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<document id="3F513B558C98B6FB25A5948DCF3CB51C" ID-CLB-Dataset="22877" ID-DOI="10.5852/ejt.2020.629" ID-GBIF-Dataset="3722aa65-b935-4294-ac20-0d1179da9015" ID-Zenodo-Dep="3762280" ID-ZooBank="ECEBD539-6E3E-45BE-A0CB-264DF3270CC0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587578640316" checkinUser="valdenar" docAuthor="Herbert, David G." docDate="2020" docId="03C78785462BFFEA422040F83237FC6D" docLanguage="en" docName="ejt-629_herbert.pdf.imf" docOrigin="European Journal of Taxonomy 629" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Phialodonta atromontana Herbert 2020, gen. et sp. nov." docType="treatment" docUuid="4A34E0A1-99C9-4892-83DC-080F239C0921" docUuidSource="ZooBank" docVersion="7" lastPageNumber="41" masterDocId="FFFEFFFD460CFFC2400243213138FFD9" masterDocTitle="Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies" masterLastPageNumber="55" masterPageNumber="1" pageNumber="40" updateTime="1698835565728" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="FF22F8BE1C2408661B37CB2F5FE433FF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C841267C302801D916C080C4C0B3DE2B">
<mods:title id="3762844B187ADDF246D035CE01CEB019">Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies</mods:title>
</mods:titleInfo>
<mods:name id="6D4FB5BEF1A505932ABFFA2E0B3ECBEC" type="personal">
<mods:role id="DB92EFF97337AA1A758F03E68B5324EC">
<mods:roleTerm id="3D1F01230210D9BBAA9A94F966017130">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DBC440DBDFF3832A0DE577E207643E8B">Herbert, David G.</mods:namePart>
<mods:nameIdentifier id="99707BCC55F4860676D8F3B867BD7C34" type="LSID">urn:lsid:zoobank.org:author:0C09EE45-6198-482E-857A-EF690C2A016F</mods:nameIdentifier>
<mods:affiliation id="3DB55CAF02BD779AF14F4D8C4E65B8FE">Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff, CF 10 3 NP, UK. Formerly at KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg 3201, South Africa</mods:affiliation>
<mods:nameIdentifier id="79200D49E8CB55DEDE4868A4C88DCC09" type="email">phasianella@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="E933CA3C740BFCD87299DB81D7C79C49">text</mods:typeOfResource>
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<mods:titleInfo id="A4E07D6DDF1AD96FADBFB8A5594BC615">
<mods:title id="7468BC298C1ACE5BDA3832BAA99C6D01">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="00412834BFC92B95CE57EEA4090071FE">
<mods:date id="8F317D377CCE13E196325F348712EE94">2020</mods:date>
<mods:detail id="282AC93284E36D18119A026C96F5B184" type="pubDate">
<mods:number id="63CD37FCF2C506DC839E83BE86AD91FB">2020-04-17</mods:number>
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<mods:detail id="E27271221BDD95BB90F75BB5339D04A3" type="volume">
<mods:number id="1CB66D3C6E442B286AFDD12A116C7F03">629</mods:number>
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<mods:classification id="3CC5E960D673A667B32ED1634A82FBEE">journal article</mods:classification>
<mods:identifier id="F8EABC88721390DF0007C5B5E9F026E6" type="CLB-Dataset">22877</mods:identifier>
<mods:identifier id="7BE562580DEA063F8368350BC14D540B" type="DOI">10.5852/ejt.2020.629</mods:identifier>
<mods:identifier id="B49A3DEA4D1105C979BFDDAD38AB7E46" type="GBIF-Dataset">3722aa65-b935-4294-ac20-0d1179da9015</mods:identifier>
<mods:identifier id="649C3D26F8D48D3DDD7410B58FCCECC7" type="Zenodo-Dep">3762280</mods:identifier>
<mods:identifier id="88CE2E82F02558A3074D9B8BBD1149BF" type="ZooBank">ECEBD539-6E3E-45BE-A0CB-264DF3270CC0</mods:identifier>
</mods:mods>
<treatment id="03C78785462BFFEA422040F83237FC6D" ID-DOI="http://doi.org/10.5281/zenodo.3804758" ID-GBIF-Taxon="163732359" ID-Zenodo-Dep="3804758" LSID="urn:lsid:zoobank.org:act:4A34E0A1-99C9-4892-83DC-080F239C0921" httpUri="http://treatment.plazi.org/id/03C78785462BFFEA422040F83237FC6D" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<subSubSection id="C3746518462BFFE5422040F83586FBCE" pageId="39" pageNumber="40" type="nomenclature">
<paragraph id="8BD13693462BFFE5422040F8352AFC2D" blockId="39.[373,1214,985,1084]" box="[546,1042,985,1012]" pageId="39" pageNumber="40">
<heading id="D09981FF462BFFE5422040F8352AFC2D" box="[546,1042,985,1012]" centered="true" fontSize="11" level="2" pageId="39" pageNumber="40" reason="2">
<taxonomicName id="4C6E4D10462BFFE5422040F83262FC2D" ID-CoL="7SP52" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[546,858,985,1012]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="atromontana" status="gen. et sp. nov.">
<emphasis id="B91AEA81462BFFE5422040F83262FC2D" bold="true" box="[546,858,985,1012]" italics="true" pageId="39" pageNumber="40">Phialodonta atromontana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA462BFFE5436040FB352AFC2D" box="[866,1042,986,1012]" pageId="39" pageNumber="40" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<uri id="FFFF3A91462BFFE5417740DC3586FBCE" box="[373,1214,1021,1047]" pageId="39" pageNumber="40">
urn:lsid:zoobank.org:act:
<uuid id="FFC80C46462BFFE5429440DC3586FBCE" box="[662,1214,1021,1047]" pageId="39" pageNumber="40">4A34E0A1-99C9-4892-83DC-080F239C0921</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3746518462BFFE5428E47003290FBE2" box="[652,936,1057,1084]" pageId="39" pageNumber="40" type="description">
<paragraph id="8BD13693462BFFE5428E47003290FBE2" blockId="39.[373,1214,985,1084]" box="[652,936,1057,1084]" pageId="39" pageNumber="40">
<figureCitation id="13552A16462BFFE5428E470033DDFBE2" box="[652,741,1057,1083]" captionStart="Fig" captionStartId="36.[189,232,1860,1886]" captionTargetBox="[302,1285,1199,1820]" captionTargetId="figure@36.[302,1285,1199,1820]" captionTargetPageId="36" captionText="Fig. 14. Distribution of Iterodonta gen. nov. and Phialodonta gen. nov. species. Iterodonta ammonita gen. et sp. nov. (brown triangles), Phialodonta agulhasae gen. et sp. nov. (orange square), P. atromontana gen. et sp. nov. (turquoise square), P. aviana gen. et sp. nov. (royal blue squares), P. introtuberculata (Connolly, 1933) gen. et comb. nov. (red squares), P. perfida (Burnup, 1912) gen. et comb. nov. (green square), P. rivalalea gen. et sp. nov. (pink squares). Contour at 1000 m." figureDoi="http://doi.org/10.5281/zenodo.3762308" httpUri="https://zenodo.org/record/3762308/files/figure.png" pageId="39" pageNumber="40">Figs 14</figureCitation>
,
<figureCitation id="13552A16462BFFE542F347003234FBE5" box="[753,780,1057,1084]" captionStart="Fig" captionStartId="38.[189,232,1860,1886]" captionTargetBox="[192,1391,514,1814]" captionTargetId="figure@38.[189,1399,499,1820]" captionTargetPageId="38" captionText="Fig. 15. Shells of Phialodonta gen. nov. species. AD. P. agulhasae gen. et sp. nov., holotype, diameter 1.30 mm (NMSA W7816/T4257). EH. P. atromontana gen. et sp. nov., holotype, diameter 1.40 mm (NMSA W9580/T4254). IL. P. aviana gen. et sp. nov., holotype, diameter 1.40 mm (NMSA W624/ T4246). M. P. aviana gen. et sp. nov., specimen with palatal region broken back to reveal apertural dentition, paratype, Touwsrivier, Wilderness, W. Cape, diameter 1.32 mm (NMSA W3511/T4248)." figureDoi="http://doi.org/10.5281/zenodo.3762310" httpUri="https://zenodo.org/record/3762310/files/figure.png" pageId="39" pageNumber="40">15</figureCitation>
EH, 19CD
</paragraph>
</subSubSection>
<subSubSection id="C3746518462BFFE540BF47433005FA87" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="8BD13693462BFFE540BF4743300EFBA5" blockId="39.[189,310,1122,1148]" box="[189,310,1122,1148]" pageId="39" pageNumber="40">
<heading id="D09981FF462BFFE540BF4743300EFBA5" bold="true" box="[189,310,1122,1148]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
<emphasis id="B91AEA81462BFFE540BF4743300EFBA5" bold="true" box="[189,310,1122,1148]" pageId="39" pageNumber="40">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693462BFFE540BF47B03005FA87" blockId="39.[189,1399,1169,1374]" pageId="39" pageNumber="40">
Shell very small, spire slightly raised; protoconch lacking axial sculpture; teleoconch sculpture of curved, close-set, compound, axial riblets, with 45 finer, intermediary axial threads; spiral sculpture of extremely fine, close-set threads; aperture with a low, rounded, in-running parietal ridge, lacking basocolumellar dentition; palatal region with two pairs of close-set, spirally aligned denticles, one pair close to mid-whorl, the other basal; umbilicus wide. Shell pale corneous-brown to honey-brown; diameter up to
<quantity id="4C969B76462BFFE540D946653000FA87" box="[219,312,1348,1374]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="39" pageNumber="40" unit="mm" value="1.4">1.4 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3746518462BFFE540BF46A73418FA16" box="[189,1312,1414,1487]" pageId="39" pageNumber="40" type="etymology">
<paragraph id="8BD13693462BFFE540BF46A7307CFA79" blockId="39.[189,324,1414,1440]" box="[189,324,1414,1440]" pageId="39" pageNumber="40">
<heading id="D09981FF462BFFE540BF46A7307CFA79" bold="true" box="[189,324,1414,1440]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
<emphasis id="B91AEA81462BFFE540BF46A7307CFA79" bold="true" box="[189,324,1414,1440]" pageId="39" pageNumber="40">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693462BFFE540BF46943418FA16" blockId="39.[189,1312,1460,1487]" box="[189,1312,1460,1487]" pageId="39" pageNumber="40">
From the Latin
<emphasis id="B91AEA81462BFFE541764694309BFA17" box="[372,419,1461,1486]" italics="true" pageId="39" pageNumber="40">ater</emphasis>
: black, and
<emphasis id="B91AEA81462BFFE5423146943349FA17" box="[563,625,1461,1486]" italics="true" pageId="39" pageNumber="40">mons</emphasis>
: a mountain; with reference to the Groot Swartberg range.
</paragraph>
</subSubSection>
<subSubSection id="C3746518462BFFE540BF46D735C7F8C9" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="8BD13693462BFFE540BF46D73092F9C9" blockId="39.[189,426,1526,1552]" box="[189,426,1526,1552]" pageId="39" pageNumber="40">
<heading id="D09981FF462BFFE540BF46D73092F9C9" bold="true" box="[189,426,1526,1552]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
<emphasis id="B91AEA81462BFFE540BF46D73092F9C9" bold="true" box="[189,426,1526,1552]" pageId="39" pageNumber="40">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693462BFFE540E845043064F9E6" blockId="39.[189,1399,1573,1707]" box="[234,348,1573,1599]" pageId="39" pageNumber="40">
<emphasis id="B91AEA81462BFFE540E845043064F9E6" bold="true" box="[234,348,1573,1599]" pageId="39" pageNumber="40">
<typeStatus id="54D58831462BFFE540E845043064F9E6" box="[234,348,1573,1599]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD13693462BFFE540BF456832DDF972" blockId="39.[189,1399,1573,1707]" pageId="39" pageNumber="40">
<materialsCitation id="3B063CCE462BFFE540BF456832DDF972" ID-GBIF-Occurrence="2605305607" collectedFrom="rocky hillside with succulents and aloes, in leaf-litter" collectingDate="2013-11-04" collectionCode="NMSA" collectorName="D. G. Herbert &amp; L. S. Davis &amp; M. and K. Cole" country="South Africa" elevation="360" latitude="-33.44525" location="Matjiesvlei" longLatPrecision="1" longitude="21.65388" municipality="Calitzdorp area" pageId="39" pageNumber="40" specimenCode="W9580/T4254" specimenCount="1" stateProvince="W. Cape" typeStatus="holotype">
<collectingCountry id="F3797603462BFFE540BF456830AAF9BA" box="[189,402,1609,1635]" name="South Africa" pageId="39" pageNumber="40">SOUTH AFRICA</collectingCountry>
<collectingRegion id="49AAF871462BFFE541B045683324F9BA" box="[434,540,1609,1635]" pageId="39" pageNumber="40">W. Cape</collectingRegion>
,
<collectingMunicipality id="6BB5ACE9462BFFE5422E456833DFF9BA" box="[556,743,1609,1635]" pageId="39" pageNumber="40">Calitzdorp area</collectingMunicipality>
,
<location id="8EB16048462BFFE542F445683246F9BA" LSID="urn:lsid:plazi:treatment:03C78785462BFFEA422040F83237FC6D:8EB16048462BFFE542F445683246F9BA" box="[758,894,1609,1635]" country="South Africa" latitude="-33.44525" longLatPrecision="1" longitude="21.65388" municipality="Calitzdorp area" name="Matjiesvlei" pageId="39" pageNumber="40" stateProvince="W. Cape">Matjiesvlei</location>
;
<geoCoordinate id="EE5A5054462BFFE5438F4569351CF9BA" box="[909,1060,1608,1635]" degrees="33.44525" direction="south" orientation="latitude" pageId="39" pageNumber="40" precision="1" value="-33.44525">33.44525° S</geoCoordinate>
,
<geoCoordinate id="EE5A5054462BFFE54437456935F7F9BA" box="[1077,1231,1608,1635]" degrees="21.65388" direction="east" orientation="longitude" pageId="39" pageNumber="40" precision="1" value="21.65388">21.65388° E</geoCoordinate>
;
<elevation id="0043D1A0462BFFE544DD45693448F9BA" box="[1247,1392,1608,1635]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="39" pageNumber="40" unit="m" value="360.0">
<quantity id="4C969B76462BFFE544DD45693415F9BA" box="[1247,1325,1608,1635]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="39" pageNumber="40" unit="m" value="360.0">360 m</quantity>
a.s.l.
</elevation>
;
<date id="FFD01053462BFFE540BF454D3077F95F" box="[189,335,1644,1671]" pageId="39" pageNumber="40" value="2013-11-04">
<collectingDate id="EF94E9BB462BFFE540BF454D3077F95F" box="[189,335,1644,1671]" pageId="39" pageNumber="40" value="2013-11-04">4 Nov. 2013</collectingDate>
</date>
;
<collectorName id="269B5345462BFFE54159454D30C0F95F" box="[347,504,1644,1670]" pageId="39" pageNumber="40">D.G. Herbert</collectorName>
,
<collectorName id="269B5345462BFFE54200454D33B9F95F" box="[514,641,1644,1670]" pageId="39" pageNumber="40">L.S. Davis</collectorName>
,
<collectorName id="269B5345462BFFE5428E454C3278F95F" box="[652,832,1644,1671]" pageId="39" pageNumber="40">M. and K. Cole</collectorName>
leg.;
<collectedFrom id="3872756E462BFFE54383454D3016F973" pageId="39" pageNumber="40">rocky hillside with succulents and aloes, in leaf-litter</collectedFrom>
; diameter
<quantity id="4C969B76462BFFE541AA45B1333DF973" box="[424,517,1680,1706]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="39" pageNumber="40" unit="mm" value="1.4">1.4 mm</quantity>
, height
<quantity id="4C969B76462BFFE5426345B133F4F973" box="[609,716,1680,1706]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="39" pageNumber="40" unit="mm" value="0.61">0.61 mm</quantity>
;
<collectionCode id="ED7FAE56462BFFE542DA45B13213F973" box="[728,811,1680,1706]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="39" pageNumber="40" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE8462BFFE5433045B132D9F972" box="[818,993,1680,1707]" pageId="39" pageNumber="40">W9580/T4254</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BD13693462BFFE540E845F3305FF935" blockId="39.[189,1279,1746,1808]" box="[234,359,1746,1772]" pageId="39" pageNumber="40">
<emphasis id="B91AEA81462BFFE540E845F3305FF935" bold="true" box="[234,359,1746,1772]" pageId="39" pageNumber="40">
<typeStatus id="54D58831462BFFE540E845F3305FF935" box="[234,359,1746,1772]" pageId="39" pageNumber="40" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD13693462BFFE540BF45D435C7F8C9" blockId="39.[189,1279,1746,1808]" box="[189,1279,1781,1808]" pageId="39" pageNumber="40">
<materialsCitation id="3B063CCE462BFFE540BF45D435C3F8C9" ID-GBIF-Occurrence="2605305609" box="[189,1275,1781,1808]" collectingDate="2013-11-04" collectionCode="NMSA" collectorName="D. G. Herbert &amp; L. S. Davis &amp; M. and K. Cole" country="South Africa" elevation="360" latitude="-33.44525" location="SOUTH AFRICA" longLatPrecision="1" longitude="21.65388" municipality="Calitzdorp area" pageId="39" pageNumber="40" specimenCode="P1016/T4255" specimenCount="7" stateProvince="Western Cape" typeStatus="paratype">
<collectingCountry id="F3797603462BFFE540BF45D430B6F8D6" box="[189,398,1781,1807]" name="South Africa" pageId="39" pageNumber="40">SOUTH AFRICA</collectingCountry>
<specimenCount id="9D68FD1A462BFFE541A545D73303F8D6" box="[423,571,1781,1808]" pageId="39" pageNumber="40" type="generic">7 specimens</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED7FAE56462BFFE543FF45D43568F8D6" box="[1021,1104,1781,1807]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="39" pageNumber="40" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE8462BFFE5445A45D435C3F8C9" box="[1112,1275,1781,1808]" pageId="39" pageNumber="40">P1016/T4255</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3746518462BFFEA40BF44163351FE25" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="description">
<paragraph id="8BD13693462BFFE540BF44163076F888" blockId="39.[189,334,1847,1873]" box="[189,334,1847,1873]" pageId="39" pageNumber="40">
<heading id="D09981FF462BFFE540BF44163076F888" bold="true" box="[189,334,1847,1873]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
<emphasis id="B91AEA81462BFFE540BF44163076F888" bold="true" box="[189,334,1847,1873]" pageId="39" pageNumber="40">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693462BFFEA40BF44473351FE25" blockId="39.[189,1399,1894,2027]" lastBlockId="40.[189,1399,267,508]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
Shell very small, diameter up to
<quantity id="4C969B76462BFFE542474447339BF859" box="[581,675,1894,1920]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="39" pageNumber="40" unit="mm" value="1.4">1.4 mm</quantity>
, H/D ratio ±0.43; spire slightly raised; whorls tightly coiled, relatively narrow; last adult whorl descending slightly below penultimate whorl; suture narrowly indented, periphery evenly convex. Protoconch comprising apical cap plus approx. 0.67 whorl; diameter ±410 μm; smooth to microscopically shagreened, lacking axial sculpture. Teleoconch of up to 2.25 whorls; sculptured by distinct, curved, close-set, compound axial riblets, with 45 finer, uneven, intermediary axial threads; intervals between riblets approx. twice riblet width at whorl periphery; spiral sculpture of extremely fine, close-set threads more or less throughout. Umbilicus wide. Aperture broadly lunate to almost D-shaped; parietal region with a low, rounded, in-running ridge just below mid-whorl; baso-columellar region lacking dentition; palatal region with two pairs of close-set, spirally aligned denticles, one pair close to mid-whorl, the other basal. Shell pale corneous-brown to honey-brown, usually encrusted with soil particles.
</paragraph>
</subSubSection>
<subSubSection id="C37465184624FFEA40BF410235EAFD25" pageId="40" pageNumber="41" type="distribution">
<paragraph id="8BD136934624FFEA40BF4102330CFDE4" blockId="40.[189,564,547,574]" box="[189,564,547,574]" pageId="40" pageNumber="41">
<heading id="D09981FF4624FFEA40BF4102330CFDE4" bold="true" box="[189,564,547,574]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="B91AEA814624FFEA40BF4102330CFDE4" bold="true" box="[189,564,547,574]" pageId="40" pageNumber="41">Distribution and conservation</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136934624FFEA40BF417235EAFD25" blockId="40.[189,1399,595,764]" pageId="40" pageNumber="41">
A narrow-range endemic (
<figureCitation id="13552A164624FFEA41F141723372FDB4" box="[499,586,595,621]" captionStart="Fig" captionStartId="36.[189,232,1860,1886]" captionTargetBox="[302,1285,1199,1820]" captionTargetId="figure@36.[302,1285,1199,1820]" captionTargetPageId="36" captionText="Fig. 14. Distribution of Iterodonta gen. nov. and Phialodonta gen. nov. species. Iterodonta ammonita gen. et sp. nov. (brown triangles), Phialodonta agulhasae gen. et sp. nov. (orange square), P. atromontana gen. et sp. nov. (turquoise square), P. aviana gen. et sp. nov. (royal blue squares), P. introtuberculata (Connolly, 1933) gen. et comb. nov. (red squares), P. perfida (Burnup, 1912) gen. et comb. nov. (green square), P. rivalalea gen. et sp. nov. (pink squares). Contour at 1000 m." figureDoi="http://doi.org/10.5281/zenodo.3762308" httpUri="https://zenodo.org/record/3762308/files/figure.png" pageId="40" pageNumber="41">Fig. 14</figureCitation>
), known only from the southern edge of the Groot Swartberge in the region of Calitzdorp, at
<quantity id="4C969B764624FFEA41E241573315FD48" box="[480,557,630,657]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="40" pageNumber="41" unit="m" value="360.0">360 m</quantity>
a.s.l.; in accumulations of leaf-litter in sheltered microhabitats within Gamka Thicket. The
<typeStatus id="54D588314624FFEA41B141BA30DEFD6C" box="[435,486,667,693]" pageId="40" pageNumber="41">type</typeStatus>
locality lies close to the protected Klein Swartberg Mountain Catchment Area and there are also several private and provincial nature reserves in the vicinity. Further sites in these areas need to be surveyed in the hope of finding additional extant colonies of the species.
</paragraph>
</subSubSection>
<subSubSection id="C37465184624FFEA40BF40053237FC6D" pageId="40" pageNumber="41" type="discussion">
<paragraph id="8BD136934624FFEA40BF40053017FCE7" blockId="40.[189,303,804,830]" box="[189,303,804,830]" pageId="40" pageNumber="41">
<heading id="D09981FF4624FFEA40BF40053017FCE7" bold="true" box="[189,303,804,830]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="B91AEA814624FFEA40BF40053017FCE7" bold="true" box="[189,303,804,830]" pageId="40" pageNumber="41">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136934624FFEA40BF40723237FC6D" blockId="40.[189,1399,851,949]" pageId="40" pageNumber="41">
Conchologically closest to
<taxonomicName id="4C6E4D104624FFEA41F7407233E4FCB4" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[501,732,851,877]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="aviana" status="gen. et sp. nov.">
<emphasis id="B91AEA814624FFEA41F7407233E4FCB4" box="[501,732,851,877]" italics="true" pageId="40" pageNumber="41">Phialodonta aviana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4624FFEA42E3407232B3FCB4" box="[737,907,851,877]" pageId="40" pageNumber="41" rank="species">gen. et sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C6E4D104624FFEA43BD4075357CFCB4" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[959,1092,851,877]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="rivalalea" status="gen. et sp. nov.">
<emphasis id="B91AEA814624FFEA43BD4075357CFCB4" box="[959,1092,851,877]" italics="true" pageId="40" pageNumber="41">P. rivalalea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4624FFEA444B407235CEFCB4" box="[1097,1270,851,877]" pageId="40" pageNumber="41" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, but differs in the form of the palatal dentition. In specimens at intermediate growth stages the palatal denticles may be single rather than paired, or even totally absent.
</paragraph>
</subSubSection>
</treatment>
</document>