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<document id="A62A74B499A0A84E3D0E15EC699C5A58" ID-CLB-Dataset="29293" ID-DOI="10.11646/zootaxa.4451.1.1" ID-GBIF-Dataset="62552028-dd96-4a85-bd83-5e86bd957be4" ID-ISSN="1175-5326" ID-Zenodo-Dep="1445080" ID-ZooBank="5DCE18B3-9DBA-48F5-AEF7-90A50B984EA5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538635919757" checkinUser="plazi" docAuthor="Bosmans, Robert, Kherbouche-Abrous, Ourida, Benhalima, Souâd &amp; Hervé, Christophe" docDate="2018" docId="DE5087A6C010B330FF39708CFD5FFBE5" docLanguage="en" docName="zootaxa.4451.1.1.pdf" docOrigin="Zootaxa 4451 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Haplodrassus Chamberlin 1922" docType="treatment" docVersion="8" lastPageNumber="7" masterDocId="2269FFDEC015B336FFAE7052FFCFFFF5" masterDocTitle="The genus Haplodrassus Chamberlin, 1922 in the Mediterranean and the Maghreb in particular (Araneae: Gnaphosidae)" masterLastPageNumber="67" masterPageNumber="1" pageNumber="6" updateTime="1698564719883" updateUser="ExternalLinkService">
<mods:mods id="D148D411481DBF31E58B436F8BCA4576" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="D784B6060664B80DE6CFEB1DEC7D2CEC">The genus Haplodrassus Chamberlin, 1922 in the Mediterranean and the Maghreb in particular (Araneae: Gnaphosidae)</mods:title>
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<mods:namePart id="3D8F10FC1CA0C4AE7F60B9FDEA6531CE">Bosmans, Robert</mods:namePart>
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<mods:namePart id="A73A2024EE4C46E190904F4EEC4DB828">Kherbouche-Abrous, Ourida</mods:namePart>
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<mods:namePart id="459A8389391B7934CDC4983A35BFFF57">Benhalima, Souâd</mods:namePart>
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<mods:namePart id="1E9EEA0E137E8EA625B35509126C7E79">Hervé, Christophe</mods:namePart>
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<mods:title id="347B158D3B17B899B670FD6152411780">Zootaxa</mods:title>
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<mods:date id="DB9BF79E6EC50FAA7DF2C23B0EF53C6B">2018</mods:date>
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<mods:number id="AA1397FB283C0DC78E74ED87DA863B44">2018-07-30</mods:number>
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<mods:number id="C3F4E9672755220AF5C1AC8D10525591">4451</mods:number>
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<mods:classification id="A8E377CDB1C5A006216A9C0EBDDC2047">journal article</mods:classification>
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<mods:identifier id="4232466C807C38E872AFCB444B425795" type="DOI">10.11646/zootaxa.4451.1.1</mods:identifier>
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<treatment id="DE5087A6C010B330FF39708CFD5FFBE5" ID-DOI="http://doi.org/10.5281/zenodo.5996930" ID-GBIF-Taxon="148462569" ID-Zenodo-Dep="5996930" LSID="urn:lsid:plazi:treatment:DE5087A6C010B330FF39708CFD5FFBE5" httpUri="http://treatment.plazi.org/id/DE5087A6C010B330FF39708CFD5FFBE5" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<subSubSection id="1EE3653BC010B333FF39708CFD4EFF0D" box="[151,641,222,248]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="564636B0C010B333FF39708CFD4EFF0D" blockId="5.[151,641,222,248]" box="[151,641,222,248]" pageId="5" pageNumber="6">
<heading id="0D0E81DCC010B333FF39708CFD4EFF0D" bold="true" box="[151,641,222,248]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="648DEAA2C010B333FF39708CFD4EFF0D" bold="true" box="[151,641,222,248]" pageId="5" pageNumber="6">
Genus
<taxonomicName id="91F94D33C010B333FF40708CFD4EFF0D" ID-CoL="4TF8" ID-ENA="1107783" authority="Chamberlin, 1922" authorityName="Chamberlin" authorityYear="1922" box="[238,641,222,248]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Haplodrassus Chamberlin, 1922</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="1EE3653BC010B333FF397174FD0CFECE" box="[151,707,293,315]" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="564636B0C010B333FF397174FD0CFECE" blockId="5.[151,707,293,315]" box="[151,707,293,315]" pageId="5" pageNumber="6">
Type species:
<taxonomicName id="91F94D33C010B333FE887177FD71FECE" authority="(Emerton, 1909)" baseAuthorityName="Emerton" baseAuthorityYear="1909" box="[294,702,293,315]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="hiemalis">
<emphasis id="648DEAA2C010B333FE887177FDC0FECE" box="[294,527,293,315]" italics="true" pageId="5" pageNumber="6">Haplodrassus hiemalis</emphasis>
(Emerton, 1909)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="1EE3653BC010B333FF397135FCC9FDC1" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="564636B0C010B333FF397135FC1EFE50" blockId="5.[151,1436,359,565]" pageId="5" pageNumber="6">
<emphasis id="648DEAA2C010B333FF397135FEDEFE75" bold="true" box="[151,273,359,384]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
<bibRefCitation id="32684B41C010B333FEB27135FDE6FE75" box="[284,553,359,384]" pageId="5" pageNumber="6" refId="ref44463" refString="Kovblyuk, M. M., Kastrygina, Z. A. &amp; Omelko, M. M. (2012) A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae, Gnaphosidae). ZooKeys, 205, 59 - 89 https: // doi. org / 10.3897 / zookeys. 205.3491" type="journal article">
Kovblyuk
<emphasis id="648DEAA2C010B333FE39713BFE1BFE75" box="[407,468,359,384]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
(2012)
</bibRefCitation>
provided an excellent diagnosis of the genus. It actually counts 72 species (
<bibRefCitation id="32684B41C010B333FF3071DEFE18FE51" box="[158,471,396,421]" pageId="5" pageNumber="6" refId="ref46054" refString="World Spider Catalog (2017) World Spider Catalog. Version 18.0. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed 15 June 2017)" type="url">World Spider Catalog 2017</bibRefCitation>
), of which 8 have been cited in the Maghreb.
</paragraph>
<paragraph id="564636B0C010B333FF6971FDFCC9FDC1" blockId="5.[151,1436,359,565]" pageId="5" pageNumber="6">
<emphasis id="648DEAA2C010B333FF6971FDFC38FE3D" bold="true" box="[199,1015,431,456]" pageId="5" pageNumber="6">
Diagnostic characters of the species of the genus
<taxonomicName id="91F94D33C010B333FCBD71FDFC7DFE3D" authorityName="Chamberlin" authorityYear="1922" box="[787,946,431,456]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="648DEAA2C010B333FCBD71FDFC7DFE3D" bold="true" box="[787,946,431,456]" italics="true" pageId="5" pageNumber="6">Haplodrassus</emphasis>
</taxonomicName>
. Size.
</emphasis>
Small to medium-sized, often robust gnaphosids. This is a very variable character in
<taxonomicName id="91F94D33C010B333FD077186FC8AFE18" authorityName="Chamberlin" authorityYear="1922" box="[681,837,468,493]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="648DEAA2C010B333FD077186FC8AFE18" box="[681,837,468,493]" italics="true" pageId="5" pageNumber="6">Haplodrassus</emphasis>
</taxonomicName>
. Size categories are small (
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), medium-sized (
<quantity id="91019B55C010B333FF3171AAFF31FDE5" box="[159,254,504,528]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="5" pageNumber="6" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">4‒7 mm</quantity>
) or medium sized to large
<quantity id="91019B55C010B333FD8571A5FD56FDFA" box="[555,665,503,528]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="5" pageNumber="6" unit="mm" value="10.5" valueMax="15.0" valueMin="6.0">6‒15 mm</quantity>
. The largest species are the most variable in size. Variation in size of the different species of the Maghreb is listed in
<tableCitation id="1B7B030BC010B333FD03724EFCCFFDC1" box="[685,768,540,565]" captionStart="TABLE 1" captionStartId="5.[151,239,608,631]" captionText="TABLE 1. Variation in total size and carapace length in Haplodrassus species. Smallest species are listed in top of the table." pageId="5" pageNumber="6">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="02866638C010B333FF397232FF1DFD62" pageId="5" pageNumber="6">
<paragraph id="564636B0C010B333FF397232FF1DFD62" blockId="5.[151,1436,607,663]" pageId="5" pageNumber="6">
<emphasis id="648DEAA2C010B333FF397232FEC4FD82" bold="true" box="[151,267,608,631]" pageId="5" pageNumber="6">TABLE 1.</emphasis>
Variation in total size and carapace length in
<taxonomicName id="91F94D33C010B333FD52720DFC5FFD82" authorityName="Chamberlin" authorityYear="1922" box="[764,912,607,631]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="648DEAA2C010B333FD52720DFC5FFD82" box="[764,912,607,631]" italics="true" pageId="5" pageNumber="6">Haplodrassus</emphasis>
</taxonomicName>
species. Smallest species are listed in top of the table.
</paragraph>
</caption>
<paragraph id="564636B0C010B333FE1D72E0FB21FAC1" pageId="5" pageNumber="6">
<table id="24F9C410C0104CC9FF3172E0FA89FAC0" box="[159,1350,690,1333]" gridcols="5" gridrows="16" pageId="5" pageNumber="6">
<tr id="E8C934F2C0104CC9FF3172E0FA89FD3D" box="[159,1350,690,712]" gridrow="0" pageId="5" pageNumber="6" rowspan-0="1" rowspan-2="1" rowspan-4="1">
<th id="AB185D8EC0104CC9FE1D72E0FDE3FD3D" box="[435,556,690,712]" gridcol="1" gridrow="0" pageId="5" pageNumber="6">Male</th>
<th id="AB185D8EC0104CC9FC0F72E0FBD5FD3D" box="[929,1050,690,712]" gridcol="3" gridrow="0" pageId="5" pageNumber="6">Female</th>
</tr>
<tr id="E8C934F2C0104CC9FF31728EFA89FD07" box="[159,1350,732,754]" gridrow="1" pageId="5" pageNumber="6" rowspan-0="1">
<td id="AB185D8EC0104CC9FE1D728EFDE3FD07" box="[435,556,732,754]" gridcol="1" gridrow="1" pageId="5" pageNumber="6">Total length</td>
<td id="AB185D8EC0104CC9FD0F728EFC8BFD07" box="[673,836,732,754]" gridcol="2" gridrow="1" pageId="5" pageNumber="6">Carapace length</td>
<td id="AB185D8EC0104CC9FC0F728EFBD5FD07" box="[929,1050,732,754]" gridcol="3" gridrow="1" pageId="5" pageNumber="6">Total lenght</td>
<td id="AB185D8EC0104CC9FB0C728EFA89FD07" box="[1186,1350,732,754]" gridcol="4" gridrow="1" pageId="5" pageNumber="6">Carapace length</td>
</tr>
<tr id="E8C934F2C0104CC9FF317357FA89FCEE" box="[159,1350,773,795]" gridrow="2" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF317357FEBAFCEE" box="[159,373,773,795]" gridcol="0" gridrow="2" pageId="5" pageNumber="6">
<taxonomicName id="91F94D33C010B333FF317354FEFFFCEE" authorityName="Bosmans &amp; Kherbouche-Abrous &amp; Benhalima &amp; Hervé" authorityYear="2018" baseAuthorityName="Simon" baseAuthorityYear="1914" box="[159,304,773,795]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rhodanicus">
<emphasis id="648DEAA2C010B333FF317354FEFFFCEE" box="[159,304,773,795]" italics="true" pageId="5" pageNumber="6">H. rhodanicus</emphasis>
</taxonomicName>
</th>
<td id="AB185D8EC0104CC9FE1D7357FDE3FCEE" box="[435,556,773,795]" gridcol="1" gridrow="2" pageId="5" pageNumber="6">2.3‒5.2</td>
<td id="AB185D8EC0104CC9FD0F7357FC8BFCEE" box="[673,836,773,795]" gridcol="2" gridrow="2" pageId="5" pageNumber="6">1.20‒2.12</td>
<td id="AB185D8EC0104CC9FC0F7357FBD5FCEE" box="[929,1050,773,795]" gridcol="3" gridrow="2" pageId="5" pageNumber="6">2.9‒3.4</td>
<td id="AB185D8EC0104CC9FB0C7357FA89FCEE" box="[1186,1350,773,795]" gridcol="4" gridrow="2" pageId="5" pageNumber="6">1.14‒1.33</td>
</tr>
<tr id="E8C934F2C0104CC9FF31737CFA89FCB0" box="[159,1350,814,837]" gridrow="3" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF31737CFEBAFCB0" box="[159,373,814,837]" gridcol="0" gridrow="3" pageId="5" pageNumber="6">
<taxonomicName id="91F94D33C010B333FF317362FEE6FCB1" authority="Bosmans &amp; Kherbouche-Abrous &amp; Benhalima &amp; Hervé, 2018" authorityName="Bosmans &amp; Kherbouche-Abrous &amp; Benhalima &amp; Hervé" authorityYear="2018" box="[159,297,814,837]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="longivulva" status="sp. nov.">
<emphasis id="648DEAA2C010B333FF317362FEE6FCB1" box="[159,297,814,837]" italics="true" pageId="5" pageNumber="6">H. longivulva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FBE57D9C010B333FE81737DFEA8FCB1" box="[303,359,815,836]" pageId="5" pageNumber="6" rank="species">
<emphasis id="648DEAA2C010B333FE81737DFEA8FCB1" bold="true" box="[303,359,815,836]" pageId="5" pageNumber="6">sp. n.</emphasis>
</taxonomicNameLabel>
</th>
<td id="AB185D8EC0104CC9FE1D737CFDE3FCB0" box="[435,556,814,837]" gridcol="1" gridrow="3" pageId="5" pageNumber="6">2.5‒3.6</td>
<td id="AB185D8EC0104CC9FD0F737CFC8BFCB0" box="[673,836,814,837]" gridcol="2" gridrow="3" pageId="5" pageNumber="6">1.53‒1.94</td>
<td id="AB185D8EC0104CC9FC0F737CFBD5FCB0" box="[929,1050,814,837]" gridcol="3" gridrow="3" pageId="5" pageNumber="6">4.0‒6.6</td>
<td id="AB185D8EC0104CC9FB0C737CFA89FCB0" box="[1186,1350,814,837]" gridcol="4" gridrow="3" pageId="5" pageNumber="6">1.44‒2.79</td>
</tr>
<tr id="E8C934F2C0104CC9FF31730AFA89FC9B" box="[159,1350,856,878]" gridrow="4" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF31730AFEBAFC9B" box="[159,373,856,878]" gridcol="0" gridrow="4" pageId="5" pageNumber="6">
<taxonomicName id="91F94D33C010B333FF31730BFEF8FC9B" authority="Bosmans &amp; Kherbouche-Abrous &amp; Benhalima &amp; Hervé, 2018" authorityName="Bosmans &amp; Kherbouche-Abrous &amp; Benhalima &amp; Hervé" authorityYear="2018" box="[159,311,856,878]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="triangularis" status="sp. nov.">
<emphasis id="648DEAA2C010B333FF31730BFEF8FC9B" box="[159,311,856,878]" italics="true" pageId="5" pageNumber="6">H. triangularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FBE57D9C010B333FE93730AFEBAFC98" box="[317,373,856,877]" pageId="5" pageNumber="6" rank="species">
<emphasis id="648DEAA2C010B333FE93730AFEBAFC98" bold="true" box="[317,373,856,877]" pageId="5" pageNumber="6">sp. n.</emphasis>
</taxonomicNameLabel>
</th>
<td id="AB185D8EC0104CC9FE1D730AFDE3FC9B" box="[435,556,856,878]" gridcol="1" gridrow="4" pageId="5" pageNumber="6">2.9‒3.1</td>
<td id="AB185D8EC0104CC9FD0F730AFC8BFC9B" box="[673,836,856,878]" gridcol="2" gridrow="4" pageId="5" pageNumber="6">1.23‒1.46</td>
<td id="AB185D8EC0104CC9FC0F730AFBD5FC9B" box="[929,1050,856,878]" gridcol="3" gridrow="4" pageId="5" pageNumber="6">3.1‒4.8</td>
<td id="AB185D8EC0104CC9FB0C730AFA89FC9B" box="[1186,1350,856,878]" gridcol="4" gridrow="4" pageId="5" pageNumber="6">1.31‒1.6</td>
</tr>
<tr id="E8C934F2C0104CC9FF3173D3FA89FC62" box="[159,1350,897,919]" gridrow="5" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF3173D3FEBAFC62" box="[159,373,897,919]" gridcol="0" gridrow="5" pageId="5" pageNumber="6">
<taxonomicName id="91F94D33C010B333FF3173D0FECDFC62" authority="Bosmans &amp; Kherbouche-Abrous &amp; Benhalima &amp; Hervé, 2018" authorityName="Bosmans &amp; Kherbouche-Abrous &amp; Benhalima &amp; Hervé" authorityYear="2018" box="[159,258,897,919]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lyndae" status="sp. nov.">
<emphasis id="648DEAA2C010B333FF3173D0FECDFC62" box="[159,258,897,919]" italics="true" pageId="5" pageNumber="6">H. lyndae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FBE57D9C010B333FEA973D3FE8FFC63" box="[263,320,897,918]" pageId="5" pageNumber="6" rank="species">
<emphasis id="648DEAA2C010B333FEA973D3FE8FFC63" bold="true" box="[263,320,897,918]" pageId="5" pageNumber="6">sp. n.</emphasis>
</taxonomicNameLabel>
</th>
<td id="AB185D8EC0104CC9FE1D73D3FDE3FC62" box="[435,556,897,919]" gridcol="1" gridrow="5" pageId="5" pageNumber="6">2.9‒5.7</td>
<td id="AB185D8EC0104CC9FD0F73D3FC8BFC62" box="[673,836,897,919]" gridcol="2" gridrow="5" pageId="5" pageNumber="6">1.4‒2.7</td>
<td id="AB185D8EC0104CC9FC0F73D3FBD5FC62" box="[929,1050,897,919]" gridcol="3" gridrow="5" pageId="5" pageNumber="6">3.7‒6.4</td>
<td id="AB185D8EC0104CC9FB0C73D3FA89FC62" box="[1186,1350,897,919]" gridcol="4" gridrow="5" pageId="5" pageNumber="6">1.7‒2.7</td>
</tr>
<tr id="E8C934F2C0104CC9FF3173F8FA89FC35" box="[159,1350,938,960]" gridrow="6" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF3173F8FEBAFC35" box="[159,373,938,960]" gridcol="0" gridrow="6" pageId="5" pageNumber="6">
<taxonomicName id="91F94D33C010B333FF3173F9FEF8FC35" baseAuthorityName="L. Koch" baseAuthorityYear="1866" box="[159,311,938,960]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="dalmatensis">
<emphasis id="648DEAA2C010B333FF3173F9FEF8FC35" box="[159,311,938,960]" italics="true" pageId="5" pageNumber="6">H. dalmatensis</emphasis>
</taxonomicName>
</th>
<td id="AB185D8EC0104CC9FE1D73F8FDE3FC35" box="[435,556,938,960]" gridcol="1" gridrow="6" pageId="5" pageNumber="6">3.8‒6.5</td>
<td id="AB185D8EC0104CC9FD0F73F8FC8BFC35" box="[673,836,938,960]" gridcol="2" gridrow="6" pageId="5" pageNumber="6">1.61‒2.32</td>
<td id="AB185D8EC0104CC9FC0F73F8FBD5FC35" box="[929,1050,938,960]" gridcol="3" gridrow="6" pageId="5" pageNumber="6">3.7‒8.0</td>
<td id="AB185D8EC0104CC9FB0C73F8FA89FC35" box="[1186,1350,938,960]" gridcol="4" gridrow="6" pageId="5" pageNumber="6">1.62‒2.89</td>
</tr>
<tr id="E8C934F2C0104CC9FF317386FA89FC1F" box="[159,1350,980,1002]" gridrow="7" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF317386FEBAFC1F" box="[159,373,980,1002]" gridcol="0" gridrow="7" pageId="5" pageNumber="6">
<taxonomicName id="91F94D33C010B333FF317387FECCFC1F" authorityName="Bosmans &amp; Kherbouche-Abrous &amp; Benhalima &amp; Hervé" authorityYear="2018" baseAuthorityName="Simon" baseAuthorityYear="1878" box="[159,259,980,1002]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="typhon">
<emphasis id="648DEAA2C010B333FF317387FECCFC1F" box="[159,259,980,1002]" italics="true" pageId="5" pageNumber="6">H. typhon</emphasis>
</taxonomicName>
</th>
<td id="AB185D8EC0104CC9FE1D7386FDE3FC1F" box="[435,556,980,1002]" gridcol="1" gridrow="7" pageId="5" pageNumber="6">4.7‒6.0</td>
<td id="AB185D8EC0104CC9FD0F7386FC8BFC1F" box="[673,836,980,1002]" gridcol="2" gridrow="7" pageId="5" pageNumber="6">2.18‒2.70</td>
<td id="AB185D8EC0104CC9FC0F7386FBD5FC1F" box="[929,1050,980,1002]" gridcol="3" gridrow="7" pageId="5" pageNumber="6">6.2‒8.2</td>
<td id="AB185D8EC0104CC9FB0C7386FA89FC1F" box="[1186,1350,980,1002]" gridcol="4" gridrow="7" pageId="5" pageNumber="6">2.10‒2.90</td>
</tr>
<tr id="E8C934F2C0104CC9FF3173AFFA89FBE6" box="[159,1350,1021,1043]" gridrow="8" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF3173AFFEBAFBE6" box="[159,373,1021,1043]" gridcol="0" gridrow="8" pageId="5" pageNumber="6">
<taxonomicName id="91F94D33C010B333FF3173ACFEC4FBE6" baseAuthorityName="C. L. Koch" baseAuthorityYear="1839" box="[159,267,1021,1043]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="signifer">
<emphasis id="648DEAA2C010B333FF3173ACFEC4FBE6" box="[159,267,1021,1043]" italics="true" pageId="5" pageNumber="6">H. signifer</emphasis>
</taxonomicName>
</th>
<td id="AB185D8EC0104CC9FE1D73AFFDE3FBE6" box="[435,556,1021,1043]" gridcol="1" gridrow="8" pageId="5" pageNumber="6">4.7‒7.4</td>
<td id="AB185D8EC0104CC9FD0F73AFFC8BFBE6" box="[673,836,1021,1043]" gridcol="2" gridrow="8" pageId="5" pageNumber="6">2.1‒3.4</td>
<td id="AB185D8EC0104CC9FC0F73AFFBD5FBE6" box="[929,1050,1021,1043]" gridcol="3" gridrow="8" pageId="5" pageNumber="6">5.1‒11.8</td>
<td id="AB185D8EC0104CC9FB0C73AFFA89FBE6" box="[1186,1350,1021,1043]" gridcol="4" gridrow="8" pageId="5" pageNumber="6">2.1‒5.5</td>
</tr>
<tr id="E8C934F2C0104CC9FF317474FA89FBC8" box="[159,1350,1062,1085]" gridrow="9" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF317474FEBAFBC8" box="[159,373,1062,1085]" gridcol="0" gridrow="9" pageId="5" pageNumber="6">
<taxonomicName id="91F94D33C010B333FF31747AFED7FBC9" authority="Bosmans &amp; Kherbouche-Abrous &amp; Benhalima &amp; Hervé, 2018" authorityName="Bosmans &amp; Kherbouche-Abrous &amp; Benhalima &amp; Hervé" authorityYear="2018" box="[159,280,1062,1085]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="securifer" status="sp. nov.">
<emphasis id="648DEAA2C010B333FF31747AFED7FBC9" box="[159,280,1062,1085]" italics="true" pageId="5" pageNumber="6">H. securifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FBE57D9C010B333FEB07475FE98FBC9" box="[286,343,1063,1084]" pageId="5" pageNumber="6" rank="species">
<emphasis id="648DEAA2C010B333FEB07475FE98FBC9" bold="true" box="[286,343,1063,1084]" pageId="5" pageNumber="6">sp. n.</emphasis>
</taxonomicNameLabel>
</th>
<td id="AB185D8EC0104CC9FE1D7474FDE3FBC8" box="[435,556,1062,1085]" gridcol="1" gridrow="9" pageId="5" pageNumber="6">5.2‒6.5</td>
<td id="AB185D8EC0104CC9FD0F7474FC8BFBC8" box="[673,836,1062,1085]" gridcol="2" gridrow="9" pageId="5" pageNumber="6">2.1‒2.6</td>
<td id="AB185D8EC0104CC9FC0F7474FBD5FBC8" box="[929,1050,1062,1085]" gridcol="3" gridrow="9" pageId="5" pageNumber="6">6.0‒10.0</td>
<td id="AB185D8EC0104CC9FB0C7474FA89FBC8" box="[1186,1350,1062,1085]" gridcol="4" gridrow="9" pageId="5" pageNumber="6">2.8‒2.9</td>
</tr>
<tr id="E8C934F2C0104CC9FF317402FA89FB93" box="[159,1350,1104,1126]" gridrow="10" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF317402FEBAFB93" box="[159,373,1104,1126]" gridcol="0" gridrow="10" pageId="5" pageNumber="6">
<taxonomicName id="91F94D33C010B333FF317403FEC2FB93" authority="Bosmans &amp; Kherbouche-Abrous &amp; Benhalima &amp; Hervé, 2018" authorityName="Bosmans &amp; Kherbouche-Abrous &amp; Benhalima &amp; Hervé" authorityYear="2018" box="[159,269,1104,1126]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="dentifer" status="sp. nov.">
<emphasis id="648DEAA2C010B333FF317403FEC2FB93" box="[159,269,1104,1126]" italics="true" pageId="5" pageNumber="6">H. dentifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FBE57D9C010B333FEBD7402FE83FB90" box="[275,332,1104,1125]" pageId="5" pageNumber="6" rank="species">
<emphasis id="648DEAA2C010B333FEBD7402FE83FB90" bold="true" box="[275,332,1104,1125]" pageId="5" pageNumber="6">sp. n.</emphasis>
</taxonomicNameLabel>
</th>
<td id="AB185D8EC0104CC9FE1D7402FDE3FB93" box="[435,556,1104,1126]" gridcol="1" gridrow="10" pageId="5" pageNumber="6">5.6‒9.6</td>
<td id="AB185D8EC0104CC9FD0F7402FC8BFB93" box="[673,836,1104,1126]" gridcol="2" gridrow="10" pageId="5" pageNumber="6">1.22‒2.02</td>
<td id="AB185D8EC0104CC9FC0F7402FBD5FB93" box="[929,1050,1104,1126]" gridcol="3" gridrow="10" pageId="5" pageNumber="6">7.6‒9.6</td>
<td id="AB185D8EC0104CC9FB0C7402FA89FB93" box="[1186,1350,1104,1126]" gridcol="4" gridrow="10" pageId="5" pageNumber="6">1.96‒2.44</td>
</tr>
<tr id="E8C934F2C0104CC9FF31742BFA89FB7A" box="[159,1350,1145,1167]" gridrow="11" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF31742BFEBAFB7A" box="[159,373,1145,1167]" gridcol="0" gridrow="11" pageId="5" pageNumber="6">
<taxonomicName id="91F94D33C010B333FF317428FE8BFB7A" authorityName="Simon" authorityYear="1909" box="[159,324,1146,1167]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nigroscriptus">
<emphasis id="648DEAA2C010B333FF317428FE8BFB7A" box="[159,324,1146,1167]" italics="true" pageId="5" pageNumber="6">H. nigroscriptus</emphasis>
</taxonomicName>
</th>
<td id="AB185D8EC0104CC9FE1D742BFDE3FB7A" box="[435,556,1145,1167]" gridcol="1" gridrow="11" pageId="5" pageNumber="6">5.6‒7.2</td>
<td id="AB185D8EC0104CC9FD0F742BFC8BFB7A" box="[673,836,1145,1167]" gridcol="2" gridrow="11" pageId="5" pageNumber="6">2.68‒3.76</td>
<td id="AB185D8EC0104CC9FC0F742BFBD5FB7A" box="[929,1050,1145,1167]" gridcol="3" gridrow="11" pageId="5" pageNumber="6">4.8‒10.4</td>
<td id="AB185D8EC0104CC9FB0C742BFA89FB7A" box="[1186,1350,1145,1167]" gridcol="4" gridrow="11" pageId="5" pageNumber="6">2.4‒4.22</td>
</tr>
<tr id="E8C934F2C0104CC9FF3174F0FA89FB4D" box="[159,1350,1186,1208]" gridrow="12" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF3174F0FEBAFB4D" box="[159,373,1186,1208]" gridcol="0" gridrow="12" pageId="5" pageNumber="6">
<taxonomicName id="91F94D33C010B333FF3174F1FED3FB4D" authorityName="Lucas" authorityYear="1846" box="[159,284,1187,1208]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="648DEAA2C010B333FF3174F1FED3FB4D" box="[159,284,1187,1208]" italics="true" pageId="5" pageNumber="6">H. crassipes</emphasis>
</taxonomicName>
</th>
<td id="AB185D8EC0104CC9FE1D74F0FDE3FB4D" box="[435,556,1186,1208]" gridcol="1" gridrow="12" pageId="5" pageNumber="6">6.1‒10.3</td>
<td id="AB185D8EC0104CC9FD0F74F0FC8BFB4D" box="[673,836,1186,1208]" gridcol="2" gridrow="12" pageId="5" pageNumber="6">2.8‒5.9</td>
<td id="AB185D8EC0104CC9FC0F74F0FBD5FB4D" box="[929,1050,1186,1208]" gridcol="3" gridrow="12" pageId="5" pageNumber="6">10.2‒16.4</td>
<td id="AB185D8EC0104CC9FB0C74F0FA89FB4D" box="[1186,1350,1186,1208]" gridcol="4" gridrow="12" pageId="5" pageNumber="6">3.4‒6.4</td>
</tr>
<tr id="E8C934F2C0104CC9FF31749EFA89FB17" box="[159,1350,1228,1250]" gridrow="13" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF31749EFEBAFB17" box="[159,373,1228,1250]" gridcol="0" gridrow="13" pageId="5" pageNumber="6">
<taxonomicName id="91F94D33C010B333FF31749FFECBFB17" authorityName="Lucas" authorityYear="1846" box="[159,260,1228,1250]" class="Arachnida" family="Clubionidae" genus="Clubiona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="648DEAA2C010B333FF31749FFECBFB17" box="[159,260,1228,1250]" italics="true" pageId="5" pageNumber="6">H. rufipes</emphasis>
</taxonomicName>
</th>
<td id="AB185D8EC0104CC9FE1D749EFDE3FB17" box="[435,556,1228,1250]" gridcol="1" gridrow="13" pageId="5" pageNumber="6">6.1‒8.9</td>
<td id="AB185D8EC0104CC9FD0F749EFC8BFB17" box="[673,836,1228,1250]" gridcol="2" gridrow="13" pageId="5" pageNumber="6">2.7‒3.9</td>
<td id="AB185D8EC0104CC9FC0F749EFBD5FB17" box="[929,1050,1228,1250]" gridcol="3" gridrow="13" pageId="5" pageNumber="6">7.2‒10.0</td>
<td id="AB185D8EC0104CC9FB0C749EFA89FB17" box="[1186,1350,1228,1250]" gridcol="4" gridrow="13" pageId="5" pageNumber="6">2.4‒4.4</td>
</tr>
<tr id="E8C934F2C0104CC9FF3174A7FA89FAFE" box="[159,1350,1269,1291]" gridrow="14" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF3174A7FEBAFAFE" box="[159,373,1269,1291]" gridcol="0" gridrow="14" pageId="5" pageNumber="6">
<taxonomicName id="91F94D33C010B333FF3174A4FECFFAFE" authority="Bosmans &amp; Kherbouche-Abrous &amp; Benhalima &amp; Hervé, 2018" authorityName="Bosmans &amp; Kherbouche-Abrous &amp; Benhalima &amp; Hervé" authorityYear="2018" box="[159,256,1270,1291]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ovatus" status="sp. nov.">
<emphasis id="648DEAA2C010B333FF3174A4FECFFAFE" box="[159,256,1270,1291]" italics="true" pageId="5" pageNumber="6">H. ovatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FBE57D9C010B333FEA874A4FEF0FAFE" box="[262,319,1270,1291]" pageId="5" pageNumber="6" rank="species">
<emphasis id="648DEAA2C010B333FEA874A4FEF0FAFE" bold="true" box="[262,319,1270,1291]" pageId="5" pageNumber="6">sp. n.</emphasis>
</taxonomicNameLabel>
</th>
<td id="AB185D8EC0104CC9FE1D74A7FDE3FAFE" box="[435,556,1269,1291]" gridcol="1" gridrow="14" pageId="5" pageNumber="6">6.2‒8.0</td>
<td id="AB185D8EC0104CC9FD0F74A7FC8BFAFE" box="[673,836,1269,1291]" gridcol="2" gridrow="14" pageId="5" pageNumber="6">2.66‒3.44</td>
<td id="AB185D8EC0104CC9FC0F74A7FBD5FAFE" box="[929,1050,1269,1291]" gridcol="3" gridrow="14" pageId="5" pageNumber="6">6.8‒8.8</td>
<td id="AB185D8EC0104CC9FB0C74A7FA89FAFE" box="[1186,1350,1269,1291]" gridcol="4" gridrow="14" pageId="5" pageNumber="6">2.16‒3.14</td>
</tr>
<tr id="E8C934F2C0104CC9FF31754CFA89FAC0" box="[159,1350,1310,1333]" gridrow="15" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF31754CFEBAFAC0" box="[159,373,1310,1333]" gridcol="0" gridrow="15" pageId="5" pageNumber="6">
<taxonomicName id="91F94D33C010B333FF317572FEC1FAC0" authorityName="Cambridge" authorityYear="1872" box="[159,270,1312,1333]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="omissus">
<emphasis id="648DEAA2C010B333FF317572FEC1FAC0" box="[159,270,1312,1333]" italics="true" pageId="5" pageNumber="6">H. omissus</emphasis>
</taxonomicName>
</th>
<td id="AB185D8EC0104CC9FE1D754CFDE3FAC0" box="[435,556,1310,1333]" gridcol="1" gridrow="15" pageId="5" pageNumber="6">6.6‒7.5</td>
<td id="AB185D8EC0104CC9FD0F754CFC8BFAC0" box="[673,836,1310,1333]" gridcol="2" gridrow="15" pageId="5" pageNumber="6">2.9‒3.3</td>
<td id="AB185D8EC0104CC9FC0F754CFBD5FAC0" box="[929,1050,1310,1333]" gridcol="3" gridrow="15" pageId="5" pageNumber="6">4.9‒6.7</td>
<td id="AB185D8EC0104CC9FB0C754CFA89FAC0" box="[1186,1350,1310,1333]" gridcol="4" gridrow="15" pageId="5" pageNumber="6">2.0‒2.8</td>
</tr>
</table>
</paragraph>
<subSubSection id="1EE3653BC010B330FF697527FD5FFBE5" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph id="564636B0C010B333FF697527FAF2FA7B" blockId="5.[199,1341,1397,1422]" box="[199,1341,1397,1422]" pageId="5" pageNumber="6">
<emphasis id="648DEAA2C010B333FF697527FE97FA7B" bold="true" box="[199,344,1397,1422]" pageId="5" pageNumber="6">Spinulation.</emphasis>
Generally, the spinulation of the legs is as in
<tableCitation id="1B7B030BC010B333FCF97527FC65FA7B" box="[855,938,1397,1422]" captionStart="TABLE 2" captionStartId="5.[151,239,1465,1488]" captionText="TABLE 2. General pattern of spination of Haplodrassus species." pageId="5" pageNumber="6">Table 2</tableCitation>
(grouped spines between brackets):
</paragraph>
<caption id="02866638C010B333FF3975EBFC81FA25" box="[151,846,1464,1488]" pageId="5" pageNumber="6">
<paragraph id="564636B0C010B333FF3975EBFC81FA25" blockId="5.[151,846,1464,1488]" box="[151,846,1464,1488]" pageId="5" pageNumber="6">
<emphasis id="648DEAA2C010B333FF3975EBFEC6FA25" bold="true" box="[151,265,1465,1488]" pageId="5" pageNumber="6">TABLE 2.</emphasis>
General pattern of spination of
<taxonomicName id="91F94D33C010B333FDF075EAFD3DFA25" authorityName="Chamberlin" authorityYear="1922" box="[606,754,1464,1488]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="648DEAA2C010B333FDF075EAFD3DFA25" box="[606,754,1464,1488]" italics="true" pageId="5" pageNumber="6">Haplodrassus</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
<paragraph id="564636B0C010B333FF7475BFFAB8F952" pageId="5" pageNumber="6">
<table id="24F9C410C0104CC9FF3175BEFAB8F952" box="[159,1399,1516,1703]" gridcols="4" gridrows="5" pageId="5" pageNumber="6">
<tr id="E8C934F2C0104CC9FF3175BEFAB8F9F7" box="[159,1399,1516,1538]" gridrow="0" pageId="5" pageNumber="6" rowspan-0="1">
<th id="AB185D8EC0104CC9FF7475BEFEF7F9F7" box="[218,312,1516,1538]" gridcol="1" gridrow="0" pageId="5" pageNumber="6">Femur</th>
<th id="AB185D8EC0104CC9FE2675BEFD1DF9F7" box="[392,722,1516,1538]" gridcol="2" gridrow="0" pageId="5" pageNumber="6">Tibia</th>
<th id="AB185D8EC0104CC9FC6875BEFAB8F9F7" box="[966,1399,1516,1538]" gridcol="3" gridrow="0" pageId="5" pageNumber="6">Metatarsus</th>
</tr>
<tr id="E8C934F2C0104CC9FF317647FAB8F9DE" box="[159,1399,1557,1579]" gridrow="1" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF317647FF75F9DE" box="[159,186,1557,1579]" gridcol="0" gridrow="1" pageId="5" pageNumber="6">I</th>
<td id="AB185D8EC0104CC9FF747647FEF7F9DE" box="[218,312,1557,1579]" gridcol="1" gridrow="1" pageId="5" pageNumber="6">2d-pl</td>
<td id="AB185D8EC0104CC9FE267647FD1DF9DE" box="[392,722,1557,1579]" gridcol="2" gridrow="1" pageId="5" pageNumber="6">-</td>
<td id="AB185D8EC0104CC9FC687647FAB8F9DE" box="[966,1399,1557,1579]" gridcol="3" gridrow="1" pageId="5" pageNumber="6">(pv-rv)</td>
</tr>
<tr id="E8C934F2C0104CC9FF31766CFAB8F9A1" box="[159,1399,1598,1620]" gridrow="2" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF31766CFF75F9A1" box="[159,186,1598,1620]" gridcol="0" gridrow="2" pageId="5" pageNumber="6">II</th>
<td id="AB185D8EC0104CC9FF74766CFEF7F9A1" box="[218,312,1598,1620]" gridcol="1" gridrow="2" pageId="5" pageNumber="6">2d-pl</td>
<td id="AB185D8EC0104CC9FE26766CFD1DF9A1" box="[392,722,1598,1620]" gridcol="2" gridrow="2" pageId="5" pageNumber="6">-</td>
<td id="AB185D8EC0104CC9FC68766CFAB8F9A1" box="[966,1399,1598,1620]" gridcol="3" gridrow="2" pageId="5" pageNumber="6">(pv-rv)</td>
</tr>
<tr id="E8C934F2C0104CC9FF31763AFAB8F98B" box="[159,1399,1640,1662]" gridrow="3" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF31763AFF75F98B" box="[159,186,1640,1662]" gridcol="0" gridrow="3" pageId="5" pageNumber="6">III</th>
<td id="AB185D8EC0104CC9FF74763AFEF7F98B" box="[218,312,1640,1662]" gridcol="1" gridrow="3" pageId="5" pageNumber="6">2d-(pl-rl)</td>
<td id="AB185D8EC0104CC9FE26763AFD1DF98B" box="[392,722,1640,1662]" gridcol="2" gridrow="3" pageId="5" pageNumber="6">(pl-rl-pv-rv)-(pl-rl-pv-rl)</td>
<td id="AB185D8EC0104CC9FC68763AFAB8F98B" box="[966,1399,1640,1662]" gridcol="3" gridrow="3" pageId="5" pageNumber="6">(pl-rl-pv-rl)-(pd-rl-pl-rl-pvrv)</td>
</tr>
<tr id="E8C934F2C0104CC9FF3176C3FAB8F952" box="[159,1399,1681,1703]" gridrow="4" pageId="5" pageNumber="6">
<th id="AB185D8EC0104CC9FF3176C3FF75F952" box="[159,186,1681,1703]" gridcol="0" gridrow="4" pageId="5" pageNumber="6">IV</th>
<td id="AB185D8EC0104CC9FF7476C3FEF7F952" box="[218,312,1681,1703]" gridcol="1" gridrow="4" pageId="5" pageNumber="6">2d-(pl-rl)</td>
<td id="AB185D8EC0104CC9FE2676C3FD1DF952" box="[392,722,1681,1703]" gridcol="2" gridrow="4" pageId="5" pageNumber="6">(pl-rl-pv-rv)-(pl-rl-pv-rl)-(pv-rv)</td>
<td id="AB185D8EC0104CC9FC6876C3FAB8F952" box="[966,1399,1681,1703]" gridcol="3" gridrow="4" pageId="5" pageNumber="6">(pl-rl-pv-rl) (pl-rl-pv-rl)-(pd-rl-pl-rl-pv-rv)</td>
</tr>
</table>
</paragraph>
<paragraph id="564636B0C010B333FF6976BFFE78F828" blockId="5.[151,1437,1772,2013]" pageId="5" pageNumber="6">
Femora I‒II with 2 dorsal and 1 prolateral spines, femora III‒IV with supplementary pro- and retrolateral spines. Tibia I‒II spineless, tibiae III‒IV with numerous spines. Metatarsi I‒II with 1 proventral and 1 retroventral spine, metatarsi III‒IV with abundant spines. It appeared that even in the same population, variations can occur. The ventral spines on Mt I‒II are often absent, some additional spines can be present on TiI-II and there is variation in the number and disposition of the spines on Tibiae III‒IV and Metatarsi III‒IV. Spinulation thus cannot be used to separate species,
<taxonomicName id="91F94D33C010B333FE2F77F3FDF3F84D" authorityName="Simon" authorityYear="1909" box="[385,572,1953,1976]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nigroscriptus">
<emphasis id="648DEAA2C010B333FE2F77F3FDF3F84D" box="[385,572,1953,1976]" italics="true" pageId="5" pageNumber="6">H. nigroscriptus</emphasis>
</taxonomicName>
being the only exception with additional spines present on the palpi (see diagnosis of this species).
</paragraph>
<paragraph id="564636B0C013B330FF6970C5FB9AFD70" blockId="6.[151,1437,151,1040]" pageId="6" pageNumber="7">
<emphasis id="648DEAA2C013B330FF6970C5FEB6FF45" bold="true" box="[199,377,151,176]" pageId="6" pageNumber="7">Genital organs</emphasis>
. The morphological characters of male and female copulatory organs are the most important diagnostic characters, as in most other spiders. The most important ones are shown in
<figureCitation id="CEC22A35C013B330FBCC70EEFB2FFF21" box="[1122,1248,188,213]" captionStart="FIGURES 10–17" captionStartId="10.[151,264,1884,1907]" captionTargetBox="[225,1362,193,1868]" captionTargetId="figure@10.[225,1364,193,1868]" captionTargetPageId="10" captionText="FIGURES 10–17. Haplodrassus dalmatensis (France, male from Saint-André, female from Île de Bagaud). 10. Male, dorsal aspect; 11. Female, dorsal aspect; 12. Male palpal tibia, dorsolateral view; 13. Male left palp, retrolateral view; 14. Idem, ventral view; 15. Idem, prolateral view; 16. Epigyne, ventral view; 17. Vulva, ventral view. Male palp (Fig. 14): E=Embolus; MA=Median apophysis; RTA: Retrolateral tibial apophysis; EA=Embolic apophysis. Epigyne (Figs 16, 17): A=Areola; F=Fovea; H=Hood; LP=Lateral pocket; PF=Pro-fovea; S=Septum; Sp=Spermatheca." httpUri="https://zenodo.org/record/1445084/files/figure.png" pageId="6" pageNumber="7">
Figs
<quantity id="91019B55C013B330FB3770EEFB2FFF21" box="[1177,1248,188,212]" metricMagnitude="-1" metricUnit="m" metricValue="3.429" metricValueMax="3.556" metricValueMin="3.302" pageId="6" pageNumber="7" unit="in" value="13.5" valueMax="14.0" valueMin="13.0">13‒14</quantity>
</figureCitation>
in the male palp and
<figureCitation id="CEC22A35C013B330FF69708DFE8AFF0D" box="[199,325,223,248]" captionStart="FIGURES 10–17" captionStartId="10.[151,264,1884,1907]" captionTargetBox="[225,1362,193,1868]" captionTargetId="figure@10.[225,1364,193,1868]" captionTargetPageId="10" captionText="FIGURES 10–17. Haplodrassus dalmatensis (France, male from Saint-André, female from Île de Bagaud). 10. Male, dorsal aspect; 11. Female, dorsal aspect; 12. Male palpal tibia, dorsolateral view; 13. Male left palp, retrolateral view; 14. Idem, ventral view; 15. Idem, prolateral view; 16. Epigyne, ventral view; 17. Vulva, ventral view. Male palp (Fig. 14): E=Embolus; MA=Median apophysis; RTA: Retrolateral tibial apophysis; EA=Embolic apophysis. Epigyne (Figs 16, 17): A=Areola; F=Fovea; H=Hood; LP=Lateral pocket; PF=Pro-fovea; S=Septum; Sp=Spermatheca." httpUri="https://zenodo.org/record/1445084/files/figure.png" pageId="6" pageNumber="7">
Figs
<quantity id="91019B55C013B330FF51708DFE8AFF0D" box="[255,325,223,248]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.318" metricValueMin="4.064" pageId="6" pageNumber="7" unit="in" value="16.5" valueMax="17.0" valueMin="16.0">
16
<emphasis id="648DEAA2C013B330FEB5708DFEE6FF0D" bold="true" box="[283,297,223,248]" pageId="6" pageNumber="7">–</emphasis>
17
</quantity>
</figureCitation>
in the epigyne and vulva. The nomenclature follows
<bibRefCitation id="32684B41C013B330FC0F708DFB1BFF0D" box="[929,1236,223,248]" pageId="6" pageNumber="7" refId="ref44909" refString="Omelko, M. M. &amp; Marusik, Y. M. (2012) A review of the Haplodrassus montanus - group (Aranei: Gnaphosidae) in the east Paleartic and preliminary grouping of the genus. Arthropoda Selecta, 21, 339 - 348." type="journal article">Omelko &amp; Marusik (2012)</bibRefCitation>
. The
<emphasis id="648DEAA2C013B330FABD708DFE9EFEE8" bold="true" pageId="6" pageNumber="7">retrolateral tibial apophysis</emphasis>
(
<emphasis id="648DEAA2C013B330FECF7156FE57FEE8" bold="true" box="[353,408,260,285]" pageId="6" pageNumber="7">RTA</emphasis>
) of the male palp is a very important character. The relative length, the presence of dorsal or ventral concavities at pro- or retrolateral side, the presence of keels and the shape of the tip are important. The tegulum always carries a
<emphasis id="648DEAA2C013B330FE64711EFD69FE90" bold="true" box="[458,678,332,357]" pageId="6" pageNumber="7">median apophysis</emphasis>
(
<emphasis id="648DEAA2C013B330FD12711EFD24FE90" bold="true" box="[700,747,332,357]" pageId="6" pageNumber="7">MA</emphasis>
) at its prolateral side. It is connected to the bulbus by a membrane. Its orientation can therefore be variable and can depend on the fixation conditions of the spider. Its general shape is hook-like and it shows practically no variation throughout the genus. The
<emphasis id="648DEAA2C013B330FB1071C6FA53FE58" bold="true" box="[1214,1436,404,429]" pageId="6" pageNumber="7">embolic apophysis</emphasis>
(
<emphasis id="648DEAA2C013B330FF0E71E5FF23FE25" bold="true" box="[160,236,439,464]" pageId="6" pageNumber="7">EA)—</emphasis>
Simon’s “lame sigmoide” (1914)
<emphasis id="648DEAA2C013B330FDC871E5FD4DFE25" bold="true" box="[614,642,439,464]" pageId="6" pageNumber="7">—</emphasis>
has a medio-distal position in the palp and is another very important diagnostic character. It is an elongated, more or less curved sclerite with pro- or retrolateral concavities, eventually provided with teeth or keels and with a more or less pointed tip. At its prolateral side, the embolic apophyis is connected to the embolus. The
<emphasis id="648DEAA2C013B330FE557276FD90FDC8" bold="true" box="[507,607,548,573]" pageId="6" pageNumber="7">embolus</emphasis>
(
<emphasis id="648DEAA2C013B330FDDE7276FD4CFDC8" bold="true" box="[624,643,548,573]" pageId="6" pageNumber="7">E</emphasis>
) is a gently curved sclerite at the retrolateral side of the bulbus, often partly covered by the embolic apophysis.
<collectingRegion id="943DF852C013B330FDDB721AFD46FD95" box="[629,649,584,608]" country="Gambia" name="Upper River" pageId="6" pageNumber="7">A</collectingRegion>
basal tooth can be present and at its retrolateral margin, subterminal denticles can be present.
<collectingRegion id="943DF852C013B330FE01723EFE0CFD71" box="[431,451,620,644]" country="Gambia" name="Upper River" pageId="6" pageNumber="7">A</collectingRegion>
detailed study of the embolus is not included in this work.
</paragraph>
<paragraph id="564636B0C013B330FF6972DDFD5FFBE5" blockId="6.[151,1437,151,1040]" pageId="6" pageNumber="7">
Females have an epigyne with a plate that can be as wide as long or longer than wide, expressed as the ratio length/width. The
<emphasis id="648DEAA2C013B330FEC972E6FDE7FD38" bold="true" box="[359,552,692,717]" pageId="6" pageNumber="7">anterior margin</emphasis>
can be well sclerotized or poorly sclerotized, and can be straight or curved; it covers a
<emphasis id="648DEAA2C013B330FF557285FEF9FD05" bold="true" box="[251,310,727,752]" pageId="6" pageNumber="7">hood</emphasis>
(
<emphasis id="648DEAA2C013B330FEE97285FE92FD05" bold="true" box="[327,349,727,752]" pageId="6" pageNumber="7">H</emphasis>
) of variable size. The epigyne has a large
<emphasis id="648DEAA2C013B330FC927285FC41FD05" bold="true" box="[828,910,727,752]" pageId="6" pageNumber="7">atrium</emphasis>
which is divided into an unchitinised upper or anterior part which is laterally in open connection with the integument of the abdomen, called
<emphasis id="648DEAA2C013B330FB1E72AEFAECFCE0" bold="true" box="[1200,1315,764,789]" pageId="6" pageNumber="7">pro-fovea</emphasis>
(
<emphasis id="648DEAA2C013B330FA9D72AEFA9AFCE0" bold="true" box="[1331,1365,764,789]" pageId="6" pageNumber="7">PF</emphasis>
) here, and a well-defined basal or posterior part,
<emphasis id="648DEAA2C013B330FDDE734DFCC9FCCD" bold="true" box="[624,774,799,824]" pageId="6" pageNumber="7">the fovea (F)</emphasis>
, and its ratio can be relevant. The upper part is whitish and can be wrinkled, the basal part is a generally pale depression with lateral margins which can be denticulate. Laterally, the fovea is flanked by a brown to dark brown area, the
<emphasis id="648DEAA2C013B330FC3F7335FC14FC75" bold="true" box="[913,987,871,896]" pageId="6" pageNumber="7">areola</emphasis>
(
<emphasis id="648DEAA2C013B330FC407335FBE7FC75" bold="true" box="[1006,1064,871,896]" pageId="6" pageNumber="7">
<collectingRegion id="943DF852C013B330FC407335FBCDFC75" box="[1006,1026,871,896]" country="Gambia" name="Upper River" pageId="6" pageNumber="7">A</collectingRegion>
)—
</emphasis>
Simon’s “pièce en fer-à-cheval”- that can be denticulate. The vulva consists of basal
<emphasis id="648DEAA2C013B330FD4D73DEFCA8FC50" bold="true" box="[739,871,908,933]" pageId="6" pageNumber="7">receptacles</emphasis>
(
<emphasis id="648DEAA2C013B330FCD673DEFC56FC50" bold="true" box="[888,921,908,933]" pageId="6" pageNumber="7">Re</emphasis>
),
<emphasis id="648DEAA2C013B330FC1C73DEFBADFC50" bold="true" box="[946,1122,908,933]" pageId="6" pageNumber="7">lateral pockets</emphasis>
(
<emphasis id="648DEAA2C013B330FBDA73DEFB57FC50" bold="true" box="[1140,1176,908,933]" pageId="6" pageNumber="7">LP</emphasis>
) bordering the median fovea and copulatory and fertilisation ducts. The presence of an exudate in the lateral part of the fovea can deform the whole appearance of the epigyne and vulva. It can have the same colour of the areola and may obscure teeth or denticles, often leading to misidentifications.
</paragraph>
</subSubSection>
</treatment>
</document>
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