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<document ID-DOI="http://dx.doi.org/10.3897/herpetozoa.35.e89981" ID-Pensoft-Pub="2682-955X-35-165" ID-Pensoft-UUID="1ED262552EBA5895BADC209AC1568367" ID-ZooBank="A229882246FC40778F667A0E7840C174" ModsDocID="2682-955X-35-165" checkinTime="1664443303874" checkinUser="pensoft" docAuthor="Lin, Shi-Shi, Li, Yuan-Hang, Lu, Yu-Hong, Su, Hong-Lin, Wu, Shi-Bin, Zhang, Qi-Qi, Mo, Mei-Juan, Xiao, Shao-Jun, Pan, Zhong, Pan, Hu-Jun, Zeng, Zhao-Chi &amp; Wang, Jian" docDate="2022" docId="C9D5F78A54F95D30A081D3E79EDF2E9D" docLanguage="en" docName="Herpetozoa 35: 165-178" docOrigin="Herpetozoa 35" docPubDate="2022-09-28" docSource="http://dx.doi.org/10.3897/herpetozoa.35.e89981" docTitle="Leptobrachella verrucosa Wang, Zeng, Lin &amp; Li 2022, sp. nov." docType="treatment" docUuid="D968B6DE-22B4-48D3-B3EC-DAFDB328BBD2" docUuidSource="ZooBank" docVersion="2" id="1ED262552EBA5895BADC209AC1568367" lastPageNumber="165" masterDocId="1ED262552EBA5895BADC209AC1568367" masterDocTitle="A new species of the genus Leptobrachella (Anura, Megophryidae) from northwestern Guangdong Province, China" masterLastPageNumber="178" masterPageNumber="165" pageNumber="165" updateTime="1664443605831" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of the genus Leptobrachella (Anura, Megophryidae) from northwestern Guangdong Province, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lin, Shi-Shi</mods:namePart>
<mods:affiliation>Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Yuan-Hang</mods:namePart>
<mods:affiliation>Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lu, Yu-Hong</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6498-4155</mods:nameIdentifier>
<mods:affiliation>Nanfang College · GuangZhou, Guangzhou 510970, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Su, Hong-Lin</mods:namePart>
<mods:affiliation>Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wu, Shi-Bin</mods:namePart>
<mods:affiliation>Guangdong Zhongke Environmental Technology Development Co., Ltd, Guangzhou 510275, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhang, Qi-Qi</mods:namePart>
<mods:affiliation>Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mo, Mei-Juan</mods:namePart>
<mods:affiliation>Guangdong Lianshan Bijiashan Provincial Nature Reserve, Qingyuan 513200, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Xiao, Shao-Jun</mods:namePart>
<mods:affiliation>Guangdong Lianshan Bijiashan Provincial Nature Reserve, Qingyuan 513200, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pan, Zhong</mods:namePart>
<mods:affiliation>Guangzhou Shengheng Forestry Co., Ltd, Guangzhou 510663, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pan, Hu-Jun</mods:namePart>
<mods:affiliation>Guangzhou Shengheng Forestry Co., Ltd, Guangzhou 510663, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zeng, Zhao-Chi</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4054-8399</mods:nameIdentifier>
<mods:affiliation>Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
<mods:nameIdentifier type="email">zhaochizeng@outlook.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wang, Jian</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4249-7767</mods:nameIdentifier>
<mods:affiliation>Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
<mods:nameIdentifier type="email">wangj-1994@outlook.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Herpetozoa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-09-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>35</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>165</mods:start>
<mods:end>178</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/herpetozoa.35.e89981</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/herpetozoa.35.e89981</mods:identifier>
<mods:identifier type="Pensoft-Pub">2682-955X-35-165</mods:identifier>
<mods:identifier type="ZooBank">A229882246FC40778F667A0E7840C174</mods:identifier>
<mods:identifier type="Pensoft-UUID">1ED262552EBA5895BADC209AC1568367</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:D968B6DE-22B4-48D3-B3EC-DAFDB328BBD2" httpUri="http://treatment.plazi.org/id/C9D5F78A54F95D30A081D3E79EDF2E9D" lastPageNumber="165" pageId="0" pageNumber="165">
<subSubSection pageId="0" pageNumber="165" type="nomenclature">
<paragraph pageId="0" pageNumber="165">
<taxonomicName LSID="https://zoobank.org/D968B6DE-22B4-48D3-B3EC-DAFDB328BBD2" authority="Wang, Zeng, Lin &amp; Li" authorityName="Wang, Zeng, Lin &amp; Li" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella verrucosa" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="verrucosa" status="sp. nov.">Leptobrachella verrucosa Wang, Zeng, Lin &amp; Li</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="165">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="description">
<paragraph pageId="0" pageNumber="165">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphological features of the holotype of Leptobrachella verrucosa sp. nov., GEP a 062 in life: A. Dorsal view; B. Dorsolateral view; C. Lateral view; D. Ventral view; E. Ventral view of hand; F. Ventral view of foot." figureDoi="10.3897/herpetozoa.35.e89981.figure3" httpUri="https://binary.pensoft.net/fig/750072" pageId="0" pageNumber="165">Figs 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Morphological features of the paratypes of Leptobrachella verrucosa sp. nov., in life: A. GEP a 059 (A 1 dorsal view, A 2 lateral view, A 3 ventral view); B. GEP a 060 (B 1 dorsal view, B 2 dorsolateral view, B 3 ventral view); C. GEP a 061 (C 1 dorsal view, C 2 dorsolateral view, C 3 ventral view); D. GEP a 063 (D 1 dorsal view, D 2 dorsolateral view, D 3 ventral view)." figureDoi="10.3897/herpetozoa.35.e89981.figure4" httpUri="https://binary.pensoft.net/fig/750073" pageId="0" pageNumber="165">, 4 Verrucous Leaf Litter Toad (in English) / Mi You Zhang Tu Chan (密疣掌突蟾 in Chinese)</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="type material">
<paragraph pageId="0" pageNumber="165">Type material.</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Holotype</emphasis>
♂. GEP a062, collected by Shi-Shi Lin and Qi-Qi Zhang on 31 May 2022 from Lianshan Bijiashan Nature Reserve (
<geoCoordinate degrees="24" direction="north" minutes="12" orientation="latitude" precision="1" seconds="49.63" value="24.213787">24°12'49.63&quot;N</geoCoordinate>
,
<geoCoordinate degrees="111" direction="east" minutes="59" orientation="longitude" precision="1" seconds="39.31" value="111.99425">111°59'39.31&quot;E</geoCoordinate>
; ca. 550 m a.s.l.), Lianshan County, Qingyuan City, Guangdong Province, China.
</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">
<emphasis italics="true" pageId="0" pageNumber="165">Paratypes</emphasis>
.
</emphasis>
4 ♂: GEP a059-061, 063, the same collection data as the holotype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="etymology">
<paragraph pageId="0" pageNumber="165">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="165">
The specific epithet &quot;
<taxonomicName authorityName="Wang, Zeng, Lin &amp; Li" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella verrucosa" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="verrucosa">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella verrucosa</emphasis>
</taxonomicName>
&quot; is an adjective meaning
<normalizedToken originalValue="“verrucous”">&quot;verrucous&quot;</normalizedToken>
, referring to the tuberculate dorsal skin texture of the new species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="diagnosis">
<paragraph pageId="0" pageNumber="165">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="165">(1) Small body size [SVL 23.2-25.9 mm in five adult males], (2) iris bicolored, upper half coppery orange and lower half grayish brown, (3) tympanum distinct, (4) supratympanic line black, (5) fingers without lateral fringes, (6) toes with rudimentary webbing, lateral fringes narrow, (7) longitudinal ridges under toes continuous, (8) heels not meeting when adpressed, tibial-tarsal articulation reaching anterior corner of eye, (9) dorsal surface shagreened with numerous conical tubercles, lacking spines, enlarged warts or skin ridges, (10) ventral surface smooth, (11) dorsum grayish brown, with distinct darker brown scattered markings and irregular light orange pigmentations, (12) flanks with several dark spots, (13) ventral surface creamy white with grayish white and dark brown spots.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="description">
<paragraph pageId="0" pageNumber="165">Description of holotype.</paragraph>
<paragraph pageId="0" pageNumber="165">Adult male. Body size small, SVL 23.2 mm. Head length slightly larger than head width, HDW/HDL 0.95; snout slightly protruding, projecting slightly beyond margin of lower jaw; nostril closer to snout than eye; canthus rostralis gently rounded; loreal region slightly concave; interorbital space flat, internarial distance slightly larger than interorbital distance, IND/IOD 1.05; pineal ocellus absent; pupil vertical; snout slightly longer than eye diameter, SNT/EYE 1.03; tympanum distinct, rounded, diameter smaller than that of eye and larger than tympanum-eye distance, TMP/EYE 0.54, TEY/TMP 0.46; upper margin of tympanum in contact with supratympanic ridge; vomerine teeth absent; a single vocal sac; vocal sac openings slit-like, paired, located posterolaterally on floor of mouth, close to margins of mandible; tongue deeply notched distally; supratympanic ridge distinct, extending from posterior corner of eye to supra-axillary gland.</paragraph>
<paragraph pageId="0" pageNumber="165">Tips of fingers rounded, slightly swollen; relative finger lengths I = II = IV &lt;III; nuptial pad absent; subarticular tubercles absent; inner palmar tubercle large, rounded, distinctly separated from outer palmar tubercle; outer palmar tubercle small, rounded; fingers lacking interdigital webbing and lateral fringes. Tips of toes rounded, slightly swollen; relative toe length I &lt;II &lt;V &lt;III &lt;IV; longitudinal ridges under toes continuous; inner metatarsal tubercle large, oval; outer metatarsal tubercle absent; interdigital webbing between toes rudimentary; narrow lateral fringes present on all toes. Tibia 49% of snout-vent length; tibiotarsal articulation reaches to anterior corner of eye; heels not meeting when thighs are appressed at right angles with respect to the body.</paragraph>
<paragraph pageId="0" pageNumber="165">Dorsal surface shagreened with numerous conical tubercles, lacking spines, enlarged warts or skin ridges; ventral skin smooth; pectoral and femoral glands oval; both larger in diameter than tip of fingers; femoral glands larger in diameter than tip of toes; femoral gland situated on posteroventral surface of thigh, closer to knee than to vent; supra-axillary gland raised. Ventrolateral gland distinctly visible, forming a longitudinal uncontinuous series.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="coloration of holotype in life">
<paragraph pageId="0" pageNumber="165">Coloration of holotype in life.</paragraph>
<paragraph pageId="0" pageNumber="165">Dorsal background color grayish brown, with distinct dark brown markings, and rounded spots, and scattered with irregular light orange pigmentation. A dark brown inverted triangular marking in interorbital region, connecting to dark brown W-shaped marking on occipital region. Tympanum dark brown, lower margin grayish white. Supratympanic line black. A pair of dark brown vertical bars present under eyes; transverse dark brown bars on dorsal surfaces of distal limbs and digits; distinct dark brown blotches on flanks; surfaces of elbows and upper arms coppery orange, with dark markings and spots.</paragraph>
<paragraph pageId="0" pageNumber="165">Surface of throat, chest, belly and underside of forelimbs creamy white, with distinct grayish white and dark brown spots; under side of hindlimbs grayish purple with numerous white spots. Supra-axillary gland coppery orange, pectoral glands grayish white, femoral glands coppery orange, anterior half of ventrolateral glands coppery and posterior half grayish white. Iris bicolored, upper half coppery orange, lower half grayish brown.</paragraph>
<caption doi="10.3897/herpetozoa.35.e89981.figure3" httpUri="https://binary.pensoft.net/fig/750072" pageId="0" pageNumber="165" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figure 3.</emphasis>
Morphological features of the holotype of
<taxonomicName authorityName="Wang, Zeng, Lin &amp; Li" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella verrucosa" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="verrucosa">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella verrucosa</emphasis>
</taxonomicName>
sp. nov., GEP a062 in life:
<emphasis bold="true" pageId="0" pageNumber="165">A.</emphasis>
Dorsal view;
<emphasis bold="true" pageId="0" pageNumber="165">B.</emphasis>
Dorsolateral view;
<emphasis bold="true" pageId="0" pageNumber="165">C.</emphasis>
Lateral view;
<emphasis bold="true" pageId="0" pageNumber="165">D.</emphasis>
Ventral view;
<emphasis bold="true" pageId="0" pageNumber="165">E.</emphasis>
Ventral view of hand;
<emphasis bold="true" pageId="0" pageNumber="165">F.</emphasis>
Ventral view of foot.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="coloration of holotype in preservative">
<paragraph pageId="0" pageNumber="165">Coloration of holotype in preservative.</paragraph>
<paragraph pageId="0" pageNumber="165">Dorsal background color dark brown scattered with irregular gray pigmentations. All markings, bars and spots become indistinct. Tympanum dark brown.</paragraph>
<paragraph pageId="0" pageNumber="165">Ventral surface grayish white; grayish white spots absent, dark brown spots become more distinct. Supra-axillary, pectoral, and ventrolateral glands greyish white.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="variation">
<paragraph pageId="0" pageNumber="165">Variation.</paragraph>
<paragraph pageId="0" pageNumber="165">
Measurements and body proportions are listed in Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Measurements (minimum-maximum (mean + / - SD); in mm), and body proportions of Leptobrachella verrucosa sp. nov." httpUri="http://table.plazi.org/id/2C1C34C3DBB6703267F334C059C0BA57" pageId="0" pageNumber="165" tableUuid="2C1C34C3DBB6703267F334C059C0BA57">2</tableCitation>
. The paratypes, GEP a059 (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Morphological features of the paratypes of Leptobrachella verrucosa sp. nov., in life: A. GEP a 059 (A 1 dorsal view, A 2 lateral view, A 3 ventral view); B. GEP a 060 (B 1 dorsal view, B 2 dorsolateral view, B 3 ventral view); C. GEP a 061 (C 1 dorsal view, C 2 dorsolateral view, C 3 ventral view); D. GEP a 063 (D 1 dorsal view, D 2 dorsolateral view, D 3 ventral view)." figureDoi="10.3897/herpetozoa.35.e89981.figure4" httpUri="https://binary.pensoft.net/fig/750073" pageId="0" pageNumber="165">4A</figureCitation>
), GEP a060 (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Morphological features of the paratypes of Leptobrachella verrucosa sp. nov., in life: A. GEP a 059 (A 1 dorsal view, A 2 lateral view, A 3 ventral view); B. GEP a 060 (B 1 dorsal view, B 2 dorsolateral view, B 3 ventral view); C. GEP a 061 (C 1 dorsal view, C 2 dorsolateral view, C 3 ventral view); D. GEP a 063 (D 1 dorsal view, D 2 dorsolateral view, D 3 ventral view)." figureDoi="10.3897/herpetozoa.35.e89981.figure4" httpUri="https://binary.pensoft.net/fig/750073" pageId="0" pageNumber="165">4B</figureCitation>
), and GEP a063 (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Morphological features of the paratypes of Leptobrachella verrucosa sp. nov., in life: A. GEP a 059 (A 1 dorsal view, A 2 lateral view, A 3 ventral view); B. GEP a 060 (B 1 dorsal view, B 2 dorsolateral view, B 3 ventral view); C. GEP a 061 (C 1 dorsal view, C 2 dorsolateral view, C 3 ventral view); D. GEP a 063 (D 1 dorsal view, D 2 dorsolateral view, D 3 ventral view)." figureDoi="10.3897/herpetozoa.35.e89981.figure4" httpUri="https://binary.pensoft.net/fig/750073" pageId="0" pageNumber="165">4D</figureCitation>
), show lighter coloration of markings and pigmentations on dorsum. The paratype, GEP a061 (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Morphological features of the paratypes of Leptobrachella verrucosa sp. nov., in life: A. GEP a 059 (A 1 dorsal view, A 2 lateral view, A 3 ventral view); B. GEP a 060 (B 1 dorsal view, B 2 dorsolateral view, B 3 ventral view); C. GEP a 061 (C 1 dorsal view, C 2 dorsolateral view, C 3 ventral view); D. GEP a 063 (D 1 dorsal view, D 2 dorsolateral view, D 3 ventral view)." figureDoi="10.3897/herpetozoa.35.e89981.figure4" httpUri="https://binary.pensoft.net/fig/750073" pageId="0" pageNumber="165">4C</figureCitation>
), possesses a more tuberculate dorsal skin texture. Pectoral glands much more distinct in all paratypes.
</paragraph>
<caption ID-Table-UUID="2C1C34C3DBB6703267F334C059C0BA57" httpUri="http://table.plazi.org/id/2C1C34C3DBB6703267F334C059C0BA57" pageId="0" pageNumber="165" start="Table 2" startId="T2">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Table 2.</emphasis>
Measurements (minimum-maximum (mean
<normalizedToken originalValue="±">+/-</normalizedToken>
SD); in mm), and body proportions of
<taxonomicName authorityName="Wang, Zeng, Lin &amp; Li" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella verrucosa" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="verrucosa">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella verrucosa</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="165">
<table pageId="0" pageNumber="165">
<tr pageId="0" pageNumber="165">
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">Vocher</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">GEP a059</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">GEP a060</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">GEP a061</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">GEP a062</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">GEP a063</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">Range</th>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">Sex</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Male</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Male</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Male</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Male</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Male</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Males (n = 5)</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">24.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">25.9</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">24.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">23.2</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">23.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
23.2-25.9 (24.4
<normalizedToken originalValue="±">+/-</normalizedToken>
1.1)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">HDL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">9.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">10.1</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">9.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">9.2</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">9.3</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
9.2-10.1 (9.6
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">HDW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">9.1</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">9.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">9.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">8.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">9.0</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
8.8-9.7 (9.2
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">SNT</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">4.0</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">4.1</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">3.9</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">3.1</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">3.2</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
3.1-4.1 (3.6
<normalizedToken originalValue="±">+/-</normalizedToken>
0.5)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">IND</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
2.5-2.8 (2.7
<normalizedToken originalValue="±">+/-</normalizedToken>
0.1)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">IOD</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
2.4-2.7 (2.5
<normalizedToken originalValue="±">+/-</normalizedToken>
0.1)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">EYE</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">3.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">3.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">3.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">3.0</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">3.1</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
3.0-3.7 (3.4
<normalizedToken originalValue="±">+/-</normalizedToken>
0.3)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">TMP</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.9</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
1.6-1.9 (1.7
<normalizedToken originalValue="±">+/-</normalizedToken>
0.1)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">TEY</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.9</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.0</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.8-1.0 (0.8
<normalizedToken originalValue="±">+/-</normalizedToken>
0.1)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">ML</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">6.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">6.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">6.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">6.3</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">6.2</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
6.2-6.8 (6.5
<normalizedToken originalValue="±">+/-</normalizedToken>
0.3)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">LAHL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">12.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">12.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">12.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">11.9</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">12.0</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
11.9-12.8 (12.4
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">PL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">11.2</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">11.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">11.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">10.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">10.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
10.7-11.7 (11.2
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">TIB</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">11.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">12.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">11.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">11.3</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">12.0</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
11.3-12.4 (11.8
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">HLL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">38.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">40.0</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">36.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">36.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">38.1</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
36.6-40.0 (38.0
<normalizedToken originalValue="±">+/-</normalizedToken>
1.4)
</td>
</tr>
</table>
</paragraph>
<caption doi="10.3897/herpetozoa.35.e89981.figure4" httpUri="https://binary.pensoft.net/fig/750073" pageId="0" pageNumber="165" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figure 4.</emphasis>
Morphological features of the paratypes of
<taxonomicName authorityName="Wang, Zeng, Lin &amp; Li" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella verrucosa" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="verrucosa">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella verrucosa</emphasis>
</taxonomicName>
sp. nov., in life:
<emphasis bold="true" pageId="0" pageNumber="165">A.</emphasis>
GEP a059 (
<emphasis bold="true" pageId="0" pageNumber="165">A1</emphasis>
dorsal view,
<emphasis bold="true" pageId="0" pageNumber="165">A2</emphasis>
lateral view,
<emphasis bold="true" pageId="0" pageNumber="165">A3</emphasis>
ventral view);
<emphasis bold="true" pageId="0" pageNumber="165">B.</emphasis>
GEP a060 (
<emphasis bold="true" pageId="0" pageNumber="165">B1</emphasis>
dorsal view,
<emphasis bold="true" pageId="0" pageNumber="165">B2</emphasis>
dorsolateral view,
<emphasis bold="true" pageId="0" pageNumber="165">B3</emphasis>
ventral view);
<emphasis bold="true" pageId="0" pageNumber="165">C.</emphasis>
GEP a061 (
<emphasis bold="true" pageId="0" pageNumber="165">C1</emphasis>
dorsal view,
<emphasis bold="true" pageId="0" pageNumber="165">C2</emphasis>
dorsolateral view,
<emphasis bold="true" pageId="0" pageNumber="165">C3</emphasis>
ventral view);
<emphasis bold="true" pageId="0" pageNumber="165">D.</emphasis>
GEP a063 (
<emphasis bold="true" pageId="0" pageNumber="165">D1</emphasis>
dorsal view,
<emphasis bold="true" pageId="0" pageNumber="165">D2</emphasis>
dorsolateral view,
<emphasis bold="true" pageId="0" pageNumber="165">D3</emphasis>
ventral view).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="distribution">
<paragraph pageId="0" pageNumber="165">Distribution, ecology and conservation.</paragraph>
<paragraph pageId="0" pageNumber="165">
<taxonomicName authorityName="Wang, Zeng, Lin &amp; Li" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella verrucosa" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="verrucosa">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella verrucosa</emphasis>
</taxonomicName>
sp. nov. is currently only known from its type locality (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Localities of Leptobrachella species occurring in Guangdong Province and Hong Kong, China: Leptobrachella verrucosa sp. nov. (samples ID 1 - 5, Lianshan Bijiashan Nature Reserve, Lianshan county); L. mangshanensis (sample ID 19, Nanling Nature Reserve, Ruyuan county; sample ID 20, Mt. Tianjing, Ruyuan county; sample ID 21, Mt. Dadong, Lianxian county; samples ID 22 - 23, Shimentai Nature Reserve, Yingde city); L. liui (sample ID 13, Mt. Tongguzhang, Fengshun county; sample ID 14, Mt. Fenghuang, Chaozhou city; sample ID 15, Mt. Motianshi, Puning city; sample ID 16, Baipenzhu, Huidong county; sample ID 17, Mt. Nankun, Longmen county); L. laui (sample ID 24, Tai Mo Shan, Hong Kong; sample ID 25, Mt. Wutong, Shenzhen city; sample ID 26, Mt. Paiya, Shenzhen city); L. shimentaina (sample ID 65, Shimentai Nature Reserve, Yingde city; sample ID 66, Luokeng Nature Reserve, Qujiang district); and L. yunkaiensis (sample ID 73, Mt. Yunkai, Xinyi city). Numbers correspond to the ID numbers in Suppl. material 1: Table S 1." figureDoi="10.3897/herpetozoa.35.e89981.figure1" httpUri="https://binary.pensoft.net/fig/750070" pageId="0" pageNumber="165">1</figureCitation>
; ca. 500-600 m a.s.l.) which is located at the boundary of Guangxi Zhuang Autonomous Region, and Guangdong and Hunan provinces. The new species inhabits clear-water rocky streams in primary forests surrounded by broad-leaved forest in granite landforms. Males were observed calling while perched on rocks or leaves of dwarf shrubs near flowing seeps. Data of larvae and females remain unknown. The breeding season lasts from April to June according to our long-term observation. We recommend the new species be listed as LC (Least Concerned) in the IUCN categorization as the forest is well protected and the distribution of the species is potentially wider than currently known, i.e. the adjacent Hunan Province and Guangxi Zhuang Autonomous Region.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="comparisons">
<paragraph pageId="0" pageNumber="165">Comparisons.</paragraph>
<paragraph pageId="0" pageNumber="165">
In the phylogenetic tree (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bayesian inference Maximum Likelihood tree results from partial DNA sequences of the mitochondrial 16 S r RNA gene. Numbers before slashes indicate bootstrap support (BS) and numbers after slashes are Bayesian posterior probabilities (BPP). The ' - ' represent BS ≤ 70 or BPP ≤ 0.90. Voucher before species name at the branch terminal corresponds to the voucher ID in Suppl. material 1: Table S 1." figureDoi="10.3897/herpetozoa.35.e89981.figure2" httpUri="https://binary.pensoft.net/fig/750071" pageId="0" pageNumber="165">2</figureCitation>
),
<taxonomicName authorityName="Wang, Zeng, Lin &amp; Li" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella verrucosa" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="verrucosa">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella verrucosa</emphasis>
</taxonomicName>
sp. nov. is most closely related to the clade composed of
<taxonomicName lsidName="L. liui" pageId="0" pageNumber="165" rank="species" species="liui">
<emphasis italics="true" pageId="0" pageNumber="165">L. liui</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="L. mangshanensis" pageId="0" pageNumber="165" rank="species" species="mangshanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. mangshanensis</emphasis>
</taxonomicName>
, therefore, the latter two are selected for detailed comparison (Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Comparisons of selected diagnostic characters for the new species and its phylogenetically related congeners." httpUri="http://table.plazi.org/id/8E00660359C2063C1002A38AAF7D7E57" pageId="0" pageNumber="165" tableUuid="8E00660359C2063C1002A38AAF7D7E57">3</tableCitation>
). The new species differs from
<taxonomicName lsidName="L. liui" pageId="0" pageNumber="165" rank="species" species="liui">
<emphasis italics="true" pageId="0" pageNumber="165">L. liui</emphasis>
</taxonomicName>
by its narrow fringes along lateral surfaces of toes (vs. wide), continuous longitudinal ridges under toes (vs. interrupted), dorsal surface with numerous conical tubercles and lacking skin ridges (vs. dorsal surface with rounded warts and sparse short skin ridges), heels not meeting when adpressed (vs. heels meeting). The new species differs from
<taxonomicName lsidName="L. mangshanensis" pageId="0" pageNumber="165" rank="species" species="mangshanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. mangshanensis</emphasis>
</taxonomicName>
by its shagreened dorsal surface with numerous conical tubercles, lacking spines, enlarged warts or skin ridges (vs. dorsal surface almost smooth with tiny transparent spines, small tubercles and sparse short skin ridges), ventral surface of thighs lacking conical spines (vs. ventral surface bearing conical spines), tibial-tarsal articulation reaching anterior corner of eye (vs. reaching tip of snout, well beyond anterior corner of eye).
</paragraph>
<caption ID-Table-UUID="8E00660359C2063C1002A38AAF7D7E57" httpUri="http://table.plazi.org/id/8E00660359C2063C1002A38AAF7D7E57" pageId="0" pageNumber="165" start="Table 3" startId="T3">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Table 3.</emphasis>
Comparisons of selected diagnostic characters for the new species and its phylogenetically related congeners.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="165">
<table pageId="0" pageNumber="165">
<tr pageId="0" pageNumber="165">
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName authorityName="Smith" authorityYear="1925" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella</emphasis>
</taxonomicName>
spp.
</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">Fringes on fingers (present + / absent -)</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">Fringes on toes</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">Longitudinal ridges under toes (interrupt + / continuous -)</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">Ventral coloration</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">Dorsal skin texture</th>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName lsidName="L. verrucosa" pageId="0" pageNumber="165" rank="species" species="verrucosa">
<emphasis italics="true" pageId="0" pageNumber="165">L. verrucosa</emphasis>
</taxonomicName>
sp. nov.
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Narrow</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Creamy white with grayish white and dark brown spots</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Shagreened with numerous conical tubercles</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName lsidName="L. bashaensis" pageId="0" pageNumber="165" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. bashaensis</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Narrow</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Creamy-white chest and belly with irregular black spots</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Slightly shagreened with small tubercles</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName lsidName="L. flaviglandulosa" pageId="0" pageNumber="165" rank="species" species="flaviglandulosa">
<emphasis italics="true" pageId="0" pageNumber="165">L. flaviglandulosa</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Narrow</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">+</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Whitish, black speckling on margins</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Shagreened with yellowish- brown tubercles</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName lsidName="L. laui" pageId="0" pageNumber="165" rank="species" species="laui">
<emphasis italics="true" pageId="0" pageNumber="165">L. laui</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">+</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Wide</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Creamy white with dark brown dusting on ventrolateral flanks</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Shagreened with round granular tubercles</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName lsidName="L. liui" pageId="0" pageNumber="165" rank="species" species="liui">
<emphasis italics="true" pageId="0" pageNumber="165">L. liui</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Wide</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">+</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Gray white to creamy white with dark brown spots on chest and ventrolateral flanks</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Shagreened with numerous small tubercles, round warts and sparse short skin ridges</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName lsidName="L. mangshanensis" pageId="0" pageNumber="165" rank="species" species="mangshanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. mangshanensis</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Narrow</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Creamy white belly, scattered with white speckles</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Almost smooth with tiny transparent spines, small tubercles and sparse short skin ridges</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName lsidName="L. maoershanensis" pageId="0" pageNumber="165" rank="species" species="maoershanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. maoershanensis</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Narrow</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Creamy white chest and belly with irregular black spots</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">With longitudinal ridges</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName lsidName="L. shimentaina" pageId="0" pageNumber="165" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">L. shimentaina</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">+ (in males)</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Wide (in males)</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Grayish pink with distinct hazy brown speckling on chest and ventrolateral flanks</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Shagreened with rounded granular tubercles and glandular ridges</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName lsidName="L. yunkaiensis" pageId="0" pageNumber="165" rank="species" species="yunkaiensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. yunkaiensis</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">+</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Wide</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Belly pink with distinct or indistinct speckles</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Shagreened with short skin ridges and raised warts</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="165">
<taxonomicName authorityName="Wang, Zeng, Lin &amp; Li" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella verrucosa" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="verrucosa">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella verrucosa</emphasis>
</taxonomicName>
sp. nov. can be distinguished from
<taxonomicName lsidName="L. yunkaiensis" pageId="0" pageNumber="165" rank="species" species="yunkaiensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. yunkaiensis</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="L. shimentaina" pageId="0" pageNumber="165" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">L. shimentaina</emphasis>
</taxonomicName>
by having no fringes along lateral surfaces of fingers (vs. lateral fringes present in
<taxonomicName lsidName="L. yunkaiensis" pageId="0" pageNumber="165" rank="species" species="yunkaiensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. yunkaiensis</emphasis>
</taxonomicName>
and males of
<taxonomicName lsidName="L. shimentaina" pageId="0" pageNumber="165" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">L. shimentaina</emphasis>
</taxonomicName>
); narrow fringes along lateral surfaces of toes (vs. lateral fringes wide in
<taxonomicName lsidName="L. yunkaiensis" pageId="0" pageNumber="165" rank="species" species="yunkaiensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. yunkaiensis</emphasis>
</taxonomicName>
and males of
<taxonomicName lsidName="L. shimentaina" pageId="0" pageNumber="165" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">L. shimentaina</emphasis>
</taxonomicName>
); shagreened dorsal surface with numerous conical tubercles, lacking spines, enlarged warts or skin ridges (vs. dorsal surface with skin ridges in
<taxonomicName lsidName="L. shimentaina" pageId="0" pageNumber="165" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">L. shimentaina</emphasis>
</taxonomicName>
, dorsal surface with short skin ridges and raised warts in
<taxonomicName lsidName="L. yunkaiensis" pageId="0" pageNumber="165" rank="species" species="yunkaiensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. yunkaiensis</emphasis>
</taxonomicName>
); heels not meeting when adpressed (vs. heels slightly overlapping in
<taxonomicName lsidName="L. shimentaina" pageId="0" pageNumber="165" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">L. shimentaina</emphasis>
</taxonomicName>
, heels meeting in
<taxonomicName lsidName="L. yunkaiensis" pageId="0" pageNumber="165" rank="species" species="yunkaiensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. yunkaiensis</emphasis>
</taxonomicName>
).
<taxonomicName authorityName="Wang, Zeng, Lin &amp; Li" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella verrucosa" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="verrucosa">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella verrucosa</emphasis>
</taxonomicName>
sp. nov. can be distinguished from
<taxonomicName lsidName="L. bashaensis" pageId="0" pageNumber="165" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. bashaensis</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="L. maoershanensis" pageId="0" pageNumber="165" rank="species" species="maoershanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. maoershanensis</emphasis>
</taxonomicName>
by having shagreened dorsal surface with numerous conical tubercles, lacking spines, enlarged warts or skin ridges (vs. dorsum shagreened with small tubercles in
<taxonomicName lsidName="L. bashaensis" pageId="0" pageNumber="165" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. bashaensis</emphasis>
</taxonomicName>
, dorsum with skin ridges in
<taxonomicName lsidName="L. maoershanensis" pageId="0" pageNumber="165" rank="species" species="maoershanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. maoershanensis</emphasis>
</taxonomicName>
); tibial-tarsal articulation reaching anterior corner of eye (vs. reaching tip of snout, well beyond anterior corner of eye in
<taxonomicName lsidName="L. bashaensis" pageId="0" pageNumber="165" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. bashaensis</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="L. maoershanensis" pageId="0" pageNumber="165" rank="species" species="maoershanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. maoershanensis</emphasis>
</taxonomicName>
).
<taxonomicName authorityName="Wang, Zeng, Lin &amp; Li" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella verrucosa" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="verrucosa">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella verrucosa</emphasis>
</taxonomicName>
sp. nov. can be distinguished from
<taxonomicName lsidName="L. laui" pageId="0" pageNumber="165" rank="species" species="laui">
<emphasis italics="true" pageId="0" pageNumber="165">L. laui</emphasis>
</taxonomicName>
by having no lateral fringes along lateral surfaces of fingers (vs. moderate lateral fringes present), narrow lateral fringes along lateral surfaces of toes (vs. lateral fringes wide).
<taxonomicName authorityName="Wang, Zeng, Lin &amp; Li" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella verrucosa" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="verrucosa">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella verrucosa</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName lsidName="L. flaviglandulosa" pageId="0" pageNumber="165" rank="species" species="flaviglandulosa">
<emphasis italics="true" pageId="0" pageNumber="165">L. flaviglandulosa</emphasis>
</taxonomicName>
by having continuous longitudinal ridges under toes (vs. interrupted), heels not meeting when adpressed (vs. heels meeting).
</paragraph>
<paragraph pageId="0" pageNumber="165">
Compared with the 26 known congeners of the genus
<taxonomicName authorityName="Smith" authorityYear="1925" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella</emphasis>
</taxonomicName>
occurring south of the Isthmus of Kra, by the presence of supra-axillary and ventrolateral glands,
<taxonomicName lsidName="L. verrucosa" pageId="0" pageNumber="165" rank="species" species="verrucosa">
<emphasis italics="true" pageId="0" pageNumber="165">L. verrucosa</emphasis>
</taxonomicName>
sp. nov. can easily be distinguished from
<taxonomicName lsidName="L. arayai" pageId="0" pageNumber="165" rank="species" species="arayai">
<emphasis italics="true" pageId="0" pageNumber="165">L. arayai</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. dringi" pageId="0" pageNumber="165" rank="species" species="dringi">
<emphasis italics="true" pageId="0" pageNumber="165">L. dringi</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. fritinniens" pageId="0" pageNumber="165" rank="species" species="fritinniens">
<emphasis italics="true" pageId="0" pageNumber="165">L. fritinniens</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. gracilis" pageId="0" pageNumber="165" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="165">L. gracilis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. hamidi" pageId="0" pageNumber="165" rank="species" species="hamidi">
<emphasis italics="true" pageId="0" pageNumber="165">L. hamidi</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. heteropus" pageId="0" pageNumber="165" rank="species" species="heteropus">
<emphasis italics="true" pageId="0" pageNumber="165">L. heteropus</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. kajangensis" pageId="0" pageNumber="165" rank="species" species="kajangensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. kajangensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. kecil" pageId="0" pageNumber="165" rank="species" species="kecil">
<emphasis italics="true" pageId="0" pageNumber="165">L. kecil</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. marmorata" pageId="0" pageNumber="165" rank="species" species="marmorata">
<emphasis italics="true" pageId="0" pageNumber="165">L. marmorata</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. melanoleuca" pageId="0" pageNumber="165" rank="species" species="melanoleuca">
<emphasis italics="true" pageId="0" pageNumber="165">L. melanoleuca</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. maura" pageId="0" pageNumber="165" rank="species" species="maura">
<emphasis italics="true" pageId="0" pageNumber="165">L. maura</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. picta" pageId="0" pageNumber="165" rank="species" species="picta">
<emphasis italics="true" pageId="0" pageNumber="165">L. picta</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. platycephala" pageId="0" pageNumber="165" rank="species" species="platycephala">
<emphasis italics="true" pageId="0" pageNumber="165">L. platycephala</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. sabahmontana" pageId="0" pageNumber="165" rank="species" species="sabahmontana">
<emphasis italics="true" pageId="0" pageNumber="165">L. sabahmontana</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="L. sola" pageId="0" pageNumber="165" rank="species" species="sola">
<emphasis italics="true" pageId="0" pageNumber="165">L. sola</emphasis>
</taxonomicName>
, all of which lack supra-axillary and ventrolateral glands; and by its distinctly larger male body size, SVL 23.2-25.9 mm,
<taxonomicName lsidName="L. verrucosa" pageId="0" pageNumber="165" rank="species" species="verrucosa">
<emphasis italics="true" pageId="0" pageNumber="165">L. verrucosa</emphasis>
</taxonomicName>
sp. nov. differs from the smaller
<taxonomicName lsidName="L. baluensis" pageId="0" pageNumber="165" rank="species" species="baluensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. baluensis</emphasis>
</taxonomicName>
(SVL 14.9-15.9 mm),
<taxonomicName lsidName="L. brevicrus" pageId="0" pageNumber="165" rank="species" species="brevicrus">
<emphasis italics="true" pageId="0" pageNumber="165">L. brevicrus</emphasis>
</taxonomicName>
(SVL 17.1-17.8 mm),
<taxonomicName lsidName="L. bondangensis" pageId="0" pageNumber="165" rank="species" species="bondangensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. bondangensis</emphasis>
</taxonomicName>
(SVL 17.8 mm),
<taxonomicName lsidName="L. fusca" pageId="0" pageNumber="165" rank="species" species="fusca">
<emphasis italics="true" pageId="0" pageNumber="165">L. fusca</emphasis>
</taxonomicName>
(SVL 16.3 mm),
<taxonomicName lsidName="L. itiokai" pageId="0" pageNumber="165" rank="species" species="itiokai">
<emphasis italics="true" pageId="0" pageNumber="165">L. itiokai</emphasis>
</taxonomicName>
(SVL 15.2-16.7 mm),
<taxonomicName lsidName="L. juliandringi" pageId="0" pageNumber="165" rank="species" species="juliandringi">
<emphasis italics="true" pageId="0" pageNumber="165">L. juliandringi</emphasis>
</taxonomicName>
(SVL 17.0-17.2 mm),
<taxonomicName lsidName="L. mjobergi" pageId="0" pageNumber="165" rank="species" species="mjobergi">
<emphasis italics="true" pageId="0" pageNumber="165">L. mjobergi</emphasis>
</taxonomicName>
(SVL 15.7-19.0 mm),
<taxonomicName lsidName="L. natunae" pageId="0" pageNumber="165" rank="species" species="natunae">
<emphasis italics="true" pageId="0" pageNumber="165">L. natunae</emphasis>
</taxonomicName>
(SVL 17.6 m),
<taxonomicName lsidName="L. parva" pageId="0" pageNumber="165" rank="species" species="parva">
<emphasis italics="true" pageId="0" pageNumber="165">L. parva</emphasis>
</taxonomicName>
(SVL 15.0-16.9 mm), and
<taxonomicName lsidName="L. palmata" pageId="0" pageNumber="165" rank="species" species="palmata">
<emphasis italics="true" pageId="0" pageNumber="165">L. palmata</emphasis>
</taxonomicName>
(SVL 14.4-16.8 mm); and even distinctly larger than female
<taxonomicName lsidName="L. serasanae" pageId="0" pageNumber="165" rank="species" species="serasanae">
<emphasis italics="true" pageId="0" pageNumber="165">L. serasanae</emphasis>
</taxonomicName>
(SVL 16.9 mm).
</paragraph>
<paragraph pageId="0" pageNumber="165">
In having black spots on flanks,
<taxonomicName authorityName="Wang, Zeng, Lin &amp; Li" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella verrucosa" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="verrucosa">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella verrucosa</emphasis>
</taxonomicName>
sp. nov. further differs from
<taxonomicName lsidName="L. aerea" pageId="0" pageNumber="165" rank="species" species="aerea">
<emphasis italics="true" pageId="0" pageNumber="165">L. aerea</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. botsfordi" pageId="0" pageNumber="165" rank="species" species="botsfordi">
<emphasis italics="true" pageId="0" pageNumber="165">L. botsfordi</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. crocea" pageId="0" pageNumber="165" rank="species" species="crocea">
<emphasis italics="true" pageId="0" pageNumber="165">L. crocea</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. firthi" pageId="0" pageNumber="165" rank="species" species="firthi">
<emphasis italics="true" pageId="0" pageNumber="165">L. firthi</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. isos" pageId="0" pageNumber="165" rank="species" species="isos">
<emphasis italics="true" pageId="0" pageNumber="165">L. isos</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. pallida" pageId="0" pageNumber="165" rank="species" species="pallida">
<emphasis italics="true" pageId="0" pageNumber="165">L. pallida</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. petrops" pageId="0" pageNumber="165" rank="species" species="petrops">
<emphasis italics="true" pageId="0" pageNumber="165">L. petrops</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="L. tuberosa" pageId="0" pageNumber="165" rank="species" species="tuberosa">
<emphasis italics="true" pageId="0" pageNumber="165">L. tuberosa</emphasis>
</taxonomicName>
, all of which lack black spots on the flanks.
</paragraph>
<paragraph pageId="0" pageNumber="165">
For the remaining 54 members of the genus
<taxonomicName authorityName="Smith" authorityYear="1925" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella</emphasis>
</taxonomicName>
, males of the new species (SVL 23.2-25.9 mm)
<normalizedToken originalValue="differs">differs</normalizedToken>
from males of the larger
<taxonomicName lsidName="L. bijie" pageId="0" pageNumber="165" rank="species" species="bijie">
<emphasis italics="true" pageId="0" pageNumber="165">L. bijie</emphasis>
</taxonomicName>
(29.0-30.4 mm),
<taxonomicName lsidName="L. chishuiensis" pageId="0" pageNumber="165" rank="species" species="chishuiensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. chishuiensis</emphasis>
</taxonomicName>
(30.8-33.4 mm),
<taxonomicName lsidName="L. bourreti" pageId="0" pageNumber="165" rank="species" species="bourreti">
<emphasis italics="true" pageId="0" pageNumber="165">L. bourreti</emphasis>
</taxonomicName>
(27.4-36.2 mm),
<taxonomicName lsidName="L. damingshanensis" pageId="0" pageNumber="165" rank="species" species="damingshanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. damingshanensis</emphasis>
</taxonomicName>
(33.6-34.4 mm),
<taxonomicName lsidName="L. dorsospina" pageId="0" pageNumber="165" rank="species" species="dorsospina">
<emphasis italics="true" pageId="0" pageNumber="165">L. dorsospina</emphasis>
</taxonomicName>
(28.7-30.5 mm),
<taxonomicName lsidName="L. eos" pageId="0" pageNumber="165" rank="species" species="eos">
<emphasis italics="true" pageId="0" pageNumber="165">L. eos</emphasis>
</taxonomicName>
(33.1-34.7 mm),
<taxonomicName lsidName="L. fuliginosa" pageId="0" pageNumber="165" rank="species" species="fuliginosa">
<emphasis italics="true" pageId="0" pageNumber="165">L. fuliginosa</emphasis>
</taxonomicName>
(28.2-30.0 mm),
<taxonomicName lsidName="L. jinshaensis" pageId="0" pageNumber="165" rank="species" species="jinshaensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. jinshaensis</emphasis>
</taxonomicName>
(29.7-31.2 mm),
<taxonomicName lsidName="L. macrops" pageId="0" pageNumber="165" rank="species" species="macrops">
<emphasis italics="true" pageId="0" pageNumber="165">L. macrops</emphasis>
</taxonomicName>
(28.0-29.3 mm),
<taxonomicName lsidName="L. nahangensis" pageId="0" pageNumber="165" rank="species" species="nahangensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. nahangensis</emphasis>
</taxonomicName>
(40.8 mm),
<taxonomicName lsidName="L. namdongensis" pageId="0" pageNumber="165" rank="species" species="namdongensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. namdongensis</emphasis>
</taxonomicName>
(30.9 mm),
<taxonomicName lsidName="L. pelodytoides" pageId="0" pageNumber="165" rank="species" species="pelodytoides">
<emphasis italics="true" pageId="0" pageNumber="165">L. pelodytoides</emphasis>
</taxonomicName>
(27.5-32.3 mm),
<taxonomicName lsidName="L. purpuraventra" pageId="0" pageNumber="165" rank="species" species="purpuraventra">
<emphasis italics="true" pageId="0" pageNumber="165">L. purpuraventra</emphasis>
</taxonomicName>
(27.3-29.8 mm),
<taxonomicName lsidName="L. pyrrhops" pageId="0" pageNumber="165" rank="species" species="pyrrhops">
<emphasis italics="true" pageId="0" pageNumber="165">L. pyrrhops</emphasis>
</taxonomicName>
(30.3-33.9 mm),
<taxonomicName lsidName="L. suiyangensis" pageId="0" pageNumber="165" rank="species" species="suiyangensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. suiyangensis</emphasis>
</taxonomicName>
(28.7-29.7 mm),
<taxonomicName lsidName="L. sungi" pageId="0" pageNumber="165" rank="species" species="sungi">
<emphasis italics="true" pageId="0" pageNumber="165">L. sungi</emphasis>
</taxonomicName>
(48.3-52.7 mm),
<taxonomicName lsidName="L. yunyangensis" pageId="0" pageNumber="165" rank="species" species="yunyangensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. yunyangensis</emphasis>
</taxonomicName>
(28.3-30.6 mm), and
<taxonomicName lsidName="L. zhangyapingi" pageId="0" pageNumber="165" rank="species" species="zhangyapingi">
<emphasis italics="true" pageId="0" pageNumber="165">L. zhangyapingi</emphasis>
</taxonomicName>
(48.5-52.5 mm). Through its possession of toes with rudimentary webbing and narrow lateral fringes, the new species can be distinguished from
<taxonomicName lsidName="L. ardens" pageId="0" pageNumber="165" rank="species" species="ardens">
<emphasis italics="true" pageId="0" pageNumber="165">L. ardens</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. kalonensis" pageId="0" pageNumber="165" rank="species" species="kalonensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. kalonensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. maculosa" pageId="0" pageNumber="165" rank="species" species="maculosa">
<emphasis italics="true" pageId="0" pageNumber="165">L. maculosa</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. neangi" pageId="0" pageNumber="165" rank="species" species="neangi">
<emphasis italics="true" pageId="0" pageNumber="165">L. neangi</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. rowleyae" pageId="0" pageNumber="165" rank="species" species="rowleyae">
<emphasis italics="true" pageId="0" pageNumber="165">L. rowleyae</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. shiwandashanensis" pageId="0" pageNumber="165" rank="species" species="shiwandashanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. shiwandashanensis</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="L. tadungensis" pageId="0" pageNumber="165" rank="species" species="tadungensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. tadungensis</emphasis>
</taxonomicName>
(no webbing); male
<taxonomicName lsidName="L. alpina" pageId="0" pageNumber="165" rank="species" species="alpina">
<emphasis italics="true" pageId="0" pageNumber="165">L. alpina</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. graminicola" pageId="0" pageNumber="165" rank="species" species="graminicola">
<emphasis italics="true" pageId="0" pageNumber="165">L. graminicola</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. khasiorum" pageId="0" pageNumber="165" rank="species" species="khasiorum">
<emphasis italics="true" pageId="0" pageNumber="165">L. khasiorum</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. murphyi" pageId="0" pageNumber="165" rank="species" species="murphyi">
<emphasis italics="true" pageId="0" pageNumber="165">L. murphyi</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. purpurus" pageId="0" pageNumber="165" rank="species" species="purpurus">
<emphasis italics="true" pageId="0" pageNumber="165">L. purpurus</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="L. yingjiangensis" pageId="0" pageNumber="165" rank="species" species="yingjiangensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. yingjiangensis</emphasis>
</taxonomicName>
(wide lateral fringes);
<taxonomicName lsidName="L. applebyi" pageId="0" pageNumber="165" rank="species" species="applebyi">
<emphasis italics="true" pageId="0" pageNumber="165">L. applebyi</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. lateralis" pageId="0" pageNumber="165" rank="species" species="lateralis">
<emphasis italics="true" pageId="0" pageNumber="165">L. lateralis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. melica" pageId="0" pageNumber="165" rank="species" species="melica">
<emphasis italics="true" pageId="0" pageNumber="165">L. melica</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. minima" pageId="0" pageNumber="165" rank="species" species="minima">
<emphasis italics="true" pageId="0" pageNumber="165">L. minima</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. nyx" pageId="0" pageNumber="165" rank="species" species="nyx">
<emphasis italics="true" pageId="0" pageNumber="165">L. nyx</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. oshanensis" pageId="0" pageNumber="165" rank="species" species="oshanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. oshanensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. pluvialis" pageId="0" pageNumber="165" rank="species" species="pluvialis">
<emphasis italics="true" pageId="0" pageNumber="165">L. pluvialis</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="L. ventripunctata" pageId="0" pageNumber="165" rank="species" species="ventripunctata">
<emphasis italics="true" pageId="0" pageNumber="165">L. ventripunctata</emphasis>
</taxonomicName>
(no lateral fringes). In having shagreened dorsal surface with numerous conical tubercles, lacking spines, enlarged warts or skin ridges, it can be distinguished from
<taxonomicName lsidName="L. aspera" pageId="0" pageNumber="165" rank="species" species="aspera">
<emphasis italics="true" pageId="0" pageNumber="165">L. aspera</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. feii" pageId="0" pageNumber="165" rank="species" species="feii">
<emphasis italics="true" pageId="0" pageNumber="165">L. feii</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. nokrekensis" pageId="0" pageNumber="165" rank="species" species="nokrekensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. nokrekensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. puhoatensis" pageId="0" pageNumber="165" rank="species" species="puhoatensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. puhoatensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. tengchongensis" pageId="0" pageNumber="165" rank="species" species="tengchongensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. tengchongensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. wuhuangmontis" pageId="0" pageNumber="165" rank="species" species="wuhuangmontis">
<emphasis italics="true" pageId="0" pageNumber="165">L. wuhuangmontis</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="L. wulingensis" pageId="0" pageNumber="165" rank="species" species="wulingensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. wulingensis</emphasis>
</taxonomicName>
(dorsum with skin ridges). In having creamy white ventral surface with grayish white and dark brown spots, it can be distinguished from
<taxonomicName lsidName="L. bidoupensis" pageId="0" pageNumber="165" rank="species" species="bidoupensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. bidoupensis</emphasis>
</taxonomicName>
(ventral surface reddish brown),
<taxonomicName lsidName="L. niveimontis" pageId="0" pageNumber="165" rank="species" species="niveimontis">
<emphasis italics="true" pageId="0" pageNumber="165">L. niveimontis</emphasis>
</taxonomicName>
(ventral surface marbling),
<taxonomicName lsidName="L. shangsiensis" pageId="0" pageNumber="165" rank="species" species="shangsiensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. shangsiensis</emphasis>
</taxonomicName>
(ventral surface yellowish creamy-white with marble patterns), and
<taxonomicName lsidName="L. tamdil" pageId="0" pageNumber="165" rank="species" species="tamdil">
<emphasis italics="true" pageId="0" pageNumber="165">L. tamdil</emphasis>
</taxonomicName>
(abdominal region unpatterned). In having continuous longitudinal ridges under toes, it can be distinguished from
<taxonomicName lsidName="L. yeae" pageId="0" pageNumber="165" rank="species" species="yeae">
<emphasis italics="true" pageId="0" pageNumber="165">L. yeae</emphasis>
</taxonomicName>
(longitudinal ridges under toes interrupted).
</paragraph>
</subSubSection>
</treatment>
</document>
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