324 lines
25 KiB
XML
324 lines
25 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/evolsyst.5.70944" ID-GBIF-Dataset="d91a3554-48df-4520-8d3b-5189c20cd23b" ID-GBIF-Taxon="183925325" ID-Pensoft-Pub="2535-0730-2-189" ID-Pensoft-UUID="6B15A19DDF4E55C5A7762DF939697BEC" ID-ZooBank="B7258B7EDCAC4A6E9E1E03218501861B" LSID="urn:lsid:zoobank.org:act:E10AAAD8-94EE-4A7B-8DAC-94BB1BDA9427" ModsDocID="2535-0730-5-2-189" checkinTime="1628190608819" checkinUser="pensoft" docAuthor="Ratcliffe, Brett C. & Seidel, Matthias" docDate="2021" docId="CFF2ECFF19E057C0922B233CD4AD1BC8" docLanguage="en" docName="EvolutSyst 5(2): 189-192" docOrigin="Evolutionary Systematics 5 (2)" docPubDate="2021-08-05" docSource="http://dx.doi.org/10.3897/evolsyst.5.70944" docTitle="Chalepides pantanalensis Ratcliffe & Seidel 2021, sp. nov." docType="treatment" docUuid="E10AAAD8-94EE-4A7B-8DAC-94BB1BDA9427" docUuidSource="ZooBank" docVersion="3" id="6B15A19DDF4E55C5A7762DF939697BEC" lastPageNumber="189" masterDocId="6B15A19DDF4E55C5A7762DF939697BEC" masterDocTitle="A new species of Chalepides Casey, 1915 (Scarabaeidae, Dynastinae, Cyclocephalini) from the Pantanal of Brazil" masterLastPageNumber="192" masterPageNumber="189" pageId="0" pageNumber="189" updateTime="1643436797749" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new species of Chalepides Casey, 1915 (Scarabaeidae, Dynastinae, Cyclocephalini) from the Pantanal of Brazil</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ratcliffe, Brett C.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0589-8791</mods:nameIdentifier>
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<mods:affiliation>Systematics Research Collections, 307 Morrill Hall, University of Nebraska State Museum, Lincoln, NE 68588 - 0338, USA</mods:affiliation>
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<mods:nameIdentifier type="email">bratcliffe1@unl.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Seidel, Matthias</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4913-8778</mods:nameIdentifier>
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<mods:affiliation>Centrum fuer Naturkunde - Center of Natural History, Universitaet Hamburg - Zoological Museum, Martin-Luther-King-Platz 3, 20146, Hamburg, Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Evolutionary Systematics</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-08-05</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>189</mods:start>
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<mods:end>192</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/evolsyst.5.70944</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/evolsyst.5.70944</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2535-0730-2-189</mods:identifier>
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<mods:identifier type="ZooBank">B7258B7EDCAC4A6E9E1E03218501861B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">6B15A19DDF4E55C5A7762DF939697BEC</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="183925325" LSID="urn:lsid:zoobank.org:act:E10AAAD8-94EE-4A7B-8DAC-94BB1BDA9427" httpUri="http://treatment.plazi.org/id/CFF2ECFF19E057C0922B233CD4AD1BC8" lastPageNumber="189" pageId="0" pageNumber="189">
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<subSubSection pageId="0" pageNumber="189" type="nomenclature">
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<paragraph pageId="0" pageNumber="189">
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<taxonomicName LSID="http://zoobank.org/E10AAAD8-94EE-4A7B-8DAC-94BB1BDA9427" authority="Ratcliffe & Seidel" authorityName="Ratcliffe & Seidel" authorityYear="2021" class="Insecta" family="Scarabaeidae" genus="Chalepides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalepides pantanalensis" order="Coleoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="pantanalensis" status="sp. nov.">Chalepides pantanalensis Ratcliffe & Seidel</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="189">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="189" type="description">
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<paragraph pageId="0" pageNumber="189">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype, dorsal view, of Chalepides pantanalensis Ratcliffe & Seidel, sp. nov." httpUri="https://binary.pensoft.net/fig/574481" pageId="0" pageNumber="189">Figs 1</figureCitation>
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<figureCitation captionStart="Figures 2, 3" captionStartId="F2" captionText="Figures 2, 3. Male protarsal large claw of (2) C. pantanalensis Ratcliffe & Seidel, sp. nov. and (3) C. howdenorum Joly & Escalona." httpUri="https://binary.pensoft.net/fig/574482" pageId="0" pageNumber="189">, 2</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F3" captionText="Figure 4. Parameres, caudal and lateral views, of Chalepides pantanalensis Ratcliffe & Seidel, sp. nov." httpUri="https://binary.pensoft.net/fig/574483" pageId="0" pageNumber="189">, 4</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F4" captionText="Figure 5. Location of type localities of Fazenda 4 Cantos and Fazenda Natureza, Mato Grosso do Sul, Brazil" httpUri="https://binary.pensoft.net/fig/574484" pageId="0" pageNumber="189">, 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="189" type="type material">
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<paragraph pageId="0" pageNumber="189">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="189">
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<emphasis bold="true" pageId="0" pageNumber="189">Holotype</emphasis>
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male labeled "BRAZIL/ Pantanal-Fazenda/ Natureza 4.-8.11.2005/ lgt. Z.
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<normalizedToken originalValue="Tüdös”">Tuedoes"</normalizedToken>
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and with our red holotype label.
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<emphasis bold="true" pageId="0" pageNumber="189">Allotype</emphasis>
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female labeled "Brazil, Pantanal/ Fazenda 4 cantos, 7.-10.5.2012/
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<geoCoordinate degrees="18" direction="south" minutes="36" orientation="latitude" precision="1" seconds="54.40" value="-18.615112">18°36'54.40"S</geoCoordinate>
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,
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<geoCoordinate degrees="56" direction="west" minutes="15" orientation="longitude" precision="1" seconds="56.97" value="-56.265823">56°15'56.97"W</geoCoordinate>
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/ lgt. O. Konvicka" and with our red allotype label. One male and three female
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<emphasis bold="true" pageId="0" pageNumber="189">paratypes</emphasis>
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with same data as holotype and each with our yellow paratype label. Two male
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<emphasis bold="true" pageId="0" pageNumber="189">paratypes</emphasis>
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with same data as allotype and each with our yellow paratype label. Holotype deposited at the Zoological Museum (
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<emphasis bold="true" pageId="0" pageNumber="189">ZMH</emphasis>
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), Centrum
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<normalizedToken originalValue="für">fuer</normalizedToken>
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Naturkunde,
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<normalizedToken originalValue="Universität">Universitaet</normalizedToken>
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Hamburg, Hamburg, Germany. Allotype and one male paratype deposited at the University of Nebraska State Museum (
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<emphasis bold="true" pageId="0" pageNumber="189">UNSM</emphasis>
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), Lincoln, NE, USA; one male paratype deposited in the Matthias Seidel Collection (
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<emphasis bold="true" pageId="0" pageNumber="189">MSPC</emphasis>
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), Hamburg, Germany; and one male and three female paratypes deposited in the
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<normalizedToken originalValue="Ondřej">Ondrej</normalizedToken>
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<normalizedToken originalValue="Konvička">Konvicka</normalizedToken>
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Collection,
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<normalizedToken originalValue="Zlín">Zlin</normalizedToken>
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, Czechia (
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<emphasis bold="true" pageId="0" pageNumber="189">OKPC</emphasis>
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).
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/574481" pageId="0" pageNumber="189" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="189">
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<emphasis bold="true" pageId="0" pageNumber="189">Figure 1.</emphasis>
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Holotype, dorsal view, of
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<taxonomicName authorityName="Ratcliffe & Seidel" authorityYear="2021" class="Insecta" family="Scarabaeidae" genus="Chalepides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalepides pantanalensis" order="Coleoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="pantanalensis">
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<emphasis italics="true" pageId="0" pageNumber="189">Chalepides pantanalensis</emphasis>
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</taxonomicName>
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Ratcliffe & Seidel, sp. nov.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="189" type="diagnosis">
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<paragraph pageId="0" pageNumber="189">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="189">
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<taxonomicName authorityName="Ratcliffe & Seidel" authorityYear="2021" class="Insecta" family="Scarabaeidae" genus="Chalepides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalepides pantanalensis" order="Coleoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="pantanalensis">
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<emphasis italics="true" pageId="0" pageNumber="189">Chalepides pantanalensis</emphasis>
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</taxonomicName>
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will key to couplet 11 in
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<bibRefCitation author="Endroedi, S" journalOrPublisher="Junk Publisher, Dordrecht, The Netherlands" pageId="0" pageNumber="189" refId="B2" refString="Endroedi, S, 1985. The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, The Netherlands" title="The Dynastinae of the World. Dr. W." year="1985">
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<normalizedToken originalValue="Endrödi">Endroedi</normalizedToken>
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(1985)
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</bibRefCitation>
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which ends with
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<taxonomicName genus="C." lsidName="C. comes" pageId="0" pageNumber="189" rank="species" species="comes">
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<emphasis italics="true" pageId="0" pageNumber="189">C. comes</emphasis>
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</taxonomicName>
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Prell (Venezuela, Colombia, Brazil) where the character states do not match: degree of punctation of pronotum versus frons, pygidial sculpturing, and form of the parameres. In
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<bibRefCitation author="Joly, LJ" journalOrPublisher="Entomotropica" pageId="0" pageNumber="189" pagination="37 - 90" refId="B3" refString="Joly, LJ, Escalona, HE, 2002. Revision del genero Chalepides Casey, 1915 (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini). Entomotropica 17: 37 - 90" title="Revision del genero Chalepides Casey, 1915 (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini)." volume="17" year="2002">Joly and Escalona (2002)</bibRefCitation>
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, it will key to couplet 8 for
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<taxonomicName genus="C." lsidName="C. osunai" pageId="0" pageNumber="189" rank="species" species="osunai">
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<emphasis italics="true" pageId="0" pageNumber="189">C. osunai</emphasis>
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</taxonomicName>
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Joly & Escalona (Venezuela) and
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<taxonomicName genus="C." lsidName="C. howdenorum" pageId="0" pageNumber="189" rank="species" species="howdenorum">
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<emphasis italics="true" pageId="0" pageNumber="189">C. howdenorum</emphasis>
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</taxonomicName>
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Joly & Escalona (northern Bolivia), the only
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<taxonomicName authorityName="Casey" authorityYear="1915" class="Insecta" family="Scarabaeidae" genus="Chalepides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalepides" order="Coleoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="189">Chalepides</emphasis>
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</taxonomicName>
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species with a bulbous and rounded apex of the parameres (as in
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<taxonomicName genus="C." lsidName="C. pantanalensis" pageId="0" pageNumber="189" rank="species" species="pantanalensis">
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<emphasis italics="true" pageId="0" pageNumber="189">C. pantanalensis</emphasis>
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</taxonomicName>
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).
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<taxonomicName authorityName="Ratcliffe & Seidel" authorityYear="2021" class="Insecta" family="Scarabaeidae" genus="Chalepides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalepides pantanalensis" order="Coleoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="pantanalensis">
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<emphasis italics="true" pageId="0" pageNumber="189">Chalepides pantanalensis</emphasis>
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</taxonomicName>
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has some character states seen in both of these species. The parameres are nearly identical to those of
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<taxonomicName genus="C." lsidName="C. howdenorum" pageId="0" pageNumber="189" rank="species" species="howdenorum">
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<emphasis italics="true" pageId="0" pageNumber="189">C. howdenorum</emphasis>
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</taxonomicName>
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, but the male of
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<taxonomicName genus="C." lsidName="C. pantanalensis" pageId="0" pageNumber="189" rank="species" species="pantanalensis">
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<emphasis italics="true" pageId="0" pageNumber="189">C. pantanalensis</emphasis>
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</taxonomicName>
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has a tooth on the venter of protarsomere 5 (absent in
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<taxonomicName genus="C." lsidName="C. howdenorum" pageId="0" pageNumber="189" rank="species" species="howdenorum">
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<emphasis italics="true" pageId="0" pageNumber="189">C. howdenorum</emphasis>
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</taxonomicName>
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, Fig.
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<figureCitation captionStart="Figures 2, 3" captionStartId="F2" captionText="Figures 2, 3. Male protarsal large claw of (2) C. pantanalensis Ratcliffe & Seidel, sp. nov. and (3) C. howdenorum Joly & Escalona." httpUri="https://binary.pensoft.net/fig/574482" pageId="0" pageNumber="189">3</figureCitation>
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) and the larger protarsal claw is short and thick in
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<taxonomicName genus="C." lsidName="C. pantanalensis" pageId="0" pageNumber="189" rank="species" species="pantanalensis">
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<emphasis italics="true" pageId="0" pageNumber="189">C. pantanalensis</emphasis>
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</taxonomicName>
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(narrowly elongate in
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<taxonomicName genus="C." lsidName="C. howdenorum" pageId="0" pageNumber="189" rank="species" species="howdenorum">
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<emphasis italics="true" pageId="0" pageNumber="189">C. howdenorum</emphasis>
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</taxonomicName>
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) (compare Figs
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<figureCitation captionStart="Figures 2, 3" captionStartId="F2" captionText="Figures 2, 3. Male protarsal large claw of (2) C. pantanalensis Ratcliffe & Seidel, sp. nov. and (3) C. howdenorum Joly & Escalona." httpUri="https://binary.pensoft.net/fig/574482" pageId="0" pageNumber="189">2</figureCitation>
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and
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<figureCitation captionStart="Figures 2, 3" captionStartId="F2" captionText="Figures 2, 3. Male protarsal large claw of (2) C. pantanalensis Ratcliffe & Seidel, sp. nov. and (3) C. howdenorum Joly & Escalona." httpUri="https://binary.pensoft.net/fig/574482" pageId="0" pageNumber="189">3</figureCitation>
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).
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/574482" pageId="0" pageNumber="189" start="Figures 2, 3" startId="F2">
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<paragraph pageId="0" pageNumber="189">
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<emphasis bold="true" pageId="0" pageNumber="189">Figures 2, 3.</emphasis>
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Male protarsal large claw of
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<emphasis bold="true" pageId="0" pageNumber="189">(2)</emphasis>
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<taxonomicName genus="C." lsidName="C. pantanalensis" pageId="0" pageNumber="189" rank="species" species="pantanalensis">
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<emphasis italics="true" pageId="0" pageNumber="189">C. pantanalensis</emphasis>
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</taxonomicName>
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Ratcliffe & Seidel, sp. nov. and
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<emphasis bold="true" pageId="0" pageNumber="189">(3)</emphasis>
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<taxonomicName genus="C." lsidName="C. howdenorum" pageId="0" pageNumber="189" rank="species" species="howdenorum">
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<emphasis italics="true" pageId="0" pageNumber="189">C. howdenorum</emphasis>
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</taxonomicName>
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Joly & Escalona.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="189">
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The male of
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<taxonomicName genus="C." lsidName="C. pantanalensis" pageId="0" pageNumber="189" rank="species" species="pantanalensis">
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<emphasis italics="true" pageId="0" pageNumber="189">C. pantanalensis</emphasis>
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</taxonomicName>
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shares with
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<taxonomicName genus="C." lsidName="C. osunai" pageId="0" pageNumber="189" rank="species" species="osunai">
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<emphasis italics="true" pageId="0" pageNumber="189">C. osunai</emphasis>
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</taxonomicName>
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a tooth on the venter of protarsomere 5 and a larger short, thick protarsal claw, but
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<taxonomicName genus="C." lsidName="C. pantanalensis" pageId="0" pageNumber="189" rank="species" species="pantanalensis">
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<emphasis italics="true" pageId="0" pageNumber="189">C. pantanalensis</emphasis>
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</taxonomicName>
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has a greenish sheen (absent in
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<taxonomicName genus="C." lsidName="C. osunai" pageId="0" pageNumber="189" rank="species" species="osunai">
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<emphasis italics="true" pageId="0" pageNumber="189">C. osunai</emphasis>
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</taxonomicName>
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), a densely punctate clypeus (sparsely punctate in
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<taxonomicName genus="C." lsidName="C. osunai" pageId="0" pageNumber="189" rank="species" species="osunai">
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<emphasis italics="true" pageId="0" pageNumber="189">C. osunai</emphasis>
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</taxonomicName>
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), and shorter, suboval parameres (elongate in
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<taxonomicName genus="C." lsidName="C. osunai" pageId="0" pageNumber="189" rank="species" species="osunai">
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<emphasis italics="true" pageId="0" pageNumber="189">C. osunai</emphasis>
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="189" type="description">
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<paragraph pageId="0" pageNumber="189">Description of holotype</paragraph>
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<paragraph pageId="0" pageNumber="189">
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(Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype, dorsal view, of Chalepides pantanalensis Ratcliffe & Seidel, sp. nov." httpUri="https://binary.pensoft.net/fig/574481" pageId="0" pageNumber="189">1</figureCitation>
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). Male. Length 16.7 mm; width 6.9 mm. Color black with faint olive-green sheen.
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<emphasis bold="true" pageId="0" pageNumber="189">Head</emphasis>
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: Frons with small, moderately dense, ocellate punctures. Clypeus with small, dense, ocellate punctures; apex broadly, shallowly emarginate, slightly reflexed. Frontal suture arcuate, complete. Interocular width equals 6.0 transverse eye diameters.
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<emphasis bold="true" pageId="0" pageNumber="189">Pronotum</emphasis>
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: Surface with small, sparse, ocellate punctures.
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<emphasis bold="true" pageId="0" pageNumber="189">Elytra</emphasis>
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: Surface punctate-striate; punctures small, ocellate, separated by about 1 puncture diameter in each row, moderately dense in intervals. Sutural stria a row of small punctures.
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<emphasis bold="true" pageId="0" pageNumber="189">Pygidium</emphasis>
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: Prepygidium elongated, coarsely and longitudinally strigose, with long, dense, tawny setae. Pygidium short; surface shiny with large, dense, setigerous punctures, punctures becoming small in lateral angles; setae short, moderately dense, tawny. Surface convex in lateral view.
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<emphasis bold="true" pageId="0" pageNumber="189">Legs</emphasis>
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: Protibia tridentate, teeth subequally spaced. Protarsus enlarged; tarsomere 4 and base of tarsomere 5 densely strigulose ventrally; tarsomere 5 longitudinally carinulate on inner margin and with distinctive tooth beneath (Fig.
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<figureCitation captionStart="Figures 2, 3" captionStartId="F2" captionText="Figures 2, 3. Male protarsal large claw of (2) C. pantanalensis Ratcliffe & Seidel, sp. nov. and (3) C. howdenorum Joly & Escalona." httpUri="https://binary.pensoft.net/fig/574482" pageId="0" pageNumber="189">2</figureCitation>
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); median claw stout, short, strongly bent (Fig.
|
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<figureCitation captionStart="Figures 2, 3" captionStartId="F2" captionText="Figures 2, 3. Male protarsal large claw of (2) C. pantanalensis Ratcliffe & Seidel, sp. nov. and (3) C. howdenorum Joly & Escalona." httpUri="https://binary.pensoft.net/fig/574482" pageId="0" pageNumber="189">2</figureCitation>
|
||
). Metatarsus shorter than metatibia.
|
||
<emphasis bold="true" pageId="0" pageNumber="189">Venter</emphasis>
|
||
: Prosternal process long, thick, columnar, apex flattened with raised, round
|
||
<normalizedToken originalValue="“button”">"button"</normalizedToken>
|
||
on all but slender, setose posterior margin.
|
||
<emphasis bold="true" pageId="0" pageNumber="189">Parameres</emphasis>
|
||
: Each side of shaft at about midpoint with small tooth and then shaft expanded to rounded apex (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F3" captionText="Figure 4. Parameres, caudal and lateral views, of Chalepides pantanalensis Ratcliffe & Seidel, sp. nov." httpUri="https://binary.pensoft.net/fig/574483" pageId="0" pageNumber="189">4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption httpUri="https://binary.pensoft.net/fig/574483" pageId="0" pageNumber="189" start="Figure 4" startId="F3">
|
||
<paragraph pageId="0" pageNumber="189">
|
||
<emphasis bold="true" pageId="0" pageNumber="189">Figure 4.</emphasis>
|
||
Parameres, caudal and lateral views, of
|
||
<taxonomicName authorityName="Ratcliffe & Seidel" authorityYear="2021" class="Insecta" family="Scarabaeidae" genus="Chalepides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalepides pantanalensis" order="Coleoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="pantanalensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="189">Chalepides pantanalensis</emphasis>
|
||
</taxonomicName>
|
||
Ratcliffe & Seidel, sp. nov.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="189">
|
||
<emphasis bold="true" pageId="0" pageNumber="189">Allotype.</emphasis>
|
||
Female. Length 16.7 mm; width 6.9 mm. As holotype except in the following respects:
|
||
<emphasis bold="true" pageId="0" pageNumber="189">Legs</emphasis>
|
||
: Protarsi simple, not enlarged.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="189" type="variation">
|
||
<paragraph pageId="0" pageNumber="189">Variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="189">Male (3 paratypes). Length 15.8-16.5 mm; width 6.8-7.3 mm. The paratypes do not differ from the holotype except in size, and the small tooth on the venter of tarsomere 5 is nearly obsolete in one specimen. Female (3 paratypes). Length 15.5-16.5 mm; width 7.0-7.7 mm. The paratypes do not differ from the allotype.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="189" type="etymology">
|
||
<paragraph pageId="0" pageNumber="189">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="189">The specific epithet is an adjective and refers to the habitat type, the Pantanal, in which this species occurs.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="189" type="distribution">
|
||
<paragraph pageId="0" pageNumber="189">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="189">
|
||
<taxonomicName authorityName="Ratcliffe & Seidel" authorityYear="2021" class="Insecta" family="Scarabaeidae" genus="Chalepides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalepides pantanalensis" order="Coleoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="pantanalensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="189">Chalepides pantanalensis</emphasis>
|
||
</taxonomicName>
|
||
is known only from
|
||
<normalizedToken originalValue="Brazil’s">Brazil's</normalizedToken>
|
||
Pantanal Region (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F4" captionText="Figure 5. Location of type localities of Fazenda 4 Cantos and Fazenda Natureza, Mato Grosso do Sul, Brazil" httpUri="https://binary.pensoft.net/fig/574484" pageId="0" pageNumber="189">5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption httpUri="https://binary.pensoft.net/fig/574484" pageId="0" pageNumber="189" start="Figure 5" startId="F4">
|
||
<paragraph pageId="0" pageNumber="189">
|
||
<emphasis bold="true" pageId="0" pageNumber="189">Figure 5.</emphasis>
|
||
Location of type localities of Fazenda 4 Cantos and Fazenda Natureza, Mato Grosso do Sul, Brazil
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="189" type="locality records">
|
||
<paragraph pageId="0" pageNumber="189">Locality records.</paragraph>
|
||
<paragraph pageId="0" pageNumber="189">8 specimens examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="189">
|
||
<emphasis bold="true" pageId="0" pageNumber="189">Brazil (8)</emphasis>
|
||
• MATO GROSSO DO SUL (8): Fazenda 4 Cantos, Fazenda Natureza (140 km NW Campo Grande).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="189" type="distribution">
|
||
<paragraph pageId="0" pageNumber="189">Temporal distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="189">May (3), November (5).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="189" type="natural history">
|
||
<paragraph pageId="0" pageNumber="189">Natural history.</paragraph>
|
||
<paragraph pageId="0" pageNumber="189">
|
||
Nothing is known of the life history of
|
||
<taxonomicName genus="C." lsidName="C. pantanalensis" pageId="0" pageNumber="189" rank="species" species="pantanalensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="189">C. pantanalensis</emphasis>
|
||
</taxonomicName>
|
||
other than the specimens were presumably collected at lights as are other species of
|
||
<taxonomicName authorityName="Casey" authorityYear="1915" class="Insecta" family="Scarabaeidae" genus="Chalepides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalepides" order="Coleoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="189">Chalepides</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |