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<document id="5AE634652890577796CA9BCFDB4AE2E2" ID-CLB-Dataset="21162" ID-DOI="10.1080/00222933.2015.1009514" ID-GBIF-Dataset="8c6c97a7-1c63-41ba-b75a-217ce1a3411d" ID-ISSN="1464-5262" ID-Zenodo-Dep="3997986" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1598292250840" checkinUser="carolina" docAuthor="Rota, Emilia" docDate="2015" docId="039AAD67FFF64517FE18FCD7FCF10A21" docLanguage="en" docName="JNATHIST.49.33-34.1987-2020.pdf" docOrigin="Journal of Natural History 49 (33)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Fridericia miraflores Sesma and Dozsa-Farkas 1993" docType="treatment" docVersion="2" lastPageNumber="2018" masterDocId="FFA3D51FFFE84508FF92FFAFFFB6091F" masterDocTitle="Five new species of Enchytraeidae (Annelida: Clitellata) from Mediterranean woodlands of Italy and reaffirmed validity of Achaeta etrusca, Fridericia bulbosa and F. miraflores" masterLastPageNumber="2020" masterPageNumber="1987" pageNumber="2017" updateTime="1728498831906" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="30D1E449591D5979B28FA1B55DF4359D">Five new species of Enchytraeidae (Annelida: Clitellata) from Mediterranean woodlands of Italy and reaffirmed validity of Achaeta etrusca, Fridericia bulbosa and F. miraflores</mods:title>
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<mods:namePart id="C4DA86E2C41832E541FD20EA73957235">Rota, Emilia</mods:namePart>
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<mods:title id="BCEBE45888B37F1CF7BC4C044E124C13">Journal of Natural History</mods:title>
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<mods:date id="B04E80C4936BB01C5905AF119F1D9FAB">2015</mods:date>
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<treatment id="039AAD67FFF64517FE18FCD7FCF10A21" ID-DOI="http://doi.org/10.5281/zenodo.4330405" ID-GBIF-Taxon="167293492" ID-Zenodo-Dep="4330405" LSID="urn:lsid:plazi:treatment:039AAD67FFF64517FE18FCD7FCF10A21" httpUri="http://treatment.plazi.org/id/039AAD67FFF64517FE18FCD7FCF10A21" lastPageId="31" lastPageNumber="2018" pageId="30" pageNumber="2017">
<subSubSection id="C3294FFAFFF64516FE18FCD7FC6B0A8F" box="[394,989,888,912]" pageId="30" pageNumber="2017" type="nomenclature">
<paragraph id="8B8C1C71FFF64516FE18FCD7FC6B0A8F" blockId="30.[394,989,888,912]" box="[394,989,888,912]" pageId="30" pageNumber="2017">
<heading id="D0C4AB1DFFF64516FE18FCD7FC6B0A8F" box="[394,989,888,912]" centered="true" fontSize="10" level="2" pageId="30" pageNumber="2017" reason="2">
<taxonomicName id="4C3367F2FFF64516FE18FCD7FC6B0A8F" ID-CoL="85R5P" authority="Sesma and Dozsa-Farkas, 1993" authorityName="Sesma and Dozsa-Farkas" authorityYear="1993" box="[394,989,888,912]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="30" pageNumber="2017" phylum="Annelida" rank="species" species="miraflores">
<emphasis id="B947C063FFF64516FE18FCD7FDD80A8F" bold="true" box="[394,622,888,912]" italics="true" pageId="30" pageNumber="2017">Fridericia miraflores</emphasis>
<bibRefCitation id="EFA26180FFF64516FDE5FCD7FC6B0A8F" author="Sesma V &amp; Dozsa-Farkas K" box="[631,989,888,912]" pageId="30" pageNumber="2017" pagination="249 - 265" refId="ref17151" refString="Sesma V, Dozsa-Farkas K. 1993. Descriptions of seven new species of Enchytraeidae (Oligochaeta) from Spain. Acta Zool Hung. 39: 249 - 265." type="journal article" year="1993">Sesma and Dózsa-Farkas, 1993</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3294FFAFFF64516FF46FC17FD860D70" pageId="30" pageNumber="2017" type="reference_group">
<paragraph id="8B8C1C71FFF64516FF46FC17FBD70ACF" blockId="30.[212,1172,952,1135]" box="[212,1121,952,976]" pageId="30" pageNumber="2017">
<taxonomicName id="4C3367F2FFF64516FF46FC17FC720ACF" authority="Sesma and Dozsa-Farkas, 1993, pp. 252 - 254" authorityName="Sesma and Dozsa-Farkas" authorityPageNumber="252 - 254" authorityYear="1993" box="[212,964,952,976]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="30" pageNumber="2017" phylum="Annelida" rank="species" species="miraflores">
<emphasis id="B947C063FFF64516FF46FC17FE0E0ACF" box="[212,440,952,976]" italics="true" pageId="30" pageNumber="2017">Fridericia miraflores</emphasis>
<bibRefCitation id="EFA26180FFF64516FE52FC17FC910ACF" author="Sesma V &amp; Dozsa-Farkas K" box="[448,807,952,976]" pageId="30" pageNumber="2017" pagination="249 - 265" refId="ref17151" refString="Sesma V, Dozsa-Farkas K. 1993. Descriptions of seven new species of Enchytraeidae (Oligochaeta) from Spain. Acta Zool Hung. 39: 249 - 265." type="journal article" year="1993">Sesma and Dózsa-Farkas, 1993</bibRefCitation>
, pp. 252–254
</taxonomicName>
, figures 8–14.
</paragraph>
<paragraph id="8B8C1C71FFF64516FF46FC77FE3A0D2F" blockId="30.[212,1172,952,1135]" pageId="30" pageNumber="2017">
<taxonomicName id="4C3367F2FFF64516FF46FC77FD570AEF" authority=", Rota 1995, pp. 211 - 212" authorityName=", Rota" authorityPageNumber="211 - 212" authorityYear="1995" box="[212,737,984,1008]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="30" pageNumber="2017" phylum="Annelida" rank="species" species="miraflores">
<emphasis id="B947C063FFF64516FF46FC77FE0F0AEF" box="[212,441,984,1008]" italics="true" pageId="30" pageNumber="2017">Fridericia miraflores</emphasis>
,
<bibRefCitation id="EFA26180FFF64516FE58FC76FDF40AEF" author="Rota E" box="[458,578,984,1008]" pageId="30" pageNumber="2017" pagination="183 - 231" refId="ref16775" refString="Rota E. 1995. Italian Enchytraeidae (Oligochaeta). I. Boll Zool. 62: 183 - 231." type="journal article" year="1995">Rota 1995</bibRefCitation>
, pp. 211–212
</taxonomicName>
, tables 5–6;
<bibRefCitation id="EFA26180FFF64516FCE6FC76FBDB0AEF" author="Rota E &amp; Healy B" box="[884,1133,984,1008]" pageId="30" pageNumber="2017" pagination="29 - 64" refId="ref16915" refString="Rota E, Healy B. 1999. A taxonomic study of some Swedish Enchytraeidae (Oligochaeta), with descriptions of four new species and notes on the genus Fridericia. J Nat Hist. 33: 29 - 64." type="journal article" year="1999">Rota and Healy 1999</bibRefCitation>
, p. 58;
<bibRefCitation id="EFA26180FFF64516FE8EFC56FE790D0F" author="Rota E &amp; Caruso T &amp; Monaci F &amp; Baldantoni D &amp; De Nicola &amp; Iovieno P &amp; Bargagli R" box="[284,463,1016,1040]" pageId="30" pageNumber="2017" pagination="268 - 276" refId="ref16835" refString="Rota E, Caruso T, Monaci F, Baldantoni D, De Nicola F, Iovieno P, Bargagli R. 2013. Effects of soil pollutants, biogeochemistry and microbiology on the distribution and composition of enchytraeid communities in urban and suburban holm oak stands. Environ Poll. 179: 268 - 276." type="journal article" year="2013">Rota et al. 2013</bibRefCitation>
, table 1 and figure 3 (species ‘s25’);
<bibRefCitation id="EFA26180FFF64516FCF0FC56FBA30D0F" author="Rota E &amp; Caruso T &amp; Bargagli R" box="[866,1045,1016,1040]" pageId="30" pageNumber="2017" pagination="83 - 91" refId="ref16797" refString="Rota E, Caruso T, Bargagli R. 2014. Community structure, diversity and spatial organization of enchytraeids in Mediterranean urban holm oak stands. Eur J Soil Biol. 62: 83 - 91." type="journal article" year="2014">Rota et al. 2014</bibRefCitation>
, tables 1, 2 and Suppl. 1.
</paragraph>
<paragraph id="8B8C1C71FFF64516FF46FB97FD860D70" blockId="30.[212,1172,952,1135]" pageId="30" pageNumber="2017">
<taxonomicName id="4C3367F2FFF64516FF46FB97FC940D4F" authority=", Schmelz 2003, pp. 330 - 333" authorityName=", Schmelz" authorityPageNumber="330 - 333" authorityYear="2003" box="[212,802,1080,1104]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="30" pageNumber="2017" phylum="Annelida" rank="species" species="sylvatica">
<emphasis id="B947C063FFF64516FF46FB97FE040D4F" box="[212,434,1080,1104]" italics="true" pageId="30" pageNumber="2017">Fridericia sylvatica</emphasis>
,
<bibRefCitation id="EFA26180FFF64516FE5EFB97FDC70D4F" author="Schmelz RM" box="[460,625,1080,1104]" pageId="30" pageNumber="2017" pagination="1 - 415" refId="ref17019" refString="Schmelz RM. 2003. Taxonomy of Fridericia (Oligochaeta, Enchytraeidae). Revision of species with morphological and biochemical methods. Abh Naturwiss Ver Hamburg (N. F.). 38: 1 - 415." type="journal article" year="2003">Schmelz 2003</bibRefCitation>
, pp. 330–333
</taxonomicName>
, figure 65A–H;
<bibRefCitation id="EFA26180FFF64516FC68FB97FE3A0D70" author="Schmelz RM &amp; Collado R" pageId="30" pageNumber="2017" pagination="1 - 176" refId="ref17059" refString="Schmelz RM, Collado R. 2010. A guide to European terrestrial and freshwater species of Enchytraeidae (Oligochaeta). Soil Organ. 82: 1 - 176." type="journal article" year="2010">Schmelz and Collado 2010</bibRefCitation>
, pp. 130, 142.
</paragraph>
</subSubSection>
<subSubSection id="C3294FFAFFF64516FF46FB19FDD90C4A" pageId="30" pageNumber="2017" type="materials_examined">
<paragraph id="8B8C1C71FFF64516FF46FB19FE110DD1" blockId="30.[212,1172,1206,1365]" box="[212,423,1206,1230]" pageId="30" pageNumber="2017">
<heading id="D0C4AB1DFFF64516FF46FB19FE110DD1" box="[212,423,1206,1230]" fontSize="10" level="3" pageId="30" pageNumber="2017" reason="5">
<emphasis id="B947C063FFF64516FF46FB19FE110DD1" box="[212,423,1206,1230]" italics="true" pageId="30" pageNumber="2017">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B8C1C71FFF64516FF46FB71FBDB0DE9" blockId="30.[212,1172,1206,1365]" box="[212,1133,1246,1271]" pageId="30" pageNumber="2017">
Published material. Specimens from
<collectingRegion id="49F7D293FFF64516FDE7FB70FC630DE9" box="[629,981,1246,1271]" isEnumeration="true" pageId="30" pageNumber="2017">Tuscany, Latium and Calabria</collectingRegion>
(
<bibRefCitation id="EFA26180FFF64516FC77FB71FBEA0DE9" author="Rota E" box="[997,1116,1246,1270]" pageId="30" pageNumber="2017" pagination="183 - 231" refId="ref16775" refString="Rota E. 1995. Italian Enchytraeidae (Oligochaeta). I. Boll Zool. 62: 183 - 231." type="journal article" year="1995">Rota 1995</bibRefCitation>
).
</paragraph>
<paragraph id="8B8C1C71FFF64516FF6CFB51FDD90C4A" blockId="30.[212,1172,1206,1365]" pageId="30" pageNumber="2017">
<materialsCitation id="3B5B162CFFF64516FF6CFB51FDD50C2A" ID-GBIF-Occurrence="2848394319" collectingDate="2009-05-14" collectingDateMax="2009-10-26" country="Italy" location="Campania" pageId="30" pageNumber="2017" specimenCount="12" stateProvince="Campania">
New material (in the author’ s collection).
<specimenCount id="9D35D7F8FFF64516FD63FB51FC3F0C09" box="[753,905,1278,1302]" pageId="30" pageNumber="2017" type="generic">12 specimens</specimenCount>
from
<collectingCountry id="F3245CE1FFF64516FC45FB51FBB80C09" box="[983,1038,1278,1302]" name="Italy" pageId="30" pageNumber="2017">Italy</collectingCountry>
,
<collectingRegion id="49F7D293FFF64516FB8DFB51FB220C09" box="[1055,1172,1278,1302]" country="Italy" name="Campania" pageId="30" pageNumber="2017">Campania</collectingRegion>
(Ca-1),
<collectingDate id="EFC9C359FFF64516FEBFFAB2FDD50C2A" box="[301,611,1309,1333]" isEnumeration="true" pageId="30" pageNumber="2017">
<date id="FF8D3AB1FFF64516FEBFFAB2FDD50C2A" box="[301,611,1309,1333]" pageId="30" pageNumber="2017" value="2009-05-14" valueMax="2009-10-27">14.05.2009 and 27.10.2009</date>
</collectingDate>
</materialsCitation>
.
<materialsCitation id="3B5B162CFFF64516FDE0FAB2FDDD0C4A" ID-GBIF-Occurrence="2848394327" collectingDate="2009-05-13" collectingDateMax="2009-10-26" collectionCode="N" country="Italy" location="Campania" pageId="30" pageNumber="2017" specimenCode="N2, N3" specimenCount="32" stateProvince="Campania">
<specimenCount id="9D35D7F8FFF64516FDE0FAB2FCBE0C2A" box="[626,776,1309,1333]" pageId="30" pageNumber="2017" type="generic">32 specimens</specimenCount>
from
<collectingRegion id="49F7D293FFF64516FCC7FAB2FC7C0C2A" box="[853,970,1309,1333]" country="Italy" name="Campania" pageId="30" pageNumber="2017">Campania</collectingRegion>
(Ca-2), plots
<specimenCode id="DB95B40AFFF64516FBE2FAB2FB220C2A" box="[1136,1172,1309,1333]" collectionCode="N" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="30" pageNumber="2017" type="Herbarium">N2</specimenCode>
and
<specimenCode id="DB95B40AFFF64516FE95FA92FE9A0C4A" box="[263,300,1341,1365]" collectionCode="N" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="30" pageNumber="2017" type="Herbarium">N3</specimenCode>
,
<collectingDate id="EFC9C359FFF64516FEA8FA92FDDD0C4A" box="[314,619,1341,1365]" isEnumeration="true" pageId="30" pageNumber="2017">
<date id="FF8D3AB1FFF64516FEA8FA92FDDD0C4A" box="[314,619,1341,1365]" pageId="30" pageNumber="2017" value="2009-05-13">13.05.2009 and 26.10.2009</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3294FFAFFF64517FF46FA33FE130BA1" lastPageId="31" lastPageNumber="2018" pageId="30" pageNumber="2017" type="discussion">
<paragraph id="8B8C1C71FFF64516FF46FA33FE800CAB" blockId="30.[212,1172,1436,1691]" box="[212,310,1436,1460]" pageId="30" pageNumber="2017">
<heading id="D0C4AB1DFFF64516FF46FA33FE800CAB" box="[212,310,1436,1460]" fontSize="10" level="3" pageId="30" pageNumber="2017" reason="5">
<emphasis id="B947C063FFF64516FF46FA33FE800CAB" box="[212,310,1436,1460]" italics="true" pageId="30" pageNumber="2017">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8C1C71FFF64516FF46FA6BFC640F44" blockId="30.[212,1172,1436,1691]" pageId="30" pageNumber="2017">
This is one of few species in
<taxonomicName id="4C3367F2FFF64516FD89FA6BFD300CC3" authorityName="Michaelsen" authorityYear="1889" box="[539,646,1476,1500]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="30" pageNumber="2017" phylum="Annelida" rank="genus">
<emphasis id="B947C063FFF64516FD89FA6BFD300CC3" box="[539,646,1476,1500]" italics="true" pageId="30" pageNumber="2017">Fridericia</emphasis>
</taxonomicName>
having the clitellum interrupted both dorsally and ventrally (
<bibRefCitation id="EFA26180FFF64516FEEEFA4BFDC00CE3" author="Rota E &amp; Healy B" box="[380,630,1508,1532]" pageId="30" pageNumber="2017" pagination="29 - 64" refId="ref16915" refString="Rota E, Healy B. 1999. A taxonomic study of some Swedish Enchytraeidae (Oligochaeta), with descriptions of four new species and notes on the genus Fridericia. J Nat Hist. 33: 29 - 64." type="journal article" year="1999">Rota and Healy 1999</bibRefCitation>
, p. 58; see also
<bibRefCitation id="EFA26180FFF64516FCA6FA4BFC660CE3" author="Schmelz RM" box="[820,976,1508,1532]" pageId="30" pageNumber="2017" pagination="1 - 415" refId="ref17019" refString="Schmelz RM. 2003. Taxonomy of Fridericia (Oligochaeta, Enchytraeidae). Revision of species with morphological and biochemical methods. Abh Naturwiss Ver Hamburg (N. F.). 38: 1 - 415." type="journal article" year="2003">Schmelz 2003</bibRefCitation>
, as
<taxonomicName id="4C3367F2FFF64516FB91FA4AFB320CE3" authorityName=", Schmelz" authorityYear="2003" box="[1027,1156,1508,1532]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="30" pageNumber="2017" phylum="Annelida" rank="species" species="sylvatica">
<emphasis id="B947C063FFF64516FB91FA4AFB320CE3" box="[1027,1156,1508,1532]" italics="true" pageId="30" pageNumber="2017">F. sylvatica</emphasis>
</taxonomicName>
). This feature and the occurrence of ventral pharyngeal glands in VII (
<bibRefCitation id="EFA26180FFF64516FC46F9ACFBD80F04" author="Schmelz RM" box="[980,1134,1539,1563]" pageId="30" pageNumber="2017" pagination="1 - 415" refId="ref17019" refString="Schmelz RM. 2003. Taxonomy of Fridericia (Oligochaeta, Enchytraeidae). Revision of species with morphological and biochemical methods. Abh Naturwiss Ver Hamburg (N. F.). 38: 1 - 415." type="journal article" year="2003">Schmelz 2003</bibRefCitation>
, as
<taxonomicName id="4C3367F2FFF64516FF46F98BFEE50F23" authorityName=", Schmelz" authorityYear="2003" box="[212,339,1572,1596]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="30" pageNumber="2017" phylum="Annelida" rank="species" species="sylvatica">
<emphasis id="B947C063FFF64516FF46F98BFEE50F23" box="[212,339,1572,1596]" italics="true" pageId="30" pageNumber="2017">F. sylvatica</emphasis>
</taxonomicName>
) is confirmed for the present and previous material from
<collectingCountry id="F3245CE1FFF64516FC70F98CFBA00F24" box="[994,1046,1571,1595]" name="Italy" pageId="30" pageNumber="2017">Italy</collectingCountry>
(as well as for specimens from
<collectingCountry id="F3245CE1FFF64516FE25F9ECFDB80F44" box="[439,526,1603,1627]" name="Algeria" pageId="30" pageNumber="2017">Algeria</collectingCountry>
), including those with short diverticula.
</paragraph>
<paragraph id="8B8C1C71FFF64517FF6CF9CCFE130BA1" blockId="30.[212,1172,1436,1691]" lastBlockId="31.[144,1104,168,703]" lastPageId="31" lastPageNumber="2018" pageId="30" pageNumber="2017">
The synonymy of this species with the poorly defined
<taxonomicName id="4C3367F2FFF64516FCC5F9CBFBD40F64" authority="Healy, 1979" authorityName="Healy" authorityYear="1979" box="[855,1122,1635,1659]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="30" pageNumber="2017" phylum="Annelida" rank="species" species="sylvatica">
<emphasis id="B947C063FFF64516FCC5F9CBFC630F64" box="[855,981,1635,1659]" italics="true" pageId="30" pageNumber="2017">F. sylvatica</emphasis>
<bibRefCitation id="EFA26180FFF64516FC4EF9CCFBD40F64" author="Healy B" box="[988,1122,1635,1659]" pageId="30" pageNumber="2017" pagination="39 - 70" refId="ref16442" refString="Healy B. 1979. Records of Enchytraeidae (Oligochaeta) in Ireland. J Life Sci R Dubl Soc. 1: 39 - 70." type="journal article" year="1979">Healy, 1979</bibRefCitation>
</taxonomicName>
was established by
<bibRefCitation id="EFA26180FFF64516FE12F92CFD990F84" author="Schmelz RM" box="[384,559,1667,1691]" pageId="30" pageNumber="2017" pagination="1 - 415" refId="ref17019" refString="Schmelz RM. 2003. Taxonomy of Fridericia (Oligochaeta, Enchytraeidae). Revision of species with morphological and biochemical methods. Abh Naturwiss Ver Hamburg (N. F.). 38: 1 - 415." type="journal article" year="2003">Schmelz (2003)</bibRefCitation>
, on the basis of the similar chaetal formula and the laterally projecting spermathecal diverticula, and despite important differences in the two original descriptions (size of chaetae, origin of the dorsal vessel, variation in size of the spermathecal gland) and the lack of
<typeStatus id="5488A2D3FFF74517FD07FF46FD73081E" box="[661,709,233,257]" pageId="31" pageNumber="2018">type</typeStatus>
material of
<taxonomicName id="4C3367F2FFF74517FCCFFF46FC56081F" authorityName="Healy" authorityYear="1979" box="[861,992,232,256]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="2018" phylum="Annelida" rank="species" species="sylvatica">
<emphasis id="B947C063FFF74517FCCFFF46FC56081F" box="[861,992,232,256]" italics="true" pageId="31" pageNumber="2018">F. sylvatica</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFA26180FFF74517FC61FF47FF60083F" author="Schmelz RM" pageId="31" pageNumber="2018" pagination="1 - 415" refId="ref17019" refString="Schmelz RM. 2003. Taxonomy of Fridericia (Oligochaeta, Enchytraeidae). Revision of species with morphological and biochemical methods. Abh Naturwiss Ver Hamburg (N. F.). 38: 1 - 415." type="journal article" year="2003">Schmelz (2003)</bibRefCitation>
himself discovered that the sole specimen (submature and incomplete) labelled as
<typeStatus id="5488A2D3FFF74517FF21FE87FF55085F" box="[179,227,296,320]" pageId="31" pageNumber="2018">type</typeStatus>
of
<taxonomicName id="4C3367F2FFF74517FE81FE86FE20085F" authorityName="Healy" authorityYear="1979" box="[275,406,296,320]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="2018" phylum="Annelida" rank="species" species="sylvatica">
<emphasis id="B947C063FFF74517FE81FE86FE20085F" box="[275,406,296,320]" italics="true" pageId="31" pageNumber="2018">F. sylvatica</emphasis>
</taxonomicName>
was not compatible with any of the key characters in the original diagnosis. Although
<taxonomicName id="4C3367F2FFF74517FE4CFEE6FDD8087F" authorityName=", Rota" authorityYear="1995" box="[478,622,328,352]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="2018" phylum="Annelida" rank="species" species="miraflores">
<emphasis id="B947C063FFF74517FE4CFEE6FDD8087F" box="[478,622,328,352]" italics="true" pageId="31" pageNumber="2018">F. miraflores</emphasis>
</taxonomicName>
can be collected at the
<typeStatus id="5488A2D3FFF74517FC11FEE7FC05087F" box="[899,947,328,352]" pageId="31" pageNumber="2018">type</typeStatus>
locality of
<taxonomicName id="4C3367F2FFF74517FBABFEE6FF46089F" authorityName="Healy" authorityYear="1979" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="2018" phylum="Annelida" rank="species" species="sylvatica">
<emphasis id="B947C063FFF74517FBABFEE6FF46089F" italics="true" pageId="31" pageNumber="2018">F. sylvatica</emphasis>
</taxonomicName>
(UCD campus,
<collectingRegion id="49F7D293FFF74517FE29FEC8FDBD0860" box="[443,523,359,383]" country="Ireland" name="South Dublin" pageId="31" pageNumber="2018">Dublin</collectingRegion>
) (
<bibRefCitation id="EFA26180FFF74517FDBBFEC8FD120860" author="Rota E" box="[553,676,359,383]" pageId="31" pageNumber="2018" pagination="183 - 231" refId="ref16775" refString="Rota E. 1995. Italian Enchytraeidae (Oligochaeta). I. Boll Zool. 62: 183 - 231." type="journal article" year="1995">Rota 1995</bibRefCitation>
), I believe the circumstances rather suggest that
<taxonomicName id="4C3367F2FFF74517FEBAFE27FE1A08BF" authorityName="Healy" authorityYear="1979" box="[296,428,392,416]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="2018" phylum="Annelida" rank="species" species="sylvatica">
<emphasis id="B947C063FFF74517FEBAFE27FE1A08BF" box="[296,428,392,416]" italics="true" pageId="31" pageNumber="2018">F. sylvatica</emphasis>
</taxonomicName>
should be considered a nomen dubium, in that (1) ‘the taxonomic identity of the taxon cannot be determined from its existing name-bearing type’ (ICZN 1999: Art. 75.6), and (2) the original description of
<taxonomicName id="4C3367F2FFF74517FCF6FE67FC5508C0" authorityName="Healy" authorityYear="1979" box="[868,995,455,479]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="2018" phylum="Annelida" rank="species" species="sylvatica">
<emphasis id="B947C063FFF74517FCF6FE67FC5508C0" box="[868,995,455,479]" italics="true" pageId="31" pageNumber="2018">F. sylvatica</emphasis>
</taxonomicName>
probably involved one or more different species (in particular, the reported variation in the spermatheca towards more globular ampullar diverticula and larger ectal gland point to
<taxonomicName id="4C3367F2FFF74517FF23FD87FE6E0B20" authority="Rota, 1995" authorityName="Rota" authorityYear="1995" box="[177,472,551,575]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="2018" phylum="Annelida" rank="species" species="globuligera">
<emphasis id="B947C063FFF74517FF23FD87FEF80B20" box="[177,334,551,575]" italics="true" pageId="31" pageNumber="2018">F. globuligera</emphasis>
<bibRefCitation id="EFA26180FFF74517FECAFD88FE6E0B20" author="Rota E" box="[344,472,551,575]" pageId="31" pageNumber="2018" pagination="183 - 231" refId="ref16775" refString="Rota E. 1995. Italian Enchytraeidae (Oligochaeta). I. Boll Zool. 62: 183 - 231." type="journal article" year="1995">Rota, 1995</bibRefCitation>
</taxonomicName>
, a species also present in the UCD area; own unpublished record). In conclusion, the originally better described and type-based
<taxonomicName id="4C3367F2FFF74517FC66FDE7FF7B0B60" authorityName=", Rota" authorityYear="1995" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="2018" phylum="Annelida" rank="species" species="miraflores">
<emphasis id="B947C063FFF74517FC66FDE7FF7B0B60" italics="true" pageId="31" pageNumber="2018">F. miraflores</emphasis>
</taxonomicName>
is to be regarded as a valid name. This choice is consistent with the reasoning by
<bibRefCitation id="EFA26180FFF74517FF20FD29FEEB0B81" author="Schmelz RM" box="[178,349,646,670]" pageId="31" pageNumber="2018" pagination="1 - 415" refId="ref17019" refString="Schmelz RM. 2003. Taxonomy of Fridericia (Oligochaeta, Enchytraeidae). Revision of species with morphological and biochemical methods. Abh Naturwiss Ver Hamburg (N. F.). 38: 1 - 415." type="journal article" year="2003">Schmelz (2003)</bibRefCitation>
on the analogous cases involving
<taxonomicName id="4C3367F2FFF74517FD48FD27FBA80B81" authority="Schmidegg, 1938" authorityName="Schmidegg" authorityYear="1938" box="[730,1054,646,671]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="2018" phylum="Annelida" rank="species" species="tirolensis">
<emphasis id="B947C063FFF74517FD48FD27FCEC0B80" box="[730,858,647,671]" italics="true" pageId="31" pageNumber="2018">F. tirolensis</emphasis>
<bibRefCitation id="EFA26180FFF74517FCF3FD29FBA80B81" author="Schmidegg E" box="[865,1054,646,670]" pageId="31" pageNumber="2018" pagination="26 - 71" refId="ref17127" refString="Schmidegg E. 1938. Die Enchytraeiden des Hochgebirges der Nordtiroler Kalkalpen. Ber Naturw-med Ver Innsbruck. 45: 26 - 71." type="journal article" year="1938">Schmidegg, 1938</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C3367F2FFF74517FF02FD08FE2B0BA1" authority="Healy, 1975" authorityName="Healy" authorityYear="1975" box="[144,413,678,703]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="2018" phylum="Annelida" rank="species" species="discifera">
<emphasis id="B947C063FFF74517FF02FD08FEBA0BA0" box="[144,268,679,703]" italics="true" pageId="31" pageNumber="2018">F. discifera</emphasis>
<bibRefCitation id="EFA26180FFF74517FE87FD09FE2B0BA1" author="Healy B" box="[277,413,678,702]" pageId="31" pageNumber="2018" pagination="315 - 326" refId="ref16412" refString="Healy B. 1975. A description of five new species of Enchytraeidae (Oligochaeta) from Ireland. Zool J Linn Soc. 56: 315 - 326." type="journal article" year="1975">Healy, 1975</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3294FFAFFF74517FF02FD51FCF10A21" box="[144,839,766,830]" pageId="31" pageNumber="2018" type="distribution">
<paragraph id="8B8C1C71FFF74517FF02FD51FEA20A09" blockId="31.[144,839,766,830]" box="[144,276,766,790]" pageId="31" pageNumber="2018">
<heading id="D0C4AB1DFFF74517FF02FD51FEA20A09" box="[144,276,766,790]" fontSize="10" level="3" pageId="31" pageNumber="2018" reason="5">
<emphasis id="B947C063FFF74517FF02FD51FEA20A09" box="[144,276,766,790]" italics="true" pageId="31" pageNumber="2018">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B8C1C71FFF74517FF02FC89FCF10A21" blockId="31.[144,839,766,830]" box="[144,839,806,830]" pageId="31" pageNumber="2018">Euro-Mediterranean. Present in both seasons at Naples sites.</paragraph>
</subSubSection>
</treatment>
</document>
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