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<figureCitation id="131E3FFFFFB67747FC8493FFBA51C98D" box="[854,1024,1757,1781]" captionStart-0="Figure 96" captionStart-1="Figures 97–101" captionStart-2="Figures 102–103" captionStart-3="Figures 104–107" captionStart-4="Figures 108–111" captionStart-5="Figures 112–117" captionStart-6="Figures 118–124" captionStart-7="Figures 125–126" captionStartId-0="33.[116,184,1219,1241]" captionStartId-1="34.[116,193,1979,2001]" captionStartId-2="35.[113,190,1222,1244]" captionStartId-3="36.[113,190,1983,2005]" captionStartId-4="37.[113,190,1627,1649]" captionStartId-5="38.[113,190,1297,1319]" captionStartId-6="39.[116,193,1942,1964]" captionStartId-7="40.[113,190,1985,2007]" captionTargetBox-0="[113,1474,232,1206]" captionTargetBox-1="[130,1447,323,1890]" captionTargetBox-2="[113,1474,234,1210]" captionTargetBox-3="[183,1374,269,1965]" captionTargetBox-4="[113,1474,234,1616]" captionTargetBox-5="[113,1474,234,1286]" captionTargetBox-6="[143,1429,224,1936]" captionTargetBox-7="[113,1474,234,1965]" captionTargetId-0="figure-422@33.[113,1474,229,1206]" captionTargetId-1="figure-16@34.[113,1474,295,1890]" captionTargetId-2="figure-424@35.[113,1474,234,1210]" captionTargetId-3="figure-15@36.[113,1474,249,1965]" captionTargetId-4="figure-185@37.[113,1474,234,1616]" captionTargetId-5="figure-400@38.[113,1474,234,1286]" captionTargetId-6="figure-8@39.[113,1474,221,1938]" captionTargetId-7="figure-16@40.[110,1474,234,1965]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionTargetPageId-2="35" captionTargetPageId-3="36" captionTargetPageId-4="37" captionTargetPageId-5="38" captionTargetPageId-6="39" captionTargetPageId-7="40" captionText-0="Figure 96. Scorpiops lioneli sp. n., male, paratype BHNS SC 263 in vivo habitus." captionText-1="Figures 97–101. Scorpiops lioneli sp. n., male, holotype, dorsal (97) and ventral (98) views, sternopectinal area (99), metasomal segment V and telson in lateral view (100) and chelicera in dorsal view (101)." captionText-2="Figures 102–103. Scorpiops lioneli sp. n., male, holotype in dorsal (102) and ventral (103) views under UV light." captionText-3="Figures 104–107. Scorpiops lioneli sp. n., female, paratype, BNHS SC 264, in dorsal (104) ventral (105) views, sternopectinal area (106) and metasomal segment V and telson in lateral view (107)." captionText-4="Figures 108–111: Scorpiops lioneli sp. n. Figures 108–109. Male, holotype, carapace under white light (108) and UV fluorescence (109). Figures 110–111. Female, paratype, BHNS SC 264, carapace under white light (110) and UV fluorescence (111)." captionText-5="Figures 112–117. Scorpiops lioneli sp. n., male, holotype. Figures 112–117. Pedipalp chela dorsal (112), ventral (113) and dorsoexternal (114), patella external (115), dorsal (116) and ventral (117) views under UV fluorescence. Trichobothrial pattern indicated by yellow circles." captionText-6="Figures 118–124. Scorpiops lioneli sp. n. Figures 118–121. Scorpiops lioneli sp. n., male, holotype, left legs I-IV, retrolateral aspect. Figures 122–124. Male, paratype, BNHS SC 263, hemispermatophore internal (122), external (123) views and capsular region in internal view (124). Abbrevations: distal lamina (dl), capsule (c), trunk (t), pedicel (p), distal posterior lobe (dpl), lateral hook (lh), terminal membrane of the sperm duct (tmsd) and basal carina (bc)." captionText-7="Figures 125–126. Type locality of Scorpiops lioneli sp. n., view of mountains with semi-arid habitat and rocky out-crops at the type locality (125, 126)." figureDoi-0="http://doi.org/10.5281/zenodo.5742599" figureDoi-1="http://doi.org/10.5281/zenodo.5742601" figureDoi-2="http://doi.org/10.5281/zenodo.5742603" figureDoi-3="http://doi.org/10.5281/zenodo.5742605" figureDoi-4="http://doi.org/10.5281/zenodo.5742607" figureDoi-5="http://doi.org/10.5281/zenodo.5742609" figureDoi-6="http://doi.org/10.5281/zenodo.5742613" figureDoi-7="http://doi.org/10.5281/zenodo.5742615" httpUri-0="https://zenodo.org/record/5742599/files/figure.png" httpUri-1="https://zenodo.org/record/5742601/files/figure.png" httpUri-2="https://zenodo.org/record/5742603/files/figure.png" httpUri-3="https://zenodo.org/record/5742605/files/figure.png" httpUri-4="https://zenodo.org/record/5742607/files/figure.png" httpUri-5="https://zenodo.org/record/5742609/files/figure.png" httpUri-6="https://zenodo.org/record/5742613/files/figure.png" httpUri-7="https://zenodo.org/record/5742615/files/figure.png" pageId="33" pageNumber="32">Figures 96–126</figureCitation>
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<figureCitation id="131E3FFFFFB67747FBD993FFBAF1C98D" box="[1035,1184,1757,1781]" captionStart="Figures 127–138" captionStartId="41.[116,193,1928,1950]" captionTargetBox="[113,1474,245,1897]" captionTargetId="figure-102@41.[113,1474,234,1897]" captionTargetPageId="41" captionText="Figures 127–138: Figures 127–130: Comparison of left pedipalps. Figures 131–134: Comparison of metasomal segments II-IV in lateral view. Figures 135–138: Comparison of pedipalp patella dorsal view with internal aspect tubercles. Figures 127, 131, 135. Scorpiops vrushchik sp. n., male, holotype. Figures 128, 132, 136. Scorpiops nagphani sp. n., male, holotype. Figures 129, 133, 137. Scorpiops neera sp. n., male, holotype. Figures 130, 134, 138. Scorpiops lioneli sp. n., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.5742617" httpUri="https://zenodo.org/record/5742617/files/figure.png" pageId="33" pageNumber="32">130, 134, 138</figureCitation>
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<figureCitation id="131E3FFFFFB67747FB7993FFBB11C98D" box="[1195,1344,1757,1781]" captionStart-0="Figures 139–152" captionStart-1="Figure 153" captionStart-2="Figure 154" captionStart-3="Figure 155" captionStart-4="Figure 156" captionStart-5="Figure 157" captionStartId-0="43.[115,192,1183,1205]" captionStartId-1="45.[113,181,1216,1238]" captionStartId-2="45.[117,185,1992,2014]" captionStartId-3="46.[113,181,942,964]" captionStartId-4="47.[119,187,1584,1606]" captionStartId-5="48.[115,183,1359,1381]" captionTargetBox-0="[113,1474,234,1167]" captionTargetBox-1="[114,1474,234,1198]" captionTargetBox-2="[135,1438,1286,1971]" captionTargetBox-3="[117,1465,232,922]" captionTargetBox-4="[145,1432,286,1493]" captionTargetBox-5="[113,1474,257,1330]" captionTargetId-0="figure-329@43.[113,1474,234,1167]" captionTargetId-1="figure-7@45.[113,1474,234,1198]" captionTargetId-2="figure-9@45.[113,1474,1260,1983]" captionTargetId-3="figure-631@46.[113,1474,229,925]" captionTargetId-4="figure-220@47.[115,1476,232,1570]" captionTargetId-5="figure-208@48.[113,1474,257,1330]" captionTargetPageId-0="43" captionTargetPageId-1="45" captionTargetPageId-2="45" captionTargetPageId-3="46" captionTargetPageId-4="47" captionTargetPageId-5="48" captionText-0="Figures 139–152: Figures 139–142: Comparison of dorsolateral granules on anal rim of metasomal segment V in dorsal view. Figures 143–152: Comparison of dentition on movable finger of chela with field emission scanning electron microscopy (FESEM) images in dorsal view. Figures 139, 149. Scorpiops vrushchik sp. n., male, holotype. Figures 140, 150. Scorpiops nagphani sp. n., male, holotype. Figures 141, 151. Scorpiops neera sp. n., male, holotype. Figures 142, 152. Scorpiops lioneli sp. n., male, holotype. Figure 143. S. tenuicauda, male, INHER-214. Figure 144. S. satarensis, male, INHER-213. Figure 145. S. deccanensis, male, INHER-86. Figure 146. S. maharashtraensis, male, INHER-179. Figure 147. S. phaltanensis, male, paratype, INHER-64. Figure 148. S. telbaila, male, paratype, INHER-123." captionText-1="Figure 153. Distribution of Scorpiops species from Peninsular India. Localities with a star indicate new species described in this study." captionText-2="Figure 154. Boxplots showing variations in ratios of pedipalp chela length to width in males for all Scorpiops species from Peninsular India (except S. pachmarhicus due to unavailability of sufficient males)." captionText-3="Figure 155. Projection of first two Principal Component factors explaining 81.89% variation among the nine species of the genus Scorpiops from Peninsular India (except S. pachmarhicus and S. satarensis due to unavailability of sufficient males)." captionText-4="Figure 156. Maximum Likelihood phylogenetic tree (ML) for Scorpiops. Values along the nodes are bootstraps for 1000 iterations." captionText-5="Figure 157. Bayesian phylogenetic tree for Scorpiops. Values along the nodes are Bayesian posterior probabilities for Bayesian Inference and Bayesian Poisson Tree Process (bPTP) respectively. Vertical bars represent delimitation analyses results." figureDoi-0="http://doi.org/10.5281/zenodo.5742621" figureDoi-1="http://doi.org/10.5281/zenodo.5742623" figureDoi-2="http://doi.org/10.5281/zenodo.5742625" figureDoi-3="http://doi.org/10.5281/zenodo.5742627" figureDoi-4="http://doi.org/10.5281/zenodo.5742631" figureDoi-5="http://doi.org/10.5281/zenodo.5742633" httpUri-0="https://zenodo.org/record/5742621/files/figure.png" httpUri-1="https://zenodo.org/record/5742623/files/figure.png" httpUri-2="https://zenodo.org/record/5742625/files/figure.png" httpUri-3="https://zenodo.org/record/5742627/files/figure.png" httpUri-4="https://zenodo.org/record/5742631/files/figure.png" httpUri-5="https://zenodo.org/record/5742633/files/figure.png" pageId="33" pageNumber="32">142, 152–157</figureCitation>
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<tableCitation id="C6A716C1FFB67747FA9993FFBBF7C98D" box="[1355,1446,1757,1781]" captionStart="Table 4" captionStartId="32.[111,167,2026,2047]" captionText="Table 4. Morphometric data for Scorpiops lioneli sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to median width), depth (D)." pageId="33" pageNumber="32">Tables 4</tableCitation>
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<tableCitation id="C6A716C1FFB67747FBB393DFBADBC86D" box="[1121,1162,1789,1813]" captionStart-0="Table 5" captionStart-1="Table 6" captionStart-2="Table 7" captionStartId-0="44.[115,171,1066,1087]" captionStartId-1="44.[115,171,1470,1491]" captionStartId-2="44.[115,171,1970,1991]" captionText-0="Table 5. Pairwise uncorrected raw distances (%) expressed as minimum–maximum based on COI gene sequence for Scorpiops species from Maharashtra, India. Values in brackets are intra-clade distances." captionText-1="Table 6: Primers used for PCR amplification and sequencing of (COI) mitochondrial gene." captionText-2="Table 7: Voucher numbers and GenBank accession numbers for the sequence data used for the phylogenetic analysis." pageId="33" pageNumber="32">5–7</tableCitation>
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<location id="8EFA75A1FFB67747FBFC92BFBAB6C8CD" LSID="urn:lsid:plazi:treatment:038C926CFFB6774CFC3D9399BA8DCEBA:8EFA75A1FFB67747FBFC92BFBAB6C8CD" box="[1070,1255,1949,1973]" country="India" latitude="20.279722" longLatPrecision="21" longitude="74.8275" municipality="Nashik District" name="Pardhadi Village" pageId="33" pageNumber="32" stateProvince="Maharashtra">Pardhadi Village</location>
, 
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<geoCoordinate id="EE1145BDFFB67747FAAF92BFBD39C8AD" degrees="74" direction="east" minutes="49" orientation="longitude" pageId="33" pageNumber="32" precision="15" seconds="39" value="74.8275">74°49' 39&quot;E</geoCoordinate>
, 
<elevation id="0008C449FFB67747FCA6929FBDAFC8AD" box="[884,1022,1981,2005]" metricMagnitude="2" metricUnit="m" metricValue="5.59" pageId="33" pageNumber="32" unit="m" value="559.0">
<quantity id="4CDD8E9FFFB67747FCA6929FBDE6C8AD" box="[884,951,1981,2005]" metricMagnitude="2" metricUnit="m" metricValue="5.59" pageId="33" pageNumber="32" unit="m" value="559.0">559 m</quantity>
a. s. l.
</elevation>
</materialsCitation>
; 
<collectionCode id="ED34BBBFFFB67747FBD9929FBA07C8AD" box="[1035,1110,1981,2005]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="33" pageNumber="32">BNHS</collectionCode>
.
</paragraph>
<paragraph id="8B9A237AFFB67747FCE292FCBB93C88D" blockId="33.[816,1475,1917,2037]" box="[816,1474,2013,2037]" pageId="33" pageNumber="32">
<materialsCitation id="3B4D2927FFB67747FCE292FCBBEFC88D" box="[816,1470,2013,2037]" collectionCode="TYPE" country="India" location="Nashik District" pageId="33" pageNumber="32" specimenCount="1" stateProvince="Maharashtra" typeStatus="holotype">
<typeStatus id="549E9DD8FFB67747FCE292FCBD37C88D" box="[816,870,2014,2037]" pageId="33" pageNumber="32">TYPE</typeStatus>
MATERIAL. 
<emphasis id="B951FF68FFB67747FC3592FFBA75C88D" bold="true" box="[999,1060,2013,2037]" pageId="33" pageNumber="32">
<collectingCountry id="F33263EAFFB67747FC3592FFBA75C88D" box="[999,1060,2013,2037]" name="India" pageId="33" pageNumber="32">India</collectingCountry>
</emphasis>
, 
<emphasis id="B951FF68FFB67747FBE492FFBB52C88D" box="[1078,1283,2013,2037]" italics="true" pageId="33" pageNumber="32">
<collectingRegion id="49E1ED98FFB67747FBE492FFBB52C88D" box="[1078,1283,2013,2037]" country="India" name="Maharashtra" pageId="33" pageNumber="32">Maharashtra State</collectingRegion>
</emphasis>
, 
<location id="8EFA75A1FFB67747FAC792FFBBEFC88D" LSID="urn:lsid:plazi:treatment:038C926CFFB6774CFC3D9399BA8DCEBA:8EFA75A1FFB67747FAC792FFBBEFC88D" box="[1301,1470,2013,2037]" country="India" name="Nashik District" pageId="33" pageNumber="32" stateProvince="Maharashtra">Nashik District</location>
</materialsCitation>
,
</paragraph>
<caption id="DF5A73F2FFB57744FFA69299BCA2C896" ID-DOI="http://doi.org/10.5281/zenodo.5742601" ID-Zenodo-Dep="5742601" httpUri="https://zenodo.org/record/5742601/files/figure.png" pageId="34" pageNumber="33" startId="34.[116,193,1979,2001]" targetBox="[130,1447,323,1890]" targetPageId="34">
<paragraph id="8B9A237AFFB57744FFA69299BCA2C896" blockId="34.[116,1474,1979,2030]" pageId="34" pageNumber="33">
<emphasis id="B951FF68FFB57744FFA69299BF5EC8A8" bold="true" box="[116,271,1979,2001]" pageId="34" pageNumber="33">Figures 97–101</emphasis>
. 
<taxonomicName id="4C2558F9FFB57744FECE9299BF91C8A8" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[284,448,1979,2000]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="lioneli" status="sp. nov.">
<emphasis id="B951FF68FFB57744FECE9299BF91C8A8" box="[284,448,1979,2000]" italics="true" pageId="34" pageNumber="33">Scorpiops lioneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFB57744FE159299BFADC8A8" box="[455,508,1979,2000]" pageId="34" pageNumber="33" rank="species">
<emphasis id="B951FF68FFB57744FE159299BF8FC8A8" bold="true" box="[455,478,1979,2000]" pageId="34" pageNumber="33">sp</emphasis>
. 
<emphasis id="B951FF68FFB57744FE389299BFA6C8A8" bold="true" box="[490,503,1979,2000]" pageId="34" pageNumber="33">n</emphasis>
.
</taxonomicNameLabel>
, male, holotype, dorsal (97) and ventral (98) views, sternopectinal area (99), metasomal segment V and telson in lateral view (100) and chelicera in dorsal view (101).
</paragraph>
</caption>
<caption id="DF5A73F2FFB47745FFA391E4BAE8CBA4" ID-DOI="http://doi.org/10.5281/zenodo.5742603" ID-Zenodo-Dep="5742603" box="[113,1209,1222,1245]" httpUri="https://zenodo.org/record/5742603/files/figure.png" pageId="35" pageNumber="34" startId="35.[113,190,1222,1244]" targetBox="[113,1474,234,1210]" targetPageId="35">
<paragraph id="8B9A237AFFB47745FFA391E4BAE8CBA4" blockId="35.[113,1209,1222,1245]" box="[113,1209,1222,1245]" pageId="35" pageNumber="34">
<emphasis id="B951FF68FFB47745FFA391E4BF48CBA3" bold="true" box="[113,281,1222,1244]" pageId="35" pageNumber="34">Figures 102–103</emphasis>
. 
<taxonomicName id="4C2558F9FFB47745FEF691E5BF96CBA3" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[292,455,1222,1244]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="35" pageNumber="34" phylum="Arthropoda" rank="species" species="lioneli" status="sp. nov.">
<emphasis id="B951FF68FFB47745FEF691E5BF96CBA3" box="[292,455,1222,1244]" italics="true" pageId="35" pageNumber="34">Scorpiops lioneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFB47745FE1C91E5BC52CBA4" box="[462,515,1223,1244]" pageId="35" pageNumber="34" rank="species">
<emphasis id="B951FF68FFB47745FE1C91E5BFB4CBA4" bold="true" box="[462,485,1223,1244]" pageId="35" pageNumber="34">sp</emphasis>
. 
<emphasis id="B951FF68FFB47745FE2291E5BFACCBA4" bold="true" box="[496,509,1223,1244]" pageId="35" pageNumber="34">n</emphasis>
.
</taxonomicNameLabel>
, male, holotype in dorsal (102) and ventral (103) views under UV light.
</paragraph>
</caption>
<paragraph id="8B9A237AFFB47745FFA39020BFADCAC2" blockId="35.[113,771,1281,1466]" pageId="35" pageNumber="34">
<materialsCitation id="3B4D2927FFB47745FFA39020BCAECA22" collectingDate="2019-09-15" collectionCode="BNHS" collectorName="S. Sulakhe &amp; Sulakhe, C &amp; M. Ketkar" elevation="559" latitude="20.279722" location="Pardhadi Village" longLatPrecision="21" longitude="74.8275" pageId="35" pageNumber="34" specimenCode="SC 262" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<location id="8EFA75A1FFB47745FFA39020BF79CA62" LSID="urn:lsid:plazi:treatment:038C926CFFB6774CFC3D9399BA8DCEBA:8EFA75A1FFB47745FFA39020BF79CA62" box="[113,296,1282,1306]" latitude="20.279722" longLatPrecision="21" longitude="74.8275" name="Pardhadi Village" pageId="35" pageNumber="34">Pardhadi Village</location>
, 
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, 
<elevation id="0008C449FFB47745FD919023BC85CA62" box="[579,724,1281,1306]" metricMagnitude="2" metricUnit="m" metricValue="5.59" pageId="35" pageNumber="34" unit="m" value="559.0">
<quantity id="4CDD8E9FFFB47745FD919023BCD8CA61" box="[579,649,1281,1305]" metricMagnitude="2" metricUnit="m" metricValue="5.59" pageId="35" pageNumber="34" unit="m" value="559.0">559 m</quantity>
a. s. l.
</elevation>
, 
<specimenCount id="9D23E8F3FFB47745FD309020BD52CA62" box="[738,771,1282,1306]" pageId="35" pageNumber="34" type="male">1♂</specimenCount>
(
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, 
<collectionCode id="ED34BBBFFFB47745FF3B9000BF61CA42" box="[233,304,1314,1338]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="35" pageNumber="34">BNHS</collectionCode>
<specimenCode id="DB838B01FFB47745FEE99000BFC2CA42" box="[315,403,1314,1338]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="35" pageNumber="34">SC 262</specimenCode>
), 
<specimenCount id="9D23E8F3FFB47745FE7B9000BF9ACA42" box="[425,459,1314,1338]" pageId="35" pageNumber="34" type="male">1♂</specimenCount>
(
<typeStatus id="549E9DD8FFB47745FE0F9000BC6CCA42" box="[477,573,1314,1338]" pageId="35" pageNumber="34" type="paratype">paratype</typeStatus>
, INHER-177), 
<date id="FF9B05BAFFB47745FD3A9003BF76CA22" pageId="35" pageNumber="34" value="2019-09-15">
<collectingDate id="EFDFFC52FFB47745FD3A9003BF76CA22" pageId="35" pageNumber="34" value="2019-09-15">15 September 2019</collectingDate>
</date>
, leg. 
<collectorName id="26D046ACFFB47745FEBA9060BFB3CA22" box="[360,482,1346,1370]" pageId="35" pageNumber="34">S. Sulakhe</collectorName>
, C. Risbud &amp; 
<collectorName id="26D046ACFFB47745FD599060BCAECA22" box="[651,767,1346,1370]" pageId="35" pageNumber="34">M. Ketkar</collectorName>
</materialsCitation>
, 
<materialsCitation id="3B4D2927FFB47745FFA39040BFADCAC2" collectingDate="2021-03-16" collectionCode="BNHS" collectorName="S. Sulakhe &amp; Sulakhe, S &amp; Deshpande, A &amp; D. Pangarkar." pageId="35" pageNumber="34" specimenCode="SC 263, SC 264, 265" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D23E8F3FFB47745FFA39040BEC2CA02" box="[113,147,1378,1402]" pageId="35" pageNumber="34" type="male">1♂</specimenCount>
(
<typeStatus id="549E9DD8FFB47745FF719040BF52CA02" box="[163,259,1378,1402]" pageId="35" pageNumber="34" type="paratype">paratype</typeStatus>
, 
<collectionCode id="ED34BBBFFFB47745FEDD9040BF07CA02" box="[271,342,1378,1402]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="35" pageNumber="34">BNHS</collectionCode>
<specimenCode id="DB838B01FFB47745FE8D9040BFE5CA02" box="[351,436,1378,1402]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="35" pageNumber="34">SC 263</specimenCode>
), 
<specimenCount id="9D23E8F3FFB47745FE1B9040BFBACA02" box="[457,491,1378,1402]" pageId="35" pageNumber="34" type="female">3♀</specimenCount>
(
<typeStatus id="549E9DD8FFB47745FE289040BC35CA02" box="[506,612,1378,1402]" pageId="35" pageNumber="34" type="paratype">paratypes</typeStatus>
, INHER- 323, 
<collectionCode id="ED34BBBFFFB47745FFA390A0BEE9CAE2" box="[113,184,1410,1434]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="35" pageNumber="34">BNHS</collectionCode>
<specimenCode id="DB838B01FFB47745FF1790A0BF4DCAE2" box="[197,284,1410,1434]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="35" pageNumber="34">SC 264</specimenCode>
, 
<specimenCode id="DB838B01FFB47745FEFE90A3BF06CAE1" box="[300,343,1409,1433]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="35" pageNumber="34">265</specimenCode>
), 
<date id="FF9B05BAFFB47745FEA290A0BC71CAE2" box="[368,544,1410,1434]" pageId="35" pageNumber="34" value="2021-03-16">
<collectingDate id="EFDFFC52FFB47745FEA290A0BC71CAE2" box="[368,544,1410,1434]" pageId="35" pageNumber="34" value="2021-03-16">16 March 2021</collectingDate>
</date>
, leg. 
<collectorName id="26D046ACFFB47745FDB190A0BC8FCAE2" box="[611,734,1410,1434]" pageId="35" pageNumber="34">S. Sulakhe</collectorName>
, S. 
<collectorName id="26D046ACFFB47745FFA39080BF5BCAC2" box="[113,266,1442,1466]" pageId="35" pageNumber="34">Deshpande, A</collectorName>
. Date &amp; 
<collectorName id="26D046ACFFB47745FEBB9080BFADCAC2" box="[361,508,1442,1466]" pageId="35" pageNumber="34">D. Pangarkar.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33F70F1FFB47745FFA390C0BEA4C942" pageId="35" pageNumber="34" type="etymology">
<paragraph id="8B9A237AFFB47745FFA390C0BEA4C942" blockId="35.[113,771,1506,1594]" pageId="35" pageNumber="34">ETYMOLOGY. The specific epithet is a patronym honouring Lionel Monod for his remarkable contribution to the world of scorpiology.</paragraph>
</subSubSection>
<subSubSection id="C33F70F1FFB47745FFA39340BA13C902" pageId="35" pageNumber="34" type="diagnosis">
<paragraph id="8B9A237AFFB47745FFA39340BBD5CA02" blockId="35.[113,772,1634,2042]" lastBlockId="35.[816,1475,1282,2042]" pageId="35" pageNumber="34">
DIAGNOSIS (
<specimenCount id="9D23E8F3FFB47745FF3D9340BF57C902" box="[239,262,1634,1658]" pageId="35" pageNumber="34" type="male">♂</specimenCount>
<specimenCount id="9D23E8F3FFB47745FED49340BF47C902" box="[262,278,1634,1658]" pageId="35" pageNumber="34" type="female">♀</specimenCount>
). Total length 
<quantity id="4CDD8E9FFFB47745FE7F9340BC71C901" box="[429,544,1634,1658]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="4.9" metricValueMin="4.1" pageId="35" pageNumber="34" unit="mm" value="45.0" valueMax="49.0" valueMin="41.0">41–49 mm</quantity>
. Base color yellowish brown. Pectinal teeth number 
<quantity id="4CDD8E9FFFB47745FE7893A0BFBCC9E2" box="[426,493,1666,1690]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.778" pageId="35" pageNumber="34" unit="in" value="7.5" valueMax="8.0" valueMin="7.0">7–8 in</quantity>
both sexes, fulcra reduced to absent. Pectine morphology of 
<typeStatus id="549E9DD8FFB47745FE0E9380BC58C9C2" box="[476,521,1698,1722]" pageId="35" pageNumber="34">type</typeStatus>
P4. Patella of pedipalp with 23–27 (5 
<emphasis id="B951FF68FFB47745FEC393E0BF78C9A2" box="[273,297,1730,1754]" italics="true" pageId="35" pageNumber="34">eb</emphasis>
, 2 
<emphasis id="B951FF68FFB47745FE9D93E0BF20C9A2" box="[335,369,1730,1754]" italics="true" pageId="35" pageNumber="34">esb</emphasis>
, 2 
<emphasis id="B951FF68FFB47745FE4593E0BFEDC9A1" box="[407,444,1730,1753]" italics="true" pageId="35" pageNumber="34">em,</emphasis>
8–11 
<emphasis id="B951FF68FFB47745FDD393E0BC4CC9A1" box="[513,541,1730,1753]" italics="true" pageId="35" pageNumber="34">est</emphasis>
, 6–7 
<emphasis id="B951FF68FFB47745FD8F93E0BC3EC9A1" box="[605,623,1730,1753]" italics="true" pageId="35" pageNumber="34">et</emphasis>
) external and 15–18 ventral trichobothria. Chela of pedipalp with 4 ventral trichobothria located on ventral surface. Chelal trichobothrium 
<emphasis id="B951FF68FFB47745FFA39200BEC7C838" box="[113,150,1826,1856]" italics="true" pageId="35" pageNumber="34">
Eb 
<subScript id="17A1213FFFB47745FF5C9210BEC7C838" attach="left" box="[142,150,1842,1856]" fontSize="6" pageId="35" pageNumber="34">3</subScript>
</emphasis>
is located in proximal half of manus between trichobothria Dt and Db. Fingers of pedipalps strongly undulate in male and margins undulate in female. Chela length to width ratio 
<quantity id="4CDD8E9FFFB47745FD149240BD55C802" box="[710,772,1890,1914]" metricMagnitude="-1" metricUnit="m" metricValue="1.1176" pageId="35" pageNumber="34" unit="in" value="4.4">4.4 in</quantity>
males. Pedipalp movable finger with ca 65–70 IAD, which form second row, parallel with MD (ca 
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number); there are also 3–4 ID and 11–12 OD present. Internal surface of patella with small anterio-ventral tubercle. Carapace with deep U shaped anterior median notch.Tarsomere II of legs with 4–6 stout median ventral spinules. Metasoma I–III with ten, metasoma IV with 8 and metasoma V with 7 carinae. Telson elongate and smooth, length to depth ratio 2.8–3.5; annular ring absent.
</paragraph>
<paragraph id="8B9A237AFFB47745FCE290A0BB0BCAE2" blockId="35.[816,1475,1282,2042]" box="[816,1370,1410,1434]" pageId="35" pageNumber="34">
Description (
<specimenCount id="9D23E8F3FFB47745FC6E90A0BD81CAE2" box="[956,976,1410,1434]" pageId="35" pageNumber="34" type="male">♂</specimenCount>
<typeStatus id="549E9DD8FFB47745FC0590A0BA66CAE2" box="[983,1079,1410,1434]" pageId="35" pageNumber="34" type="holotype">holotype</typeStatus>
, measurements in 
<tableCitation id="C6A716C1FFB47745FB2E90A0BB1BCAE2" box="[1276,1354,1410,1434]" captionStart="Table 1" captionStartId="5.[113,169,1994,2015]" captionText="Table 1. Morphometric data for Scorpiops vrushchik sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to median width), depth (D)." pageId="35" pageNumber="34">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8B9A237AFFB47745FCE29083BA13C902" blockId="35.[816,1475,1282,2042]" pageId="35" pageNumber="34">
<emphasis id="B951FF68FFB47745FCE29083BDFACAC1" bold="true" box="[816,939,1441,1465]" pageId="35" pageNumber="34">Coloration</emphasis>
(in preservation) (
<figureCitation id="131E3FFFFFB47745FBA09080BAA5CAC2" box="[1138,1268,1442,1466]" captionStart-0="Figure 96" captionStart-1="Figures 97–101" captionStartId-0="33.[116,184,1219,1241]" captionStartId-1="34.[116,193,1979,2001]" captionTargetBox-0="[113,1474,232,1206]" captionTargetBox-1="[130,1447,323,1890]" captionTargetId-0="figure-422@33.[113,1474,229,1206]" captionTargetId-1="figure-16@34.[113,1474,295,1890]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionText-0="Figure 96. Scorpiops lioneli sp. n., male, paratype BHNS SC 263 in vivo habitus." captionText-1="Figures 97–101. Scorpiops lioneli sp. n., male, holotype, dorsal (97) and ventral (98) views, sternopectinal area (99), metasomal segment V and telson in lateral view (100) and chelicera in dorsal view (101)." figureDoi-0="http://doi.org/10.5281/zenodo.5742599" figureDoi-1="http://doi.org/10.5281/zenodo.5742601" httpUri-0="https://zenodo.org/record/5742599/files/figure.png" httpUri-1="https://zenodo.org/record/5742601/files/figure.png" pageId="35" pageNumber="34">Figs. 96–98</figureCitation>
, 
<figureCitation id="131E3FFFFFB47745FB2C9080BB31CAC1" box="[1278,1376,1441,1466]" captionStart="Figures 104–107" captionStartId="36.[113,190,1983,2005]" captionTargetBox="[183,1374,269,1965]" captionTargetId="figure-15@36.[113,1474,249,1965]" captionTargetPageId="36" captionText="Figures 104–107. Scorpiops lioneli sp. n., female, paratype, BNHS SC 264, in dorsal (104) ventral (105) views, sternopectinal area (106) and metasomal segment V and telson in lateral view (107)." figureDoi="http://doi.org/10.5281/zenodo.5742605" httpUri="https://zenodo.org/record/5742605/files/figure.png" pageId="35" pageNumber="34">104, 105</figureCitation>
). Overall body color yellowish brown. Legs yellowish brown. Telson brownish yellow on vesicle and dark brown on aculeus. Ventral portion of body yellowish brown. Carapace and fingers of manus dark brown. Pedipalps yellowish brown, darker on carinae. Chelicera basal segment dark brown. Fingers of chelicera brownish black.
</paragraph>
</subSubSection>
<subSubSection id="C33F70F1FFB4774CFCE293A3BEA8CCDB" lastPageId="42" lastPageNumber="41" pageId="35" pageNumber="34" type="description">
<paragraph id="8B9A237AFFB47745FCE293A3BA1EC8A2" blockId="35.[816,1475,1282,2042]" pageId="35" pageNumber="34">
<emphasis id="B951FF68FFB47745FCE293A3BDCFC9E1" bold="true" box="[816,926,1665,1689]" pageId="35" pageNumber="34">Carapace</emphasis>
(
<figureCitation id="131E3FFFFFB47745FC6593A0BA0DC9E2" box="[951,1116,1666,1690]" captionStart="Figures 108–111" captionStartId="37.[113,190,1627,1649]" captionTargetBox="[113,1474,234,1616]" captionTargetId="figure-185@37.[113,1474,234,1616]" captionTargetPageId="37" captionText="Figures 108–111: Scorpiops lioneli sp. n. Figures 108–109. Male, holotype, carapace under white light (108) and UV fluorescence (109). Figures 110–111. Female, paratype, BHNS SC 264, carapace under white light (110) and UV fluorescence (111)." figureDoi="http://doi.org/10.5281/zenodo.5742607" httpUri="https://zenodo.org/record/5742607/files/figure.png" pageId="35" pageNumber="34">Figs. 108–111</figureCitation>
). Anterior margin of carapace strongly granular with deep U shaped anterior median notch. Anterior sub margin coarsely granular. Anterior margin of carapace between lateral eyes and U shaped anterior median notch curved. Entire surface of carapace with coarse and fine granules. Anterio-lateral ocular tubercles granular provided with 
<typeStatus id="549E9DD8FFB47745FCB49260BDC5C822" box="[870,916,1858,1882]" pageId="35" pageNumber="34">type</typeStatus>
3A lateral ocelli. Two pairs of large major ocelli and one pair of minor ocelli. Median ocular tubercle smooth on dorsal portion with a pair of median ocelli situated in the ratio of 1:2 (ratio of median ocelli to anterior margin and median ocelli to posterior margin).
</paragraph>
<paragraph id="8B9A237AFFB47743FCE292C3BA81C881" blockId="35.[816,1475,1282,2042]" lastBlockId="37.[113,1475,1627,2041]" lastPageId="37" lastPageNumber="36" pageId="35" pageNumber="34">
<emphasis id="B951FF68FFB47745FCE292C3BDF8C881" bold="true" box="[816,937,2017,2041]" pageId="35" pageNumber="34">Chelicerae</emphasis>
(
<figureCitation id="131E3FFFFFB47745FC1092C0BA79C882" box="[962,1064,2018,2042]" captionStart="Figures 97–101" captionStartId="34.[116,193,1979,2001]" captionTargetBox="[130,1447,323,1890]" captionTargetId="figure-16@34.[113,1474,295,1890]" captionTargetPageId="34" captionText="Figures 97–101. Scorpiops lioneli sp. n., male, holotype, dorsal (97) and ventral (98) views, sternopectinal area (99), metasomal segment V and telson in lateral view (100) and chelicera in dorsal view (101)." figureDoi="http://doi.org/10.5281/zenodo.5742601" httpUri="https://zenodo.org/record/5742601/files/figure.png" pageId="35" pageNumber="34">Fig. 101</figureCitation>
). Proximal portion with reticulated mosaic design. Fixed finger of chelicera with 3 large triangular retrolateral dorsosubmedian, ventral retrolateral and ventral teeth on inner margin. Ventral fang of movable finger with a prolateral). Internal surface of patella with one large posteriorow of 7 minute teeth on inner margin. Dorsal fang of movable ventral tubercle with one small, thick adjacent bulge and one finger with 4 teeth on inner margin. adjacent small posterio-dorsal tubercle and small anterio- 
<emphasis id="B951FF68FFB27743FFA39202BE87C840" bold="true" box="[113,214,1824,1848]" pageId="37" pageNumber="36">Pedipalp</emphasis>
(
<figureCitation id="131E3FFFFFB27743FF359203BFD2C841" box="[231,387,1825,1849]" captionStart="Figures 112–117" captionStartId="38.[113,190,1297,1319]" captionTargetBox="[113,1474,234,1286]" captionTargetId="figure-400@38.[113,1474,234,1286]" captionTargetPageId="38" captionText="Figures 112–117. Scorpiops lioneli sp. n., male, holotype. Figures 112–117. Pedipalp chela dorsal (112), ventral (113) and dorsoexternal (114), patella external (115), dorsal (116) and ventral (117) views under UV fluorescence. Trichobothrial pattern indicated by yellow circles." figureDoi="http://doi.org/10.5281/zenodo.5742609" httpUri="https://zenodo.org/record/5742609/files/figure.png" pageId="37" pageNumber="36">Figs. 112–117</figureCitation>
, 
<figureCitation id="131E3FFFFFB27743FE429203BFA0C841" box="[400,497,1825,1849]" captionStart="Figures 127–138" captionStartId="41.[116,193,1928,1950]" captionTargetBox="[113,1474,245,1897]" captionTargetId="figure-102@41.[113,1474,234,1897]" captionTargetPageId="41" captionText="Figures 127–138: Figures 127–130: Comparison of left pedipalps. Figures 131–134: Comparison of metasomal segments II-IV in lateral view. Figures 135–138: Comparison of pedipalp patella dorsal view with internal aspect tubercles. Figures 127, 131, 135. Scorpiops vrushchik sp. n., male, holotype. Figures 128, 132, 136. Scorpiops nagphani sp. n., male, holotype. Figures 129, 133, 137. Scorpiops neera sp. n., male, holotype. Figures 130, 134, 138. Scorpiops lioneli sp. n., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.5742617" httpUri="https://zenodo.org/record/5742617/files/figure.png" pageId="37" pageNumber="36">130, 138</figureCitation>
, 
<figureCitation id="131E3FFFFFB27743FE2C9203BC7BC841" box="[510,554,1825,1849]" captionStart="Figures 139–152" captionStartId="43.[115,192,1183,1205]" captionTargetBox="[113,1474,234,1167]" captionTargetId="figure-329@43.[113,1474,234,1167]" captionTargetPageId="43" captionText="Figures 139–152: Figures 139–142: Comparison of dorsolateral granules on anal rim of metasomal segment V in dorsal view. Figures 143–152: Comparison of dentition on movable finger of chela with field emission scanning electron microscopy (FESEM) images in dorsal view. Figures 139, 149. Scorpiops vrushchik sp. n., male, holotype. Figures 140, 150. Scorpiops nagphani sp. n., male, holotype. Figures 141, 151. Scorpiops neera sp. n., male, holotype. Figures 142, 152. Scorpiops lioneli sp. n., male, holotype. Figure 143. S. tenuicauda, male, INHER-214. Figure 144. S. satarensis, male, INHER-213. Figure 145. S. deccanensis, male, INHER-86. Figure 146. S. maharashtraensis, male, INHER-179. Figure 147. S. phaltanensis, male, paratype, INHER-64. Figure 148. S. telbaila, male, paratype, INHER-123." figureDoi="http://doi.org/10.5281/zenodo.5742621" httpUri="https://zenodo.org/record/5742621/files/figure.png" pageId="37" pageNumber="36">152</figureCitation>
). Femur and patella ventral tubercle (
<figureCitation id="131E3FFFFFB27743FC3E9203BA1DC841" box="[1004,1100,1825,1849]" captionStart="Figures 127–138" captionStartId="41.[116,193,1928,1950]" captionTargetBox="[113,1474,245,1897]" captionTargetId="figure-102@41.[113,1474,234,1897]" captionTargetPageId="41" captionText="Figures 127–138: Figures 127–130: Comparison of left pedipalps. Figures 131–134: Comparison of metasomal segments II-IV in lateral view. Figures 135–138: Comparison of pedipalp patella dorsal view with internal aspect tubercles. Figures 127, 131, 135. Scorpiops vrushchik sp. n., male, holotype. Figures 128, 132, 136. Scorpiops nagphani sp. n., male, holotype. Figures 129, 133, 137. Scorpiops neera sp. n., male, holotype. Figures 130, 134, 138. Scorpiops lioneli sp. n., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.5742617" httpUri="https://zenodo.org/record/5742617/files/figure.png" pageId="37" pageNumber="36">Fig. 138</figureCitation>
). Manus elongated with 5 carinae dorsoventrally flattened. Femur with 6 carinae (dorsal dorsal retrolateral, retrolateral median, ventral retrolateral, retrolateral, dorsal prolateral, retrolateral dorsosubmedian, dorsal prolateral and ventral prolateral). Dorsal retrolateral prolateral ventrosubmedian, ventral retrolateral and ventral and dorsal prolateral carinae running anteriorly up to the tip prolateral). Ventral prolateral carina weak and present only of fixed finger. Intercarinal space finely granular on inner on 1/3rd proximal portion. Intercarinal space with fine and outer surface. Both fingers scalloped deeply at the base. granules. Patella with 5 (dorsal retrolateral, dorsal prolateral, Trichobothrial pattern neobothriotaxic.
</paragraph>
<caption id="DF5A73F2FFB37742FFA3929DBC2FC88A" ID-DOI="http://doi.org/10.5281/zenodo.5742605" ID-Zenodo-Dep="5742605" httpUri="https://zenodo.org/record/5742605/files/figure.png" pageId="36" pageNumber="35" startId="36.[113,190,1983,2005]" targetBox="[183,1374,269,1965]" targetPageId="36">
<paragraph id="8B9A237AFFB37742FFA3929DBC2FC88A" blockId="36.[113,1472,1983,2034]" pageId="36" pageNumber="35">
<emphasis id="B951FF68FFB37742FFA3929DBF49C8AC" bold="true" box="[113,280,1983,2005]" pageId="36" pageNumber="35">Figures 104–107</emphasis>
. 
<taxonomicName id="4C2558F9FFB37742FEF692E2BF97C8AC" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[292,454,1983,2005]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="35" phylum="Arthropoda" rank="species" species="lioneli" status="sp. nov.">
<emphasis id="B951FF68FFB37742FEF692E2BF97C8AC" box="[292,454,1983,2005]" italics="true" pageId="36" pageNumber="35">Scorpiops lioneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFB37742FE1E929DBC50C8AD" box="[460,513,1983,2005]" pageId="36" pageNumber="35" rank="species">
<emphasis id="B951FF68FFB37742FE1E929DBFB2C8AC" bold="true" box="[460,483,1983,2004]" pageId="36" pageNumber="35">sp</emphasis>
. 
<emphasis id="B951FF68FFB37742FE3D929DBFADC8AC" bold="true" box="[495,508,1983,2004]" pageId="36" pageNumber="35">n</emphasis>
.
</taxonomicNameLabel>
, female, paratype, BNHS SC 264, in dorsal (104) ventral (105) views, sternopectinal area (106) and metasomal segment V and telson in lateral view (107).
</paragraph>
</caption>
<caption id="DF5A73F2FFB27743FFA39379BAE1C9F6" ID-DOI="http://doi.org/10.5281/zenodo.5742607" ID-Zenodo-Dep="5742607" httpUri="https://zenodo.org/record/5742607/files/figure.png" pageId="37" pageNumber="36" startId="37.[113,190,1627,1649]" targetBox="[113,1474,234,1616]" targetPageId="37">
<paragraph id="8B9A237AFFB27743FFA39379BAE1C9F6" blockId="37.[113,1475,1627,2041]" pageId="37" pageNumber="36">
<emphasis id="B951FF68FFB27743FFA39379BF46C908" bold="true" box="[113,279,1627,1649]" pageId="37" pageNumber="36">Figures 108–111</emphasis>
: 
<taxonomicName id="4C2558F9FFB27743FEF49379BF9AC908" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[294,459,1627,1648]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="lioneli" status="sp. nov.">
<emphasis id="B951FF68FFB27743FEF49379BF9AC908" box="[294,459,1627,1648]" italics="true" pageId="37" pageNumber="36">Scorpiops lioneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFB27743FE019379BC5AC908" box="[467,523,1627,1648]" pageId="37" pageNumber="36" rank="species">
<emphasis id="B951FF68FFB27743FE019379BFBBC908" bold="true" box="[467,490,1627,1648]" pageId="37" pageNumber="36">sp</emphasis>
. 
<emphasis id="B951FF68FFB27743FE259379BC55C908" bold="true" box="[503,516,1627,1648]" pageId="37" pageNumber="36">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B951FF68FFB27743FDC19379BCEDC908" bold="true" box="[531,700,1627,1649]" pageId="37" pageNumber="36">Figures 108–109</emphasis>
. Male, holotype, carapace under white light (108) and UV fluorescence (109). 
<emphasis id="B951FF68FFB27743FFA39355BF44C9F4" bold="true" box="[113,277,1655,1677]" pageId="37" pageNumber="36">Figures 110–111</emphasis>
. Female, paratype, BHNS SC 264, carapace under white light (110) and UV fluorescence (111).
</paragraph>
</caption>
<caption id="DF5A73F2FFB17740FFA39033BBE4CA3C" ID-DOI="http://doi.org/10.5281/zenodo.5742609" ID-Zenodo-Dep="5742609" httpUri="https://zenodo.org/record/5742609/files/figure.png" pageId="38" pageNumber="37" startId="38.[113,190,1297,1319]" targetBox="[113,1474,234,1286]" targetPageId="38">
<paragraph id="8B9A237AFFB17740FFA39033BBE4CA3C" blockId="38.[113,1472,1297,1348]" pageId="38" pageNumber="37">
<emphasis id="B951FF68FFB17740FFA39033BF49CA5E" bold="true" box="[113,280,1297,1319]" pageId="38" pageNumber="37">Figures 112–117</emphasis>
. 
<taxonomicName id="4C2558F9FFB17740FEF49030BF9DCA5E" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[294,460,1297,1319]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="lioneli" status="sp. nov.">
<emphasis id="B951FF68FFB17740FEF49030BF9DCA5E" box="[294,460,1297,1319]" italics="true" pageId="38" pageNumber="37">Scorpiops lioneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFB17740FE069033BC5DCA5F" box="[468,524,1297,1319]" pageId="38" pageNumber="37" rank="species">
<emphasis id="B951FF68FFB17740FE069033BFBACA5E" bold="true" box="[468,491,1297,1318]" pageId="38" pageNumber="37">sp</emphasis>
. 
<emphasis id="B951FF68FFB17740FE2B9033BC57CA5E" bold="true" box="[505,518,1297,1318]" pageId="38" pageNumber="37">n</emphasis>
.
</taxonomicNameLabel>
, male, holotype. 
<emphasis id="B951FF68FFB17740FD699033BD33CA5E" bold="true" box="[699,866,1297,1319]" pageId="38" pageNumber="37">Figures 112–117</emphasis>
. Pedipalp chela dorsal (112), ventral (113) and dorsoexternal (114), patella external (115), dorsal (116) and ventral (117) views under UV fluorescence. Trichobothrial pattern indicated by yellow circles.
</paragraph>
</caption>
<paragraph id="8B9A237AFFB17740FFA39041BC22CAA3" blockId="38.[113,772,1378,2043]" pageId="38" pageNumber="37">
<emphasis id="B951FF68FFB17740FFA39041BEF7CA03" bold="true" box="[113,166,1379,1403]" pageId="38" pageNumber="37">Legs</emphasis>
(
<figureCitation id="131E3FFFFFB17740FF689041BF15CA03" box="[186,324,1379,1403]" captionStart="Figures 97–101" captionStartId="34.[116,193,1979,2001]" captionTargetBox="[130,1447,323,1890]" captionTargetId="figure-16@34.[113,1474,295,1890]" captionTargetPageId="34" captionText="Figures 97–101. Scorpiops lioneli sp. n., male, holotype, dorsal (97) and ventral (98) views, sternopectinal area (99), metasomal segment V and telson in lateral view (100) and chelicera in dorsal view (101)." figureDoi="http://doi.org/10.5281/zenodo.5742601" httpUri="https://zenodo.org/record/5742601/files/figure.png" pageId="38" pageNumber="37">Figs. 97, 98</figureCitation>
, 
<figureCitation id="131E3FFFFFB17740FE819040BFE2CA02" box="[339,435,1378,1402]" captionStart-0="Figures 102–103" captionStart-1="Figures 104–107" captionStartId-0="35.[113,190,1222,1244]" captionStartId-1="36.[113,190,1983,2005]" captionTargetBox-0="[113,1474,234,1210]" captionTargetBox-1="[183,1374,269,1965]" captionTargetId-0="figure-424@35.[113,1474,234,1210]" captionTargetId-1="figure-15@36.[113,1474,249,1965]" captionTargetPageId-0="35" captionTargetPageId-1="36" captionText-0="Figures 102–103. Scorpiops lioneli sp. n., male, holotype in dorsal (102) and ventral (103) views under UV light." captionText-1="Figures 104–107. Scorpiops lioneli sp. n., female, paratype, BNHS SC 264, in dorsal (104) ventral (105) views, sternopectinal area (106) and metasomal segment V and telson in lateral view (107)." figureDoi-0="http://doi.org/10.5281/zenodo.5742603" figureDoi-1="http://doi.org/10.5281/zenodo.5742605" httpUri-0="https://zenodo.org/record/5742603/files/figure.png" httpUri-1="https://zenodo.org/record/5742605/files/figure.png" pageId="38" pageNumber="37">102–105</figureCitation>
, 
<figureCitation id="131E3FFFFFB17740FE119041BC72CA03" box="[451,547,1379,1403]" captionStart="Figures 118–124" captionStartId="39.[116,193,1942,1964]" captionTargetBox="[143,1429,224,1936]" captionTargetId="figure-8@39.[113,1474,221,1938]" captionTargetPageId="39" captionText="Figures 118–124. Scorpiops lioneli sp. n. Figures 118–121. Scorpiops lioneli sp. n., male, holotype, left legs I-IV, retrolateral aspect. Figures 122–124. Male, paratype, BNHS SC 263, hemispermatophore internal (122), external (123) views and capsular region in internal view (124). Abbrevations: distal lamina (dl), capsule (c), trunk (t), pedicel (p), distal posterior lobe (dpl), lateral hook (lh), terminal membrane of the sperm duct (tmsd) and basal carina (bc)." figureDoi="http://doi.org/10.5281/zenodo.5742613" httpUri="https://zenodo.org/record/5742613/files/figure.png" pageId="38" pageNumber="37">118–121</figureCitation>
). Femur and patella carinated, intercarinal space with coarse and fine granules. Tarsomere I provided with three to four rows of spinules and tarsomere II with single ventral row of spinules.
</paragraph>
<paragraph id="8B9A237AFFB17740FFA390C0BFC8C863" blockId="38.[113,772,1378,2043]" pageId="38" pageNumber="37">
<emphasis id="B951FF68FFB17740FFA390C0BE85CA82" bold="true" box="[113,212,1506,1530]" pageId="38" pageNumber="37">Sternum</emphasis>
, 
<emphasis id="B951FF68FFB17740FF3790C0BC01CA82" bold="true" box="[229,592,1506,1531]" pageId="38" pageNumber="37">genital operculum and pectines</emphasis>
(
<figureCitation id="131E3FFFFFB17740FDB390C1BCEFCA83" box="[609,702,1507,1531]" captionStart="Figures 97–101" captionStartId="34.[116,193,1979,2001]" captionTargetBox="[130,1447,323,1890]" captionTargetId="figure-16@34.[113,1474,295,1890]" captionTargetPageId="34" captionText="Figures 97–101. Scorpiops lioneli sp. n., male, holotype, dorsal (97) and ventral (98) views, sternopectinal area (99), metasomal segment V and telson in lateral view (100) and chelicera in dorsal view (101)." figureDoi="http://doi.org/10.5281/zenodo.5742601" httpUri="https://zenodo.org/record/5742601/files/figure.png" pageId="38" pageNumber="37">Figs. 99</figureCitation>
, 
<figureCitation id="131E3FFFFFB17740FD1E90C1BCA9CA83" box="[716,760,1507,1531]" captionStart="Figures 104–107" captionStartId="36.[113,190,1983,2005]" captionTargetBox="[183,1374,269,1965]" captionTargetId="figure-15@36.[113,1474,249,1965]" captionTargetPageId="36" captionText="Figures 104–107. Scorpiops lioneli sp. n., female, paratype, BNHS SC 264, in dorsal (104) ventral (105) views, sternopectinal area (106) and metasomal segment V and telson in lateral view (107)." figureDoi="http://doi.org/10.5281/zenodo.5742605" httpUri="https://zenodo.org/record/5742605/files/figure.png" pageId="38" pageNumber="37">106</figureCitation>
). Broad, pentagonal and finely granular only on anterior middle portion. Genital operculum with a pair of strongly protruding genital papillae. Basal piece without depression on middle portion. Pectine morphology of 
<typeStatus id="549E9DD8FFB17740FE129341BFBFC903" box="[448,494,1635,1659]" pageId="38" pageNumber="37">type</typeStatus>
P4 with 8/7 pectinal teeth. 
<emphasis id="B951FF68FFB17740FFA393A1BEB8C9E3" bold="true" box="[113,233,1667,1691]" pageId="38" pageNumber="37">Mesosoma</emphasis>
(
<figureCitation id="131E3FFFFFB17740FF2D93A1BFC1C9E3" box="[255,400,1667,1691]" captionStart="Figures 97–101" captionStartId="34.[116,193,1979,2001]" captionTargetBox="[130,1447,323,1890]" captionTargetId="figure-16@34.[113,1474,295,1890]" captionTargetPageId="34" captionText="Figures 97–101. Scorpiops lioneli sp. n., male, holotype, dorsal (97) and ventral (98) views, sternopectinal area (99), metasomal segment V and telson in lateral view (100) and chelicera in dorsal view (101)." figureDoi="http://doi.org/10.5281/zenodo.5742601" httpUri="https://zenodo.org/record/5742601/files/figure.png" pageId="38" pageNumber="37">Figs. 97, 98</figureCitation>
, 
<figureCitation id="131E3FFFFFB17740FE7093A0BC55C9E2" box="[418,516,1666,1690]" captionStart-0="Figures 102–103" captionStart-1="Figures 104–107" captionStartId-0="35.[113,190,1222,1244]" captionStartId-1="36.[113,190,1983,2005]" captionTargetBox-0="[113,1474,234,1210]" captionTargetBox-1="[183,1374,269,1965]" captionTargetId-0="figure-424@35.[113,1474,234,1210]" captionTargetId-1="figure-15@36.[113,1474,249,1965]" captionTargetPageId-0="35" captionTargetPageId-1="36" captionText-0="Figures 102–103. Scorpiops lioneli sp. n., male, holotype in dorsal (102) and ventral (103) views under UV light." captionText-1="Figures 104–107. Scorpiops lioneli sp. n., female, paratype, BNHS SC 264, in dorsal (104) ventral (105) views, sternopectinal area (106) and metasomal segment V and telson in lateral view (107)." figureDoi-0="http://doi.org/10.5281/zenodo.5742603" figureDoi-1="http://doi.org/10.5281/zenodo.5742605" httpUri-0="https://zenodo.org/record/5742603/files/figure.png" httpUri-1="https://zenodo.org/record/5742605/files/figure.png" pageId="38" pageNumber="37">102–105</figureCitation>
).). All tergites finely granular, with median carina absent on segment I. Tergite VII additionally with two pairs of lateral granular carinae present only on half posterior portion. Sternites III-VI entirely smooth. Sternite VII finely granular.
</paragraph>
<paragraph id="8B9A237AFFB17740FFA39201BA2FC963" blockId="38.[113,772,1378,2043]" lastBlockId="38.[816,1475,1379,2043]" pageId="38" pageNumber="37">
<emphasis id="B951FF68FFB17740FFA39201BEB6C843" bold="true" box="[113,231,1827,1851]" pageId="38" pageNumber="37">Metasoma</emphasis>
(
<figureCitation id="131E3FFFFFB17740FF2F9201BFDBC843" box="[253,394,1827,1851]" captionStart="Figures 97–101" captionStartId="34.[116,193,1979,2001]" captionTargetBox="[130,1447,323,1890]" captionTargetId="figure-16@34.[113,1474,295,1890]" captionTargetPageId="34" captionText="Figures 97–101. Scorpiops lioneli sp. n., male, holotype, dorsal (97) and ventral (98) views, sternopectinal area (99), metasomal segment V and telson in lateral view (100) and chelicera in dorsal view (101)." figureDoi="http://doi.org/10.5281/zenodo.5742601" httpUri="https://zenodo.org/record/5742601/files/figure.png" pageId="38" pageNumber="37">Figs. 97, 98</figureCitation>
, 
<figureCitation id="131E3FFFFFB17740FE4E9200BFADC842" box="[412,508,1826,1850]" captionStart-0="Figures 102–103" captionStart-1="Figures 104–107" captionStartId-0="35.[113,190,1222,1244]" captionStartId-1="36.[113,190,1983,2005]" captionTargetBox-0="[113,1474,234,1210]" captionTargetBox-1="[183,1374,269,1965]" captionTargetId-0="figure-424@35.[113,1474,234,1210]" captionTargetId-1="figure-15@36.[113,1474,249,1965]" captionTargetPageId-0="35" captionTargetPageId-1="36" captionText-0="Figures 102–103. Scorpiops lioneli sp. n., male, holotype in dorsal (102) and ventral (103) views under UV light." captionText-1="Figures 104–107. Scorpiops lioneli sp. n., female, paratype, BNHS SC 264, in dorsal (104) ventral (105) views, sternopectinal area (106) and metasomal segment V and telson in lateral view (107)." figureDoi-0="http://doi.org/10.5281/zenodo.5742603" figureDoi-1="http://doi.org/10.5281/zenodo.5742605" httpUri-0="https://zenodo.org/record/5742603/files/figure.png" httpUri-1="https://zenodo.org/record/5742605/files/figure.png" pageId="38" pageNumber="37">102–105</figureCitation>
, 
<figureCitation id="131E3FFFFFB17740FDDF9201BC66C843" box="[525,567,1827,1851]" captionStart="Figures 127–138" captionStartId="41.[116,193,1928,1950]" captionTargetBox="[113,1474,245,1897]" captionTargetId="figure-102@41.[113,1474,234,1897]" captionTargetPageId="41" captionText="Figures 127–138: Figures 127–130: Comparison of left pedipalps. Figures 131–134: Comparison of metasomal segments II-IV in lateral view. Figures 135–138: Comparison of pedipalp patella dorsal view with internal aspect tubercles. Figures 127, 131, 135. Scorpiops vrushchik sp. n., male, holotype. Figures 128, 132, 136. Scorpiops nagphani sp. n., male, holotype. Figures 129, 133, 137. Scorpiops neera sp. n., male, holotype. Figures 130, 134, 138. Scorpiops lioneli sp. n., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.5742617" httpUri="https://zenodo.org/record/5742617/files/figure.png" pageId="38" pageNumber="37">134</figureCitation>
, 
<figureCitation id="131E3FFFFFB17740FD9A9201BC25C843" box="[584,628,1827,1851]" captionStart="Figures 139–152" captionStartId="43.[115,192,1183,1205]" captionTargetBox="[113,1474,234,1167]" captionTargetId="figure-329@43.[113,1474,234,1167]" captionTargetPageId="43" captionText="Figures 139–152: Figures 139–142: Comparison of dorsolateral granules on anal rim of metasomal segment V in dorsal view. Figures 143–152: Comparison of dentition on movable finger of chela with field emission scanning electron microscopy (FESEM) images in dorsal view. Figures 139, 149. Scorpiops vrushchik sp. n., male, holotype. Figures 140, 150. Scorpiops nagphani sp. n., male, holotype. Figures 141, 151. Scorpiops neera sp. n., male, holotype. Figures 142, 152. Scorpiops lioneli sp. n., male, holotype. Figure 143. S. tenuicauda, male, INHER-214. Figure 144. S. satarensis, male, INHER-213. Figure 145. S. deccanensis, male, INHER-86. Figure 146. S. maharashtraensis, male, INHER-179. Figure 147. S. phaltanensis, male, paratype, INHER-64. Figure 148. S. telbaila, male, paratype, INHER-123." figureDoi="http://doi.org/10.5281/zenodo.5742621" httpUri="https://zenodo.org/record/5742621/files/figure.png" pageId="38" pageNumber="37">142</figureCitation>
). Metasomal segments I with 5 pairs of carinae (dorsal lateral, lateral supramedian, lateral inframedian, ventral lateral and ventral submedian), metasomal segments II, III and IV with 4 pairs of carinae (dorsal lateral, lateral supramedian, ventral lateral and ventral submedian), segment II and III with additional lateral inframedian carina present only on posterior portion and segment V with 3 pairs (dorsal lateral, lateral inframedian and ventral lateral) and 1 single ventral median carinae. Intercarinal space with fine granules. Dorsal lateral carination on segments III and IV ending posteriorly into a short spine. Anal rim of segment V evenly crenulated. A pair of dorsolateral granules of anal rim weakly tuberculate.
</paragraph>
<paragraph id="8B9A237AFFB17740FCE29301BD9CC8A3" blockId="38.[816,1475,1379,2043]" pageId="38" pageNumber="37">
<emphasis id="B951FF68FFB17740FCE29301BD28C943" bold="true" box="[816,889,1571,1595]" pageId="38" pageNumber="37">Telson</emphasis>
(
<figureCitation id="131E3FFFFFB17740FC549301BDBBC943" box="[902,1002,1571,1595]" captionStart="Figures 97–101" captionStartId="34.[116,193,1979,2001]" captionTargetBox="[130,1447,323,1890]" captionTargetId="figure-16@34.[113,1474,295,1890]" captionTargetPageId="34" captionText="Figures 97–101. Scorpiops lioneli sp. n., male, holotype, dorsal (97) and ventral (98) views, sternopectinal area (99), metasomal segment V and telson in lateral view (100) and chelicera in dorsal view (101)." figureDoi="http://doi.org/10.5281/zenodo.5742601" httpUri="https://zenodo.org/record/5742601/files/figure.png" pageId="38" pageNumber="37">Figs. 100</figureCitation>
, 
<figureCitation id="131E3FFFFFB17740FC219301BA4EC943" box="[1011,1055,1571,1595]" captionStart="Figures 104–107" captionStartId="36.[113,190,1983,2005]" captionTargetBox="[183,1374,269,1965]" captionTargetId="figure-15@36.[113,1474,249,1965]" captionTargetPageId="36" captionText="Figures 104–107. Scorpiops lioneli sp. n., female, paratype, BNHS SC 264, in dorsal (104) ventral (105) views, sternopectinal area (106) and metasomal segment V and telson in lateral view (107)." figureDoi="http://doi.org/10.5281/zenodo.5742605" httpUri="https://zenodo.org/record/5742605/files/figure.png" pageId="38" pageNumber="37">107</figureCitation>
). Elongated and almost entirely smooth. 
<emphasis id="B951FF68FFB17740FCE29360BA4AC922" bold="true" box="[816,1051,1602,1626]" pageId="38" pageNumber="37">Hemispermatophore</emphasis>
<specimenCount id="9D23E8F3FFB17740FBFE9361BA11C922" box="[1068,1088,1603,1626]" pageId="38" pageNumber="37" type="male">♂</specimenCount>
<typeStatus id="549E9DD8FFB17740FB829361BAFEC923" box="[1104,1199,1603,1627]" pageId="38" pageNumber="37" type="paratype">paratype</typeStatus>
, BNHS SC 263 (
<figureCitation id="131E3FFFFFB17740FA5E9361BDC3C903" captionStart="Figures 118–124" captionStartId="39.[116,193,1942,1964]" captionTargetBox="[143,1429,224,1936]" captionTargetId="figure-8@39.[113,1474,221,1938]" captionTargetPageId="39" captionText="Figures 118–124. Scorpiops lioneli sp. n. Figures 118–121. Scorpiops lioneli sp. n., male, holotype, left legs I-IV, retrolateral aspect. Figures 122–124. Male, paratype, BNHS SC 263, hemispermatophore internal (122), external (123) views and capsular region in internal view (124). Abbrevations: distal lamina (dl), capsule (c), trunk (t), pedicel (p), distal posterior lobe (dpl), lateral hook (lh), terminal membrane of the sperm duct (tmsd) and basal carina (bc)." figureDoi="http://doi.org/10.5281/zenodo.5742613" httpUri="https://zenodo.org/record/5742613/files/figure.png" pageId="38" pageNumber="37">Figs. 122–124</figureCitation>
). Lamelliform; distal 
<emphasis id="B951FF68FFB17740FBAC9341BB56C902" box="[1150,1287,1635,1659]" italics="true" pageId="38" pageNumber="37">basal carina</emphasis>
(
<emphasis id="B951FF68FFB17740FACA9341BB60C903" box="[1304,1329,1635,1659]" italics="true" pageId="38" pageNumber="37">bc</emphasis>
) with a plate like structure present, sclerotized, crown-like structure (
<emphasis id="B951FF68FFB17740FA4E93A1BBEBC9E3" box="[1436,1466,1667,1691]" italics="true" pageId="38" pageNumber="37">cls</emphasis>
) with distal margin with 5–6 denticles whose internal surface is attached to terminal membrane of the sperm duct (
<emphasis id="B951FF68FFB17740FAB793E1BBC6C9A3" box="[1381,1431,1731,1755]" italics="true" pageId="38" pageNumber="37">tmsd</emphasis>
) on the inner concave side of the capsule with numerous spicules; pointed lateral hook (
<emphasis id="B951FF68FFB17740FBFF9221BA13C863" box="[1069,1090,1795,1819]" italics="true" pageId="38" pageNumber="37">lh</emphasis>
); a long, inwardly twisted distal posterior lobe (
<emphasis id="B951FF68FFB17740FC009201BDA5C843" box="[978,1012,1827,1851]" italics="true" pageId="38" pageNumber="37">dpl</emphasis>
) close to the distal lamina, with denticulate margin; short and stout trunk; long and slender distal lamina (
<emphasis id="B951FF68FFB17740FCEB9241BD1FC803" box="[825,846,1891,1915]" italics="true" pageId="38" pageNumber="37">dl</emphasis>
) narrow at the base and sharply bent and tapered apex. Total length 8.0 mm; pedicel 
<quantity id="4CDD8E9FFFB17740FBA492A1BA87C8E2" box="[1142,1238,1923,1947]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" pageId="38" pageNumber="37" unit="mm" value="1.14">1.14 mm</quantity>
long; trunk 
<quantity id="4CDD8E9FFFB17740FAB392A1BB93C8E2" box="[1377,1474,1923,1947]" metricMagnitude="-3" metricUnit="m" metricValue="1.66" pageId="38" pageNumber="37" unit="mm" value="1.66">1.66 mm</quantity>
long; capsule 
<quantity id="4CDD8E9FFFB17740FC189281BA7DC8C2" box="[970,1068,1955,1979]" metricMagnitude="-3" metricUnit="m" metricValue="1.12" pageId="38" pageNumber="37" unit="mm" value="1.12">1.12 mm</quantity>
long; 
<emphasis id="B951FF68FFB17740FBA49281BAC9C8C3" box="[1142,1176,1955,1979]" italics="true" pageId="38" pageNumber="37">dpl</emphasis>
length 
<quantity id="4CDD8E9FFFB17740FB3C9281BB02C8C2" box="[1262,1363,1955,1979]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="38" pageNumber="37" unit="mm" value="0.43">0.43 mm</quantity>
; 
<emphasis id="B951FF68FFB17740FAB39281BB27C8C3" box="[1377,1398,1955,1979]" italics="true" pageId="38" pageNumber="37">dl</emphasis>
length 
<quantity id="4CDD8E9FFFB17740FCE292E1BDDEC8A2" box="[816,911,1987,2011]" metricMagnitude="-3" metricUnit="m" metricValue="4.08" pageId="38" pageNumber="37" unit="mm" value="4.08">4.08 mm</quantity>
long.
</paragraph>
<paragraph id="8B9A237AFFB1774CFCE292C1BF89CE3B" blockId="38.[816,1475,1379,2043]" lastBlockId="42.[113,771,235,323]" lastPageId="42" lastPageNumber="41" pageId="38" pageNumber="37">
SEXUAL DIMORPHISM. Male genital operculum with pair of genital papillae. In females, the genital operculum medially sutured. Pedipalp manus slender in males and stouter in females (
<figureCitation id="131E3FFFFFBD774CFF019409BF30CE3B" box="[211,353,299,323]" captionStart="Figures 97–101" captionStartId="34.[116,193,1979,2001]" captionTargetBox="[130,1447,323,1890]" captionTargetId="figure-16@34.[113,1474,295,1890]" captionTargetPageId="34" captionText="Figures 97–101. Scorpiops lioneli sp. n., male, holotype, dorsal (97) and ventral (98) views, sternopectinal area (99), metasomal segment V and telson in lateral view (100) and chelicera in dorsal view (101)." figureDoi="http://doi.org/10.5281/zenodo.5742601" httpUri="https://zenodo.org/record/5742601/files/figure.png" pageId="42" pageNumber="41">Figs. 97–100</figureCitation>
, 
<figureCitation id="131E3FFFFFBD774CFEBE9409BF9DCE3B" box="[364,460,299,323]" captionStart-0="Figures 102–103" captionStart-1="Figures 104–107" captionStart-2="Figures 108–111" captionStartId-0="35.[113,190,1222,1244]" captionStartId-1="36.[113,190,1983,2005]" captionStartId-2="37.[113,190,1627,1649]" captionTargetBox-0="[113,1474,234,1210]" captionTargetBox-1="[183,1374,269,1965]" captionTargetBox-2="[113,1474,234,1616]" captionTargetId-0="figure-424@35.[113,1474,234,1210]" captionTargetId-1="figure-15@36.[113,1474,249,1965]" captionTargetId-2="figure-185@37.[113,1474,234,1616]" captionTargetPageId-0="35" captionTargetPageId-1="36" captionTargetPageId-2="37" captionText-0="Figures 102–103. Scorpiops lioneli sp. n., male, holotype in dorsal (102) and ventral (103) views under UV light." captionText-1="Figures 104–107. Scorpiops lioneli sp. n., female, paratype, BNHS SC 264, in dorsal (104) ventral (105) views, sternopectinal area (106) and metasomal segment V and telson in lateral view (107)." captionText-2="Figures 108–111: Scorpiops lioneli sp. n. Figures 108–109. Male, holotype, carapace under white light (108) and UV fluorescence (109). Figures 110–111. Female, paratype, BHNS SC 264, carapace under white light (110) and UV fluorescence (111)." figureDoi-0="http://doi.org/10.5281/zenodo.5742603" figureDoi-1="http://doi.org/10.5281/zenodo.5742605" figureDoi-2="http://doi.org/10.5281/zenodo.5742607" httpUri-0="https://zenodo.org/record/5742603/files/figure.png" httpUri-1="https://zenodo.org/record/5742605/files/figure.png" httpUri-2="https://zenodo.org/record/5742607/files/figure.png" pageId="42" pageNumber="41">102–111</figureCitation>
).
</paragraph>
<caption id="DF5A73F2FFB07741FFA692B4BFEBC77B" ID-DOI="http://doi.org/10.5281/zenodo.5742613" ID-Zenodo-Dep="5742613" httpUri="https://zenodo.org/record/5742613/files/figure.png" pageId="39" pageNumber="38" startId="39.[116,193,1942,1964]" targetBox="[143,1429,224,1936]" targetPageId="39">
<paragraph id="8B9A237AFFB07741FFA692B4BFEBC77B" blockId="39.[116,1475,1942,2051]" pageId="39" pageNumber="38">
<emphasis id="B951FF68FFB07741FFA692B4BF48C8D4" bold="true" box="[116,281,1942,1964]" pageId="39" pageNumber="38">Figures 118–124</emphasis>
. 
<taxonomicName id="4C2558F9FFB07741FEF692B5BF96C8D4" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[292,455,1943,1964]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="lioneli" status="sp. nov.">
<emphasis id="B951FF68FFB07741FEF692B5BF96C8D4" box="[292,455,1943,1964]" italics="true" pageId="39" pageNumber="38">Scorpiops lioneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFB07741FE1F92B5BC53C8D4" box="[461,514,1943,1964]" pageId="39" pageNumber="38" rank="species">
<emphasis id="B951FF68FFB07741FE1F92B5BFB5C8D4" bold="true" box="[461,484,1943,1964]" pageId="39" pageNumber="38">sp</emphasis>
. 
<emphasis id="B951FF68FFB07741FE3D92B5BFADC8D4" bold="true" box="[495,508,1943,1964]" pageId="39" pageNumber="38">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B951FF68FFB07741FDDA92B4BCFCC8D4" bold="true" box="[520,685,1942,1964]" pageId="39" pageNumber="38">Figures 118–121</emphasis>
. 
<taxonomicName id="4C2558F9FFB07741FD6A92B5BD0AC8D4" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[696,859,1943,1964]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="lioneli" status="sp. nov.">
<emphasis id="B951FF68FFB07741FD6A92B5BD0AC8D4" box="[696,859,1943,1964]" italics="true" pageId="39" pageNumber="38">Scorpiops lioneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFB07741FCB392B5BDC4C8D4" box="[865,917,1943,1964]" pageId="39" pageNumber="38" rank="species">
<emphasis id="B951FF68FFB07741FCB392B5BD29C8D4" bold="true" box="[865,888,1943,1964]" pageId="39" pageNumber="38">sp</emphasis>
. 
<emphasis id="B951FF68FFB07741FC5192B5BDC1C8D4" bold="true" box="[899,912,1943,1964]" pageId="39" pageNumber="38">n</emphasis>
.
</taxonomicNameLabel>
, male, holotype, left legs I-IV, retrolateral aspect. 
<emphasis id="B951FF68FFB07741FAA792B4BE99C8B0" bold="true" pageId="39" pageNumber="38">Figures 122–124</emphasis>
. Male, paratype, BNHS SC 263, hemispermatophore internal (122), external (123) views and capsular region in internal view (124). Abbrevations: distal lamina (
<emphasis id="B951FF68FFB07741FE5B92F2BFCDC89D" box="[393,412,2000,2021]" italics="true" pageId="39" pageNumber="38">dl</emphasis>
), capsule (
<emphasis id="B951FF68FFB07741FDD692F3BC5EC89E" box="[516,527,2001,2022]" italics="true" pageId="39" pageNumber="38">c</emphasis>
), trunk (
<emphasis id="B951FF68FFB07741FDB392F3BC39C89E" box="[609,616,2001,2022]" italics="true" pageId="39" pageNumber="38">t</emphasis>
), pedicel (
<emphasis id="B951FF68FFB07741FD1F92F3BC88C89E" box="[717,729,2001,2022]" italics="true" pageId="39" pageNumber="38">p</emphasis>
), distal posterior lobe (
<emphasis id="B951FF68FFB07741FC6492F2BD84C89D" box="[950,981,2000,2021]" italics="true" pageId="39" pageNumber="38">dpl</emphasis>
), lateral hook (
<emphasis id="B951FF68FFB07741FBB492F2BA28C89D" box="[1126,1145,2000,2021]" italics="true" pageId="39" pageNumber="38">lh</emphasis>
), terminal membrane of the sperm duct (
<emphasis id="B951FF68FFB07741FF7992CFBE89C77A" box="[171,216,2029,2050]" italics="true" pageId="39" pageNumber="38">tmsd</emphasis>
) and 
<emphasis id="B951FF68FFB07741FEDD92CFBFD9C77B" box="[271,392,2029,2051]" italics="true" pageId="39" pageNumber="38">basal carina</emphasis>
(
<emphasis id="B951FF68FFB07741FE4492CFBFFCC77A" box="[406,429,2029,2050]" italics="true" pageId="39" pageNumber="38">bc</emphasis>
).
</paragraph>
</caption>
<caption id="DF5A73F2FFBF774EFFA392E3BE8DC88C" ID-DOI="http://doi.org/10.5281/zenodo.5742615" ID-Zenodo-Dep="5742615" httpUri="https://zenodo.org/record/5742615/files/figure.png" pageId="40" pageNumber="39" startId="40.[113,190,1985,2007]" targetBox="[113,1474,234,1965]" targetPageId="40">
<paragraph id="8B9A237AFFBF774EFFA392E3BE8DC88C" blockId="40.[113,1471,1985,2036]" pageId="40" pageNumber="39">
<emphasis id="B951FF68FFBF774EFFA392E3BF48C8AE" bold="true" box="[113,281,1985,2007]" pageId="40" pageNumber="39">Figures 125–126</emphasis>
. Type locality of 
<taxonomicName id="4C2558F9FFBF774EFE1692E0BC36C8AE" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[452,615,1985,2007]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="lioneli" status="sp. nov.">
<emphasis id="B951FF68FFBF774EFE1692E0BC36C8AE" box="[452,615,1985,2007]" italics="true" pageId="40" pageNumber="39">Scorpiops lioneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFBF774EFDBC92E0BCF5C8AF" box="[622,676,1986,2007]" pageId="40" pageNumber="39" rank="species">
<emphasis id="B951FF68FFBF774EFDBC92E0BCD4C8AF" bold="true" box="[622,645,1986,2007]" pageId="40" pageNumber="39">sp</emphasis>
. 
<emphasis id="B951FF68FFBF774EFD4392E0BCCFC8AF" bold="true" box="[657,670,1986,2007]" pageId="40" pageNumber="39">n</emphasis>
.
</taxonomicNameLabel>
, 
<emphasis id="B951FF68FFBF774EFD6392E0BCECC8AF" bold="true" box="[689,701,1986,2007]" pageId="40" pageNumber="39">v</emphasis>
iew of mountains with semi-arid habitat and rocky out-crops at the type locality (125, 126).
</paragraph>
</caption>
<caption id="DF5A73F2FFBE774FFFA692AABD69C88C" ID-DOI="http://doi.org/10.5281/zenodo.5742617" ID-Zenodo-Dep="5742617" httpUri="https://zenodo.org/record/5742617/files/figure.png" pageId="41" pageNumber="40" startId="41.[116,193,1928,1950]" targetBox="[113,1474,245,1897]" targetPageId="41">
<paragraph id="8B9A237AFFBE774FFFA692AABD69C88C" blockId="41.[116,1474,1928,2036]" pageId="41" pageNumber="40">
<emphasis id="B951FF68FFBE774FFFA692AABF4CC8E5" bold="true" box="[116,285,1928,1950]" pageId="41" pageNumber="40">Figures 127–138</emphasis>
: 
<emphasis id="B951FF68FFBE774FFEFE92AABF84C8E5" bold="true" box="[300,469,1928,1950]" pageId="41" pageNumber="40">Figures 127–130</emphasis>
: Comparison of left pedipalps. 
<emphasis id="B951FF68FFBE774FFCDD92AABDE9C8E5" bold="true" box="[783,952,1928,1950]" pageId="41" pageNumber="40">Figures 131–134</emphasis>
: Comparison of metasomal segments II-IV in lateral view. 
<emphasis id="B951FF68FFBE774FFF7E9286BF03C8C2" bold="true" box="[172,338,1956,1978]" pageId="41" pageNumber="40">Figures 135–138</emphasis>
: Comparison of pedipalp patella dorsal view with internal aspect tubercles. 
<emphasis id="B951FF68FFBE774FFBF39286BAC6C8C2" bold="true" box="[1057,1175,1956,1978]" pageId="41" pageNumber="40">Figures 127</emphasis>
, 
<emphasis id="B951FF68FFBE774FFB709287BA97C8C2" bold="true" box="[1186,1222,1957,1978]" pageId="41" pageNumber="40">131</emphasis>
, 
<emphasis id="B951FF68FFBE774FFB039287BAA4C8C2" bold="true" box="[1233,1269,1957,1978]" pageId="41" pageNumber="40">135</emphasis>
. 
<taxonomicName id="4C2558F9FFBE774FFAD29287BB93C8C2" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[1280,1474,1957,1978]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="41" pageNumber="40" phylum="Arthropoda" rank="species" species="vrushchik" status="sp. nov.">
<emphasis id="B951FF68FFBE774FFAD29287BB93C8C2" box="[1280,1474,1957,1978]" italics="true" pageId="41" pageNumber="40">Scorpiops vrushchik</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFBE774FFFA692E0BEF9C8AF" box="[116,168,1986,2007]" pageId="41" pageNumber="40" rank="species">
<emphasis id="B951FF68FFBE774FFFA692E0BEDAC8AF" bold="true" box="[116,139,1986,2007]" pageId="41" pageNumber="40">sp</emphasis>
. 
<emphasis id="B951FF68FFBE774FFF4492E0BEF2C8AF" bold="true" box="[150,163,1986,2007]" pageId="41" pageNumber="40">n</emphasis>
.
</taxonomicNameLabel>
, male, holotype. 
<emphasis id="B951FF68FFBE774FFE9C92E3BF95C8AE" bold="true" box="[334,452,1985,2007]" pageId="41" pageNumber="40">Figures 128</emphasis>
, 
<emphasis id="B951FF68FFBE774FFE1D92E3BFA2C8AE" bold="true" box="[463,499,1985,2006]" pageId="41" pageNumber="40">132</emphasis>
, 
<emphasis id="B951FF68FFBE774FFE2C92E3BC73C8AE" bold="true" box="[510,546,1985,2006]" pageId="41" pageNumber="40">136</emphasis>
. 
<taxonomicName id="4C2558F9FFBE774FFDFF92E0BCBFC8AE" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[557,750,1985,2007]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="41" pageNumber="40" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FFBE774FFDFF92E0BCBFC8AE" box="[557,750,1985,2007]" italics="true" pageId="41" pageNumber="40">Scorpiops nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFBE774FFD2092E0BD76C8AF" box="[754,807,1986,2007]" pageId="41" pageNumber="40" rank="species">
<emphasis id="B951FF68FFBE774FFD2092E0BD58C8AF" bold="true" box="[754,777,1986,2007]" pageId="41" pageNumber="40">sp</emphasis>
. 
<emphasis id="B951FF68FFBE774FFCC692E0BD70C8AF" bold="true" box="[788,801,1986,2007]" pageId="41" pageNumber="40">n</emphasis>
.
</taxonomicNameLabel>
, male, holotype. 
<emphasis id="B951FF68FFBE774FFC1E92E3BA13C8AE" bold="true" box="[972,1090,1985,2007]" pageId="41" pageNumber="40">Figures 129</emphasis>
, 
<emphasis id="B951FF68FFBE774FFB9F92E3BA20C8AE" bold="true" box="[1101,1137,1985,2006]" pageId="41" pageNumber="40">133</emphasis>
, 
<emphasis id="B951FF68FFBE774FFBAF92E3BAF0C8AE" bold="true" box="[1149,1185,1985,2006]" pageId="41" pageNumber="40">137</emphasis>
. 
<taxonomicName id="4C2558F9FFBE774FFB7E92E0BB19C8AF" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[1196,1352,1986,2007]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="41" pageNumber="40" phylum="Arthropoda" rank="species" species="neera" status="sp. nov.">
<emphasis id="B951FF68FFBE774FFB7E92E0BB19C8AF" box="[1196,1352,1986,2007]" italics="true" pageId="41" pageNumber="40">Scorpiops neera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFBE774FFA9F92E0BBD0C8AF" box="[1357,1409,1986,2007]" pageId="41" pageNumber="40" rank="species">
<emphasis id="B951FF68FFBE774FFA9F92E0BB35C8AF" bold="true" box="[1357,1380,1986,2007]" pageId="41" pageNumber="40">sp</emphasis>
. 
<emphasis id="B951FF68FFBE774FFABD92E0BB2DC8AF" bold="true" box="[1391,1404,1986,2007]" pageId="41" pageNumber="40">n</emphasis>
.
</taxonomicNameLabel>
, male, holotype. 
<emphasis id="B951FF68FFBE774FFF0692FCBF1AC88B" bold="true" box="[212,331,2014,2036]" pageId="41" pageNumber="40">Figures 130</emphasis>
, 
<emphasis id="B951FF68FFBE774FFE8592FCBF2AC88B" bold="true" box="[343,379,2014,2035]" pageId="41" pageNumber="40">134</emphasis>
, 
<emphasis id="B951FF68FFBE774FFE5592FCBFFAC88B" bold="true" box="[391,427,2014,2035]" pageId="41" pageNumber="40">138</emphasis>
. 
<taxonomicName id="4C2558F9FFBE774FFE6592FDBC0BC88B" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[439,602,2014,2036]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="41" pageNumber="40" phylum="Arthropoda" rank="species" species="lioneli" status="sp. nov.">
<emphasis id="B951FF68FFBE774FFE6592FDBC0BC88B" box="[439,602,2014,2036]" italics="true" pageId="41" pageNumber="40">Scorpiops lioneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFBE774FFDB292FCBCC4C88C" box="[608,661,2014,2036]" pageId="41" pageNumber="40" rank="species">
<emphasis id="B951FF68FFBE774FFDB292FCBC26C88B" bold="true" box="[608,631,2014,2035]" pageId="41" pageNumber="40">sp</emphasis>
. 
<emphasis id="B951FF68FFBE774FFD5192FCBCC1C88B" bold="true" box="[643,656,2014,2035]" pageId="41" pageNumber="40">n</emphasis>
.
</taxonomicNameLabel>
, male, holotype.
</paragraph>
</caption>
<paragraph id="8B9A237AFFBD774CFFA39449BCFECD9A" blockId="42.[113,771,363,931]" pageId="42" pageNumber="41">
AFFINITIES. 
<taxonomicName id="4C2558F9FFBD774CFF269449BFFFCEFB" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[244,430,363,387]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="lioneli" status="sp. nov.">
<emphasis id="B951FF68FFBD774CFF269449BFFFCEFB" box="[244,430,363,387]" italics="true" pageId="42" pageNumber="41">Scorpiops lioneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFBD774CFE689449BFAACEFA" box="[442,507,363,387]" pageId="42" pageNumber="41" rank="species">
<emphasis id="B951FF68FFBD774CFE689449BF82CEFB" bold="true" box="[442,467,363,387]" pageId="42" pageNumber="41">sp</emphasis>
. 
<emphasis id="B951FF68FFBD774CFE379449BFA5CEFB" bold="true" box="[485,500,363,387]" pageId="42" pageNumber="41">n</emphasis>
.
</taxonomicNameLabel>
is closely related to 
<taxonomicName id="4C2558F9FFBD774CFD3D9449BE87CEDB" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FFBD774CFD3D9449BE87CEDB" italics="true" pageId="42" pageNumber="41">S. nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFBD774CFF3294A9BF4ECEDA" box="[224,287,395,419]" pageId="42" pageNumber="41" rank="species">
<emphasis id="B951FF68FFBD774CFF3294A9BEA8CEDB" bold="true" box="[224,249,395,419]" pageId="42" pageNumber="41">sp</emphasis>
. 
<emphasis id="B951FF68FFBD774CFEDB94A9BF49CEDB" bold="true" box="[265,280,395,419]" pageId="42" pageNumber="41">n</emphasis>
.
</taxonomicNameLabel>
and 
<emphasis id="B951FF68FFBD774CFE8A94A9BC6DCEDB" box="[344,572,395,419]" italics="true" pageId="42" pageNumber="41">
<taxonomicName id="4C2558F9FFBD774CFE8A94A9BC68CEDB" box="[344,569,395,419]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="maharashtraensis">S. maharashtraensis</taxonomicName>
.
</emphasis>
It is separated by a raw genetic divergence of 6.7% from 
<taxonomicName id="4C2558F9FFBD774CFDE69489BCE9CEBB" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[564,696,427,451]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FFBD774CFDE69489BCE9CEBB" box="[564,696,427,451]" italics="true" pageId="42" pageNumber="41">S. nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFBD774CFD119489BD52CEBA" box="[707,771,427,451]" pageId="42" pageNumber="41" rank="species">
<emphasis id="B951FF68FFBD774CFD119489BC8DCEBB" bold="true" box="[707,732,427,451]" pageId="42" pageNumber="41">sp</emphasis>
. 
<emphasis id="B951FF68FFBD774CFD3F9489BCADCEBB" bold="true" box="[749,764,427,451]" pageId="42" pageNumber="41">n</emphasis>
.
</taxonomicNameLabel>
and 11% from 
<emphasis id="B951FF68FFBD774CFEF294E9BC57CE9B" box="[288,518,459,483]" italics="true" pageId="42" pageNumber="41">
<taxonomicName id="4C2558F9FFBD774CFEF294E9BC53CE9B" box="[288,514,459,483]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="maharashtraensis">S. maharashtraensis</taxonomicName>
.
</emphasis>
The new species also differs from its close congeners based on following set of morphological characters: Trichobothria on patella ventral 15–18 as opposed to 
<quantity id="4CDD8E9FFFBD774CFEBF9709BF83CD3B" box="[365,466,555,579]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="3.556" metricValueMin="3.048" pageId="42" pageNumber="41" unit="in" value="13.0" valueMax="14.0" valueMin="12.0">12–14 in</quantity>
<taxonomicName id="4C2558F9FFBD774CFE329709BC37CD3B" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[480,614,555,579]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FFBD774CFE329709BC37CD3B" box="[480,614,555,579]" italics="true" pageId="42" pageNumber="41">S. nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFBD774CFDA19709BCE4CD3A" box="[627,693,555,579]" pageId="42" pageNumber="41" rank="species">
<emphasis id="B951FF68FFBD774CFDA19709BCDDCD3B" bold="true" box="[627,652,555,579]" pageId="42" pageNumber="41">sp</emphasis>
. 
<emphasis id="B951FF68FFBD774CFD729709BCFECD3B" bold="true" box="[672,687,555,579]" pageId="42" pageNumber="41">n</emphasis>
.
</taxonomicNameLabel>
; chela length to width ratio in males 4.4 as opposed to 
<quantity id="4CDD8E9FFFBD774CFDA99769BCB9CD1B" box="[635,744,587,611]" metricMagnitude="-2" metricUnit="m" metricValue="9.906" metricValueMax="10.414" metricValueMin="9.398" pageId="42" pageNumber="41" unit="in" value="3.9" valueMax="4.1" valueMin="3.7">3.7–4.1 in</quantity>
<taxonomicName id="4C2558F9FFBD774CFD3D9769BE87CDFB" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FFBD774CFD3D9769BE87CDFB" italics="true" pageId="42" pageNumber="41">S. nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFBD774CFF0F9749BF49CDFA" box="[221,280,619,643]" pageId="42" pageNumber="41" rank="species">
<emphasis id="B951FF68FFBD774CFF0F9749BEA7CDFB" bold="true" box="[221,246,619,643]" pageId="42" pageNumber="41">sp</emphasis>
. 
<emphasis id="B951FF68FFBD774CFED19749BF43CDFB" bold="true" box="[259,274,619,643]" pageId="42" pageNumber="41">n</emphasis>
.
</taxonomicNameLabel>
; body coloration yellowish brown as opposed to blackish brown in 
<taxonomicName id="4C2558F9FFBD774CFE8F97A9BFB1CDDB" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[349,480,651,675]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FFBD774CFE8F97A9BFB1CDDB" box="[349,480,651,675]" italics="true" pageId="42" pageNumber="41">S. nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFBD774CFE3B97A9BC70CDDB" box="[489,545,651,675]" pageId="42" pageNumber="41" rank="species">
<emphasis id="B951FF68FFBD774CFE3B97A9BC53CDDB" bold="true" box="[489,514,651,675]" pageId="42" pageNumber="41">sp</emphasis>
. 
<emphasis id="B951FF68FFBD774CFDC097A9BC70CDDB" bold="true" box="[530,545,651,675]" pageId="42" pageNumber="41">n</emphasis>
</taxonomicNameLabel>
; carapace with deep U shaped anterior median notch as opposed to moderate U shaped anterior median notch in 
<taxonomicName id="4C2558F9FFBD774CFE1C97E9BCF9CD9B" box="[462,680,715,739]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="maharashtraensis">
<emphasis id="B951FF68FFBD774CFE1C97E9BCF9CD9B" box="[462,680,715,739]" italics="true" pageId="42" pageNumber="41">S. maharashtraensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9A237AFFBD774CFF7397C9BEA8CCDB" blockId="42.[113,771,363,931]" pageId="42" pageNumber="41">
It differs from all the remaining species of 
<taxonomicName id="4C2558F9FFBD774CFD4A97C9BD52CC7A" authorityName="Peters" authorityYear="1861" box="[664,771,747,770]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="42" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B951FF68FFBD774CFD4A97C9BD52CC7A" box="[664,771,747,770]" italics="true" pageId="42" pageNumber="41">Scorpiops</emphasis>
</taxonomicName>
from the northern Western Ghats and northern 
<collectingRegion id="49E1ED98FFBD774CFDAC9629BD52CC5B" box="[638,771,779,803]" country="India" name="Maharashtra" pageId="42" pageNumber="41">Maharashtra</collectingRegion>
(
<collectingCountry id="F33263EAFFBD774CFFAB9609BE99CC3B" box="[121,200,811,835]" name="India" pageId="42" pageNumber="41">INDIA</collectingCountry>
) by a raw genetic divergence of 9.4–12.4 % (
<tableCitation id="C6A716C1FFBD774CFD769609BCA4CC3A" box="[676,757,810,835]" captionStart="Table 5" captionStartId="44.[115,171,1066,1087]" captionText="Table 5. Pairwise uncorrected raw distances (%) expressed as minimum–maximum based on COI gene sequence for Scorpiops species from Maharashtra, India. Values in brackets are intra-clade distances." pageId="42" pageNumber="41">Table 5</tableCitation>
). It is also distinguished from all the species of 
<taxonomicName id="4C2558F9FFBD774CFD8C9669BC98CC1A" authorityName="Peters" authorityYear="1861" box="[606,713,843,866]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="42" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B951FF68FFBD774CFD8C9669BC98CC1A" box="[606,713,843,866]" italics="true" pageId="42" pageNumber="41">Scorpiops</emphasis>
</taxonomicName>
from peninsular India based on the key of morphological characters (See below).
</paragraph>
</subSubSection>
<subSubSection id="C33F70F1FFBD774CFFA396E9BC2DC81B" pageId="42" pageNumber="41" type="discussion">
<paragraph id="8B9A237AFFBD774CFFA396E9BC2DC81B" blockId="42.[113,772,971,1891]" pageId="42" pageNumber="41">
<materialsCitation id="3B4D2927FFBD774CFFA396E9BC29C81B" collectionCode="BNHS" country="India" county="Pardhadi Village" elevation="586" latitude="20.279722" location="Shidi Ghat'" longLatPrecision="21" longitude="74.8275" municipality="Nashik District" pageId="42" pageNumber="41" specimenCode="SC-62, SC-63, 64, SC-65, SC-66" specimenCount="6" specimenCount-female="3" specimenCount-male="2" stateProvince="Maharashtra" typeStatus="holotype">
COMMENTS. 
<taxonomicName id="4C2558F9FFBD774CFF2196E9BF9BCC9B" box="[243,458,971,995]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="maharashtraensis">
<emphasis id="B951FF68FFBD774CFF2196E9BF9BCC9B" box="[243,458,971,995]" italics="true" pageId="42" pageNumber="41">S. maharashtraensis</emphasis>
</taxonomicName>
, 
<typeStatus id="549E9DD8FFBD774CFE0B96E9BC66CC9B" box="[473,567,971,995]" pageId="42" pageNumber="41" type="holotype">holotype</typeStatus>
, 
<specimenCount id="9D23E8F3FFBD774CFD9696E9BC09CC9A" box="[580,600,971,994]" pageId="42" pageNumber="41" type="male">♂</specimenCount>
(
<collectionCode id="ED34BBBFFFBD774CFDB596E9BCFECC9B" box="[615,687,971,995]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="42" pageNumber="41">BNHS</collectionCode>
<specimenCode id="DB838B01FFBD774CFD6596E9BCADCC9B" box="[695,764,971,995]" collectionCode="SC-" pageId="42" pageNumber="41">SC-62</specimenCode>
) is described from the type locality ‘‘ 
<location id="8EFA75A1FFBD774CFE2796C9BC26CB7B" LSID="urn:lsid:plazi:treatment:038C926CFFB6774CFC3D9399BA8DCEBA:8EFA75A1FFBD774CFE2796C9BC26CB7B" box="[501,631,1003,1027]" country="India" county="Pardhadi Village" latitude="20.279722" longLatPrecision="21" longitude="74.8275" municipality="Nashik District" name="Shidi Ghat'" pageId="42" pageNumber="41" stateProvince="Maharashtra">Shidi Ghat’</location>
’ near 
<location id="8EFA75A1FFBD774CFD6896C9BEEECB5B" LSID="urn:lsid:plazi:treatment:038C926CFFB6774CFC3D9399BA8DCEBA:8EFA75A1FFBD774CFD6896C9BEEECB5B" country="India" county="Pardhadi Village" latitude="20.279722" longLatPrecision="21" longitude="74.8275" municipality="Nashik District" name="Wadali Village" pageId="42" pageNumber="41" stateProvince="Maharashtra">Wadali Village</location>
, 
<location id="8EFA75A1FFBD774CFF1B9129BF03CB5B" LSID="urn:lsid:plazi:treatment:038C926CFFB6774CFC3D9399BA8DCEBA:8EFA75A1FFBD774CFF1B9129BF03CB5B" box="[201,338,1035,1059]" country="India" county="Pardhadi Village" latitude="20.279722" longLatPrecision="21" longitude="74.8275" municipality="Nashik District" name="Tehsil-Sillod" pageId="42" pageNumber="41" stateProvince="Maharashtra">Tehsil-Sillod</location>
, 
<location id="8EFA75A1FFBD774CFE899129BC62CB5B" LSID="urn:lsid:plazi:treatment:038C926CFFB6774CFC3D9399BA8DCEBA:8EFA75A1FFBD774CFE899129BC62CB5B" box="[347,563,1035,1059]" country="India" county="Pardhadi Village" latitude="20.279722" longLatPrecision="21" longitude="74.8275" municipality="Nashik District" name="Aurangabad District" pageId="42" pageNumber="41" stateProvince="Maharashtra">Aurangabad District</location>
, 
<collectingRegion id="49E1ED98FFBD774CFDEC9129BC92CB5B" box="[574,707,1035,1059]" country="India" name="Maharashtra" pageId="42" pageNumber="41">Maharashtra</collectingRegion>
, 
<collectingCountry id="F33263EAFFBD774CFD1F9129BD52CB5B" box="[717,771,1035,1059]" name="India" pageId="42" pageNumber="41">India</collectingCountry>
(
<geoCoordinate id="EE1145BDFFBD774CFFAA9108BEA9CB3A" box="[120,248,1066,1090]" degrees="20.5321808" direction="north" orientation="latitude" pageId="42" pageNumber="41" precision="1" value="20.53218">20.5321808</geoCoordinate>
, 
<geoCoordinate id="EE1145BDFFBD774CFEDF9108BF2ECB3A" box="[269,383,1066,1090]" degrees="75.879628" direction="east" orientation="longitude" pageId="42" pageNumber="41" precision="1" value="75.87963">75.879628</geoCoordinate>
, 
<elevation id="0008C449FFBD774CFE409109BC00CB3B" box="[402,593,1066,1091]" metricMagnitude="2" metricUnit="m" metricValue="5.86" pageId="42" pageNumber="41" unit="m" value="586.0">
elevation 
<quantity id="4CDD8E9FFFBD774CFDD09108BC00CB3B" box="[514,593,1066,1091]" metricMagnitude="2" metricUnit="m" metricValue="5.86" pageId="42" pageNumber="41" unit="m" value="586.0">586 m</quantity>
</elevation>
). 
<typeStatus id="549E9DD8FFBD774CFDBF9109BC82CB3B" box="[621,723,1067,1091]" pageId="42" pageNumber="41" type="paratype">Paratypes</typeStatus>
<specimenCount id="9D23E8F3FFBD774CFD309109BD52CB3B" box="[738,771,1067,1091]" pageId="42" pageNumber="41" type="female">2♀</specimenCount>
(
<collectionCode id="ED34BBBFFFBD774CFFAB9169BE90CB1B" box="[121,193,1099,1123]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="42" pageNumber="41">BNHS</collectionCode>
<specimenCode id="DB838B01FFBD774CFF189169BF5ECB1B" box="[202,271,1099,1123]" collectionCode="SC-" pageId="42" pageNumber="41">SC-63</specimenCode>
, 
<specimenCode id="DB838B01FFBD774CFECC9169BF68CB1B" box="[286,313,1099,1123]" collectionCode="SC-" pageId="42" pageNumber="41">64</specimenCode>
) are also collected from the type locality. 
<specimenCount id="9D23E8F3FFBD774CFFA39149BEC2CBFB" box="[113,147,1131,1155]" pageId="42" pageNumber="41" type="male">1♂</specimenCount>
(
<collectionCode id="ED34BBBFFFBD774CFF7A9149BEA1CBFB" box="[168,240,1131,1155]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="42" pageNumber="41">BNHS</collectionCode>
<specimenCode id="DB838B01FFBD774CFF2D9148BF12CBFA" box="[255,323,1130,1154]" collectionCode="SC-" pageId="42" pageNumber="41">SC-65</specimenCode>
) and 
<specimenCount id="9D23E8F3FFBD774CFE5C9149BFFECBFB" box="[398,431,1131,1155]" pageId="42" pageNumber="41" type="female">1♀</specimenCount>
(
<collectionCode id="ED34BBBFFFBD774CFE179149BC5CCBFB" box="[453,525,1131,1155]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="42" pageNumber="41">BNHS</collectionCode>
<specimenCode id="DB838B01FFBD774CFDCE9149BC31CBFB" box="[540,608,1131,1155]" collectionCode="SC-" pageId="42" pageNumber="41">SC-66</specimenCode>
) are collected from Chalisgaon Taluka, near Ghodegaon, 
<location id="8EFA75A1FFBD774CFD8091A9BCAECBDB" LSID="urn:lsid:plazi:treatment:038C926CFFB6774CFC3D9399BA8DCEBA:8EFA75A1FFBD774CFD8091A9BCAECBDB" box="[594,767,1163,1187]" country="India" county="Pardhadi Village" latitude="20.279722" longLatPrecision="21" longitude="74.8275" municipality="Nashik District" name="Jalgaon District" pageId="42" pageNumber="41" stateProvince="Maharashtra">Jalgaon District</location>
, 
<collectingRegion id="49E1ED98FFBD774CFFA39189BEA5CBBB" box="[113,244,1195,1219]" country="India" name="Maharashtra" pageId="42" pageNumber="41">Maharashtra</collectingRegion>
(
<geoCoordinate id="EE1145BDFFBD774CFED39189BFD0CBBB" box="[257,385,1195,1219]" degrees="20.3127998" direction="north" orientation="latitude" pageId="42" pageNumber="41" precision="1" value="20.3128">20.3127998</geoCoordinate>
, 
<geoCoordinate id="EE1145BDFFBD774CFE599189BC5BCBBB" box="[395,522,1195,1219]" degrees="74.8622418" direction="east" orientation="longitude" pageId="42" pageNumber="41" precision="1" value="74.86224">74.8622418</geoCoordinate>
, 
<elevation id="0008C449FFBD774CFDC19189BCEFCBBB" box="[531,702,1195,1219]" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="42" pageNumber="41" unit="m" value="460.0">
elevation 
<quantity id="4CDD8E9FFFBD774CFDAB9189BCEFCBBB" box="[633,702,1195,1219]" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="42" pageNumber="41" unit="m" value="460.0">460 m</quantity>
</elevation>
). Type locality of 
<taxonomicName id="4C2558F9FFBD774CFF3991E9BFF0CB9B" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[235,417,1227,1251]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="lioneli" status="sp. nov.">
<emphasis id="B951FF68FFBD774CFF3991E9BFF0CB9B" box="[235,417,1227,1251]" italics="true" pageId="42" pageNumber="41">Scorpiops lioneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFBD774CFE7991E9BFBBCB9A" box="[427,490,1227,1251]" pageId="42" pageNumber="41" rank="species">
<emphasis id="B951FF68FFBD774CFE7991E9BF95CB9B" bold="true" box="[427,452,1227,1251]" pageId="42" pageNumber="41">sp</emphasis>
. 
<emphasis id="B951FF68FFBD774CFE0791E9BFB5CB9B" bold="true" box="[469,484,1227,1251]" pageId="42" pageNumber="41">n</emphasis>
.
</taxonomicNameLabel>
<collectingCountry id="F33263EAFFBD774CFE2791E9BC7CCB9B" box="[501,557,1227,1251]" name="India" pageId="42" pageNumber="41">India</collectingCountry>
, 
<collectingRegion id="49E1ED98FFBD774CFDE991E9BCAECB9B" box="[571,767,1227,1251]" country="India" name="Maharashtra" pageId="42" pageNumber="41">Maharashtra State</collectingRegion>
, 
<collectingMunicipality id="6BFEB900FFBD774CFFA391C9BF47CA7B" box="[113,278,1259,1283]" pageId="42" pageNumber="41">Nashik District</collectingMunicipality>
, 
<collectingCounty id="62FB5BF6FFBD774CFEF091C9BF87CA7B" box="[290,470,1259,1283]" pageId="42" pageNumber="41">Pardhadi Village</collectingCounty>
, 
<geoCoordinate id="EE1145BDFFBD774CFE3091C9BC2DCA7A" box="[482,636,1259,1283]" degrees="20" direction="north" minutes="16" orientation="latitude" pageId="42" pageNumber="41" precision="15" seconds="47" value="20.279722">20° 16’ 47” N</geoCoordinate>
<geoCoordinate id="EE1145BDFFBD774CFD5791C9BED2CA5A" degrees="74" direction="east" minutes="49" orientation="longitude" pageId="42" pageNumber="41" precision="15" seconds="39" value="74.8275">74° 49’ 39” E</geoCoordinate>
, 
<elevation id="0008C449FFBD774CFF429028BF4DCA5B" box="[144,284,1290,1315]" metricMagnitude="2" metricUnit="m" metricValue="5.59" pageId="42" pageNumber="41" unit="m" value="559.0">
<quantity id="4CDD8E9FFFBD774CFF429028BE85CA5A" box="[144,212,1290,1314]" metricMagnitude="2" metricUnit="m" metricValue="5.59" pageId="42" pageNumber="41" unit="m" value="559.0">559 m</quantity>
a. s. l.
</elevation>
(locality data is same for all type material) is ca 10 kms from the locality of 
<taxonomicName id="4C2558F9FFBD774CFE7D9009BCD5CA3B" box="[431,644,1323,1347]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="maharashtraensis">
<emphasis id="B951FF68FFBD774CFE7D9009BCD5CA3B" box="[431,644,1323,1347]" italics="true" pageId="42" pageNumber="41">S. maharashtraensis</emphasis>
</taxonomicName>
from where the 
<typeStatus id="549E9DD8FFBD774CFF499069BEAECA1B" box="[155,255,1355,1379]" pageId="42" pageNumber="41" type="paratype">paratypes</typeStatus>
<collectionCode id="ED34BBBFFFBD774CFED89069BF01CA1B" box="[266,336,1355,1379]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="42" pageNumber="41">BNHS</collectionCode>
<specimenCode id="DB838B01FFBD774CFE889068BFCCCA1A" box="[346,413,1354,1378]" collectionCode="SC-" pageId="42" pageNumber="41">SC-65</specimenCode>
and 
<collectionCode id="ED34BBBFFFBD774CFE059069BC4CCA1B" box="[471,541,1355,1379]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="42" pageNumber="41">BNHS</collectionCode>
<specimenCode id="DB838B01FFBD774CFDF59069BC3BCA1B" box="[551,618,1355,1379]" collectionCode="SC-" pageId="42" pageNumber="41">SC-66</specimenCode>
are collected. Our surveys did not yield any success from this locality. Type specimens of 
<taxonomicName id="4C2558F9FFBD774CFEC290A9BFBCCADB" box="[272,493,1419,1443]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="maharashtraensis">
<emphasis id="B951FF68FFBD774CFEC290A9BFBCCADB" box="[272,493,1419,1443]" italics="true" pageId="42" pageNumber="41">S. maharashtraensis</emphasis>
</taxonomicName>
are deposited at 
<collectionCode id="ED34BBBFFFBD774CFD6F90A9BD55CADB" box="[701,772,1419,1443]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="42" pageNumber="41">BNHS</collectionCode>
without accession numbers assigned to individual specimens and preserved in single preservation jar with more specimens apart from the type series. Sulakhe et al. (2020a) were able to identify only the 
<typeStatus id="549E9DD8FFBD774CFEF29329BF2FC95B" box="[288,382,1547,1571]" pageId="42" pageNumber="41" type="holotype">holotype</typeStatus>
, 
<collectionCode id="ED34BBBFFFBD774CFE559329BF9CC95B" box="[391,461,1547,1571]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="42" pageNumber="41">BNHS</collectionCode>
<specimenCode id="DB838B01FFBD774CFE009329BC44C95B" box="[466,533,1547,1571]" collectionCode="SC-" pageId="42" pageNumber="41">SC-62</specimenCode>
and 
<specimenCount id="9D23E8F3FFBD774CFD979329BCE5C95B" box="[581,692,1547,1571]" pageId="42" pageNumber="41" type="generic" typeStatus="paratype">1 paratype</specimenCount>
, 
<collectionCode id="ED34BBBFFFBD774CFD6F9329BD55C95B" box="[701,772,1547,1571]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="42" pageNumber="41">BNHS</collectionCode>
<specimenCode id="DB838B01FFBD774CFFA39309BEE4C93B" box="[113,181,1579,1603]" collectionCode="SC-" pageId="42" pageNumber="41">SC-63</specimenCode>
collected from the type locality based on the photographs published by 
<bibRefCitation id="EFB45E8BFFBD774CFED69369BF85C91B" author="MIRZA" box="[260,468,1611,1635]" pageId="42" pageNumber="41" pagination="143 - 149" refId="ref19790" refString="MIRZA, Z. A., R. SANAP &amp; R. UPADHYE. 2014. A new species of scorpion of the genus Neoscorpiops Vachon, 1980 (Scorpiones: Euscorpiidae) from India. Comptes Rendus Biologies, 337 (2): 143 - 149." type="journal article" year="2014">Mirza et al. (2014)</bibRefCitation>
. The deteriorating condition of the preserved specimens, ambiguity in assigning accession numbers to individual specimens (type material is collected from one more locality apart from the type locality) and inability of extracting DNA due to preservation in 70% ethanol, the status of 
<typeStatus id="549E9DD8FFBD774CFF2193C9BF06C87B" box="[243,343,1771,1795]" pageId="42" pageNumber="41" type="paratype">paratypes</typeStatus>
<collectionCode id="ED34BBBFFFBD774CFE8E93C9BFF3C87B" box="[348,418,1771,1795]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="42" pageNumber="41">BNHS</collectionCode>
<specimenCode id="DB838B01FFBD774CFE7A93C8BFBAC87A" box="[424,491,1770,1794]" collectionCode="SC-" pageId="42" pageNumber="41">SC-65</specimenCode>
and 66 could not be tested. Considering the geographical proximity these specimens could be 
<taxonomicName id="4C2558F9FFBD774CFF469209BEA2C83B" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[148,243,1835,1859]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="lioneli" status="sp. nov.">
<emphasis id="B951FF68FFBD774CFF469209BEA2C83B" box="[148,243,1835,1859]" italics="true" pageId="42" pageNumber="41">S. lioneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFBD774CFF2E9209BF68C83A" box="[252,313,1835,1859]" pageId="42" pageNumber="41" rank="species">
<emphasis id="B951FF68FFBD774CFF2E9209BF44C83B" bold="true" box="[252,277,1835,1859]" pageId="42" pageNumber="41">sp</emphasis>
. 
<emphasis id="B951FF68FFBD774CFEF79209BF65C83B" bold="true" box="[293,308,1835,1859]" pageId="42" pageNumber="41">n</emphasis>
.
</taxonomicNameLabel>
, however this needs to be confirmed with more sampling and genetic data from this locality
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F70F1FFBD774CFFA392A6BA8DCEBA" pageId="42" pageNumber="41" type="distribution">
<paragraph id="8B9A237AFFBD774CFFA392A6BA8DCEBA" blockId="42.[113,771,1924,2044]" lastBlockId="42.[816,1474,235,451]" pageId="42" pageNumber="41">
<materialsCitation id="3B4D2927FFBD774CFFA392A6BA89CEBA" collectionCode="AND" country="India" county="Locations" location="Presently" municipality="Any" pageId="42" pageNumber="41" specimenCount="4" stateProvince="Maharashtra">
DISTRIBUTION, HABITAT 
<collectionCode id="ED34BBBFFFBD774CFEA892A6BFF6C8E4" box="[378,423,1924,1948]" pageId="42" pageNumber="41">AND</collectionCode>
ECOLOGY. 
<location id="8EFA75A1FFBD774CFDF592A6BCDBC8E4" LSID="urn:lsid:plazi:treatment:038C926CFFB6774CFC3D9399BA8DCEBA:8EFA75A1FFBD774CFDF592A6BCDBC8E4" box="[551,650,1924,1948]" country="India" county="Locations" municipality="Any" name="Presently" pageId="42" pageNumber="41" stateProvince="Maharashtra">Presently</location>
<taxonomicName id="4C2558F9FFBD774CFD4A92A7BEE4C8C4" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="lioneli" status="sp. nov.">
<emphasis id="B951FF68FFBD774CFD4A92A7BEE4C8C4" italics="true" pageId="42" pageNumber="41">Scorpiops lioneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FFBD774CFF6D9286BEAFC8C4" box="[191,254,1956,1980]" pageId="42" pageNumber="41" rank="species">
<emphasis id="B951FF68FFBD774CFF6D9286BE89C8C4" bold="true" box="[191,216,1956,1980]" pageId="42" pageNumber="41">sp</emphasis>
. 
<emphasis id="B951FF68FFBD774CFF3B9286BEA9C8C4" bold="true" box="[233,248,1956,1980]" pageId="42" pageNumber="41">n</emphasis>
.
</taxonomicNameLabel>
is known only from its 
<typeStatus id="549E9DD8FFBD774CFDDC9287BC6DC8C5" box="[526,572,1957,1981]" pageId="42" pageNumber="41">type</typeStatus>
locality, 
<location id="8EFA75A1FFBD774CFD769286BE90C8A4" LSID="urn:lsid:plazi:treatment:038C926CFFB6774CFC3D9399BA8DCEBA:8EFA75A1FFBD774CFD769286BE90C8A4" country="India" county="Locations" municipality="Any" name="Pardhadi Village" pageId="42" pageNumber="41" stateProvince="Maharashtra">Pardhadi Village</location>
, 
<location id="8EFA75A1FFBD774CFF0892E6BFDCC8A4" LSID="urn:lsid:plazi:treatment:038C926CFFB6774CFC3D9399BA8DCEBA:8EFA75A1FFBD774CFF0892E6BFDCC8A4" box="[218,397,1988,2012]" country="India" county="Locations" municipality="Any" name="Nashik District" pageId="42" pageNumber="41" stateProvince="Maharashtra">Nashik District</location>
, 
<collectingRegion id="49E1ED98FFBD774CFE7792E6BC7CC8A4" box="[421,557,1988,2012]" country="India" name="Maharashtra" pageId="42" pageNumber="41">Maharashtra</collectingRegion>
, 
<collectingCountry id="F33263EAFFBD774CFD9792E6BC2EC8A4" box="[581,639,1988,2012]" name="India" pageId="42" pageNumber="41">India</collectingCountry>
. 
<location id="8EFA75A1FFBD774CFD4592E6BC91C8A4" LSID="urn:lsid:plazi:treatment:038C926CFFB6774CFC3D9399BA8DCEBA:8EFA75A1FFBD774CFD4592E6BC91C8A4" box="[663,704,1988,2012]" country="India" county="Locations" municipality="Any" name="The" pageId="42" pageNumber="41" stateProvince="Maharashtra">The</location>
<typeStatus id="549E9DD8FFBD774CFD0792E7BD52C8A5" box="[725,771,1989,2013]" pageId="42" pageNumber="41">type</typeStatus>
series was collected from rock boulders on a hill side of the road leading from 
<location id="8EFA75A1FFBD774CFBD795C9BAEDCE7B" LSID="urn:lsid:plazi:treatment:038C926CFFB6774CFC3D9399BA8DCEBA:8EFA75A1FFBD774CFBD795C9BAEDCE7B" box="[1029,1212,235,259]" country="India" county="Locations" municipality="Any" name="Pardhadi Village" pageId="42" pageNumber="41" stateProvince="Maharashtra">Pardhadi Village</location>
to 
<location id="8EFA75A1FFBD774CFB3A95C9BBDBCE7B" LSID="urn:lsid:plazi:treatment:038C926CFFB6774CFC3D9399BA8DCEBA:8EFA75A1FFBD774CFB3A95C9BBDBCE7B" box="[1256,1418,235,259]" country="India" county="Locations" municipality="Any" name="Dheku Village" pageId="42" pageNumber="41" stateProvince="Maharashtra">Dheku Village</location>
. The 
<typeStatus id="549E9DD8FFBD774CFCE29429BD0FCE5B" box="[816,862,267,291]" pageId="42" pageNumber="41">type</typeStatus>
locality is surrounded by scrub forest in deep valleys interspersed with semi-arid grasslands. 
<collectingMunicipality id="6BFEB900FFBD774CFB299409BB78CE3B" box="[1275,1321,299,323]" pageId="42" pageNumber="41">Any</collectingMunicipality>
further road widening work may be a direct threat to the species. 
<collectingCounty id="62FB5BF6FFBD774CFA8B9469BB93CE1B" box="[1369,1474,331,355]" pageId="42" pageNumber="41">Locations</collectingCounty>
around the 
<typeStatus id="549E9DD8FFBD774CFC7C9449BD8DCEFB" box="[942,988,363,387]" pageId="42" pageNumber="41">type</typeStatus>
locality in the deep valleys may have more populations of this species and need to be confirmed with dedicated surveys (
<figureCitation id="131E3FFFFFBD774CFC2F9489BAC8CEBA" box="[1021,1177,426,451]" captionStart="Figures 125–126" captionStartId="40.[113,190,1985,2007]" captionTargetBox="[113,1474,234,1965]" captionTargetId="figure-16@40.[110,1474,234,1965]" captionTargetPageId="40" captionText="Figures 125–126. Type locality of Scorpiops lioneli sp. n., view of mountains with semi-arid habitat and rocky out-crops at the type locality (125, 126)." figureDoi="http://doi.org/10.5281/zenodo.5742615" httpUri="https://zenodo.org/record/5742615/files/figure.png" pageId="42" pageNumber="41">Figs. 125–126</figureCitation>
, 
<figureCitation id="131E3FFFFFBD774CFB769488BA9ECEBA" box="[1188,1231,426,450]" captionStart="Figure 153" captionStartId="45.[113,181,1216,1238]" captionTargetBox="[114,1474,234,1198]" captionTargetId="figure-7@45.[113,1474,234,1198]" captionTargetPageId="45" captionText="Figure 153. Distribution of Scorpiops species from Peninsular India. Localities with a star indicate new species described in this study." figureDoi="http://doi.org/10.5281/zenodo.5742623" httpUri="https://zenodo.org/record/5742623/files/figure.png" pageId="42" pageNumber="41">153</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>