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<document id="62D9B3CC732315180FD46C1DC7B3C37E" ID-CLB-Dataset="30484" ID-DOI="10.11646/zootaxa.4396.1.1" ID-GBIF-Dataset="918f2145-a708-460e-b19b-4f67e6c99b30" ID-ISSN="1175-5326" ID-Zenodo-Dep="1202723" ID-ZooBank="8EEBE6DE-0ECC-4B9C-AD14-01438291782B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1521441296340" checkinUser="plazi" docAuthor="Smith, Patrick M., Paterson, John R. &amp; Brock, Glenn A." docDate="2018" docId="038CC426FFA0FF99FF3999D8FC9CFAB6" docLanguage="en" docName="zootaxa.4396.1.1.pdf" docOrigin="Zootaxa 4396 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Trephina Smith &amp; Paterson &amp; Brock 2018, gen. nov." docType="treatment" docUuid="E424E3C1-FC17-47E9-AD00-2DE030DFC7AE" docUuidSource="ZooBank" docVersion="8" lastPageNumber="17" masterDocId="FFB5BC5EFFAFFF89FFAE9D71FF99FFE3" masterDocTitle="Trilobites and agnostids from the Goyder Formation (Cambrian Series 3, Guzhangian; Mindyallan), Amadeus Basin, central Australia" masterLastPageNumber="67" masterPageNumber="1" pageNumber="16" updateTime="1698532001681" updateUser="ExternalLinkService">
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<mods:title id="8D8483185503ECDF1841EBE70785A082">Trilobites and agnostids from the Goyder Formation (Cambrian Series 3, Guzhangian; Mindyallan), Amadeus Basin, central Australia</mods:title>
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<paragraph id="8B9A7530FFA0FF86FF3999D8FEE2FB20" blockId="15.[151,379,1193,1219]" box="[151,379,1193,1219]" pageId="15" pageNumber="16">
<heading id="D0D2C25CFFA0FF86FF3999D8FEE2FB20" bold="true" box="[151,379,1193,1219]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="B951A922FFA0FF86FF3999D8FEE2FB20" bold="true" box="[151,379,1193,1219]" pageId="15" pageNumber="16">
<taxonomicName id="4C250EB3FFA0FF86FF3999D8FE9FFB20" authority="Smith &amp; Paterson &amp; Brock, 2018" authorityName="Smith &amp; Paterson &amp; Brock" authorityYear="2018" box="[151,262,1193,1219]" class="Trilobita" family="Proasaphiscidae" genus="Trephina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B951A922FFA0FF86FF3999D8FE9FFB20" bold="true" box="[151,262,1193,1219]" italics="true" pageId="15" pageNumber="16">Trephina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2621459FFA0FF86FEA399D8FEE2FB20" box="[269,379,1193,1219]" pageId="15" pageNumber="16" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C33F26BBFFA0FF86FF399981FD18FAE5" box="[151,641,1264,1286]" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8B9A7530FFA0FF86FF399981FD18FAE5" blockId="15.[151,641,1264,1286]" box="[151,641,1264,1286]" pageId="15" pageNumber="16">
<emphasis id="B951A922FFA0FF86FF399981FEBBFAE5" bold="true" box="[151,290,1264,1286]" pageId="15" pageNumber="16">Type species.</emphasis>
<taxonomicName id="4C250EB3FFA0FF86FE879981FE47FAE5" authority="Smith &amp; Paterson &amp; Brock, 2018" authorityName="Smith &amp; Paterson &amp; Brock" authorityYear="2018" box="[297,478,1264,1286]" class="Trilobita" family="Proasaphiscidae" genus="Trephina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ranfordi" status="gen. et sp. nov.">
<emphasis id="B951A922FFA0FF86FE879981FE47FAE5" box="[297,478,1264,1286]" italics="true" pageId="15" pageNumber="16">Trephina ranfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2621459FFA0FF86FE4A9981FD18FAE5" box="[484,641,1264,1286]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B951A922FFA0FF86FE4A9981FD18FAE5" bold="true" box="[484,641,1264,1286]" pageId="15" pageNumber="16">gen. et sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
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<subSubSection id="C33F26BBFFA0FF86FF399843FCB0FAA8" box="[151,809,1330,1355]" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8B9A7530FFA0FF86FF399843FCB0FAA8" blockId="15.[151,1437,1330,2003]" box="[151,809,1330,1355]" pageId="15" pageNumber="16">
<emphasis id="B951A922FFA0FF86FF399843FE87FAA8" bold="true" box="[151,286,1330,1355]" pageId="15" pageNumber="16">Etymology.</emphasis>
After
<taxonomicName id="4C250EB3FFA0FF86FEC89843FE53FAA8" authority="Smith &amp; Paterson &amp; Brock, 2018" authorityName="Smith &amp; Paterson &amp; Brock" authorityYear="2018" box="[358,458,1330,1355]" class="Trilobita" family="Proasaphiscidae" genus="Trephina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="15" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">Trephina</taxonomicName>
Creek, near Ross River Gorge.
</paragraph>
</subSubSection>
<subSubSection id="C33F26BBFFA0FF99FF699827FACEFF53" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8B9A7530FFA0FF99FF699827FACEFF53" blockId="15.[151,1437,1330,2003]" lastBlockId="16.[151,1437,151,1365]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="B951A922FFA0FF86FF699827FED8FA8C" bold="true" box="[199,321,1366,1391]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
Cranidium subtrapezoidal in outline, length:width ratio of 60%. Glabella anteriorly truncate, subquadrate in outline, width:length ratio of 60% to 68%. Anterior of glabella merges with anterior border, preglabellar furrow effaced; axial furrow narrow (tr.) and deep. S1 moderately well defined and bifurcated, with the anterior branch directed anteromedially and the posterior branch directed posteromedially; S2 faintly defined and bifurcated, with the anterior branch directed slightly anteromedially and the posterior branch directed slightly posteromedially; S3 relatively faint, directed anteromedially; S4 variably developed, directed anteromedially. SO bowed backwards. Anterior cranidial border narrow, 5% to 10% of sagittal cranidial length. Preocular field 28% of sagittal cranidial length between the anterior border and γ. Palpebral lobes 33% cranidial length, reniform in outline. Palpebral area of fixigena 60% of adjacent glabellar width. Posterolateral projections at posterior of palpebral lobe 13% total cranidial length. Genal spine blade-like, length 41% librigenal length. Pygidium up to
<quantity id="4CDDD8D5FFA0FF86FA209BEAFF5AF935" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="16" unit="mm" value="8.0">8 mm</quantity>
long (sag.), subtriangular in outline slightly convex, length:width ratio of 64% to 69% excluding spines. Axis prominent, moderately narrow (tr.), width:length ratio of 75% to 81%, occupying about 65% of sagittal length of pygidium. Three well-defined axial rings and a faint fourth present. Small terminal piece. A single pair of broadbased spines attached to pygidium anterolateral corners, spines are broad (tr.) and blade-like, curved posteriorly. Prosopon comprising closely spaced granules over most of the cephalon, except the posterolateral corners of the glabella, palpebral lobes and librigenal borders that are covered by small, closely spaced pits. Anterior cranidial border and the librigenal border also covered by well-defined horizontal terrace ridges. Pygidium covered by small, closely spaced pits over pygidial axis; pleural field with moderately sized pustules on pleural ribs. Spines and posterior margin covered with fine horizontal terrace ridges and evenly spaced pustules near the spine bases.
</paragraph>
</subSubSection>
<subSubSection id="C33F26BBFFBFFF99FF699DCDFC9CFAB6" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8B9A7530FFBFFF99FF699DCDFDD5FD83" blockId="16.[151,1437,151,1365]" pageId="16" pageNumber="17">
<emphasis id="B951A922FFBFFF99FF699DCDFED5FF36" bold="true" box="[199,332,188,213]" pageId="16" pageNumber="17">Discussion.</emphasis>
This new genus has similarities to various leiostegioids, such as
<taxonomicName id="4C250EB3FFBFFF99FBB09DCDFABAFF36" authority="Walcott, 1905" authorityName="Walcott" authorityYear="1905" box="[1054,1315,188,213]" class="Trilobita" family="Pagodiidae" genus="Pagodia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B951A922FFBFFF99FBB09DCDFBE4FF36" box="[1054,1149,188,213]" italics="true" pageId="16" pageNumber="17">Pagodia</emphasis>
<bibRefCitation id="EFB408C1FFBFFF99FB2A9DCDFABAFF36" box="[1156,1315,188,213]" pageId="16" pageNumber="17" refId="ref46138" refString="Walcott, C. D. (1905) Cambrian faunas of China. Proceedings of the U. S. National Museum, 29, 1 - 106. https: // doi. org / 10.5479 / si. 00963801.1415" type="journal article">Walcott, 1905</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C250EB3FFBFFF99FA839DCDFEABFF1B" authority="Walcott, 1911" authorityName="Walcott" authorityYear="1911" class="Trilobita" family="Leiostegiidae" genus="Chuangia" higherTaxonomySource="CoL" kingdom="Animalia" order="Ptychopariida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B951A922FFBFFF99FA839DCDFA02FF36" box="[1325,1435,188,213]" italics="true" pageId="16" pageNumber="17">Chuangia</emphasis>
<bibRefCitation id="EFB408C1FFBFFF99FF399DAEFEABFF1B" box="[151,306,223,248]" pageId="16" pageNumber="17" refId="ref46220" refString="Walcott, C. D. (1911) Cambrian faunas of China. Smithsonian Miscellaneous Collections, 57, 69 - 109." type="journal article">Walcott, 1911</bibRefCitation>
</taxonomicName>
and
<emphasis id="B951A922FFBFFF99FEC69DAEFE60FF1B" box="[360,505,223,248]" italics="true" pageId="16" pageNumber="17">Lophoholcus</emphasis>
Öpik, 1967. Cranidia of
<taxonomicName id="4C250EB3FFBFFF99FCBE9DAEFCEDFF1B" authority="Smith &amp; Paterson &amp; Brock, 2018" authorityName="Smith &amp; Paterson &amp; Brock" authorityYear="2018" box="[784,884,223,248]" class="Trilobita" family="Proasaphiscidae" genus="Trephina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B951A922FFBFFF99FCBE9DAEFCEDFF1B" box="[784,884,223,248]" italics="true" pageId="16" pageNumber="17">Trephina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2621459FFBFFF99FCD59DAEFC7DFF1B" box="[891,996,223,248]" pageId="16" pageNumber="17" rank="genus">
<emphasis id="B951A922FFBFFF99FCD59DAEFC7DFF1B" bold="true" box="[891,996,223,248]" pageId="16" pageNumber="17">gen. nov.</emphasis>
</taxonomicNameLabel>
and some species of
<taxonomicName id="4C250EB3FFBFFF99FB7F9DAEFAA9FF1B" authorityName="Walcott" authorityYear="1905" box="[1233,1328,223,248]" class="Trilobita" family="Pagodiidae" genus="Pagodia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B951A922FFBFFF99FB7F9DAEFAA9FF1B" box="[1233,1328,223,248]" italics="true" pageId="16" pageNumber="17">Pagodia</emphasis>
</taxonomicName>
(cf. Öpik 1967, pl. 17, figs 1, 2, 4;
<bibRefCitation id="EFB408C1FFBFFF99FE1F9C75FDCEFEFE" box="[433,599,260,285]" pageId="16" pageNumber="17" refId="ref45167" refString="Shergold, J. H. (1975) Late Cambrian and Early Ordovician trilobites from the Burke River Structural Belt, western Queensland. Bulletin of the Bureau of Mineral Resources, Geology &amp; Geophysics, 153, 1 - 251." type="journal article">Shergold 1975</bibRefCitation>
, pl. 29, figs 5, 6;
<bibRefCitation id="EFB408C1FFBFFF99FCB49C75FC58FEFF" box="[794,961,260,285]" pageId="16" pageNumber="17" refId="ref45210" refString="Shergold, J. H. (1982) Idamean (Late Cambrian) trilobites, Burke River Structural Belt, western Queensland. Bulletin of the Bureau of Mineral Resources, Geology &amp; Geophysics, 187, 1 - 69." type="journal article">Shergold 1982</bibRefCitation>
, pl. 13, figs 1, 2, 4; Zhang &amp; Jell 1987, pl. 94, figs 8, 10, 11, pl. 95, figs 2, 4, 5) possess an anteriorly truncate, subquadrate glabella that slightly constricts at or anterior to the mid-length, a short (sag., exsag.) anterior border, and prominent reniform palpebral lobes. Both genera also possess an anterior border furrow that is often obscured by the encroaching anterior glabellar lobe. However, the Goyder cranidia mainly differ in possessing medial effacement of the anterior border furrow through fusion of the anterior glabellar lobe with the anterior border, and in having distinctly incised glabellar furrows (particularly the bifid S1).
<taxonomicName id="4C250EB3FFBFFF99FE679CADFDAEFE16" authorityName="Walcott" authorityYear="1911" box="[457,567,476,501]" class="Trilobita" family="Leiostegiidae" genus="Chuangia" higherTaxonomySource="CoL" kingdom="Animalia" order="Ptychopariida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B951A922FFBFFF99FE679CADFDAEFE16" box="[457,567,476,501]" italics="true" pageId="16" pageNumber="17">Chuangia</emphasis>
</taxonomicName>
possesses some of the cranidial features mentioned above, although species typically have an effaced and more anteriorly tapered glabella (cf. Zhang &amp; Jell 1987, pl. 89, figs 8, 9, pl. 90, figs 1, 3, 5, 12, pl. 91, figs 2, 6, 7, 8, pl. 92, figs 1, 3, 5, 7, 11, 12, pl. 93, figs 18; Peng
<emphasis id="B951A922FFBFFF99FBB39F54FBCCFDDE" box="[1053,1109,548,573]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2004a, pl. 20, figs 18; Choi
<emphasis id="B951A922FFBFFF99FF399F38FF56FD83" box="[151,207,583,608]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2008, figs 15.1, 15.2, 15.415.7).
</paragraph>
<paragraph id="8B9A7530FFBFFF99FF699F1DFDDFFCDB" blockId="16.[151,1437,151,1365]" pageId="16" pageNumber="17">
<taxonomicName id="4C250EB3FFBFFF99FF699F1DFEB2FD66" authority="Smith &amp; Paterson &amp; Brock, 2018" authorityName="Smith &amp; Paterson &amp; Brock" authorityYear="2018" box="[199,299,620,645]" class="Trilobita" family="Proasaphiscidae" genus="Trephina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B951A922FFBFFF99FF699F1DFEB2FD66" box="[199,299,620,645]" italics="true" pageId="16" pageNumber="17">Trephina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2621459FFBFFF99FE9A9F1DFE07FD66" box="[308,414,620,645]" pageId="16" pageNumber="17" rank="genus">
<emphasis id="B951A922FFBFFF99FE9A9F1DFE07FD66" bold="true" box="[308,414,620,645]" pageId="16" pageNumber="17">gen. nov.</emphasis>
</taxonomicNameLabel>
shares features with the monotypic genus
<emphasis id="B951A922FFBFFF99FC299F1DFB81FD66" box="[903,1048,620,645]" italics="true" pageId="16" pageNumber="17">Lophoholcus</emphasis>
(type species:
<emphasis id="B951A922FFBFFF99FB6A9F1DFA05FD67" box="[1220,1436,620,645]" italics="true" pageId="16" pageNumber="17">Lophoholcus asper</emphasis>
Öpik, 1967) from the Mindyallan OHara Shale in the
<taxonomicName id="4C250EB3FFBFFF99FCA49FFEFB98FD4B" authority="Basin. Both" authorityName="Basin. Both" box="[778,1025,655,680]" class="Insecta" family="Morabidae" genus="Georgina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Georgina Basin. Both</taxonomicName>
have similar cranidial and glabellar outlines, development of glabellar furrows, and shape of the anterior border.
<emphasis id="B951A922FFBFFF99FC5B9FC5FB1FFD2E" box="[1013,1158,692,717]" italics="true" pageId="16" pageNumber="17">Lophoholcus</emphasis>
differs from
<taxonomicName id="4C250EB3FFBFFF99FAB59FC5FAE6FD2E" authority="Smith &amp; Paterson &amp; Brock, 2018" authorityName="Smith &amp; Paterson &amp; Brock" authorityYear="2018" box="[1307,1407,692,717]" class="Trilobita" family="Proasaphiscidae" genus="Trephina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B951A922FFBFFF99FAB59FC5FAE6FD2E" box="[1307,1407,692,717]" italics="true" pageId="16" pageNumber="17">Trephina</emphasis>
</taxonomicName>
in having an anterior border separated from the glabella by a deep border furrow, lower glabellar convexity, presence of an occipital node, narrower (tr.) palpebral areas, longer (exsag.) and narrower (tr.) palpebral lobes, and a pygidium with a much wider (tr.) axis.
</paragraph>
<paragraph id="8B9A7530FFBFFF99FF699E35FC9CFAB6" blockId="16.[151,1437,151,1365]" pageId="16" pageNumber="17">
Pygidia of
<taxonomicName id="4C250EB3FFBFFF99FEE89E35FD96FCBE" authorityName="Smith &amp; Paterson &amp; Brock" authorityYear="2018" box="[326,527,836,861]" class="Trilobita" family="Proasaphiscidae" genus="Trephina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ranfordi">
<emphasis id="B951A922FFBFFF99FEE89E35FD96FCBE" box="[326,527,836,861]" italics="true" pageId="16" pageNumber="17">Trephina ranfordi</emphasis>
</taxonomicName>
were originally reported by Öpik (1967, appendix 2, p. 16) as
<emphasis id="B951A922FFBFFF99FB5E9E35FAC5FCBE" box="[1264,1372,836,861]" italics="true" pageId="16" pageNumber="17">Plectrifer</emphasis>
?
<emphasis id="B951A922FFBFFF99FAD49E35FF51FC63" bold="true" pageId="16" pageNumber="17">sp. nov.</emphasis>
(pygidia), but they show little similarity to this genus.
<emphasis id="B951A922FFBFFF99FC949E16FC3CFC63" box="[826,933,871,896]" italics="true" pageId="16" pageNumber="17">Plectrifer</emphasis>
Öpik, 1967 differs in having a long axis that extends more than 90% the length of the pygidium and a distinct pygidial border furrow. The material from the Goyder Formation more closely resembles other leiostegioids, in possessing a single pair of posteriorly directed spines along the lateral border (e.g.
<taxonomicName id="4C250EB3FFBFFF99FD889EA5FCE2FC0E" authority="Kobayashi, 1935" authorityName="Kobayashi" authorityYear="1935" box="[550,891,980,1005]" class="Trilobita" family="Leiostegiidae" genus="Prochuangia" higherTaxonomySource="CoL" kingdom="Animalia" order="Ptychopariida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B951A922FFBFFF99FD889EA5FD2EFC0E" box="[550,695,980,1005]" italics="true" pageId="16" pageNumber="17">Prochuangia</emphasis>
Kobayashi, 1935
</taxonomicName>
and
<emphasis id="B951A922FFBFFF99FC1F9EA5FBB3FC0E" box="[945,1066,980,1005]" italics="true" pageId="16" pageNumber="17">Lotosoides</emphasis>
<bibRefCitation id="EFB408C1FFBFFF99FB9F9EA5FB46FC0E" box="[1073,1247,980,1005]" pageId="16" pageNumber="17" refId="ref45167" refString="Shergold, J. H. (1975) Late Cambrian and Early Ordovician trilobites from the Burke River Structural Belt, western Queensland. Bulletin of the Bureau of Mineral Resources, Geology &amp; Geophysics, 153, 1 - 251." type="journal article">Shergold, 1975</bibRefCitation>
). However, most of these taxa can be distinguished because their spines emerge from behind the anterolateral corners of the pygidium.
<emphasis id="B951A922FFBFFF99FEBD996DFD80FBD6" box="[275,537,1052,1077]" italics="true" pageId="16" pageNumber="17">Perischodory grandgei</emphasis>
<bibRefCitation id="EFB408C1FFBFFF99FD8C996DFD41FBD7" box="[546,728,1052,1077]" pageId="16" pageNumber="17" refId="ref44955" refString="Raymond, P. E. (1937) Upper Cambrian and Lower Ordovician Trilobita and Ostracoda from Vermont. Geological Society of America Bulletin, 48, 1079 - 1146. https: // doi. org / 10.1130 / GSAB- 48 - 1079" type="journal article">Raymond, 1937</bibRefCitation>
from the Lower Ordovician Highgate Formation of the USA (see
<bibRefCitation id="EFB408C1FFBFFF99FF61994EFE1BFBBB" box="[207,386,1087,1112]" pageId="16" pageNumber="17" refId="ref44955" refString="Raymond, P. E. (1937) Upper Cambrian and Lower Ordovician Trilobita and Ostracoda from Vermont. Geological Society of America Bulletin, 48, 1079 - 1146. https: // doi. org / 10.1130 / GSAB- 48 - 1079" type="journal article">Raymond 1937</bibRefCitation>
, pl. 4 fig. 28;
<bibRefCitation id="EFB408C1FFBFFF99FD9C994EFD2FFBBB" box="[562,694,1087,1112]" pageId="16" pageNumber="17" refId="ref45134" refString="Shaw, A. B. (1966) Paleontology of Northwestern Vermont. XII. Fossils from the Ordovician Highgate Formation. Journal of Paleontology, 40, 1312 - 1330." type="journal article">Shaw 1966</bibRefCitation>
, p1. 161, figs 3, 4) is similar to the new taxon in that it has a comparable subtriangular pygidium with a single pair of spines placed at the anterolateral extremities. However, the Goyder Formation material differs in having a shorter (sag.) axis, deeper pleural furrows, fainter posterior border, more posteriorly directed spines, and a pleural field with minute granules on the pleural ribs. Placement of spines in the anterolateral corners of
<taxonomicName id="4C250EB3FFBFFF99FDEA99BEFD31FB0B" authority="Smith &amp; Paterson &amp; Brock, 2018" authorityName="Smith &amp; Paterson &amp; Brock" authorityYear="2018" box="[580,680,1231,1256]" class="Trilobita" family="Proasaphiscidae" genus="Trephina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B951A922FFBFFF99FDEA99BEFD31FB0B" box="[580,680,1231,1256]" italics="true" pageId="16" pageNumber="17">Trephina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2621459FFBFFF99FD1F99BEFC84FB0B" box="[689,797,1231,1256]" pageId="16" pageNumber="17" rank="genus">
<emphasis id="B951A922FFBFFF99FD1F99BEFC84FB0B" bold="true" box="[689,797,1231,1256]" pageId="16" pageNumber="17">gen. nov.</emphasis>
</taxonomicNameLabel>
is also comparable to material described as Pagodiinae gen. et sp. indet. from the Scopus Formation at Christmas Hills in Tasmania (Jago &amp; Bentley 2007, p. 292, fig. 4D J), which is Boomerangian in age (Jago
<emphasis id="B951A922FFBFFF99FDF09868FD09FAD3" box="[606,656,1303,1328]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
. 2016). The Goyder Formation taxon differs in having a proportially shorter (sag.) axis and more posteriorly directed spines.
</paragraph>
</subSubSection>
</treatment>
</document>