, a few small tubercles are present on the pedipalpal femora. The same characteristics were also found in the specimens identified by Beier and deposited at the Natural History Museum Vienna (Inventory numbers 27067 and 27102).
</paragraph>
<paragraph id="8BD136A3B945FF8426C4F961FE90F80E" blockId="4.[151,1437,258,1939]" pageId="4" pageNumber="398">
Considering the important differences between
<taxonomicName id="4C6E4D20B945FF8424DFF960FC6AF89E" box="[732,1009,1771,1795]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="398" phylum="Arthropoda" rank="subSpecies" species="brevipes" subGenus="Blothrus" subSpecies="brevipes">
<emphasis id="B91AEAB1B945FF8424DFF960FC6AF89E" box="[732,1009,1771,1795]" italics="true" pageId="4" pageNumber="398">N. (B.) brevipes brevipes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6E4D20B945FF84222BF960FACEF89F" box="[1064,1365,1771,1795]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="398" phylum="Arthropoda" rank="subSpecies" species="brevipes" subGenus="Blothrus" subSpecies="montanum">
<emphasis id="B91AEAB1B945FF84222BF960FACEF89F" box="[1064,1365,1771,1795]" italics="true" pageId="4" pageNumber="398">N. (B.) brevipes montanum</emphasis>
</taxonomicName>
in the measurements and ratios of the pedipalps, and the arrangement of the trichobothria of the fixed chelal finger, it is warranted to elevate
<taxonomicName id="4C6E4D20B945FF842791F8B8FD4BF8D7" box="[402,720,1843,1867]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="398" phylum="Arthropoda" rank="subSpecies" species="brevipes" subGenus="Blothrus" subSpecies="montanum">
<emphasis id="B91AEAB1B945FF842791F8B8FD4BF8D7" box="[402,720,1843,1867]" italics="true" pageId="4" pageNumber="398">N. (B.) brevipes montanum</emphasis>
</taxonomicName>
to full species rank. Consequently, the author of this paper proposes using the names
<taxonomicName id="4C6E4D20B945FF8427BEF8DDFC6DF8F2" authority="Frivaldszky, 1865" authorityName="Frivaldszky" authorityYear="1865" box="[445,1014,1878,1903]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="398" phylum="Arthropoda" rank="species" species="brevipes" subGenus="Blothrus">
<emphasis id="B91AEAB1B945FF8427BEF8DDFC8FF8F3" box="[445,788,1878,1902]" italics="true" pageId="4" pageNumber="398">Neobisium (Blothrus) brevipes</emphasis>
(
<bibRefCitation id="EFFF4B52B945FF842520F8DDFC75F8F2" author="Frivaldszky" box="[803,1006,1878,1903]" pageId="4" pageNumber="398" refString="Frivaldszky, J. (1865) Adatok a magyarhoni barlangok faunajahoz. Mathematikai es Termeszettudomanyi Kozlemenyek, Vonatkozolag a Hazai Viszonyokra, 3, 17 - 53." type="journal article" year="1865">Frivaldszky, 1865</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C6E4D20B945FF84222EF8DCFAE4F8F2" authority="Beier, 1939" authorityName="Beier" authorityYear="1939" box="[1069,1407,1878,1903]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="398" phylum="Arthropoda" rank="species" species="montanum" subGenus="Blothrus">
<emphasis id="B91AEAB1B945FF84222EF8DCFB6DF8F3" box="[1069,1270,1879,1902]" italics="true" pageId="4" pageNumber="398">N. (B.) montanum</emphasis>
<bibRefCitation id="EFFF4B52B945FF8422FEF8DDFAE4F8F2" author="Beier" box="[1277,1407,1878,1903]" pageId="4" pageNumber="398" refString="Beier, M. (1939) Pseudoscorpionidea de Roumanie. Bulletin du Musee Royal d'Histoire Naturelle de Belgique, 15 (39), 1 - 21." type="journal article" year="1939">
Beier,
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</bibRefCitation>
</taxonomicName>
in the future.
</paragraph>
</subSubSection>
</treatment>
</document>