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<document id="51B81596FB0E7368FE69C020A86D0F15" ID-CLB-Dataset="30484" ID-DOI="10.11646/zootaxa.4396.1.1" ID-GBIF-Dataset="918f2145-a708-460e-b19b-4f67e6c99b30" ID-ISSN="1175-5326" ID-Zenodo-Dep="1202723" ID-ZooBank="8EEBE6DE-0ECC-4B9C-AD14-01438291782B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1521441296340" checkinUser="plazi" docAuthor="Smith, Patrick M., Paterson, John R. &amp; Brock, Glenn A." docDate="2018" docId="038CC426FFA4FF84FF399DE9FEE4FDFB" docLanguage="en" docName="zootaxa.4396.1.1.pdf" docOrigin="Zootaxa 4396 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ammagnostus antarcticus Bentley, Jago &amp; Cooper 2009" docType="treatment" docVersion="7" lastPageNumber="14" masterDocId="FFB5BC5EFFAFFF89FFAE9D71FF99FFE3" masterDocTitle="Trilobites and agnostids from the Goyder Formation (Cambrian Series 3, Guzhangian; Mindyallan), Amadeus Basin, central Australia" masterLastPageNumber="67" masterPageNumber="1" pageNumber="12" updateTime="1698532001681" updateUser="ExternalLinkService">
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<mods:title id="491320217E00E31E9CF50EEAB8C021FE">Trilobites and agnostids from the Goyder Formation (Cambrian Series 3, Guzhangian; Mindyallan), Amadeus Basin, central Australia</mods:title>
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<mods:namePart id="96521647846F282CDB654D51718BFF59">Smith, Patrick M.</mods:namePart>
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<mods:namePart id="CA9FF745EA2DFF56335A1EFFEC93AEF9">Paterson, John R.</mods:namePart>
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<mods:namePart id="544D9F97CCE8B33F482A98E1EE722AA2">Brock, Glenn A.</mods:namePart>
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<subSubSection id="C33F26BBFFA4FF82FF399DE9FF40FF30" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8B9A7530FFA4FF82FF399DE9FCD0FF51" blockId="11.[151,841,152,211]" box="[151,841,152,178]" pageId="11" pageNumber="12">
<heading id="D0D2C25CFFA4FF82FF399DE9FCD0FF51" bold="true" box="[151,841,152,178]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<taxonomicName id="4C250EB3FFA4FF82FF399DE9FCD0FF51" authority="Bentley, Jago &amp; Cooper, 2009" authorityName="Bentley, Jago &amp; Cooper" authorityYear="2009" box="[151,841,152,178]" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="antarcticus">
<emphasis id="B951A922FFA4FF82FF399DE9FCD0FF51" bold="true" box="[151,841,152,178]" pageId="11" pageNumber="12">
<emphasis id="B951A922FFA4FF82FF399DE9FE4BFF51" bold="true" box="[151,466,152,178]" italics="true" pageId="11" pageNumber="12">Ammagnostus antarcticus</emphasis>
Bentley, Jago &amp; Cooper, 2009
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B9A7530FFA4FF82FF399DCBFF40FF30" blockId="11.[151,841,152,211]" box="[151,217,186,211]" pageId="11" pageNumber="12">
<figureCitation id="131E69B5FFA4FF82FF399DCBFF40FF30" box="[151,217,186,211]" captionStart="FIGURE 5" captionStartId="12.[151,250,1722,1744]" captionTargetBox="[191,1404,204,1700]" captionTargetId="figure@12.[178,1414,193,1711]" captionTargetPageId="12" captionText="FIGURE 5. Ammagnostus antarcticus Bentley, Jago &amp; Cooper, 2009. All specimens come from sample GOY/173.6. A, S, T, U, CPC42177, partial cephalon; A, dorsal view; S, oblique anterolateral view; T, lateral view; U, anterior view.B, CPC42178, partial cephalon. C, CPC42179, partial cephalon. D, CPC42180, partial cephalon. E, CPC42181, partial cephalon. F, CPC42182, partial cephalon.G, CPC42183, partial cephalon.H, CPC42184, partial cephalon.I, CPC42185,partial cephalon. J, CPC42186, partial cephalon. K, CPC42187, partial cephalon.L, CPC42188, partial cephalon. M, CPC42189, partial cephalon. N, CPC42190, partial cephalon. O, CPC42191, partial cephalon. P, CPC42192, partial cephalon. Q, CPC42193, partial cephalon. R, CPC42194, partial cephalon. V, CPC42195, partial cephalon. W, Z, A 1, CPC42196, partial pygidium; W, dorsal view; Z, lateral view; A 1, posterior view. X, CPC42197, partial pygidium. Y, CPC42198, partial pygidium. B 1, CPC42199, partial pygidium. C 1, CPC42200, partial pygidium. D 1, CPC42201, partial pygidium. E 1, CPC42202, partial pygidium. F 1, CPC42203, partial pygidium.All scale bars are 1 mm." httpUri="https://zenodo.org/record/1202733/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33F26BBFFA4FF82FF399C73FC5DFEB5" pageId="11" pageNumber="12" type="description">
<paragraph id="8B9A7530FFA4FF82FF399C73FCE8FEFB" blockId="11.[151,1001,258,342]" box="[151,881,258,280]" pageId="11" pageNumber="12">
?1967
<emphasis id="B951A922FFA4FF82FF759C73FE36FEFB" box="[219,431,258,280]" italics="true" pageId="11" pageNumber="12">
Agnostoglossa
<taxonomicName id="4C250EB3FFA4FF82FEDB9C73FE36FEFB" authorityName="Opik" authorityYear="1967" box="[373,431,258,280]" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bassa">bassa</taxonomicName>
</emphasis>
; Öpik, p. 146, pl. 60, figs 614, text-fig. 44.
</paragraph>
<paragraph id="8B9A7530FFA4FF82FF399C50FC5DFEB5" blockId="11.[151,1001,258,342]" pageId="11" pageNumber="12">
?1989
<taxonomicName id="4C250EB3FFA4FF82FF759C53FEF0FED4" authorityName="Opik" authorityYear="1967" box="[219,361,290,311]" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B951A922FFA4FF82FF759C53FEF0FED4" box="[219,361,290,311]" italics="true" pageId="11" pageNumber="12">Ammagnostus</emphasis>
</taxonomicName>
sp. indet.; Wang, Mills, Webby &amp; Shergold, p. 111, fig. 4AD. 2009
<taxonomicName id="4C250EB3FFA4FF82FF7E9C30FE4CFEB5" authorityName="Bentley, Jago &amp; Cooper" authorityYear="2009" box="[208,469,321,342]" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="antarcticus">
<emphasis id="B951A922FFA4FF82FF7E9C30FE4CFEB5" box="[208,469,321,342]" italics="true" pageId="11" pageNumber="12">Ammagnostus antarcticus</emphasis>
</taxonomicName>
; Bentley, Jago &amp; Cooper, p. 171, 172, fig. 4AI.
</paragraph>
</subSubSection>
<subSubSection id="C33F26BBFFA4FF82FF399CF3FE55FE5C" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8B9A7530FFA4FF82FF399CF3FE55FE5C" blockId="11.[151,1437,386,2031]" pageId="11" pageNumber="12">
<emphasis id="B951A922FFA4FF82FF399CF3FE9FFE78" bold="true" box="[151,262,386,411]" pageId="11" pageNumber="12">Material.</emphasis>
Nineteen cephala and eight pygidia figured, CPC42177CPC42203. 26 cephala and eleven pygidia not figured (mostly fragments).
</paragraph>
</subSubSection>
<subSubSection id="C33F26BBFFA4FF82FF699CBBFDA4FB34" pageId="11" pageNumber="12" type="description">
<paragraph id="8B9A7530FFA4FF82FF699CBBFE3DFCC4" blockId="11.[151,1437,386,2031]" pageId="11" pageNumber="12">
<emphasis id="B951A922FFA4FF82FF699CBBFEC0FE00" bold="true" box="[199,345,458,483]" pageId="11" pageNumber="12">Description.</emphasis>
Cephalon and pygidium each attaining about
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in length; nonscrobiculate; surface smooth. Cephalon subquadrate in outline, about 94% as long as wide, nonspinose, moderately convex; border reasonably wide (sag., exsag.), flattened; border furrow well developed; preglabellar median furrow faintly developed or absent. Glabella 7073% (mean 71%, n = 4) as long (sag.) as cephalon. Small anterior lobe clearly defined, semiovate in shape, obtusely angular to rounded anteriorly, occupying 2930% (mean 30%, n = 4) of glabellar length. F3 deeply incised, strongly bowed rearward in a V-shape, bowing is weaker in smaller specimens. Posterior glabellar lobe wider (tr.) than anterior lobe, slightly tapers forward, convexity similar in both lobes. F2 furrows at glabellar mid-length (sag.), moderately developed as distinct notches, gently directed anteriorly for a short distance. F1 furrows effaced. Glabellar node indistinct, located slightly posterior of F2 furrows; basal lobes small, subtriangular in outline.
</paragraph>
<paragraph id="8B9A7530FFA4FF82FF699E43FE87FB50" blockId="11.[151,1437,386,2031]" pageId="11" pageNumber="12">Pygidium rounded to weakly quadrate in outline, 90% as long as wide, moderately convex, with short (exsag.) posterolateral spines, situated level with widest point in pygidial axis; border flattened, wide (sag., exsag.) and narrows distinctly toward anterior; border furrow moderately deep and wide (sag., exsag.). Axis wide (tr.) and long (sag., exsag.), reaching posterior border furrow; about 64 to 70% (mean 67%, n = 5) as wide as long and occupying 73% of pygidial length (sag.); slightly constricted across M2 and expanding to become wider behind F2; posteriorly rounded to obtusely rounded and wide (tr.), moderately convex. F1 and F2 furrows effaced. M1 lobe broad (tr.), lateral margins slightly convex. M2 lobe only distinguished from M1 by constriction. Axial node well developed, of moderate size, extending a short distance onto posteroaxis. Posteroaxis occupying about 58 to 63% (mean 60%, n = 3) of axial length, with maximum width slightly posterior of lobe mid-length. Small secondary axial node developed on some specimens, located a short distance posterior to the maximum width of the posteroaxis.</paragraph>
<paragraph id="8B9A7530FFA4FF82FF6999CEFDA4FB34" blockId="11.[151,1437,386,2031]" box="[199,573,1214,1239]" pageId="11" pageNumber="12">Hypostome and thorax unknown.</paragraph>
</subSubSection>
<subSubSection id="C33F26BBFFA4FF82FF699993FCCBF80C" pageId="11" pageNumber="12" type="discussion">
<paragraph id="8B9A7530FFA4FF82FF699993FF63F8D8" blockId="11.[151,1437,386,2031]" pageId="11" pageNumber="12">
<emphasis id="B951A922FFA4FF82FF699993FED5FB18" bold="true" box="[199,332,1250,1275]" pageId="11" pageNumber="12">Discussion.</emphasis>
The larger cephala and pygidia from the Goyder Formation most closely resemble those of
<taxonomicName id="4C250EB3FFA4FF82FF399876FCAAFAFC" authority="Bentley, Jago &amp; Cooper, 2009" authorityName="Bentley, Jago &amp; Cooper" authorityYear="2009" box="[151,819,1286,1311]" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="antarcticus">
<emphasis id="B951A922FFA4FF82FF399876FE27FAFD" box="[151,446,1287,1310]" italics="true" pageId="11" pageNumber="12">Ammagnostus antarcticus</emphasis>
Bentley, Jago &amp; Cooper, 2009
</taxonomicName>
described from the Mindyallan Spurs Formation in
<collectingCountry id="F33235A0FFA4FF82FF39985BFE96FAA0" box="[151,271,1322,1347]" name="Antarctica" pageId="11" pageNumber="12">Antarctica</collectingCountry>
. Similarities include: a long (sag.) anterior border with a shallow border furrow; a glabella that is approximately 70% the length (sag.) of the cephalon; F3 that is strongly bowed rearward in a V-shape; and an indistinct glabellar node. They also share a pygidium that is wider than long with a well developed axial node that encroaches onto the posteroaxis, and a posterior border that is very wide (sag.), with small posterolateral spines. The
<typeStatus id="549ECB92FFA4FF82FF6598CBFEB4FA30" box="[203,301,1466,1491]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
cephalon of
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<emphasis id="B951A922FFA4FF82FE6A98CAFE4CFA31" box="[452,469,1467,1490]" italics="true" pageId="11" pageNumber="12">A</emphasis>
.
<emphasis id="B951A922FFA4FF82FE4898CDFDFAFA30" box="[486,611,1468,1491]" italics="true" pageId="11" pageNumber="12">antarcticus</emphasis>
</taxonomicName>
(Bentley
<emphasis id="B951A922FFA4FF82FD7B98CDFC91FA30" box="[725,776,1466,1491]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
. 2009, fig. 4E) is
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in length (sag.), which is more than 3 times larger than the largest specimen from the Goyder Formation (at
<quantity id="4CDDD8D5FFA4FF82FC5398AFFBCEFA15" box="[1021,1111,1502,1527]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="11" pageNumber="12" unit="mm" value="2.6">2.6 mm</quantity>
;
<figureCitation id="131E69B5FFA4FF82FBCD98AFFB27FA14" box="[1123,1214,1502,1527]" captionStart="FIGURE 5" captionStartId="12.[151,250,1722,1744]" captionTargetBox="[191,1404,204,1700]" captionTargetId="figure@12.[178,1414,193,1711]" captionTargetPageId="12" captionText="FIGURE 5. Ammagnostus antarcticus Bentley, Jago &amp; Cooper, 2009. All specimens come from sample GOY/173.6. A, S, T, U, CPC42177, partial cephalon; A, dorsal view; S, oblique anterolateral view; T, lateral view; U, anterior view.B, CPC42178, partial cephalon. C, CPC42179, partial cephalon. D, CPC42180, partial cephalon. E, CPC42181, partial cephalon. F, CPC42182, partial cephalon.G, CPC42183, partial cephalon.H, CPC42184, partial cephalon.I, CPC42185,partial cephalon. J, CPC42186, partial cephalon. K, CPC42187, partial cephalon.L, CPC42188, partial cephalon. M, CPC42189, partial cephalon. N, CPC42190, partial cephalon. O, CPC42191, partial cephalon. P, CPC42192, partial cephalon. Q, CPC42193, partial cephalon. R, CPC42194, partial cephalon. V, CPC42195, partial cephalon. W, Z, A 1, CPC42196, partial pygidium; W, dorsal view; Z, lateral view; A 1, posterior view. X, CPC42197, partial pygidium. Y, CPC42198, partial pygidium. B 1, CPC42199, partial pygidium. C 1, CPC42200, partial pygidium. D 1, CPC42201, partial pygidium. E 1, CPC42202, partial pygidium. F 1, CPC42203, partial pygidium.All scale bars are 1 mm." httpUri="https://zenodo.org/record/1202733/files/figure.png" pageId="11" pageNumber="12">Fig. 5A</figureCitation>
). The other cephala figured by Bentley
<emphasis id="B951A922FFA4FF82FED89B75FE33F9F8" box="[374,426,1538,1563]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
. (2009, fig. 4AC, F) are closer in size to the specimens reported here. Given the close morphological similarity to
<taxonomicName id="4C250EB3FFA4FF82FE7A9B56FDE8F9DD" authorityName="Bentley, Jago &amp; Cooper" authorityYear="2009" box="[468,625,1575,1598]" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="antarcticus">
<emphasis id="B951A922FFA4FF82FE7A9B56FE7CF9DD" box="[468,485,1575,1598]" italics="true" pageId="11" pageNumber="12">A</emphasis>
.
<emphasis id="B951A922FFA4FF82FE5A9B56FDE8F9DD" box="[500,625,1575,1598]" italics="true" pageId="11" pageNumber="12">antarcticus</emphasis>
</taxonomicName>
, it is considered likely that the Goyder Formation material represents a collection of mostly juvenile specimens or perhaps a geographic variant. Smaller cephala from the Goyder Formation are reminiscent of
<taxonomicName id="4C250EB3FFA4FF82FE479B1EFC2FF964" authority="(Lorenz, 1906)" authorityName="Lorenz" authorityYear="1906" baseAuthorityName="Lorenz" baseAuthorityYear="1906" box="[489,950,1646,1671]" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="laiwuensis">
<emphasis id="B951A922FFA4FF82FE479B1EFC9DF964" box="[489,772,1646,1671]" italics="true" pageId="11" pageNumber="12">Ammagnostus laiwuensis</emphasis>
(Lorenz, 1906)
</taxonomicName>
and
<taxonomicName id="4C250EB3FFA4FF82FC419B1EFE22F948" authority="(Guo &amp; Luo in Guo et al., 1996)" authorityName="Guo &amp; Luo in Guo et al." authorityYear="1996" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="quadratiformis">
<emphasis id="B951A922FFA4FF82FC419B1EFAA5F964" box="[1007,1340,1646,1671]" italics="true" pageId="11" pageNumber="12">Ammagnostus quadratiformis</emphasis>
(Guo &amp; Luo in Guo
<emphasis id="B951A922FFA4FF82FE839BE5FEFAF948" box="[301,355,1682,1707]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
., 1996)
</taxonomicName>
—both of which are synonyms according to
<bibRefCitation id="EFB408C1FFA4FF82FC7C9BE2FB71F948" box="[978,1256,1682,1707]" pageId="11" pageNumber="12" refId="ref44413" refString="Peng, S. C. &amp; Robison, R. A. (2000) Agnostoid biostratigraphy across the Middle-Upper Cambrian boundary in Hunan, China. Paleontological Society Memoir, 53, 1 - 104. https: // doi. org / 10.1666 / 0022 - 3360 (2000) 53 [1: ABATMC] 2.0. CO; 2" type="journal article">Peng &amp; Robison (2000)</bibRefCitation>
—based on the presence of a shorter (sag.) anterior border and a transverse F3 furrow (e.g. compare
<figureCitation id="131E69B5FFA4FF82FB2C9BC7FB64F92D" box="[1154,1277,1718,1743]" captionStart="FIGURE 5" captionStartId="12.[151,250,1722,1744]" captionTargetBox="[191,1404,204,1700]" captionTargetId="figure@12.[178,1414,193,1711]" captionTargetPageId="12" captionText="FIGURE 5. Ammagnostus antarcticus Bentley, Jago &amp; Cooper, 2009. All specimens come from sample GOY/173.6. A, S, T, U, CPC42177, partial cephalon; A, dorsal view; S, oblique anterolateral view; T, lateral view; U, anterior view.B, CPC42178, partial cephalon. C, CPC42179, partial cephalon. D, CPC42180, partial cephalon. E, CPC42181, partial cephalon. F, CPC42182, partial cephalon.G, CPC42183, partial cephalon.H, CPC42184, partial cephalon.I, CPC42185,partial cephalon. J, CPC42186, partial cephalon. K, CPC42187, partial cephalon.L, CPC42188, partial cephalon. M, CPC42189, partial cephalon. N, CPC42190, partial cephalon. O, CPC42191, partial cephalon. P, CPC42192, partial cephalon. Q, CPC42193, partial cephalon. R, CPC42194, partial cephalon. V, CPC42195, partial cephalon. W, Z, A 1, CPC42196, partial pygidium; W, dorsal view; Z, lateral view; A 1, posterior view. X, CPC42197, partial pygidium. Y, CPC42198, partial pygidium. B 1, CPC42199, partial pygidium. C 1, CPC42200, partial pygidium. D 1, CPC42201, partial pygidium. E 1, CPC42202, partial pygidium. F 1, CPC42203, partial pygidium.All scale bars are 1 mm." httpUri="https://zenodo.org/record/1202733/files/figure.png" pageId="11" pageNumber="12">Fig. 5K, P</figureCitation>
with
<bibRefCitation id="EFB408C1FFA4FF82FAEA9BC6FEA0F910" pageId="11" pageNumber="12" refId="ref44413" refString="Peng, S. C. &amp; Robison, R. A. (2000) Agnostoid biostratigraphy across the Middle-Upper Cambrian boundary in Hunan, China. Paleontological Society Memoir, 53, 1 - 104. https: // doi. org / 10.1666 / 0022 - 3360 (2000) 53 [1: ABATMC] 2.0. CO; 2" type="journal article">Peng &amp; Robison 2000</bibRefCitation>
, fig. 20.120.5 and
<bibRefCitation id="EFB408C1FFA4FF82FD8C9BAAFD7DF910" box="[546,740,1754,1779]" pageId="11" pageNumber="12" refId="ref46836" refString="Yuan, J. L., Li, Y., Mu, X. N., Lin, J. P. &amp; Zhu, X. J. (2012) Trilobite fauna of the Changhia Formation (Cambrian Series 3) from Shandong and adjacent area, North China. Palaeontologica Sinica, New Series B, 35, 758 pp. [in Chinese with English summary]" type="book">
Yuan
<emphasis id="B951A922FFA4FF82FDCA9BADFD01F910" box="[612,664,1754,1779]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
. 2012
</bibRefCitation>
, pl. 2, figs. 7, 12). This suggests it is difficult to distinguish species of
<taxonomicName id="4C250EB3FFA4FF82FEB79B8EFDD1F8F4" authority="Opik, 1967" authorityName="Opik" authorityYear="1967" box="[281,584,1790,1815]" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B951A922FFA4FF82FEB79B8EFE2FF8F5" box="[281,438,1791,1814]" italics="true" pageId="11" pageNumber="12">Ammagnostus</emphasis>
Öpik, 1967
</taxonomicName>
based on small specimens, prompting taxonomic caution with juvenile material.
</paragraph>
<paragraph id="8B9A7530FFA4FF82FF699A36FCCBF80C" blockId="11.[151,1437,386,2031]" pageId="11" pageNumber="12">
<taxonomicName id="4C250EB3FFA4FF82FF699A36FE35F8BC" authorityName="Opik" authorityYear="1967" box="[199,428,1862,1887]" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bassa">
<emphasis id="B951A922FFA4FF82FF699A36FE35F8BC" box="[199,428,1862,1887]" italics="true" pageId="11" pageNumber="12">Ammagnostus bassa</emphasis>
</taxonomicName>
(=
<emphasis id="B951A922FFA4FF82FE609A37FD22F8BC" box="[462,699,1862,1887]" italics="true" pageId="11" pageNumber="12">
Agnostoglossa
<taxonomicName id="4C250EB3FFA4FF82FDD59A37FD22F8BC" authorityName="Opik" authorityYear="1967" box="[635,699,1862,1887]" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bassa">bassa</taxonomicName>
</emphasis>
) Öpik, 1967 from the Mindyallan Mungerebar Limestone of the
<taxonomicName id="4C250EB3FFA4FF82FF399A1BFEDEF860" authority="Basin" authorityName="Basin" box="[151,327,1898,1923]" class="Insecta" family="Morabidae" genus="Georgina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Georgina Basin</taxonomicName>
resembles the Goyder Formation material in sharing a moderately long (sag.) anterior border, F3 that is slightly bowed rearward, an indistinct glabellar node, and a pygidium with a posterior border that is relatively long (sag.). However, the coarsely silicified material of
<taxonomicName id="4C250EB3FFA4FF82FCDD9AC2FC4BF828" authorityName="Opik" authorityYear="1967" box="[883,978,1970,1995]" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bassa">
<emphasis id="B951A922FFA4FF82FCDD9AC2FC1DF829" box="[883,900,1971,1994]" italics="true" pageId="11" pageNumber="12">A</emphasis>
.
<emphasis id="B951A922FFA4FF82FC3C9AC3FC4BF828" box="[914,978,1970,1995]" italics="true" pageId="11" pageNumber="12">bassa</emphasis>
</taxonomicName>
makes it otherwise difficult to determine if
<taxonomicName id="4C250EB3FFA4FF82FF019AA6FED2F80D" authorityName="Bentley, Jago &amp; Cooper" authorityYear="2009" box="[175,331,2007,2030]" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="antarcticus">
<emphasis id="B951A922FFA4FF82FF019AA6FF59F80D" box="[175,192,2007,2030]" italics="true" pageId="11" pageNumber="12">A</emphasis>
.
<emphasis id="B951A922FFA4FF82FF609AA6FED2F80D" box="[206,331,2007,2030]" italics="true" pageId="11" pageNumber="12">antarcticus</emphasis>
</taxonomicName>
is synonymous (Öpik 1967, pl. 60, fig. 614).
</paragraph>
</subSubSection>
<caption id="DF5A25B8FFA3FF85FF399BCBFD27F808" httpUri="https://zenodo.org/record/1202733/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,250,1722,1744]" targetBox="[191,1404,204,1700]" targetPageId="12">
<paragraph id="8B9A7530FFA3FF85FF399BCBFD27F808" blockId="12.[151,1437,1722,2027]" pageId="12" pageNumber="13">
<emphasis id="B951A922FFA3FF85FF399BCBFE8AF933" bold="true" box="[151,275,1722,1744]" pageId="12" pageNumber="13">FIGURE 5.</emphasis>
<taxonomicName id="4C250EB3FFA3FF85FEB59BCBFCF9F933" authority="Bentley, Jago &amp; Cooper, 2009" authorityName="Bentley, Jago &amp; Cooper" authorityYear="2009" box="[283,864,1722,1744]" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="antarcticus">
<emphasis id="B951A922FFA3FF85FEB59BCBFDB8F933" box="[283,545,1722,1744]" italics="true" pageId="12" pageNumber="13">Ammagnostus antarcticus</emphasis>
Bentley, Jago &amp; Cooper, 2009
</taxonomicName>
. All specimens come from sample GOY/173.6. A, S, T, U, CPC42177, partial cephalon; A, dorsal view; S, oblique anterolateral view; T, lateral view; U, anterior view. B, CPC42178, partial cephalon. C, CPC42179, partial cephalon. D, CPC42180, partial cephalon. E, CPC42181, partial cephalon. F, CPC42182, partial cephalon. G, CPC42183, partial cephalon. H, CPC42184, partial cephalon. I, CPC42185, partial cephalon. J, CPC42186, partial cephalon. K, CPC42187, partial cephalon. L, CPC42188, partial cephalon. M, CPC42189, partial cephalon. N, CPC42190, partial cephalon. O, CPC42191, partial cephalon. P, CPC42192, partial cephalon. Q, CPC42193, partial cephalon. R, CPC42194, partial cephalon. V, CPC42195, partial cephalon. W, Z, A1, CPC42196, partial pygidium; W, dorsal view; Z, lateral view; A1, posterior view. X, CPC42197, partial pygidium. Y, CPC42198, partial pygidium. B1, CPC42199, partial pygidium. C1, CPC42200, partial pygidium. D1, CPC42201, partial pygidium. E1, CPC42202, partial pygidium. F1, CPC42203, partial pygidium. All scale bars are 1 mm.
</paragraph>
</caption>
<subSubSection id="C33F26BBFFA2FF84FF699DE9FC8DFE6B" pageId="13" pageNumber="14" type="description">
<paragraph id="8B9A7530FFA2FF84FF699DE9FAFBFE86" blockId="13.[151,1436,151,536]" pageId="13" pageNumber="14">
<taxonomicName id="4C250EB3FFA2FF84FF699DE9FEFDFF4C" authorityName="Opik" authorityYear="1967" box="[199,356,152,175]" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B951A922FFA2FF84FF699DE9FEFDFF4C" box="[199,356,152,175]" italics="true" pageId="13" pageNumber="14">Ammagnostus</emphasis>
</taxonomicName>
sp. indet. from the Mindyallan Boshy Formation of N.S.W. (
<bibRefCitation id="EFB408C1FFA2FF84FBB59DE9FB7AFF53" box="[1051,1251,151,176]" pageId="13" pageNumber="14" refId="ref46334" refString="Wang, Q., Mills, K. J., Webby, B. D. &amp; Shergold, J. H. (1989) Upper Cambrian (Mindyallan) trilobites and stratigraphy of the Kayrunnera Group, western New South Wales. BMR Journal of Australian Geology &amp; Geophysics, 11, 107 - 118." type="journal article">
Wang
<emphasis id="B951A922FFA2FF84FBC89DE8FB00FF53" box="[1126,1177,151,176]" italics="true" pageId="13" pageNumber="14">et al</emphasis>
. 1989
</bibRefCitation>
, fig. 4A-D) also resembles
<taxonomicName id="4C250EB3FFA2FF84FEBF9DCDFE29FF37" authorityName="Bentley, Jago &amp; Cooper" authorityYear="2009" box="[273,432,188,212]" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="antarcticus">
<emphasis id="B951A922FFA2FF84FEBF9DCDFEBBFF30" box="[273,290,188,211]" italics="true" pageId="13" pageNumber="14">A</emphasis>
.
<emphasis id="B951A922FFA2FF84FE9D9DCCFE29FF37" box="[307,432,189,212]" italics="true" pageId="13" pageNumber="14">antarcticus</emphasis>
</taxonomicName>
in having a posterior glabellar lobe that is wider (tr.) than the anterior lobe, F3 that is slightly bowed rearward, an indistinct glabellar node, and a long (sag.) posterior border on the pygidium. The limited amount of poorly preserved material from the Boshy Formation only permits a tentative assignment to
<taxonomicName id="4C250EB3FFA2FF84FA2A9C75FE8DFEA3" authorityName="Bentley, Jago &amp; Cooper" authorityYear="2009" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="antarcticus">
<emphasis id="B951A922FFA2FF84FA2A9C75FE8DFEA3" italics="true" pageId="13" pageNumber="14">A. antarcticus</emphasis>
</taxonomicName>
. All other known species of
<taxonomicName id="4C250EB3FFA2FF84FDF79C59FD6FFEDC" authorityName="Opik" authorityYear="1967" box="[601,758,296,319]" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B951A922FFA2FF84FDF79C59FD6FFEDC" box="[601,758,296,319]" italics="true" pageId="13" pageNumber="14">Ammagnostus</emphasis>
</taxonomicName>
differ from
<taxonomicName id="4C250EB3FFA2FF84FC2F9C59FB84FEA3" authorityName="Bentley, Jago &amp; Cooper" authorityYear="2009" box="[897,1053,296,320]" class="Trilobita" family="Agnostidae" genus="Ammagnostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="antarcticus">
<emphasis id="B951A922FFA2FF84FC2F9C59FC0BFEDC" box="[897,914,296,319]" italics="true" pageId="13" pageNumber="14">A</emphasis>
.
<emphasis id="B951A922FFA2FF84FC0E9C58FB84FEA3" box="[928,1053,297,320]" italics="true" pageId="13" pageNumber="14">antarcticus</emphasis>
</taxonomicName>
in having a shorter (sag.) anterior border, more transverse (or only slightly posteriorly bowed) F3, and a shorter (sag.) posterior pygidial border.
</paragraph>
<paragraph id="8B9A7530FFA2FF84FF699C1EFC8DFE6B" blockId="13.[151,1436,151,536]" box="[199,788,367,392]" pageId="13" pageNumber="14">
<emphasis id="B951A922FFA2FF84FF699C1EFEC0FE6B" bold="true" box="[199,345,367,392]" pageId="13" pageNumber="14">Occurrence.</emphasis>
GOY section horizon
<quantity id="4CDDD8D5FFA2FF84FDF99C01FD2AFE64" box="[599,691,368,392]" metricMagnitude="2" metricUnit="m" metricValue="1.327" pageId="13" pageNumber="14" unit="m" value="132.7">132.7 m</quantity>
(
<figureCitation id="131E69B5FFA2FF84FD6C9C1EFC9FFE6B" box="[706,774,367,392]" captionStart="FIGURE 3" captionStartId="9.[151,250,1832,1854]" captionTargetBox="[189,1384,215,1785]" captionTargetId="figure@9.[174,1416,195,1820]" captionTargetPageId="9" captionText="FIGURE 3. Generalised lithostratigraphy and taxon ranges through the Goyder Formation at the GOY section. Sampled horizons are indicated on the right side of the section, with sample numbers representing tape meterage along the section line (see Fig.2). Measurements on the left side of the section are in meters (true thickness) from the base of the Goyder Formation. SF = Shannon Formation, PSS = Pacoota Sandstone. Solid black circles represent the occurrences of taxa present in the GOY section. Hollow circles represent the approximate stratigraphic occurrences of taxa found at nearby spot localities. Ranges of taxa found only atspot localities approximate, and has been inferred using lithological, faunal and geographical information." httpUri="https://zenodo.org/record/1202729/files/figure.png" pageId="13" pageNumber="14">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F26BBFFA2FF84FF699CE5FEE4FDFB" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8B9A7530FFA2FF84FF699CE5FEE4FDFB" blockId="13.[151,1436,151,536]" pageId="13" pageNumber="14">
<emphasis id="B951A922FFA2FF84FF699CE5FEF9FE4E" bold="true" box="[199,352,404,429]" pageId="13" pageNumber="14">Distribution.</emphasis>
Goyder Formation, Amadeus Basin,
<collectingRegion id="49E1BBD2FFA2FF84FCB79CE5FC69FE4E" box="[793,1008,404,429]" country="Australia" name="Northern Territory" pageId="13" pageNumber="14">Northern Territory</collectingRegion>
; Spurs Formation, Northern
<collectingRegion id="49E1BBD2FFA2FF84FAEC9CE5FA05FE4E" box="[1346,1436,404,429]" country="Australia" name="Victoria" pageId="13" pageNumber="14">Victoria</collectingRegion>
Land,
<collectingCountry id="F33235A0FFA2FF84FF4F9CC6FEC0FE33" box="[225,345,439,464]" name="Antarctica" pageId="13" pageNumber="14">Antarctica</collectingCountry>
. Possibly the Mungerebar Limestone,
<taxonomicName id="4C250EB3FFA2FF84FCBC9CC6FC5CFE33" authority="Basin" authorityName="Basin" box="[786,965,439,464]" class="Insecta" family="Morabidae" genus="Georgina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">Georgina Basin</taxonomicName>
,
<collectingRegion id="49E1BBD2FFA2FF84FC7D9CC6FB3DFE33" box="[979,1188,439,464]" country="Australia" name="Northern Territory" pageId="13" pageNumber="14">Northern Territory</collectingRegion>
and
<collectingRegion id="49E1BBD2FFA2FF84FB719CC6FAFFFE33" box="[1247,1382,439,464]" country="Australia" name="Queensland" pageId="13" pageNumber="14">Queensland</collectingRegion>
, and the Boshy Formation, Koonenberry Belt,
<collectingRegion id="49E1BBD2FFA2FF84FDDF9CACFCD8FE16" box="[625,833,476,501]" country="Australia" name="New South Wales" pageId="13" pageNumber="14">
New South
<collectingCountry id="F33235A0FFA2FF84FD579CADFCD8FE16" box="[761,833,476,501]" name="United Kingdom" pageId="13" pageNumber="14">Wales</collectingCountry>
</collectingRegion>
. All occurrences are Cambrian Series 3, Guzhangian (Mindyallan) in age.
</paragraph>
</subSubSection>
</treatment>
</document>