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<document id="0033E3BC41B3D1814D417902CD3BB543" ID-DOI="10.5281/zenodo.179195" ID-GBIF-Dataset="e0a7d367-be4e-4031-acad-42a17507fadf" ID-ISSN="1175-5326" ID-Zenodo-Dep="179195" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459941676950" checkinUser="plazi" docAuthor="Naruse, Tohru &amp; Ng, Peter K. L." docDate="2007" docId="BB10DE286C372C644485FE68FC8B29F6" docLanguage="en" docName="zt01621p031.pdf" docOrigin="Zootaxa 1621" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Limnopilos Chuang &amp; Ng 1991" docType="treatment" docVersion="9" lastPageNumber="26" masterDocId="4729A6506C3E2C6D4412FF97FF8C2C32" masterDocTitle="On the taxonomy of the genus Hymenicoides Kemp, 1917 (Crustacea: Decapoda: Brachyura: Hymenosomatidae), with resurrection of Limnopilos Chuang &amp; Ng, 1991, and descriptions of two new species" masterLastPageNumber="31" masterPageNumber="17" pageNumber="26" updateTime="1698230522269" updateUser="plazi">
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<mods:title id="5A082C934443551EEDCAF88AEF59A7BE">On the taxonomy of the genus Hymenicoides Kemp, 1917 (Crustacea: Decapoda: Brachyura: Hymenosomatidae), with resurrection of Limnopilos Chuang &amp; Ng, 1991, and descriptions of two new species</mods:title>
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<mods:namePart id="7CD76760446589F46A0EAD4421658384">Naruse, Tohru</mods:namePart>
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<mods:namePart id="FD86613FAE04C88A07E74FB7EEBE68F7">Ng, Peter K. L.</mods:namePart>
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<mods:date id="D190AE217EDFE02D8C1290ED42CC4896">2007</mods:date>
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<mods:number id="BBCB84A6F0398925273B029C627709C7">1621</mods:number>
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<treatment id="BB10DE286C372C644485FE68FC8B29F6" ID-DOI="http://doi.org/10.5281/zenodo.6251676" ID-GBIF-Taxon="119375999" ID-Zenodo-Dep="6251676" LSID="urn:lsid:plazi:treatment:BB10DE286C372C644485FE68FC8B29F6" httpUri="http://treatment.plazi.org/id/BB10DE286C372C644485FE68FC8B29F6" lastPageNumber="26" pageId="9" pageNumber="26">
<subSubSection id="7BA33CB56C372C644485FE68FD912E2B" box="[151,541,511,537]" pageId="9" pageNumber="26" type="nomenclature">
<paragraph id="33066F3E6C372C644485FE68FD912E2B" blockId="9.[151,541,511,537]" box="[151,541,511,537]" pageId="9" pageNumber="26">
<heading id="684ED8526C372C644485FE68FD912E2B" bold="true" box="[151,541,511,537]" fontSize="11" level="1" pageId="9" pageNumber="26" reason="1">
<taxonomicName id="F4B914BD6C372C644485FE68FD912E2B" ID-CoL="7PMTG" authority="Chuang &amp; Ng, 1991" authorityName="Chuang &amp; Ng" authorityYear="1991" box="[151,541,511,537]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C372C644485FE68FD912E2B" bold="true" box="[151,541,511,537]" pageId="9" pageNumber="26">
<emphasis id="01CDB32C6C372C644485FE68FEAC2E2B" bold="true" box="[151,288,511,537]" italics="true" pageId="9" pageNumber="26">Limnopilos</emphasis>
<bibRefCitation id="572812CF6C372C644534FE68FD912E2B" author="Chuang" box="[294,541,511,537]" pageId="9" pageNumber="31" refString="Chuang, C. T. N. &amp; Ng, P. K. L. (1991) Preliminary descriptions of one new genus and three new species of hymenosomatid crabs from Southeast Asia (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 39 (2), 363 - 368." type="journal article" year="1991">Chuang &amp; Ng, 1991</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C372C644485FDDDFB0D2EB3" pageId="9" pageNumber="26" type="reference_group">
<paragraph id="33066F3E6C372C644485FDDDFDA72E53" blockId="9.[151,1153,586,641]" box="[151,555,586,609]" pageId="9" pageNumber="26">
<treatmentCitationGroup id="13A948106C372C644485FDDDFDA72E53" box="[151,555,586,609]" pageId="9" pageNumber="26">
<taxonomicName id="F4B914BD6C372C644485FDDDFDAB2E53" ID-CoL="7PMTG" authority="Chuang &amp; Ng, 1991: 363" authorityName="Chuang &amp; Ng" authorityPageNumber="363" authorityYear="1991" box="[151,551,586,609]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C372C644485FDDDFE832E53" box="[151,271,586,609]" italics="true" pageId="9" pageNumber="26">Limnopilos</emphasis>
<treatmentCitation id="B218492F6C372C644507FDDDFDAB2E53" author="Chuang" box="[277,551,586,609]" page="363" pageId="9" pageNumber="26" year="1991">
<bibRefCitation id="572812CF6C372C644507FDDDFE7D2E53" author="Chuang" box="[277,497,586,609]" pageId="9" pageNumber="31" refString="Chuang, C. T. N. &amp; Ng, P. K. L. (1991) Preliminary descriptions of one new genus and three new species of hymenosomatid crabs from Southeast Asia (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 39 (2), 363 - 368." type="journal article" year="1991">Chuang &amp; Ng, 1991</bibRefCitation>
: 363
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="33066F3E6C372C644485FDFDFB0D2EB3" blockId="9.[151,1153,586,641]" box="[151,1153,618,641]" pageId="9" pageNumber="26">
<emphasis id="01CDB32C6C372C644485FDFDFECA2EB3" box="[151,326,618,641]" italics="true" pageId="9" pageNumber="26">Hymenicoides—</emphasis>
<bibRefCitation id="572812CF6C372C64455EFDFDFDA42EB3" author="Ng" box="[332,552,618,641]" pageId="9" pageNumber="31" refString="Ng, P. K. L. &amp; Chuang, C. T. N. (1996) The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology, Supplement, 3, 1 - 82." type="journal article" year="1996">Ng &amp; Chuang, 1996</bibRefCitation>
: 50 (part); Guinot &amp; Richer de
<bibRefCitation id="572812CF6C372C644765FDFDFB892EB3" author="Forges" box="[887,1029,618,641]" pageId="9" pageNumber="31" refString="Guinot, D. &amp; Richer de Forges, B. (1997) Affinites entre les Hymenosomatidae MacLeay ' 1838 et les Inachoididae Dana ' 1851 (Crustacea ' Decapoda ' Brachyura). Zoosystema, 19 (2 / 3), 453 - 502." type="journal article" year="1997">Forges, 1997</bibRefCitation>
: 460 (part).
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C372C644485FD25FEED2EC6" pageId="9" pageNumber="26" type="materials_examined">
<paragraph id="33066F3E6C372C644485FD25FEED2EC6" blockId="9.[151,1437,690,1476]" pageId="9" pageNumber="26">
<emphasis id="01CDB32C6C372C644485FD25FEBF2EFE" bold="true" box="[151,307,690,716]" pageId="9" pageNumber="26">
<typeStatus id="EC02D19C6C372C644485FD25FF582EFE" box="[151,212,690,716]" pageId="9" pageNumber="26">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="F4B914BD6C372C644556FD25FCCB2EFE" authority="Chuang &amp; Ng, 1991" authorityName="Chuang &amp; Ng" authorityYear="1991" box="[324,839,690,716]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="naiyanetri">
<emphasis id="01CDB32C6C372C644556FD25FDC62EF9" box="[324,586,690,715]" italics="true" pageId="9" pageNumber="26">Limnopilos naiyanetri</emphasis>
<bibRefCitation id="572812CF6C372C644640FD25FCCB2EFE" author="Chuang" box="[594,839,690,716]" pageId="9" pageNumber="31" refString="Chuang, C. T. N. &amp; Ng, P. K. L. (1991) Preliminary descriptions of one new genus and three new species of hymenosomatid crabs from Southeast Asia (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 39 (2), 363 - 368." type="journal article" year="1991">Chuang &amp; Ng, 1991</bibRefCitation>
</taxonomicName>
, by original designation and monotypy; gender of genus masculine.
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C372C6444D4FC95FC6028E6" pageId="9" pageNumber="26" type="diagnosis">
<paragraph id="33066F3E6C372C6444D4FC95FC6028E6" blockId="9.[151,1437,690,1476]" pageId="9" pageNumber="26">
<emphasis id="01CDB32C6C372C6444D4FC95FEB32F2E" bold="true" box="[198,319,770,796]" pageId="9" pageNumber="26">Diagnosis</emphasis>
. Carapace circular, pilose, dorsal surface concave; grooves distinct; rostrum absent or very weak; antenna with proximal portion of basal article posterior to inner section of proximal portion of eye; eyes, antennae, antennules visible dorsally; third maxillipeds narrow, not covering more than three-quarters of mouth field when closed, merus rectangular, dactylus styliform, approximately twice length of propodus; male chelae relatively stout, outer surface evenly convex, partially covered by dense setae, without tubercle. Female vulva placed on imaginary line joining inner ends of sutures between sternites 5, 6 on medial fused plate of thoracic sternum, vulva with basal mount. Male abdomen-pleotelson 6 segmented, pleotelson slightly trilobed, inner surface thickened distally, forming socket for sternal button (
<figureCitation id="AB8273BB6C372C64403AFB8DFBE12806" box="[1064,1133,1050,1076]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="9" pageNumber="26">Fig. 1</figureCitation>
c, d). G1 stout, bent outwards medially; distal part with distal inner processes, tuberculate distal outer angle. Female abdomen-pleotelson with six distinctly demarcated segments, boundary between first and second (
<taxonomicName id="F4B914BD6C372C6440AFFBFCFAE028B6" box="[1213,1388,1131,1156]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="01CDB32C6C372C6440AFFBFCFAE028B6" box="[1213,1388,1131,1156]" italics="true" pageId="9" pageNumber="26">L. sumatranus</emphasis>
</taxonomicName>
) or between second and third segments (
<taxonomicName id="F4B914BD6C372C644658FB04FD6E289E" box="[586,738,1171,1196]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="naiyanetri">
<emphasis id="01CDB32C6C372C644658FB04FD6E289E" box="[586,738,1171,1196]" italics="true" pageId="9" pageNumber="26">L. naiyanetri</emphasis>
</taxonomicName>
) movable; long, biramous pleopods on second to fifth segments, developed from distal outer end of inner surface of each segment.
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C372C6444D4FB75FC8B29F6" pageId="9" pageNumber="26" type="discussion">
<paragraph id="33066F3E6C372C6444D4FB75FEAE2946" blockId="9.[151,1437,690,1476]" pageId="9" pageNumber="26">
<emphasis id="01CDB32C6C372C6444D4FB75FEB428CE" bold="true" box="[198,312,1250,1276]" pageId="9" pageNumber="26">Remarks</emphasis>
. Several characters clearly distinguish
<taxonomicName id="F4B914BD6C372C64473DFB74FC3F28CE" box="[815,947,1251,1276]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C372C64473DFB74FC3F28CE" box="[815,947,1251,1276]" italics="true" pageId="9" pageNumber="26">Limnopilos</emphasis>
</taxonomicName>
from
<taxonomicName id="F4B914BD6C372C64401EFB74FB3E28CE" box="[1036,1202,1251,1276]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C372C64401EFB74FB3E28CE" box="[1036,1202,1251,1276]" italics="true" pageId="9" pageNumber="26">Hymenicoides</emphasis>
</taxonomicName>
(see Remarks for
<taxonomicName id="F4B914BD6C372C644485FA9CFEB12916" box="[151,317,1291,1316]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C372C644485FA9CFEB12916" box="[151,317,1291,1316]" italics="true" pageId="9" pageNumber="26">Hymenicoides</emphasis>
</taxonomicName>
above). In addition to
<taxonomicName id="F4B914BD6C372C644672FA9CFD702916" box="[608,764,1291,1316]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="naiyanetri">
<emphasis id="01CDB32C6C372C644672FA9CFD702916" box="[608,764,1291,1316]" italics="true" pageId="9" pageNumber="26">L. naiyanetri</emphasis>
</taxonomicName>
, the
<typeStatus id="EC02D19C6C372C64472DFA9DFCFE2916" box="[831,882,1290,1316]" pageId="9" pageNumber="26">type</typeStatus>
species,
<taxonomicName id="F4B914BD6C372C6447F4FA9CFAB12916" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" box="[998,1341,1290,1316]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="01CDB32C6C372C6447F4FA9CFB312916" box="[998,1213,1291,1316]" italics="true" pageId="9" pageNumber="26">H. microrhynchus</emphasis>
Ng, 1995
</taxonomicName>
, and
<taxonomicName id="F4B914BD6C372C644196FA9CFEAC297E" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="sumatranus" status="sp. nov.">
<emphasis id="01CDB32C6C372C644196FA9CFEAC297E" italics="true" pageId="9" pageNumber="26">L. sumatranus</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C372C644538FAA5FE30297E" bold="true" box="[298,444,1330,1356]" pageId="9" pageNumber="26">
<taxonomicNameLabel id="1AFE0E576C372C644538FAA5FE30297E" box="[298,444,1330,1356]" pageId="9" pageNumber="26" rank="species">new species</taxonomicNameLabel>
</emphasis>
, all possess the diagnostic characters of the genus and are therefore transferred to
<taxonomicName id="F4B914BD6C372C644485FACCFE972946" box="[151,283,1371,1396]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C372C644485FACCFE972946" box="[151,283,1371,1396]" italics="true" pageId="9" pageNumber="26">Limnopilos</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="33066F3E6C372C6444D4FA15FC8B29F6" blockId="9.[151,1437,690,1476]" pageId="9" pageNumber="26">
In the case of
<taxonomicName id="F4B914BD6C372C644579FA14FDB429AE" box="[363,568,1411,1436]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="01CDB32C6C372C644579FA14FDB429AE" box="[363,568,1411,1436]" italics="true" pageId="9" pageNumber="26">L. microrhynchus</emphasis>
</taxonomicName>
, only male specimens are known thus far, and the female characters diagnosed above are therefore not known for this species.
</paragraph>
</subSubSection>
</treatment>
</document>