324 lines
31 KiB
XML
324 lines
31 KiB
XML
<document ID-DOI="10.3897/zookeys.28.325" ID-GBIF-Dataset="cac72c82-2c83-401f-b6d0-8afb5321fde3" ID-ISSN="1313–2970" ID-Zenodo-Dep="576570" ID-ZooBank="A979ECDE-871F-4AFC-9ABA-63A0FD6DC323" approvalRequired="46" approvalRequired_for_textStreams="46" checkinTime="1587979922717" checkinUser="plazi" docAuthor="Sereno, Paul & Larsson, Hans" docDate="2009" docId="039B2B68FFBB9B69A5FAB330FB93557B" docLanguage="en" docName="ZK_article_2104.pdf" docOrigin="ZooKeys 28 (28)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@fc5bfa5" docStyleName="zookeys.2008.journal_article" docTitle="Araripesuchus" docType="treatment" docVersion="6" lastPageNumber="63" masterDocId="FFA25310FF879B57A576B119FF895175" masterDocTitle="Cretaceous Crocodyliforms from the Sahara" masterLastPageNumber="143" masterPageNumber="1" pageNumber="61" updateTime="1643567712823" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Cretaceous Crocodyliforms from the Sahara</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sereno, Paul</mods:namePart>
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<mods:affiliation>University of Chicago, Chicago, United States of America</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Larsson, Hans</mods:namePart>
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<mods:affiliation>Redpath Museum, McGill University, Montreal ,, Canada</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-11-19</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>28</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>28</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>143</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.28.325</mods:identifier>
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<mods:identifier type="GBIF-Dataset">cac72c82-2c83-401f-b6d0-8afb5321fde3</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576570</mods:identifier>
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<mods:identifier type="ZooBank">A979ECDE-871F-4AFC-9ABA-63A0FD6DC323</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3790373" ID-GBIF-Taxon="163750070" ID-Zenodo-Dep="3790373" LSID="urn:lsid:plazi:treatment:039B2B68FFBB9B69A5FAB330FB93557B" httpUri="http://treatment.plazi.org/id/039B2B68FFBB9B69A5FAB330FB93557B" lastPageId="62" lastPageNumber="63" pageId="60" pageNumber="61">
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<subSubSection pageId="60" pageNumber="61" type="nomenclature">
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<paragraph blockId="60.[140,348,553,615]" box="[140,348,553,580]" pageId="60" pageNumber="61">
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||
<heading box="[140,348,553,580]" fontSize="11" level="2" pageId="60" pageNumber="61" reason="3">
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<emphasis bold="true" box="[140,348,553,580]" pageId="60" pageNumber="61">
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<taxonomicName baseAuthorityName="Pol and Apesteguia" baseAuthorityYear="2005" box="[140,308,553,580]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="60" pageNumber="61" phylum="Chordata" rank="genus">
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<emphasis bold="true" box="[140,308,553,580]" italics="true" pageId="60" pageNumber="61">Araripesuchus</emphasis>
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</taxonomicName>
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sp.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="60.[140,348,553,615]" box="[140,219,589,615]" pageId="60" pageNumber="61">Fig. 31</paragraph>
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</subSubSection>
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<subSubSection pageId="60" pageNumber="61" type="materials_examined">
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<paragraph blockId="60.[140,1108,659,1672]" box="[140,826,659,686]" pageId="60" pageNumber="61">
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<emphasis bold="true" box="[140,249,659,685]" pageId="60" pageNumber="61">
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<emphasis box="[140,244,659,685]" inLineHeading="true" pageId="60" pageNumber="61" reason="1">Material</emphasis>
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.
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</emphasis>
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<collectionCode box="[256,329,660,686]" country="Niger" httpUri="http://grbio.org/cool/fx98-05bc" name="Musee National du Niger" pageId="60" pageNumber="61">MNN</collectionCode>
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<collectionCode box="[336,402,660,686]" pageId="60" pageNumber="61">GAD</collectionCode>
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27; isolated left dentary lacking teeth.
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</paragraph>
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<paragraph blockId="60.[140,1108,659,1672]" pageId="60" pageNumber="61">
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<emphasis bold="true" box="[185,343,694,721]" pageId="60" pageNumber="61">
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||
<emphasis box="[185,243,695,721]" inLineHeading="true" pageId="60" pageNumber="61" reason="1">
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<typeStatus box="[185,243,695,721]" pageId="60" pageNumber="61">Type</typeStatus>
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</emphasis>
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locality.
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</emphasis>
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Gadoufaoua,
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<collectingRegion box="[502,680,695,721]" country="Niger" name="Agadez" pageId="60" pageNumber="61">Agadez District</collectingRegion>
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,
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<collectingCountry box="[690,754,695,721]" name="Niger" pageId="60" pageNumber="61">Niger</collectingCountry>
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Republic (more precise locality unknown) (Fig. 1A, C).
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</paragraph>
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<paragraph blockId="60.[140,1108,659,1672]" pageId="60" pageNumber="61">
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<emphasis bold="true" box="[185,294,765,791]" pageId="60" pageNumber="61">
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<emphasis box="[185,288,765,791]" inLineHeading="true" pageId="60" pageNumber="61" reason="1">Horizon</emphasis>
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.
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</emphasis>
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Elrhaz Formation, Tegama Series; Lower Cretaceous (Aptian-Albian), ca. 110 Mya (
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<bibRefCitation author="Taquet P" box="[295,446,800,827]" pageId="60" pageNumber="61" pagination="1 - 191" refId="ref47707" refString="Taquet P (1976) Geologie et paleontologie du gisement de Gadoufaoua (Aptian du Niger). Cahiers de Paleontologie 1976: 1 - 191." type="journal article" year="1976">Taquet 1976</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="discussion">
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<paragraph blockId="60.[140,1108,659,1672]" pageId="60" pageNumber="61">
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<emphasis bold="true" box="[185,324,835,861]" pageId="60" pageNumber="61">
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<emphasis box="[185,318,835,861]" inLineHeading="true" pageId="60" pageNumber="61" reason="1">Discussion</emphasis>
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.
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</emphasis>
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With the notable exception of MNN GAD27, all of the cranial remains of
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<taxonomicName baseAuthorityName="Pol and Apesteguia" baseAuthorityYear="2005" box="[251,400,871,897]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="60" pageNumber="61" phylum="Chordata" rank="genus">
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<emphasis box="[251,400,871,897]" italics="true" pageId="60" pageNumber="61">Araripesuchus</emphasis>
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</taxonomicName>
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recovered from the Elrhaz Formation pertain to subadult or adult individuals with skull lengths between
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<quantity box="[643,762,906,933]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="60" pageNumber="61" unit="cm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 cm</quantity>
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. MNN GAD27, a left dentary lacking teeth with preserved crowns (Fig. 31), is approximately twice the length of the specimens described above for
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<taxonomicName authorityName="Buffetaut & Taquet" authorityYear="1979" box="[484,609,977,1003]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="wegeneri">
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<emphasis box="[484,609,977,1003]" italics="true" pageId="60" pageNumber="61">A. wegeneri</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[664,787,976,1003]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="rattoides">
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<emphasis box="[664,787,976,1003]" italics="true" pageId="60" pageNumber="61">A. rattoides</emphasis>
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</taxonomicName>
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and would pertain to a skull approximately
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<quantity box="[308,424,1011,1038]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="60" pageNumber="61" unit="cm" value="27.5" valueMax="30.0" valueMin="25.0">25–30 cm</quantity>
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long.
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</paragraph>
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<paragraph blockId="60.[140,1108,659,1672]" pageId="60" pageNumber="61">
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In addition to its large size, the number of postcaniniform alveoli is greater than in other specimens. Probably at least three additional postcaniniform teeth are present. The largest teeth in the dentary of
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<taxonomicName authorityName="Buffetaut & Taquet" authorityYear="1979" box="[539,667,1117,1143]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="wegeneri">
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<emphasis box="[539,667,1117,1143]" italics="true" pageId="60" pageNumber="61">A. wegeneri</emphasis>
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</taxonomicName>
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are between alveoli 10 and 13. In this larger dentary, the postcaniniform alveoli increase in size markedly starting with alveolus 13, suggesting that the comparable range for the largest dentary teeth would be alveoli 13–16.
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</paragraph>
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<paragraph blockId="60.[140,1108,659,1672]" pageId="60" pageNumber="61">
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Finally, the dorsal surface of dentary medial to the postcaniniform series (Fig. 31C) is flat, horizontal, and devoid of the accessory splenial articular scar observed in
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<taxonomicName authorityName="Buffetaut & Taquet" authorityYear="1979" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="wegeneri">
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<emphasis italics="true" pageId="60" pageNumber="61">A. wegeneri</emphasis>
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</taxonomicName>
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(Fig. 18B) and
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<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[412,534,1328,1355]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="rattoides">
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<emphasis box="[412,534,1328,1355]" italics="true" pageId="60" pageNumber="61">A. rattoides</emphasis>
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</taxonomicName>
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(
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<figureCitation box="[549,651,1328,1354]" captionStart="Figure 27" captionStartId="57.[140,212,1526,1548]" captionTargetBox="[294,941,368,1483]" captionTargetId="figure@57.[268,980,351,1506]" captionTargetPageId="57" captionText="Figure 27. Right dentary of the crocodyliform Araripesuchus rattoides sp. n. Isolated right dentary lacking teeth (CMN 41893). A Lateral view (reversed). B Medial view. C Dorsal view. D Anterior view. Scale bars equal 2 cm in A-C and 1 cm in D. Abbreviations: ad1, 3, 4, 8, 11, 13, 14, alveolus for dentary tooth 1, 3, 4, 8, 11, 13, 14; asp, articular surface for the splenial; dsym, dentary symphysis; fen, fenestra; fo, foramen; Mc, Meckel’s canal; rdsym, rough dentary symphysis; sdsym, smooth dentary symphysis." figureDoi="http://doi.org/10.5281/zenodo.3768353" httpUri="https://zenodo.org/record/3768353/files/figure.png" pageId="60" pageNumber="61">Fig. 27B</figureCitation>
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). In medial view, for example, no part of the nonarticular symphyseal surface medial to the tooth row is exposed, in contrast to the other African species (
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<figureCitation box="[432,543,1399,1425]" captionStart="Figure 27" captionStartId="57.[140,212,1526,1548]" captionTargetBox="[294,941,368,1483]" captionTargetId="figure@57.[268,980,351,1506]" captionTargetPageId="57" captionText="Figure 27. Right dentary of the crocodyliform Araripesuchus rattoides sp. n. Isolated right dentary lacking teeth (CMN 41893). A Lateral view (reversed). B Medial view. C Dorsal view. D Anterior view. Scale bars equal 2 cm in A-C and 1 cm in D. Abbreviations: ad1, 3, 4, 8, 11, 13, 14, alveolus for dentary tooth 1, 3, 4, 8, 11, 13, 14; asp, articular surface for the splenial; dsym, dentary symphysis; fen, fenestra; fo, foramen; Mc, Meckel’s canal; rdsym, rough dentary symphysis; sdsym, smooth dentary symphysis." figureDoi="http://doi.org/10.5281/zenodo.3768353" httpUri="https://zenodo.org/record/3768353/files/figure.png" pageId="60" pageNumber="61">Figs. 27B</figureCitation>
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,
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<figureCitation box="[555,607,1399,1425]" captionStart="Figure 28" captionStartId="58.[140,212,774,796]" captionTargetBox="[150,1091,170,747]" captionTargetId="figure@58.[140,1108,162,757]" captionTargetPageId="58" captionText="Figure 28. Left dentary of the crocodyliform Araripesuchus rattoides sp. n. Pencil drawing of isolated left dentary ramus lacking teeth (UCRC PV3). A Dorsal view. B Ventral view. C Medial view. Scale bar equals 1 cm. Parallel lines indicate broken bone surface. Abbreviations: ad1, 4, 5, 8, alveolus for dentary tooth 1, 4, 5, 8; asp, articular surface for the splenial; fen, fenestra; fo, foramen; sym, symphysis." figureDoi="http://doi.org/10.5281/zenodo.3768355" httpUri="https://zenodo.org/record/3768355/files/figure.png" pageId="60" pageNumber="61">28C</figureCitation>
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, 31B).
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</paragraph>
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<paragraph blockId="60.[140,1108,659,1672]" pageId="60" pageNumber="61">
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The differences between this specimen and the others suggest the presence of a second, larger species of
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<taxonomicName baseAuthorityName="Pol and Apesteguia" baseAuthorityYear="2005" box="[391,542,1469,1495]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="60" pageNumber="61" phylum="Chordata" rank="genus">
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<emphasis box="[391,542,1469,1495]" italics="true" pageId="60" pageNumber="61">Araripesuchus</emphasis>
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</taxonomicName>
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in the Elrhaz Formation. A similar circumstance was recently proposed for fossil material recovered from the La Buitrera locality in the Candeleros Formation in
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<collectingCountry box="[430,543,1539,1565]" name="Argentina" pageId="60" pageNumber="61">Argentina</collectingCountry>
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(
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<bibRefCitation author="Pol D & Apesteguia S" box="[557,844,1539,1566]" pageId="60" pageNumber="61" pagination="1 - 38" refId="ref46568" refString="Pol D, Apesteguia S (2005) New Araripesuchus remains from the early Late Cretaceous (Cenomanian-Turonian) of Patagonia. American Museum Novitates 3490: 1 - 38." type="journal article" year="2005">Pol and Apesteguia 2005</bibRefCitation>
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). There, a second larger species similar in size to MNN GAD27 occurs as a contemporary of a more common smaller species of
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<taxonomicName baseAuthorityName="Pol and Apesteguia" baseAuthorityYear="2005" box="[345,496,1610,1636]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="60" pageNumber="61" phylum="Chordata" rank="genus">
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<emphasis box="[345,496,1610,1636]" italics="true" pageId="60" pageNumber="61">Araripesuchus</emphasis>
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</taxonomicName>
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(
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<taxonomicName baseAuthorityName="Pol and Apesteguia" baseAuthorityYear="2005" box="[513,683,1610,1636]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="buitreraensis">
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<emphasis box="[513,683,1610,1636]" italics="true" pageId="60" pageNumber="61">A. buitreraensis</emphasis>
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</taxonomicName>
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) with a skull length of under
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<quantity box="[1028,1104,1610,1637]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="60" pageNumber="61" unit="cm" value="15.0">15 cm</quantity>
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. On the other hand, increased tooth number and better defined or developed features
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</paragraph>
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<paragraph blockId="61.[140,1108,1044,1194]" pageId="61" pageNumber="62">
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<emphasis bold="true" box="[140,767,1044,1066]" pageId="61" pageNumber="62">
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Figure 3|. Left dentary of the crocodyliform
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<taxonomicName baseAuthorityName="Pol and Apesteguia" baseAuthorityYear="2005" box="[596,733,1044,1066]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="61" pageNumber="62" phylum="Chordata" rank="genus">
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||
<emphasis bold="true" box="[596,733,1044,1066]" italics="true" pageId="61" pageNumber="62">Araripesuchus</emphasis>
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</taxonomicName>
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sp.
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</emphasis>
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Cast (UCRC PVC7) of an isolated, edentulous left dentary (MNN GAD27).
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<emphasis bold="true" box="[534,553,1076,1098]" pageId="61" pageNumber="62">A</emphasis>
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Lateral view.
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<emphasis bold="true" box="[683,700,1076,1098]" pageId="61" pageNumber="62">B</emphasis>
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Medial view (reversed).
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||
<emphasis bold="true" box="[931,950,1076,1098]" pageId="61" pageNumber="62">C</emphasis>
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Dorsal view (reversed). Scale bar equals
|
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<quantity box="[366,416,1109,1130]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="61" pageNumber="62" unit="cm" value="1.0">1 cm</quantity>
|
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. Abbreviations:
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<emphasis box="[563,716,1108,1130]" italics="true" pageId="61" pageNumber="62">ad3–5, 9, 13, 14</emphasis>
|
||
, alveolus for dentary tooth 3–5, 9, 13, 14;
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<emphasis box="[140,168,1141,1162]" italics="true" pageId="61" pageNumber="62">asp</emphasis>
|
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, articular surface for the splenial;
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||
<emphasis box="[488,507,1140,1161]" italics="true" pageId="61" pageNumber="62">be</emphasis>
|
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, buccal emargination;
|
||
<emphasis box="[723,767,1140,1161]" italics="true" pageId="61" pageNumber="62">dsym</emphasis>
|
||
, dentary symphysis;
|
||
<emphasis box="[965,981,1141,1162]" italics="true" pageId="61" pageNumber="62">fo</emphasis>
|
||
, foramen;
|
||
<emphasis box="[1084,1101,1141,1162]" italics="true" pageId="61" pageNumber="62">gr</emphasis>
|
||
, groove;
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||
<emphasis box="[213,242,1173,1194]" italics="true" pageId="61" pageNumber="62">Mc</emphasis>
|
||
, Meckel’s canal.
|
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</paragraph>
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<paragraph blockId="61.[140,1107,1293,1390]" pageId="61" pageNumber="62">may manifest themselves in particularly large individuals within a species. We are inclined to regard MNN GAD27 as a distinct species but await confirmation from more complete remains before establishing formal taxonomic recognition.</paragraph>
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<paragraph blockId="61.[140,981,1469,1531]" pageId="61" pageNumber="62">
|
||
<emphasis bold="true" box="[140,468,1469,1495]" pageId="61" pageNumber="62">
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[140,377,1469,1495]" class="Reptilia" family="Mahajangasuchidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="61" pageNumber="62" phylum="Chordata" rank="family">Mahajangasuchidae</taxonomicName>
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fam. n.
|
||
</emphasis>
|
||
<uri box="[140,981,1504,1531]" pageId="61" pageNumber="62">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid box="[426,981,1504,1531]" pageId="61" pageNumber="62">6C28D343-821D-4CF6-B293-F12EDBB59B15</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62" type="diagnosis">
|
||
<paragraph blockId="61.[140,1108,1574,1672]" lastBlockId="62.[140,1108,167,1038]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
|
||
<emphasis bold="true" box="[140,269,1574,1600]" pageId="61" pageNumber="62">
|
||
<emphasis box="[140,264,1574,1600]" inLineHeading="true" pageId="61" pageNumber="62" reason="1">Diagnosis</emphasis>
|
||
.
|
||
</emphasis>
|
||
Mid- to large-sized (̴
|
||
<quantity box="[528,611,1575,1601]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="61" pageNumber="62" unit="m" value="5.0" valueMax="6.0" valueMin="4.0">4–6 m</quantity>
|
||
) metasuchians with fused nasals, lacrimalnasal contact absent, postorbital bearing an oval laterally facing fossa that may have served for articulation with the posterior palpebral, squamosal and parietal form a hornlike posterodorsal process, steeply arched ventral jugal margin with a ventrolateral fossa at apex, ectopterygoid vertical and flush at jugal contact rather than arching medially, jaw articulation below posterior tooth row, pterygoid choanal septum with anterior footplate for palatine, pterygoid choanal septum with ventral edge expanded to approximately 40% of septum length, and pterygoid choanal wall invaginated dorsal to posterior margin of palate, deep mandibular symphysis oriented at approximately 45° anterodorsally, dorsolateral ridge on surangular, and maxillary tooth row terminates anterior to orbit.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="62" pageNumber="63" type="etymology">
|
||
<paragraph blockId="62.[140,1108,167,1038]" pageId="62" pageNumber="63">
|
||
<emphasis bold="true" box="[185,322,448,474]" pageId="62" pageNumber="63">
|
||
<emphasis box="[185,317,448,474]" inLineHeading="true" pageId="62" pageNumber="63" reason="1">Etymology</emphasis>
|
||
.
|
||
</emphasis>
|
||
Named on the basis of
|
||
<taxonomicName authority="(Buckley and Brochu 1999)" baseAuthorityName="Buckley and Brochu" baseAuthorityYear="1999" class="Reptilia" family="Peirosauridae" genus="Mahajangasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="62" pageNumber="63" phylum="Chordata" rank="species">
|
||
<emphasis box="[601,887,448,474]" italics="true" pageId="62" pageNumber="63">Mahajangasuchus insignis</emphasis>
|
||
(
|
||
<bibRefCitation author="Buckley GA & Brochu CA" pageId="62" pageNumber="63" pagination="149 - 175" refId="ref44828" refString="Buckley GA, Brochu CA (1999) An enigmatic crocodile from the Upper Cretaceous of Madagascar. Special Papers in Palaeontology 60: 149 - 175." type="journal article" year="1999">Buckley and Brochu 1999</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, the first described member of the clade. New fossil finds continue to expand basal metasuchian diversity, although interrelationships are poorly established. Establishing this stem-based taxon for
|
||
<taxonomicName baseAuthorityName="Turner and Buckley" baseAuthorityYear="2008" box="[583,779,554,580]" class="Reptilia" family="Peirosauridae" genus="Mahajangasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="62" pageNumber="63" phylum="Chordata" rank="genus">
|
||
<emphasis box="[583,779,554,580]" italics="true" pageId="62" pageNumber="63">Mahajangasuchus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[794,930,554,580]" class="Reptilia" family="Mahajangasuchidae" genus="Kaprosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodilomorphi" pageId="62" pageNumber="63" phylum="Chordata" rank="genus">
|
||
<emphasis box="[794,930,554,580]" italics="true" pageId="62" pageNumber="63">Kaprosuchus</emphasis>
|
||
</taxonomicName>
|
||
and taxa closer to them than to several other metasuchians establishes a well known anchor among basal metasuchians, to which other taxa may eventually be assigned.
|
||
</paragraph>
|
||
<paragraph blockId="62.[140,1108,167,1038]" pageId="62" pageNumber="63">
|
||
<emphasis bold="true" box="[185,477,659,685]" pageId="62" pageNumber="63">
|
||
<emphasis box="[185,340,659,685]" inLineHeading="true" pageId="62" pageNumber="63" reason="1">Phylogenetic</emphasis>
|
||
definition.
|
||
</emphasis>
|
||
The most inclusive clade containing
|
||
<taxonomicName authority="Buckley and Brochu 1999" authorityName="Buckley and Brochu" authorityYear="1999" baseAuthorityName="Turner and Buckley" baseAuthorityYear="2008" class="Reptilia" family="Peirosauridae" genus="Mahajangasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="62" pageNumber="63" phylum="Chordata" rank="species">
|
||
<emphasis italics="true" pageId="62" pageNumber="63">Mahajangasuchus insignis</emphasis>
|
||
<bibRefCitation author="Buckley GA & Brochu CA" box="[230,532,695,721]" pageId="62" pageNumber="63" pagination="149 - 175" refId="ref44828" refString="Buckley GA, Brochu CA (1999) An enigmatic crocodile from the Upper Cretaceous of Madagascar. Special Papers in Palaeontology 60: 149 - 175." type="journal article" year="1999">Buckley and Brochu 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
but not
|
||
<taxonomicName authority="Woodward 1896" authorityName="Woodward" authorityYear="1896" box="[637,1062,695,721]" class="Reptilia" genus="Notosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="terrestris">
|
||
<emphasis box="[637,858,695,721]" italics="true" pageId="62" pageNumber="63">Notosuchus terrestris</emphasis>
|
||
Woodward 1896
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Buckley et al. 2000" authorityName="Buckley" authorityYear="2000" class="Reptilia" family="Chimaerasuchidae" genus="Simosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="62" pageNumber="63" phylum="Chordata" rank="species">
|
||
<emphasis italics="true" pageId="62" pageNumber="63">Simosuchus clarki</emphasis>
|
||
<bibRefCitation author="Buckley GA & Brochu CA & Krause DW & Pol D" box="[317,538,730,756]" pageId="62" pageNumber="63" pagination="941 - 944" refId="ref44856" refString="Buckley GA, Brochu CA, Krause DW, Pol D (2000) A pug-nosed crocodyliform from the Late Cretaceous of Madagascar. Nature 405: 941 - 944." type="journal article" year="2000">Buckley et al. 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Price 1959" authorityName="Price" authorityYear="1959" box="[549,915,730,757]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="gomesii">
|
||
<emphasis box="[549,784,730,756]" italics="true" pageId="62" pageNumber="63">Araripesuchus gomesii</emphasis>
|
||
<bibRefCitation author="Price LI" box="[791,915,730,757]" pageId="62" pageNumber="63" pagination="1 - 55" refId="ref46760" refString="Price LI (1959) Sobre um crocodilideo notosuquio do Cretaceo brasileiro. Boletim Divisao de Geologia e Mineralogia do Rio de Janeiro 188: 1 - 55." type="journal article" year="1959">Price 1959</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Price 1945" authorityName="Price" authorityYear="1945" class="Reptilia" family="Baurusuchidae" genus="Baurusuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="62" pageNumber="63" phylum="Chordata" rank="species">
|
||
<emphasis italics="true" pageId="62" pageNumber="63">Baurusuchus pachecoi</emphasis>
|
||
Price 1945
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Price 1955" authorityName="Price" authorityYear="1955" box="[348,707,765,792]" class="Reptilia" family="Peirosauridae" genus="Peirosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="62" pageNumber="63" phylum="Chordata" rank="species">
|
||
<emphasis box="[348,576,766,791]" italics="true" pageId="62" pageNumber="63">Peirosaurus torminni</emphasis>
|
||
<bibRefCitation author="Price LI" box="[583,707,765,792]" pageId="62" pageNumber="63" pagination="487 - 498" refId="ref46728" refString="Price LI (1955) Novos crocodilideos dos arenitos da Serie Bauru. Cretaceo do Estado de Minas Gerais. Anais da Academia Brasileira de Ciencias 27: 487 - 498." type="journal article" year="1955">Price 1955</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Owen 1842" authorityName="Owen" authorityYear="1842" box="[719,1102,765,791]" class="Reptilia" family="Goniopholididae" genus="Goniopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="62" pageNumber="63" phylum="Chordata" rank="species">
|
||
<emphasis box="[719,957,765,791]" italics="true" pageId="62" pageNumber="63">Goniopholis crassidens</emphasis>
|
||
Owen 1842
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Meyer 1841" authorityName="Meyer" authorityYear="1841" box="[140,619,800,827]" class="Reptilia" family="Pholidosauridae" genus="Pholidosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="62" pageNumber="63" phylum="Chordata" rank="species">
|
||
<emphasis box="[140,472,800,826]" italics="true" pageId="62" pageNumber="63">Pholidosaurus schaumbergensis</emphasis>
|
||
Meyer 1841
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="(Laurenti 1768)" baseAuthorityName="Laurenti" baseAuthorityYear="1768" box="[631,1037,800,827]" class="Reptilia" family="Crocodylidae" genus="Crocodylus" kingdom="Animalia" order="Crocodylia" pageId="62" pageNumber="63" phylum="Chordata" rank="species">
|
||
<emphasis box="[631,846,800,826]" italics="true" pageId="62" pageNumber="63">Crocodylus niloticus</emphasis>
|
||
(Laurenti 1768)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="62" pageNumber="63" type="discussion">
|
||
<paragraph blockId="62.[140,1108,167,1038]" pageId="62" pageNumber="63">
|
||
<emphasis bold="true" box="[185,323,835,861]" pageId="62" pageNumber="63">
|
||
<emphasis box="[185,318,835,861]" inLineHeading="true" pageId="62" pageNumber="63" reason="1">Discussion</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[331,572,835,861]" class="Reptilia" family="Mahajangasuchidae" genus="Kaprosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodilomorphi" pageId="62" pageNumber="63" phylum="Chordata" rank="species">
|
||
<emphasis box="[331,572,835,861]" italics="true" pageId="62" pageNumber="63">Kaprosuchus saharicus</emphasis>
|
||
</taxonomicName>
|
||
represents a distinctive new crocodyliform distinguished by numerous cranial autapomorphies. Derived characters shared with other crocodyliforms are limited, although a suite of features listed in the familial diagnosis above links
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[273,404,941,967]" class="Reptilia" family="Mahajangasuchidae" genus="Kaprosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodilomorphi" pageId="62" pageNumber="63" phylum="Chordata" rank="species">
|
||
<emphasis box="[273,404,941,967]" italics="true" pageId="62" pageNumber="63">K. saharicus</emphasis>
|
||
</taxonomicName>
|
||
to the unusual crocodyliform,
|
||
<emphasis box="[756,1044,941,967]" italics="true" pageId="62" pageNumber="63">
|
||
<taxonomicName authorityName="Buckley and Brochu" authorityYear="1999" baseAuthorityName="Turner and Buckley" baseAuthorityYear="2008" box="[756,1041,941,967]" class="Reptilia" family="Peirosauridae" genus="Mahajangasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="62" pageNumber="63" phylum="Chordata" rank="species">Mahajangasuchus insignis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
from the Upper Cretaceous of
|
||
<collectingCountry box="[424,557,976,1002]" name="Madagascar" pageId="62" pageNumber="63">Madagascar</collectingCountry>
|
||
(
|
||
<bibRefCitation author="Buckley GA & Brochu CA" box="[572,872,976,1002]" pageId="62" pageNumber="63" pagination="149 - 175" refId="ref44828" refString="Buckley GA, Brochu CA (1999) An enigmatic crocodile from the Upper Cretaceous of Madagascar. Special Papers in Palaeontology 60: 149 - 175." type="journal article" year="1999">Buckley and Brochu 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Turner AH & Buckley GA" pageId="62" pageNumber="63" pagination="382 - 408" refId="ref47861" refString="Turner AH, Buckley GA (2008) Mahajangasuchus insignis (Crocodyliformes: Mesoeucrocodylia) cranial anatomy and new data on the origin of the eusuchian-style palate. Journal of Vertebrate Paleontology 28: 382 - 408." type="journal article" year="2008">Turner and Buckley 2008</bibRefCitation>
|
||
). Th ese are described in more detail below (see Phylogenetic relationships).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |