169 lines
20 KiB
XML
169 lines
20 KiB
XML
<document id="A8366A0B8131542A08E433263A7341BF" ID-DOI="10.11646/zootaxa.1957.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5241137" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629590285433" checkinUser="felipe" docAuthor="Sartori, Michel, Peters, Janice G. & Hubbard, Michael D." docDate="2008" docId="039AE86B4423FFDBCE8E750DFAFDF85A" docLanguage="en" docName="zt01957p051.pdf" docOrigin="Zootaxa 1957 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1957.1.1" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Derlethina Sartori & Peters & Hubbard 2008, gen. nov." docType="treatment" docVersion="3" lastPageNumber="44" masterDocId="FFA390134408FFF0CE197632FFA3FFCD" masterDocTitle="A revision of Oriental Teloganodidae (Insecta, Ephemeroptera, Ephemerelloidea)" masterLastPageNumber="51" masterPageNumber="1" pageNumber="44" updateTime="1699082080968" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="AC9E17BB88C6C3531746131D3615607F">
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<mods:title id="51A41849DDD40634D1CF60A1C489BBED">A revision of Oriental Teloganodidae (Insecta, Ephemeroptera, Ephemerelloidea)</mods:title>
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<mods:name id="2C4C50E7128334B537C485B2EAC1FBFB" type="personal">
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<mods:namePart id="01834996923F99C4955370519A58CBD6">Sartori, Michel</mods:namePart>
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<mods:affiliation id="5D37760D4091E424927F4B0866B77F05">Museum of zoology, Palais de Rumine, Place Riponne 6, CH- 1014 Lausanne, Switzerland. E-mail: michel. sartori @ vd. ch Laboratory of Aquatic Entomology, Florida A & M University, Tallahassee, FL 32307, USA</mods:affiliation>
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<mods:namePart id="2151988175035474EB1EE2C5972DC4FB">Peters, Janice G.</mods:namePart>
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<mods:roleTerm id="65E9F7ADD883A4CA71BAB227C24AF946">Author</mods:roleTerm>
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<mods:namePart id="769A1EF2C2E36470FBB739F3679787F2">Hubbard, Michael D.</mods:namePart>
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<mods:title id="9683D0BA787CD57FBCD1D2295B7982EA">Zootaxa</mods:title>
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<mods:part id="C39917F942306584F063670725978019">
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<mods:date id="DDEC59323AF27B497EB2E1D3A1F152A3">2008</mods:date>
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<mods:number id="EB3F2CE7410B6B63C58EB6465E530B7F">1957</mods:number>
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<mods:url id="08075911691501C804BA3B45980D2F77">https://biotaxa.org/Zootaxa/article/view/zootaxa.1957.1.1</mods:url>
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<subSubSection id="C3290AF64423FFDBCE8E750DFE4CFC94" box="[151,495,831,857]" pageId="43" pageNumber="44" type="nomenclature">
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<paragraph id="8B8C597D4423FFDBCE8E750DFE4CFC94" blockId="43.[151,495,831,857]" box="[151,495,831,857]" pageId="43" pageNumber="44">
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<heading id="D0C4EE114423FFDBCE8E750DFE4CFC94" bold="true" box="[151,495,831,857]" fontSize="11" level="1" pageId="43" pageNumber="44" reason="1">
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<emphasis id="B947856F4423FFDBCE8E750DFE4CFC94" bold="true" box="[151,495,831,857]" pageId="43" pageNumber="44">
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<taxonomicName id="4C3322FE4423FFDBCE8E750DFEDAFC94" ID-CoL="62PLM" ID-ENA="650599" authority="Sartori" authorityName="Sartori & Peters & Hubbard" authorityYear="2008" box="[151,377,831,857]" class="Insecta" family="Teloganodidae" genus="Derlethina" kingdom="Animalia" order="Ephemeroptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B947856F4423FFDBCE8E750DFEBAFC94" bold="true" box="[151,281,831,857]" italics="true" pageId="43" pageNumber="44">Derlethina</emphasis>
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Sartori
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</taxonomicName>
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<taxonomicNameLabel id="A27438144423FFDBCF98750DFE4CFC94" box="[385,495,831,857]" pageId="43" pageNumber="44" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3290AF64423FFDBCE8E75B8FD17FC0C" pageId="43" pageNumber="44" type="materials_examined">
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<paragraph id="8B8C597D4423FFDBCE8E75B8FC32FC6C" blockId="43.[151,913,906,1025]" box="[151,913,906,929]" pageId="43" pageNumber="44">
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<typeStatus id="5488E7DF4423FFDBCE8E75B8FF6FFC6C" box="[151,204,906,929]" pageId="43" pageNumber="44">Type</typeStatus>
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species:
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<taxonomicName id="4C3322FE4423FFDBCF3475B8FDE2FC6C" authority="Sartori" authorityName="Sartori & Peters & Hubbard" authorityYear="2008" box="[301,577,906,929]" class="Insecta" family="Teloganodidae" genus="Derlethina" kingdom="Animalia" order="Ephemeroptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="eloisae" status="sp. nov.">
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<emphasis id="B947856F4423FFDBCF3475B8FE57FC6C" box="[301,500,906,929]" italics="true" pageId="43" pageNumber="44">Derlethina eloisae</emphasis>
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Sartori
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</taxonomicName>
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<taxonomicNameLabel id="A27438144423FFDBCC5075B8FD38FC6C" box="[585,667,906,929]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
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by present designation
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</paragraph>
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<paragraph id="8B8C597D4423FFDBCE8E7598FD17FC0C" blockId="43.[151,913,906,1025]" box="[151,692,938,961]" pageId="43" pageNumber="44">
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<taxonomicName id="4C3322FE4423FFDBCE8E7598FEE1FC0C" box="[151,322,938,961]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B947856F4423FFDBCE8E7598FEBEFC0C" box="[151,285,938,961]" italics="true" pageId="43" pageNumber="44">Teloganodes</emphasis>
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sp.
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</taxonomicName>
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sensu
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<bibRefCitation id="EFA2248C4423FFDBCF937598FD17FC0C" author="McCafferty, W. P. & Wang, T. - Q." box="[394,692,938,961]" pageId="43" pageNumber="44" pagination="387 - 437" refId="ref26050" refString="McCafferty, W. P. & Wang, T. - Q. (1997) Phylogenetic systematics of the family Teloganodidae (Ephemeroptera: Pannota). Annals of the Cape Provincial Museums (Natural History), 19, 387 - 437." type="journal article" year="1997">McCafferty & Wang (1997)</bibRefCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3290AF64423FFDBCE8E75F8FDF6F9B2" pageId="43" pageNumber="44" type="description">
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<paragraph id="8B8C597D4423FFDBCE8E75F8FD9EFC2C" blockId="43.[151,913,906,1025]" box="[151,573,970,993]" pageId="43" pageNumber="44">
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<taxonomicName id="4C3322FE4423FFDBCE8E75F8FEE1FC2C" box="[151,322,970,993]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B947856F4423FFDBCE8E75F8FEBEFC2C" box="[151,285,970,993]" italics="true" pageId="43" pageNumber="44">Teloganodes</emphasis>
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sp.
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</taxonomicName>
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T3 sensu
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<bibRefCitation id="EFA2248C4423FFDBCFB775F8FD9EFC2C" author="Kluge, N." box="[430,573,970,993]" pageId="43" pageNumber="44" refId="ref25872" refString="Kluge, N. (2004) The phylogenetic system of Ephemeroptera, Kluwer Academic Publishers, Dordrecht." type="book" year="2004">Kluge (2004)</bibRefCitation>
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</paragraph>
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<paragraph id="8B8C597D4423FFDBCE8E75D8FF42FBCC" blockId="43.[151,913,906,1025]" box="[151,225,1002,1025]" pageId="43" pageNumber="44">eloisae</paragraph>
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<paragraph id="8B8C597D4423FFDBCE8E7200FC49FB22" blockId="43.[151,1437,1074,1943]" pageId="43" pageNumber="44">
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<emphasis id="B947856F4423FFDBCE8E7200FE8CFB81" bold="true" box="[151,303,1074,1100]" pageId="43" pageNumber="44">Description.</emphasis>
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Adult: Forewing (
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<figureCitation id="130845F84423FFDBCC167200FDC5FB81" box="[527,614,1074,1100]" captionStart="FIGURES 14–15" captionStartId="14.[151,269,1083,1107]" captionTargetBox="[183,1409,343,1047]" captionTargetId="figure-301@14.[151,1436,279,1085]" captionTargetPageId="14" captionText="FIGURES 14–15. Derlethina eloisae. Fig. 14: Forewing; Fig. 15: Hindwing." figureDoi="http://doi.org/10.5281/zenodo.5241149" httpUri="https://zenodo.org/record/5241149/files/figure.png" pageId="43" pageNumber="44">Fig. 14</figureCitation>
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) long and slender, posterior margin regularly convex. Pterostigmatic area without crossveins; vein MP
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<subScript id="17B75B384423FFDBCC067258FD84FBB5" attach="left" box="[543,551,1130,1144]" fontSize="6" pageId="43" pageNumber="44">2</subScript>
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very short, and free; cubital field with
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<specimenCount id="9D3592F44423FFDBCDEF7268FB2AFBB9" box="[1014,1161,1114,1140]" pageId="43" pageNumber="44" type="intercalary">2 intercalary</specimenCount>
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veins. Hind wing (
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<figureCitation id="130845F84423FFDBCB747268FF16FB52" captionStart="FIGURES 14–15" captionStartId="14.[151,269,1083,1107]" captionTargetBox="[183,1409,343,1047]" captionTargetId="figure-301@14.[151,1436,279,1085]" captionTargetPageId="14" captionText="FIGURES 14–15. Derlethina eloisae. Fig. 14: Forewing; Fig. 15: Hindwing." figureDoi="http://doi.org/10.5281/zenodo.5241149" httpUri="https://zenodo.org/record/5241149/files/figure.png" pageId="43" pageNumber="44">Fig. 15</figureCitation>
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) minute with costal process acute and located almost at apex of wing; a single simple longitudinal vein. Claws of all legs dissimilar. Dorsal margin of hind femora concave. Subanal plate of female almost straight. Gill socket vestiges visible on segments II–IV. Terminal filament absent.
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</paragraph>
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<paragraph id="8B8C597D4423FFDBCEDC72CFFDAFF9E2" blockId="43.[151,1437,1074,1943]" pageId="43" pageNumber="44">
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Nymph (
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<figureCitation id="130845F84423FFDBCF3472CFFE37FADA" box="[301,404,1277,1303]" captionStart="FIGURES 144–149" captionStartId="40.[151,269,1825,1849]" captionTargetBox="[151,1436,194,1781]" captionTargetId="figure-48@40.[151,1436,194,1781]" captionTargetPageId="40" captionText="FIGURES 144–149. Holotypes of Dudgeodes and Derlethina in dorsal view (144, 147), abdomen in ventral view (145, 148) and in lateral view (146, 149). Figs. 144–146: Dudgeodes celebensis; Figs. 147–149: Derlethina eloisae" figureDoi="http://doi.org/10.5281/zenodo.5241203" httpUri="https://zenodo.org/record/5241203/files/figure.png" pageId="43" pageNumber="44">Fig. 147</figureCitation>
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): Body flattened, eyes in dorsal position blackish in male nymphs; outer margin of head fringed with a row of forked setae from behind eyes to labrum insertion. Labrum with a single row of simple setae (
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<figureCitation id="130845F84423FFDBCEF8737FFE9BFAAA" box="[225,312,1357,1383]" captionStart="FIGURES 16–25" captionStartId="15.[151,269,1790,1814]" captionTargetBox="[180,1416,190,1708]" captionTargetId="figure-93@15.[151,1436,150,1738]" captionTargetPageId="15" captionText="FIGURES 16–25. Labrum (16, 18, 20, 22, 24), with details of the dorsal setae (17, 19, 21, 23, 25), of Oriental Teloganodidae. Insertion of the setae indicated by dots on the labrum. Figs. 16–17: Teloganodes tuberculatus; Figs. 18–19: T. hubbardi; Figs. 20–21: T. jacobusi; Figs. 22–23: Dudgeodes hutanis; Figs. 24–25: Derlethina eloisae. All labra at same scale and all setae at same scale, two times enlarged compared to the labra." figureDoi="http://doi.org/10.5281/zenodo.5241153" httpUri="https://zenodo.org/record/5241153/files/figure.png" pageId="43" pageNumber="44">Fig. 24</figureCitation>
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). Galea-lacinia with 2 setae on ventral side (
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<figureCitation id="130845F84423FFDBCD24737FFC36FAAA" box="[829,917,1357,1383]" captionStart="FIGURES 40–46" captionStartId="18.[151,269,1781,1805]" captionTargetBox="[260,1412,191,1697]" captionTargetId="figure-70@18.[151,1436,156,1744]" captionTargetPageId="18" captionText="FIGURES 40–46. Maxillae (40, 41) and detail of the setae on the apico-ventral part (42–46) of Oriental Teloganodidae. Fig. 40: Teloganodes tuberculatus; Fig. 41: Dudgeodes hutanis; Fig. 42: T. jacobusi; Fig. 43: T. kodai; Fig. 44: D. pescadori; Fig. 45: D. ulmeri; Fig. 46: Derlethina eloisae." figureDoi="http://doi.org/10.5281/zenodo.5241161" httpUri="https://zenodo.org/record/5241161/files/figure.png" pageId="43" pageNumber="44">Fig. 46</figureCitation>
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). Forefemora broad and dilated with a transverse row of setae on dorsal face (Fig. 75). Hind femora with a distinct concave outer margin (
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<figureCitation id="130845F84423FFDBCAF17347FAEDFA42" box="[1256,1358,1397,1423]" captionStart="FIGURES 144–149" captionStartId="40.[151,269,1825,1849]" captionTargetBox="[151,1436,194,1781]" captionTargetId="figure-48@40.[151,1436,194,1781]" captionTargetPageId="40" captionText="FIGURES 144–149. Holotypes of Dudgeodes and Derlethina in dorsal view (144, 147), abdomen in ventral view (145, 148) and in lateral view (146, 149). Figs. 144–146: Dudgeodes celebensis; Figs. 147–149: Derlethina eloisae" figureDoi="http://doi.org/10.5281/zenodo.5241203" httpUri="https://zenodo.org/record/5241203/files/figure.png" pageId="43" pageNumber="44">Fig. 147</figureCitation>
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). Gills on abdominal segments II–IV (Figs. 95–97); gills II–III with ventral lobes, gill IV without ventral lobe and entire (not incised), gill III incised. Abdominal carina (median tubercles on terga), poorly developed. Posterolateral expansions very well developed on segments VI–IX. Lateral margins of terga with very long and thin setae. Terminal filament absent.
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</paragraph>
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<paragraph id="8B8C597D4423FFDBCEDC700FFDF6F9B2" blockId="43.[151,1437,1074,1943]" pageId="43" pageNumber="44">
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Egg: rounded, chorion without attachment structures, one polar cap, no triangular or polygonal structures at the base of the cap (
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<figureCitation id="130845F84423FFDBCF847057FDEAF9B2" box="[413,585,1637,1663]" captionStart="FIGURES 115–119" captionStartId="32.[151,269,1656,1680]" captionTargetBox="[151,1436,194,1632]" captionTargetId="figure-60@32.[151,1436,194,1632]" captionTargetPageId="32" captionText="FIGURES 115–119. Egg structure of Derlethina eloisae (SEM pictures). Fig. 115: Shape of the egg; Fig. 116: Detail of the polar cap; Fig. 117: Detail of the coiled threads of the epithema; Fig. 118: Detail of the different chorionic structures; Fig. 119: Micropyle area." figureDoi="http://doi.org/10.5281/zenodo.5241186" httpUri="https://zenodo.org/record/5241186/files/figure.png" pageId="43" pageNumber="44">Figs. 115–119</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3290AF64423FFDBCEDF70BFFAC2F93A" pageId="43" pageNumber="44" type="diagnosis">
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<paragraph id="8B8C597D4423FFDBCEDF70BFFAC2F93A" blockId="43.[151,1437,1074,1943]" pageId="43" pageNumber="44">
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<emphasis id="B947856F4423FFDBCEDF70BFFEE5F96A" bold="true" box="[198,326,1677,1703]" pageId="43" pageNumber="44">Diagnosis.</emphasis>
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<taxonomicName id="4C3322FE4423FFDBCF5470BCFE6FF96A" authorityName="Sartori & Peters & Hubbard" authorityYear="2008" box="[333,460,1678,1703]" class="Insecta" family="Teloganodidae" genus="Derlethina" kingdom="Animalia" order="Ephemeroptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="B947856F4423FFDBCF5470BCFE6FF96A" box="[333,460,1678,1703]" italics="true" pageId="43" pageNumber="44">Derlethina</emphasis>
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</taxonomicName>
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winged stages can easily be told from
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<taxonomicName id="4C3322FE4423FFDBCD8A70BCFB85F96A" authorityName="sensu Ulmer" authorityYear="1939" box="[915,1062,1678,1703]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="B947856F4423FFDBCD8A70BCFB85F96A" box="[915,1062,1678,1703]" italics="true" pageId="43" pageNumber="44">Teloganodes</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C3322FE4423FFDBCA4670BCFB40F96A" authorityName="Sartori & Peters & Hubbard" authorityYear="2008" box="[1119,1251,1678,1703]" class="Insecta" family="Teloganodidae" genus="Dudgeodes" kingdom="Animalia" order="Ephemeroptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="B947856F4423FFDBCA4670BCFB40F96A" box="[1119,1251,1678,1703]" italics="true" pageId="43" pageNumber="44">Dudgeodes</emphasis>
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</taxonomicName>
|
||
by the shape of the hindwing, the absence of crossvein in the pterostigmatic area of the forewing, and the shape of the hind femur; the nymphal stage is distinguished by the absence of gill V and the shape of the posterior femora.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3290AF64423FFDBCEDF7137FD97F8A2" pageId="43" pageNumber="44" type="etymology">
|
||
<paragraph id="8B8C597D4423FFDBCEDF7137FD97F8A2" blockId="43.[151,1437,1074,1943]" pageId="43" pageNumber="44">
|
||
<emphasis id="B947856F4423FFDBCEDF7137FEF0F8D2" bold="true" box="[198,339,1797,1823]" pageId="43" pageNumber="44">Etymology.</emphasis>
|
||
This new genus is named after Dr Pascale Derleth (Lausanne), wife of the first author, who collected most of the material during her work in
|
||
<collectingRegion id="49F7979F4423FFDBCCC6711FFC06F88A" box="[735,933,1837,1863]" country="Indonesia" name="Kalimantan Timur" pageId="43" pageNumber="44">East Kalimantan</collectingRegion>
|
||
, and was the first to recognize it as a separate genus. The gender is feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3290AF64423FFDBCEDF714FFAFDF85A" box="[198,1374,1917,1943]" pageId="43" pageNumber="44" type="distribution">
|
||
<paragraph id="8B8C597D4423FFDBCEDF714FFAFDF85A" blockId="43.[151,1437,1074,1943]" box="[198,1374,1917,1943]" pageId="43" pageNumber="44">
|
||
<emphasis id="B947856F4423FFDBCEDF714FFEC4F85A" bold="true" box="[198,359,1917,1943]" pageId="43" pageNumber="44">Distribution.</emphasis>
|
||
Known only from Borneo (East
|
||
<collectingCountry id="F32419ED4423FFDBCCF1714FFCF0F85A" box="[744,851,1917,1943]" name="Malaysia" pageId="43" pageNumber="44">Malaysia</collectingCountry>
|
||
–
|
||
<collectingRegion id="49F7979F4423FFDBCD68714FFC1AF85A" box="[881,953,1917,1943]" country="Malaysia" name="Sabah" pageId="43" pageNumber="44">Sabah</collectingRegion>
|
||
- and
|
||
<collectingCountry id="F32419ED4423FFDBCA1B714FFBD7F85A" box="[1026,1140,1917,1943]" name="Indonesia" pageId="43" pageNumber="44">Indonesia</collectingCountry>
|
||
–
|
||
<collectingRegion id="49F7979F4423FFDBCA88714FFAF5F85A" box="[1169,1366,1917,1943]" country="Indonesia" name="Kalimantan Timur" pageId="43" pageNumber="44">East Kalimantan</collectingRegion>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |