290 lines
28 KiB
XML
290 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.873.35421" ID-GBIF-Dataset="ad4b6ae3-173c-494f-b761-56211fc3fdeb" ID-PMC="PMC6728363" ID-Pensoft-Pub="1313-2970-873-25" ID-Pensoft-UUID="AB7ECFA7FA38588E89AA94B67B6C6026" ID-PubMed="31534383" ID-ZooBank="6149E4233C2847BB89FC927209B5D2DE" ModsDocID="1313-2970-873-25" checkinTime="1567102845756" checkinUser="pensoft" docAuthor="Hosie, Andrew M., Fromont, Jane, Munyard, Kylie & Jones, Diana S." docDate="2019" docId="CBA07DDC8B85516BBC8CA34CC387C3D3" docLanguage="en" docName="ZooKeys 873: 25-42" docOrigin="ZooKeys 873" docSource="http://dx.doi.org/10.3897/zookeys.873.35421" docTitle="Membranobalanus Hoek 1913" docType="treatment" docVersion="4" id="AB7ECFA7FA38588E89AA94B67B6C6026" lastPageNumber="28" masterDocId="AB7ECFA7FA38588E89AA94B67B6C6026" masterDocTitle="Description of a new species of Membranobalanus (Crustacea, Cirripedia) from southern Australia" masterLastPageNumber="42" masterPageNumber="25" pageNumber="27" updateTime="1668167761432" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Description of a new species of Membranobalanus (Crustacea, Cirripedia) from southern Australia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hosie, Andrew M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fromont, Jane</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Munyard, Kylie</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jones, Diana S.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>873</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>25</mods:start>
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<mods:end>42</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.873.35421</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.873.35421</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-873-25</mods:identifier>
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<mods:identifier type="ZooBank">6149E4233C2847BB89FC927209B5D2DE</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159319360" LSID="urn:lsid:plazi:treatment:CBA07DDC8B85516BBC8CA34CC387C3D3" httpUri="http://treatment.plazi.org/id/CBA07DDC8B85516BBC8CA34CC387C3D3" lastPageId="4" lastPageNumber="28" pageId="2" pageNumber="27">
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<subSubSection pageId="2" pageNumber="27" type="nomenclature">
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<paragraph pageId="2" pageNumber="27">
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<taxonomicName LSID="CBA07DDC8B85516BBC8CA34CC387C3D3" authority="Hoek, 1913" authorityName="Hoek" authorityYear="1913" class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus" order="Sessilia" pageId="2" pageNumber="27" phylum="Arthropoda" rank="genus">Membranobalanus Hoek, 1913</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="27" type="type species">
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<paragraph pageId="2" pageNumber="27">Type species.</paragraph>
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<paragraph pageId="2" pageNumber="27">
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<taxonomicName class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus declivis" order="Sessilia" pageId="2" pageNumber="27" phylum="Arthropoda" rank="species" species="declivis">
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<emphasis italics="true" pageId="2" pageNumber="27">Balanus declivis</emphasis>
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</taxonomicName>
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Darwin, 1854: 275, pl. 7 fig. 4
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<normalizedToken originalValue="a–d">a-d</normalizedToken>
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; by subsequent designation (
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<bibRefCitation DOI="https://doi.org/10.5479/si.03629236.93.1" author="Pilsbry, HA" journalOrPublisher="Bulletin of the United States National Museum, Washington:" pageId="15" pageNumber="40" pagination="1 - 366" refId="B28" refString="Pilsbry, HA, 1916. The sessile barnacles (Cirripedia) contained in the collections of the U.S. National Museum; including a monograph of the American species. . Bulletin of the United States National Museum, Washington: 93: 1 - 366" title="The sessile barnacles (Cirripedia) contained in the collections of the U. S. National Museum; including a monograph of the American species." url="https://doi.org/10.5479/si.03629236.93.1" volume="93" year="1916">Pilsbry 1916</bibRefCitation>
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: 229).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="27" type="species composition">
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<paragraph pageId="2" pageNumber="27">Species composition.</paragraph>
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<paragraph pageId="2" pageNumber="27">
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<taxonomicName lsidName="M. brachialis" pageId="2" pageNumber="27" rank="species" species="brachialis">
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<emphasis italics="true" pageId="2" pageNumber="27">M. brachialis</emphasis>
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</taxonomicName>
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(Rosell, 1972);
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<taxonomicName lsidName="M. costatus" pageId="2" pageNumber="27" rank="species" species="costatus">
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<emphasis italics="true" pageId="2" pageNumber="27">M. costatus</emphasis>
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</taxonomicName>
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Zullo & Standing, 1983;
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<taxonomicName lsidName="M. cuneiformis" pageId="2" pageNumber="27" rank="species" species="cuneiformis">
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<emphasis italics="true" pageId="2" pageNumber="27">M. cuneiformis</emphasis>
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</taxonomicName>
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(Hiro, 1936);
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<taxonomicName lsidName="M. declivis" pageId="2" pageNumber="27" rank="species" species="declivis">
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<emphasis italics="true" pageId="2" pageNumber="27">M. declivis</emphasis>
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</taxonomicName>
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(Darwin, 1854);
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<taxonomicName lsidName="M. koreanus" pageId="2" pageNumber="27" rank="species" species="koreanus">
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<emphasis italics="true" pageId="2" pageNumber="27">M. koreanus</emphasis>
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</taxonomicName>
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Kim & Kim, 1983;
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<taxonomicName lsidName="M. longirostrum" pageId="2" pageNumber="27" rank="species" species="longirostrum">
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<emphasis italics="true" pageId="2" pageNumber="27">M. longirostrum</emphasis>
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</taxonomicName>
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(Hoek, 1913);
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<taxonomicName lsidName="M. nebrias" pageId="2" pageNumber="27" rank="species" species="nebrias">
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<emphasis italics="true" pageId="2" pageNumber="27">M. nebrias</emphasis>
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</taxonomicName>
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(Zullo & Beach, 1973);
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<taxonomicName lsidName="M. orcutti" pageId="2" pageNumber="27" rank="species" species="orcutti">
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<emphasis italics="true" pageId="2" pageNumber="27">M. orcutti</emphasis>
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</taxonomicName>
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(Pilsbry, 1907);
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<taxonomicName lsidName="M. porphyrophilus" pageId="2" pageNumber="27" rank="species" species="porphyrophilus">
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<emphasis italics="true" pageId="2" pageNumber="27">M. porphyrophilus</emphasis>
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</taxonomicName>
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sp. nov.;
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<taxonomicName lsidName="M. robinae" pageId="2" pageNumber="27" rank="species" species="robinae">
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<emphasis italics="true" pageId="2" pageNumber="27">M. robinae</emphasis>
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</taxonomicName>
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Van Syoc, 1988.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="27" type="nomen dubium">
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<paragraph pageId="2" pageNumber="27">Nomen dubium.</paragraph>
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<paragraph pageId="2" pageNumber="27">
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<taxonomicName lsidName="M. orcuttiformis" pageId="2" pageNumber="27" rank="species" species="orcuttiformis">
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<emphasis italics="true" pageId="2" pageNumber="27">M. orcuttiformis</emphasis>
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</taxonomicName>
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(
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<normalizedToken originalValue="Kolosváry">Kolosvary</normalizedToken>
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, 1941).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="27" type="diagnosis">
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<paragraph pageId="2" pageNumber="27">Diagnosis.</paragraph>
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<paragraph pageId="2" pageNumber="27">Parietes solid, unornamented, weakly articulated, basis membranous. Rostrum scoop or boat-shaped, often elongate relative to other parietes. Tergum with spur furrow open. Cirrus IV with erect spines, with or without recurved teeth on anterior ramus.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="29" pageId="2" pageNumber="27" type="remarks">
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<paragraph pageId="2" pageNumber="27">Remarks.</paragraph>
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<paragraph lastPageId="3" lastPageNumber="28" pageId="2" pageNumber="27">
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With the addition of the below described species, there are now 10 species included within
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<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus" order="Sessilia" pageId="2" pageNumber="27" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="2" pageNumber="27">Membranobalanus</emphasis>
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</taxonomicName>
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.
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<bibRefCitation author="Utinomi, H" journalOrPublisher="Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening" pageId="15" pageNumber="40" pagination="161 - 186" refId="B39" refString="Utinomi, H, 1968. Pelagic, shelf and shallow water cirripedia from the Indo West Pacific. . Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening 131: 161 - 186" title="Pelagic, shelf and shallow water cirripedia from the Indo West Pacific." volume="131" year="1968">Utinomi (1968)</bibRefCitation>
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synonymised the taxa
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<taxonomicName class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Balanus (Membranobalanus) longirostrum var. krusadaiensis" order="Sessilia" pageId="2" pageNumber="27" phylum="Arthropoda" rank="variety" species="longirostrum" subGenus="Membranobalanus" variety="krusadaiensis">Balanus (Membranobalanus) longirostrum var. krusadaiensis</taxonomicName>
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Daniel, 1955,
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<taxonomicName lsidName="B. (M.) basicupula" pageId="2" pageNumber="27" rank="species" species="basicupula" subGenus="M.">B. (M.) basicupula</taxonomicName>
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Suhaimi, 1966, and
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<taxonomicName lsidName="B. (M.) roonwali" pageId="2" pageNumber="27" rank="species" species="roonwali" subGenus="M.">B. (M.) roonwali</taxonomicName>
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Prem-Kumar & Daniel, 1968 under
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<taxonomicName lsidName="M. longirostrum" pageId="3" pageNumber="28" rank="species" species="longirostrum">
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<emphasis italics="true" pageId="3" pageNumber="28">
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<pageBreakToken pageId="3" pageNumber="28" start="start">M</pageBreakToken>
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. longirostrum
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</emphasis>
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</taxonomicName>
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, proposing that the differences observed are within the bounds of intraspecific variability. Recently,
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<bibRefCitation pageId="3" pageNumber="28" refId="B41">van Syoc et al. (2015)</bibRefCitation>
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transferred
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<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Acasta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acasta acuta" order="Sessilia" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="acuta">
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<emphasis italics="true" pageId="3" pageNumber="28">Acasta acuta</emphasis>
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</taxonomicName>
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(Kolbasov, 1993) out of
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<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus" order="Sessilia" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="28">Membranobalanus</emphasis>
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</taxonomicName>
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based primarily on the presence of calcareous spines on the parietal wall, a character no other
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<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus" order="Sessilia" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="28">Membranobalanus</emphasis>
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</taxonomicName>
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possess and in reference to a cladistic analysis in an unpublished thesis. The fact that
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<taxonomicName lsidName="A. acuta" pageId="3" pageNumber="28" rank="species" species="acuta">
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<emphasis italics="true" pageId="3" pageNumber="28">A. acuta</emphasis>
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</taxonomicName>
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is found in sponges of the family
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<taxonomicName family="Petrosiidae" lsidName="" pageId="3" pageNumber="28" rank="family">Petrosiidae</taxonomicName>
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(order
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<taxonomicName lsidName="Balanoidea" order="Haplosclerida" pageId="3" pageNumber="28" rank="order">Haplosclerida</taxonomicName>
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), not the
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<taxonomicName family="Clionaidae" lsidName="" pageId="3" pageNumber="28" rank="family">Clionaidae</taxonomicName>
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as reported for the remaining members of the genus, separates this species ecologically from
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<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus" order="Sessilia" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="28">Membranobalanus</emphasis>
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</taxonomicName>
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, was used as further justification. The general appearance, membranous basis and elongated rostrum of
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<taxonomicName lsidName="A. acuta" pageId="3" pageNumber="28" rank="species" species="acuta">
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<emphasis italics="true" pageId="3" pageNumber="28">A. acuta</emphasis>
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</taxonomicName>
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are typical features of
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<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus" order="Sessilia" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="28">Membranobalanus</emphasis>
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</taxonomicName>
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, however. While we treat this reassignment with caution, we have no evidence with which to dispute it.
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</paragraph>
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<paragraph pageId="3" pageNumber="28">
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<bibRefCitation author="Kolosvary, G" journalOrPublisher="Monitore Zoologico Italiano" pageId="15" pageNumber="40" pagination="183 - 195" refId="B19" refString="Kolosvary, G, 1941. Revisione della collezione de Balanidi del Museo Zoologico della R. Universita de Firenze. . Monitore Zoologico Italiano 52: 183 - 195" title="Revisione della collezione de Balanidi del Museo Zoologico della R. Universita de Firenze." volume="52" year="1941">
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<normalizedToken originalValue="Kolosváry">Kolosvary</normalizedToken>
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(1941)
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</bibRefCitation>
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described
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<taxonomicName class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus (M.) orcuttiformis" order="Sessilia" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="orcuttiformis" subGenus="M.">Balanus (M.) orcuttiformis</taxonomicName>
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based on the parietes of a single empty specimen. The locality details of the specimen are vague, only given as "India Orient.", but presumably meaning eastern India. No detail regarding a host was given. The description is very brief, giving very few clues to the identification of this species, and the only illustration of the specimen, in lateral view, could belong to a number of genera, but not
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<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus" order="Sessilia" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="28">Membranobalanus</emphasis>
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</taxonomicName>
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as currently defined. Most notable is the absence of any elongation of the rostrum and the largely horizontal basal rim of the parietal wall gives the appearance that it was attached to, rather than embedded within, a substrate. Additionally, the exceptionally broad alae and absent radii are reminiscent of the
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<taxonomicName lsidName="Pachylasmatoidea" pageId="3" pageNumber="28" rank="superfamily" superfamily="Pachylasmatoidea">Pachylasmatoidea</taxonomicName>
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, which possess solid parietes and often have a membranous base (see
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<bibRefCitation author="Jones, DS" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle" pageId="15" pageNumber="40" pagination="141 - 283" refId="B16" refString="Jones, DS, 2000. CrustaceaCirripediaThoracica: Chionelasmatoidea and Pachylasmatoidea (Balanomorpha) of New Caledonia, Vanuatu and Wallis and Futuna islands, with a review of all currently assigned taxa. . Memoires du Museum National d'Histoire Naturelle 184: 141 - 283" title="CrustaceaCirripediaThoracica: Chionelasmatoidea and Pachylasmatoidea (Balanomorpha) of New Caledonia, Vanuatu and Wallis and Futuna islands, with a review of all currently assigned taxa." volume="184" year="2000">Jones 2000</bibRefCitation>
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).
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<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus orcuttiformis" order="Sessilia" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="orcuttiformis">
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<emphasis italics="true" pageId="3" pageNumber="28">Membranobalanus orcuttiformis</emphasis>
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</taxonomicName>
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has not been recorded since its description and unfortunately the specimens are missing from the Museo di Storia Naturale
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<normalizedToken originalValue="dell’Università">dell'Universita</normalizedToken>
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di Firenze, Italy and thus cannot be reexamined (
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<bibRefCitation author="Innocenti, G" journalOrPublisher="Atti della Societa Toscana di Scienze Naturali, Memorie serie B" pageId="15" pageNumber="40" pagination="1 - 11" publicationUrl="http://www.stsn.it/images/pdf/serB113/01.pdf" refId="B14" refString="Innocenti, G, 2006. Collections of the Natural History Museum, Zoological Section "La Specola" of the University of Florence. XXIII. Crustacea, Class Maxillopoda, Subclass Thecostraca, Infraclass Cirripedia. . Atti della Societa Toscana di Scienze Naturali, Memorie serie B 113: 1 - 11" title="Collections of the Natural History Museum, Zoological Section " La Specola " of the University of Florence. XXIII. Crustacea, Class Maxillopoda, Subclass Thecostraca, Infraclass Cirripedia." url="http://www.stsn.it/images/pdf/serB113/01.pdf" volume="113" year="2006">Innocenti 2006</bibRefCitation>
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). For these reasons, this species is considered herein a
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<emphasis italics="true" pageId="3" pageNumber="28">nomen dubium</emphasis>
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and has been excluded from the key below.
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</paragraph>
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<paragraph lastPageId="4" lastPageNumber="29" pageId="3" pageNumber="28">
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The remaining
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<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus" order="Sessilia" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="28">Membranobalanus</emphasis>
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</taxonomicName>
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species can be separated into two morphological lineages, approximating an American centred group and an Indo-West Pacific group. The former have recurved teeth, similar to those present in some members of the
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<taxonomicName lsidName="" pageId="3" pageNumber="28" rank="subfamily" subfamily="Acastinae">Acastinae</taxonomicName>
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, as well as erect spines on cirrus IV, smooth growth lines on the scutum and the articular ridge and groove of the scutum is prominent, extending well beyond the articular margin, with a correspondingly wide articular groove on the tergum. The latter group bears only the erect spines on cirrus IV, finely striated growth lines, and has relatively weak articular structures on the opercular plates. From a biogeographic perspective one species disrupts this pattern:
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<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus koreanus" order="Sessilia" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="koreanus">
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<emphasis italics="true" pageId="3" pageNumber="28">Membranobalanus koreanus</emphasis>
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</taxonomicName>
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from the waters around the Korean Peninsula. As described and figured by
|
||
<bibRefCitation author="Kim, HS" journalOrPublisher="Korean Journal of Zoology" pageId="15" pageNumber="40" pagination="41 - 47" refId="B17" refString="Kim, HS, Kim, IH, 1983. A new species of MembranobalanusCrustaceaCirripedia from Korean waters. . Korean Journal of Zoology 26: 41 - 47" title="A new species of MembranobalanusCrustaceaCirripedia from Korean waters." volume="26" year="1983">Kim and Kim (1983)</bibRefCitation>
|
||
,
|
||
<taxonomicName lsidName="M. koreanus" pageId="3" pageNumber="28" rank="species" species="koreanus">
|
||
<emphasis italics="true" pageId="3" pageNumber="28">M. koreanus</emphasis>
|
||
</taxonomicName>
|
||
has recurved teeth and a large articular ridge on the scutum. The records of
|
||
<taxonomicName lsidName="M. orcutti" pageId="3" pageNumber="28" rank="species" species="orcutti">
|
||
<emphasis italics="true" pageId="3" pageNumber="28">M. orcutti</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="Barnard, KH" journalOrPublisher="Annals of the South African Museum" pageId="14" pageNumber="39" pagination="1 - 103" publicationUrl="https://biodiversitylibrary.org/page/11101069" refId="B2" refString="Barnard, KH, 1924. Contributions to the crustacean fauna of South Africa. No. 7, Cirripedia. . Annals of the South African Museum 20: 1 - 103" title="Contributions to the crustacean fauna of South Africa. No. 7, Cirripedia." url="https://biodiversitylibrary.org/page/11101069" volume="20" year="1924">Barnard (1924)</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Rosell, NC" journalOrPublisher="University of the Phillippines Natural Science Research Center Technical Report" pageId="15" pageNumber="40" pagination="1 - 12" refId="B31" refString="Rosell, NC, 1973. On two less well-known balanids (Cirripedia, Thoracica) from the Sulu Archipelago, Philippines. . University of the Phillippines Natural Science Research Center Technical Report 4: 1 - 12" title="On two less well-known balanids (Cirripedia, Thoracica) from the Sulu Archipelago, Philippines." volume="4" year="1973">Rosell (1973</bibRefCitation>
|
||
,
|
||
<bibRefCitation DOI="https://doi.org/10.1163/156854075X00216" author="Rosell, NC" journalOrPublisher="Crustaceana" pageId="15" pageNumber="40" pagination="206 - 214" refId="B32" refString="Rosell, NC, 1975. On two noteworthy balanids (CirripediaThoracica) from the Sulu Archipelago, Philippines. . Crustaceana 29: 206 - 214" title="On two noteworthy balanids (CirripediaThoracica) from the Sulu Archipelago, Philippines." url="https://doi.org/10.1163/156854075X00216" volume="29" year="1975">1975</bibRefCitation>
|
||
) from South Africa and the Sulu Archipelago, respectively, were considered suspect by
|
||
<bibRefCitation author="Zullo, VA" journalOrPublisher="Los Angeles County Natural History Museum Contributions in Science" pageId="16" pageNumber="41" pagination="1 - 16" publicationUrl="https://biodiversitylibrary.org/page/52110555" refId="B46" refString="Zullo, VA, Beach, DB, 1973. New species of Membranobalanus Hoek and Hexacreusia Zullo (Cirripedia, Balanidae) from the Galapagos Archipelago. . Los Angeles County Natural History Museum Contributions in Science 249: 1 - 16" title="New species of Membranobalanus Hoek and Hexacreusia Zullo (Cirripedia, Balanidae) from the Galapagos Archipelago." url="https://biodiversitylibrary.org/page/52110555" volume="249" year="1973">Zullo and Beach (1973)</bibRefCitation>
|
||
and
|
||
<bibRefCitation DOI="https://doi.org/10.1163/156854099503528" author="Van Syoc, RJ" journalOrPublisher="Crustaceana" pageId="15" pageNumber="40" pagination="467 - 486" refId="B42" refString="Van Syoc, RJ, Winther, R, 1999. Sponge-inhabiting barnacles of the Americas: a new species of Acasta (Cirripedia, Archaeobalanidae), first record from the Eastern Pacific, including discussion of the evolution of cirral morphology. . Crustaceana 72: 467 - 486" title="Sponge-inhabiting barnacles of the Americas: a new species of Acasta (Cirripedia, Archaeobalanidae), first record from the Eastern Pacific, including discussion of the evolution of cirral morphology." url="https://doi.org/10.1163/156854099503528" volume="72" year="1999">Van Syoc and Winther (1999)</bibRefCitation>
|
||
. The Sulu Archipelago specimens lack the recurved teeth on cirrus IV of the specimens described from Catalina Island by
|
||
<bibRefCitation author="Zullo, VA" journalOrPublisher="Los Angeles County Natural History Museum Contributions in Science" pageId="16" pageNumber="41" pagination="1 - 16" publicationUrl="https://biodiversitylibrary.org/page/52110555" refId="B46" refString="Zullo, VA, Beach, DB, 1973. New species of Membranobalanus Hoek and Hexacreusia Zullo (Cirripedia, Balanidae) from the Galapagos Archipelago. . Los Angeles County Natural History Museum Contributions in Science 249: 1 - 16" title="New species of Membranobalanus Hoek and Hexacreusia Zullo (Cirripedia, Balanidae) from the Galapagos Archipelago." url="https://biodiversitylibrary.org/page/52110555" volume="249" year="1973">Zullo and Beach (1973)</bibRefCitation>
|
||
, and while
|
||
<normalizedToken originalValue="Barnard’s">Barnard's</normalizedToken>
|
||
description specifically mentions recurved teeth on cirrus IV, most of his description is deferred back to either
|
||
<bibRefCitation author="Pilsbry, HA" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="15" pageNumber="40" pagination="360 - 362" publicationUrl="https://www.jstor.org/stable/4063151" refId="B27" refString="Pilsbry, HA, 1907. Notes on some Pacific cirripedes. . Proceedings of the Academy of Natural Sciences of Philadelphia 59: 360 - 362" title="Notes on some Pacific cirripedes." url="https://www.jstor.org/stable/4063151" volume="59" year="1907">
|
||
<normalizedToken originalValue="Pilsbry’s">Pilsbry's</normalizedToken>
|
||
(1907
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation DOI="https://doi.org/10.5479/si.03629236.93.1" author="Pilsbry, HA" journalOrPublisher="Bulletin of the United States National Museum, Washington:" pageId="15" pageNumber="40" pagination="1 - 366" refId="B28" refString="Pilsbry, HA, 1916. The sessile barnacles (Cirripedia) contained in the collections of the U.S. National Museum; including a monograph of the American species. . Bulletin of the United States National Museum, Washington: 93: 1 - 366" title="The sessile barnacles (Cirripedia) contained in the collections of the U. S. National Museum; including a monograph of the American species." url="https://doi.org/10.5479/si.03629236.93.1" volume="93" year="1916">1916</bibRefCitation>
|
||
) descriptions of
|
||
<taxonomicName lsidName="M. orcutti" pageId="3" pageNumber="28" rank="species" species="orcutti">
|
||
<emphasis italics="true" pageId="3" pageNumber="28">M. orcutti</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12959" author="Hoek, PPC" journalOrPublisher="" Japan Journal of Zoology" pageId="14" pageNumber="39" refId="B12" refString="Hoek, PPC, 1913. . https://doi.org/10.5962/bhl.title.12959" url="https://doi.org/10.5962/bhl.title.12959" year="1913">
|
||
<normalizedToken originalValue="Hoek’s">Hoek's</normalizedToken>
|
||
(1913)
|
||
</bibRefCitation>
|
||
description of
|
||
<taxonomicName lsidName="M. longirostrum." pageId="3" pageNumber="28" rank="species" species="longirostrum.">
|
||
<emphasis italics="true" pageId="3" pageNumber="28">M. longirostrum.</emphasis>
|
||
</taxonomicName>
|
||
Of particular note in
|
||
<normalizedToken originalValue="Barnard’s">Barnard's</normalizedToken>
|
||
<pageBreakToken pageId="4" pageNumber="29" start="start">description</pageBreakToken>
|
||
is that the scutum has an external, setose membrane, a character seen in some species of the
|
||
<taxonomicName lsidName="" pageId="4" pageNumber="29" rank="subfamily" subfamily="Acastinae">Acastinae</taxonomicName>
|
||
, but not
|
||
<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus" order="Sessilia" pageId="4" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="4" pageNumber="29">Membranobalanus</emphasis>
|
||
</taxonomicName>
|
||
. Both reports should be considered
|
||
<emphasis italics="true" pageId="4" pageNumber="29">species inquirenda</emphasis>
|
||
, but potentially represent previously undescribed species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |