264 lines
36 KiB
XML
264 lines
36 KiB
XML
<document id="7DC130BFCA39F0082C9B71257EDF832E" ID-DOI="10.11646/phytotaxa.509.2.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="5426136" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630396608438" checkinUser="marcus" docAuthor="Simpson, Michael G. & Rebman, Jon P." docDate="2021" docId="980F534BA523FFFEFF07FF26FC2B7567" docLanguage="en" docName="phytotaxa.509.2.3.pdf" docOrigin="Phytotaxa 509 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.509.2.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Cryptantha pondii , Johnston 1928" docType="treatment" docVersion="4" lastPageNumber="196" masterDocId="64362B33A528FFF5FF8FFFB8FFDF703B" masterDocTitle="Research in Boraginaceae: A new variety of Cryptantha maritima, Cryptantha pondii resurrected, and Johnstonella echinosepala transferred back to Cryptantha" masterLastPageNumber="210" masterPageNumber="185" pageNumber="196" updateTime="1699261161300" updateUser="plazi" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="EFFF2F5C8DC70F39A2953A709F56A0B1">
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<mods:title id="7CB9599C5653CD666B3446EB8C00CEBD">Research in Boraginaceae: A new variety of Cryptantha maritima, Cryptantha pondii resurrected, and Johnstonella echinosepala transferred back to Cryptantha</mods:title>
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</mods:titleInfo>
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<mods:name id="97D789AAC302874E47AEE8B3C1C22F1A" type="personal">
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<mods:role id="8B216453B0D43FB70C7A278C9A19D843">
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<mods:roleTerm id="9F25E9FCB8C50A5C6707A0038B8B0F65">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D189E62DA1669AEB1C15F56E82CDA39F">Simpson, Michael G.</mods:namePart>
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<mods:affiliation id="9B96D852452F8B2B609A1B0869247559">Department of Biology, San Diego State University, San Diego, CA 92182, USA</mods:affiliation>
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</mods:name>
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<mods:name id="8A30EABEC59B83FEA08F00E2DF322607" type="personal">
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<mods:roleTerm id="9ECE42C2CC6A92B8BF395B227961ABE4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6E65CAF9301BF785214E7DFC75614DB6">Rebman, Jon P.</mods:namePart>
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<mods:affiliation id="CF4F52077376B4A69E302BBA4803B00E">San Diego Natural History Museum, P. O. Box 121390, San Diego, CA 92112 - 1390, USA</mods:affiliation>
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<mods:typeOfResource id="85E9B99D0DE9C2925A200CAA12B3C8D8">text</mods:typeOfResource>
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<mods:title id="BF49C8ECA0FDA2B0F5DFEF7B81B65CB6">Phytotaxa</mods:title>
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<mods:part id="3667AF533CB76C5110D538B813009C32">
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<mods:date id="D6E2AA0E159DBCDAD58BF8C935BB1D18">2021</mods:date>
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<mods:number id="F484EA07D83D6FF4BFB1A90A8FC46C7A">2021-06-29</mods:number>
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<mods:number id="001EB64EFFBD635A2C0795C81312B4A1">509</mods:number>
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<mods:number id="ED87D528EE0D790A0FEADFC9566CD87E">2</mods:number>
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<mods:start id="9BAF685AD053904804A87E01543739D5">185</mods:start>
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<mods:end id="7459F88E49547AC544F234F0159F5031">210</mods:end>
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<mods:location id="2478FDAEBA5D7C38C4A25F8E4623BE1B">
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<mods:url id="4EF5800E35A0CBC864D3EB8B3B9121B2">http://dx.doi.org/10.11646/phytotaxa.509.2.3</mods:url>
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<mods:classification id="0E2E0A4AB64B1368805D51D3F552078A">journal article</mods:classification>
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<mods:identifier id="AF33D4427C4AB056DF4CCEEE8DD20E18" type="DOI">10.11646/phytotaxa.509.2.3</mods:identifier>
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<mods:identifier id="0FA32DE061897D58CC69D79DA8065766" type="ISSN">1179-3163</mods:identifier>
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<mods:identifier id="7BAF45F116FA4BB0407E11B41A8492A7" type="Zenodo-Dep">5426136</mods:identifier>
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<treatment id="980F534BA523FFFEFF07FF26FC2B7567" ID-DOI="http://doi.org/10.5281/zenodo.5485201" ID-Zenodo-Dep="5485201" LSID="urn:lsid:plazi:treatment:980F534BA523FFFEFF07FF26FC2B7567" httpUri="http://treatment.plazi.org/id/980F534BA523FFFEFF07FF26FC2B7567" lastPageNumber="196" pageId="11" pageNumber="196">
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<subSubSection id="58BCB1D6A523FFFEFF07FF26FDC47082" box="[136,539,158,185]" pageId="11" pageNumber="196" type="nomenclature">
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<paragraph id="1019E25DA523FFFEFF07FF26FDC47082" blockId="11.[136,539,158,185]" box="[136,539,158,185]" pageId="11" pageNumber="196">
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<heading id="4B515531A523FFFEFF07FF26FDC47082" bold="true" box="[136,539,158,185]" fontSize="11" level="1" pageId="11" pageNumber="196" reason="1">
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<emphasis id="22D23E4FA523FFFEFF07FF26FDC47082" bold="true" box="[136,539,158,185]" pageId="11" pageNumber="196">
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Resurrection of
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<taxonomicName id="D7A699DEA523FFFEFEC7FF26FDC47082" ID-CoL="ZTVZ" authorityName=", Johnston" authorityYear="1928" box="[328,539,158,185]" class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" kingdom="Plantae" order="Boraginales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="species" species="pondii">
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<emphasis id="22D23E4FA523FFFEFEC7FF26FDC47082" bold="true" box="[328,539,158,185]" italics="true" pageId="11" pageNumber="196">Cryptantha pondii</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="58BCB1D6A523FFFEFF07FF5FFD4E721A" pageId="11" pageNumber="196" type="description">
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<paragraph id="1019E25DA523FFFEFF07FF5FFD4E721A" blockId="11.[136,1452,230,1373]" pageId="11" pageNumber="196">
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In our survey of
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<taxonomicName id="D7A699DEA523FFFEFECFFF5EFDF2713B" authorityName="Greene" authorityYear="1887" baseAuthorityName="Greene" baseAuthorityYear="1885" box="[320,557,230,256]" class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" kingdom="Plantae" order="Boraginales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="species" species="maritima">
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<emphasis id="22D23E4FA523FFFEFECFFF5EFDF2713B" box="[320,557,230,256]" italics="true" pageId="11" pageNumber="196">Cryptantha maritima</emphasis>
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</taxonomicName>
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, we discovered
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<specimenCount id="06A029D4A523FFFEFD6FFF5FFC59713B" box="[736,902,230,257]" pageId="11" pageNumber="196" type="generic">two specimens</specimenCount>
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that had been identified as that species but which had four, smooth nutlets, these slightly heteromorphic (
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<figureCitation id="889DFED8A523FFFEFD79FEB2FCE4711E" box="[758,827,266,293]" captionStart="FIGURE 6" captionStartId="12.[136,229,1831,1853]" captionTargetBox="[245,1343,196,1806]" captionTargetId="figure-19@12.[237,1350,190,1807]" captionTargetPageId="12" captionText="FIGURE 6. Cryptantha pondii. A. Close-up of single plant of presumed lectotype specimen (US41006), confirmed by Johnston (1928) as being correctly identified. Note linear, pustulate leaves and bracts at base of cymules. B,C. Images from Gentry 8684, 13 March 1949 (HCIB21969). B. Four nutlets removed from fruit, showing smooth surface and slight size heteromorphism. C. Close-up of cymules, showing linear bracts and relatively large corollas. D–F. Images from Philbrick B74-84, 25 March 1974 (SBBG49613). D. Fruit with four, smooth nutlets removed. Note finely hirsute calyx marginal region with ascending trichomes, as well as hispid calyx midrib of stouter trichomes. E. Close-up of four nutlets attached to gynobase. Note slight heteromorphism in size and slight extension of style tip/stigma from nutlet apices. F. Cymules, showing appressed-strigose vestiture of axes, scattered linear flower bracts, and relatively large corolla." figureDoi="http://doi.org/10.5281/zenodo.5426148" httpUri="https://zenodo.org/record/5426148/files/figure.png" pageId="11" pageNumber="196">Fig. 6</figureCitation>
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). In discovering the publication of
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<bibRefCitation id="74379FACA523FFFEFB48FEB2FAA1711F" author="Johnston, I. M." box="[1223,1406,266,292]" pageId="11" pageNumber="196" pagination="73 - 83" refId="ref13085" refString="Johnston, I. M. (1928) Studies in the Boraginaceae - VII. 2. Notes on various Boraginoideae. Contributions from the Gray Herbarium of Harvard University 81: 73 - 83." type="journal article" year="1928">Johnston (1928)</bibRefCitation>
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, we realized that this was the true
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<taxonomicName id="D7A699DEA523FFFEFE59FE97FDE07173" authorityName=", Johnston" authorityYear="1928" box="[470,575,302,329]" class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" kingdom="Plantae" order="Boraginales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="species" species="pondii">
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<emphasis id="22D23E4FA523FFFEFE59FE97FDE07173" box="[470,575,302,329]" italics="true" pageId="11" pageNumber="196">C. pondii</emphasis>
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</taxonomicName>
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. To date, we are aware of only three collections of
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<emphasis id="22D23E4FA523FFFEFBF3FE97FB347173" box="[1148,1259,302,329]" italics="true" pageId="11" pageNumber="196">
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<taxonomicName id="D7A699DEA523FFFEFBF3FE97FB387173" authorityName=", Johnston" authorityYear="1928" box="[1148,1255,302,329]" class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" kingdom="Plantae" order="Boraginales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="species" species="pondii">C. pondii</taxonomicName>
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,
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</emphasis>
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including that of the
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<typeStatus id="CF1D5CFFA523FFFEFF3DFEEBFF3A7156" box="[178,229,339,365]" pageId="11" pageNumber="196">type</typeStatus>
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, all from
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<collectingRegion id="D2622CBFA523FFFEFEDEFEEAFDF17157" box="[337,558,338,364]" country="Mexico" name="Baja California Sur" pageId="11" pageNumber="196">Baja California Sur</collectingRegion>
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,
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<collectingCountry id="68B1A2CDA523FFFEFDB6FEEAFD4D7157" box="[569,658,338,364]" name="Mexico" pageId="11" pageNumber="196">Mexico</collectingCountry>
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. One collection has duplicates for a total of five herbarium specimens, but two collections consist of single known herbarium specimens (see below; recall that we were unable to locate the presumed “
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<typeStatus id="CF1D5CFFA523FFFEFE88FE22FEE3718F" box="[263,316,410,436]" pageId="11" pageNumber="196">type</typeStatus>
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” cited by Johnston, but did locate his cited “isotype,” treated here as a designated isolectotype). Two of the collections are from the extreme western portion of the
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<taxonomicName id="D7A699DEA523FFFEFCBEFE06FC4B71E3" box="[817,916,446,472]" class="Arachnida" family="Vaejovidae" genus="Vizcaino" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="193" phylum="Arthropoda" rank="genus">Vizcaíno</taxonomicName>
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peninsula and one is from Natividad Island, just off the coast from that area (
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<figureCitation id="889DFED8A523FFFEFE4FFE5AFDEC71C7" box="[448,563,482,508]" captionStart="FIGURE 7" captionStartId="13.[136,229,950,972]" captionTargetBox="[152,1432,194,919]" captionTargetId="figure-19@13.[152,1436,190,926]" captionTargetPageId="13" captionText="FIGURE 7. A,B. Cryptantha pondii, distribution map of three known collections. Georeference point for the type, Pond 22, approximate; that of Gentry 8684 placed as a centroid of three different georeferenced estimates. Maps from ©Google 2020, INEGI Data." figureDoi="http://doi.org/10.5281/zenodo.5426150" httpUri="https://zenodo.org/record/5426150/files/figure.png" pageId="11" pageNumber="196">Fig. 7A,B</figureCitation>
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). The three collections are listed here (georeferenced coordinates and elevations indicated with “*”, estimated from label data):
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</paragraph>
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</subSubSection>
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<subSubSection id="58BCB1D6A523FFFEFF32FD92FC2B7567" pageId="11" pageNumber="196" type="materials_examined">
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<paragraph id="1019E25DA523FFFEFF32FD92FA50737B" blockId="11.[136,1452,230,1373]" pageId="11" pageNumber="196">
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<collectingCountry id="68B1A2CDA523FFFEFF32FD92FEF0727F" box="[189,303,554,580]" name="Mexico" pageId="11" pageNumber="196">MEXICO</collectingCountry>
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.
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<collectingRegion id="D2622CBFA523FFFEFEB6FD92FDCA727F" box="[313,533,554,580]" country="Mexico" name="Baja California Sur" pageId="11" pageNumber="196">Baja California Sur</collectingRegion>
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: Bay of San Bartolomé [Note: Bay of San Bartolomé is now called Bahia Tortuga], ca. 5 meters elevation, 27.6907˚
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<collectionCode id="76B77A98A523FFFEFE7CFDF6FDD77252" box="[499,520,590,617]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="11" pageNumber="196" type="Herbarium">N</collectionCode>
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, 114.8935˚
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<collectionCode id="76B77A98A523FFFEFD07FDF6FD7A7252" box="[648,677,590,617]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="11" pageNumber="196" type="Herbarium">W</collectionCode>
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*,
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<date id="6418C49DA523FFFEFD33FDF6FC977253" box="[700,840,590,616]" pageId="11" pageNumber="196" value="1889-03">March 1889</date>
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,
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<emphasis id="22D23E4FA523FFFEFCDCFDF7FC307253" box="[851,1007,590,617]" italics="true" pageId="11" pageNumber="196">
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<collectionCode id="76B77A98A523FFFEFCDCFDF7FCB67252" box="[851,873,591,617]" country="Denmark" name="University of Copenhagen" pageId="11" pageNumber="196" type="Herbarium">C</collectionCode>
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.
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<collectionCode id="76B77A98A523FFFEFCFAFDF7FC597253" box="[885,902,591,616]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="11" pageNumber="196" type="Herbarium">F</collectionCode>
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. Pond 22
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</emphasis>
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, Isolectotype:
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<collectingCountry id="68B1A2CDA523FFFEFB19FDF6FB667252" box="[1174,1209,590,617]" name="United States of America" pageId="11" pageNumber="196">US</collectingCountry>
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41006=
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<collectingCountry id="68B1A2CDA523FFFEFA9EFDF6FAEB7252" box="[1297,1332,590,617]" name="United States of America" pageId="11" pageNumber="196">US</collectingCountry>
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01050283;
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<quantity id="D75E4FB8A523FFFEFF07FDCAFF3372B7" box="[136,236,626,652]" metricMagnitude="3" metricUnit="m" metricValue="4.02336" metricValueMax="4.828032" metricValueMin="3.218688" pageId="11" pageNumber="196" unit="mi" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mile</quantity>
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<collectionCode id="76B77A98A523FFFEFF7BFDCBFEDA72B6" box="[244,261,627,653]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="11" pageNumber="196" type="Herbarium">E</collectionCode>
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of Punta Eugenia,
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<taxonomicName id="D7A699DEA523FFFEFE51FDCAFD4972B6" authority="Desert" authorityName="Desert" box="[478,662,626,653]" class="Arachnida" family="Vaejovidae" genus="Vizcaino" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="196" phylum="Arthropoda" rank="genus">Vizcaíno Desert</taxonomicName>
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, habitat: Maritime Desert, abundant,
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<quantity id="D75E4FB8A523FFFEFBB3FDCAFB0F72B7" box="[1084,1232,626,652]" metricMagnitude="1" metricUnit="m" metricValue="6.096" metricValueMax="9.144" metricValueMin="3.048" pageId="11" pageNumber="196" unit="ft" value="200.0" valueMax="300.0" valueMin="100.0">100–300 feet</quantity>
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[ca. 30–90 meters] elevation, 27.82025˚
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<collectionCode id="76B77A98A523FFFEFEFBFD2EFE56728A" box="[372,393,662,689]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="11" pageNumber="196" type="Herbarium">N</collectionCode>
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115.0502˚
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<collectionCode id="76B77A98A523FFFEFD88FD2EFDFC728A" box="[519,547,662,689]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="11" pageNumber="196" type="Herbarium">W</collectionCode>
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* (these coordinates are the centroid of three slightly varying estimates from different herbaria records),
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<date id="6418C49DA523FFFEFE3AFD02FD8072EF" box="[437,607,698,724]" pageId="11" pageNumber="196" value="1949-03-13">13 March 1949</date>
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,
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<emphasis id="22D23E4FA523FFFEFDE7FD03FCF772EF" box="[616,808,698,725]" italics="true" pageId="11" pageNumber="196">
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<collectionCode id="76B77A98A523FFFEFDE7FD03FD5E72EE" box="[616,641,699,725]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="11" pageNumber="196" type="Herbarium">H</collectionCode>
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.
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<collectionCode id="76B77A98A523FFFEFD0AFD03FD4A72EE" box="[645,661,699,725]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="11" pageNumber="196" type="Herbarium">S</collectionCode>
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. Gentry 8684
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</emphasis>
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,
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<collectionCode id="76B77A98A523FFFEFCBCFD02FCB172EF" box="[819,878,698,724]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15600" lsid="urn:lsid:biocol.org:col:15600" name="Arizona State University" pageId="11" pageNumber="196" type="Herbarium">ASU</collectionCode>
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0014603!,
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<collectionCode id="76B77A98A523FFFEFC6DFD02FBC972EF" box="[994,1046,698,724]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15533" lsid="urn:lsid:biocol.org:col:15533" name="Desert Botanical Garden" pageId="11" pageNumber="196" type="Herbarium">DES</collectionCode>
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00009392,
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<collectionCode id="76B77A98A523FFFEFB1CFD02FB0572EF" box="[1171,1242,698,724]" country="Mexico" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12778" lsid="urn:lsid:biocol.org:col:12778" name="Centro de Investigaciones Biológicas del Noroeste, S. C." pageId="11" pageNumber="196" type="Herbarium">HCIB</collectionCode>
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21969!,
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<collectionCode id="76B77A98A523FFFEFABEFD02FA8872EF" box="[1329,1367,698,724]" country="USA" lsid="urn:lsid:biocol.org:col:15659" name="San Diego Natural History Museum" pageId="11" pageNumber="196" type="Herbarium">SD</collectionCode>
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86419!, UCR-80508!; Mulegé, Isla Natividad, upper south-facing talus slope of major canyon, seventh canyon northwest of lighthouse and draining from off-shore ridge to on-shore coast, associated species:
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<taxonomicName id="D7A699DEA523FFFEFB82FCBAFBA47327" box="[1037,1147,770,796]" class="Magnoliopsida" family="Loasaceae" genus="Mentzelia" kingdom="Plantae" order="Cornales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="genus">
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<emphasis id="22D23E4FA523FFFEFB82FCBAFBA47327" box="[1037,1147,770,796]" italics="true" pageId="11" pageNumber="196">Mentzelia</emphasis>
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</taxonomicName>
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and
|
||
<taxonomicName id="D7A699DEA523FFFEFB22FCBAFACE7327" box="[1197,1297,770,796]" class="Magnoliopsida" family="Hydrophyllaceae" genus="Phacelia" kingdom="Plantae" order="Boraginales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="22D23E4FA523FFFEFB22FCBAFACE7327" box="[1197,1297,770,796]" italics="true" pageId="11" pageNumber="196">Phacelia</emphasis>
|
||
</taxonomicName>
|
||
, ca. 77 meters elevation*, 27.881926˚
|
||
<collectionCode id="76B77A98A523FFFEFE02FC9EFE7D737A" box="[397,418,806,833]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="11" pageNumber="196" type="Herbarium">N</collectionCode>
|
||
, 115.194287˚
|
||
<collectionCode id="76B77A98A523FFFEFDB0FC9EFD84737A" box="[575,603,806,833]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="11" pageNumber="196" type="Herbarium">W</collectionCode>
|
||
*,
|
||
<date id="6418C49DA523FFFEFDFEFC9EFCC0737B" box="[625,799,806,833]" pageId="11" pageNumber="196" value="1974-03-25">25 March 1974</date>
|
||
,
|
||
<emphasis id="22D23E4FA523FFFEFCA5FC9FFB38737B" box="[810,1255,806,833]" italics="true" pageId="11" pageNumber="196">
|
||
<collectionCode id="76B77A98A523FFFEFCA5FC9FFCE2737B" box="[810,829,807,832]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="196">R</collectionCode>
|
||
.
|
||
<collectionCode id="76B77A98A523FFFEFCCEFC9FFC86737B" box="[833,857,807,832]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="11" pageNumber="196" type="Herbarium">N</collectionCode>
|
||
. Philbrick &
|
||
<collectionCode id="76B77A98A523FFFEFC60FC9FFBD4737B" box="[1007,1035,807,832]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="11" pageNumber="196" type="Herbarium">M</collectionCode>
|
||
.
|
||
<collectionCode id="76B77A98A523FFFEFB80FC9FFBFD737B" box="[1039,1058,807,832]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="196">R</collectionCode>
|
||
. Benedict
|
||
<collectionCode id="76B77A98A523FFFEFB1AFC9EFB78737B" box="[1173,1191,806,832]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="11" pageNumber="196" type="Herbarium">B</collectionCode>
|
||
74-84
|
||
</emphasis>
|
||
,
|
||
<collectionCode id="76B77A98A523FFFEFB79FC9EFA9E737B" box="[1270,1345,806,832]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15722" lsid="urn:lsid:biocol.org:col:15722" name="Santa Barbara Botanic Garden" pageId="11" pageNumber="196" type="Herbarium">SBBG</collectionCode>
|
||
49613!
|
||
</paragraph>
|
||
<paragraph id="1019E25DA523FFFEFF32FCF3FD597406" blockId="11.[136,1452,230,1373]" pageId="11" pageNumber="196">
|
||
We find
|
||
<taxonomicName id="D7A699DEA523FFFEFE97FCF2FE3A735F" authorityName=", Johnston" authorityYear="1928" box="[280,485,842,868]" class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" kingdom="Plantae" order="Boraginales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="species" species="pondii">
|
||
<emphasis id="22D23E4FA523FFFEFE97FCF2FE3A735F" box="[280,485,842,868]" italics="true" pageId="11" pageNumber="196">Cryptantha pondii</emphasis>
|
||
</taxonomicName>
|
||
to be quite distinctive from other taxa in several features. It has linear bracts sporadically subtending peduncle bases of the cymules as well as flowers (
|
||
<figureCitation id="889DFED8A523FFFEFCB2FCD6FC0973B2" box="[829,982,878,905]" captionStart="FIGURE 6" captionStartId="12.[136,229,1831,1853]" captionTargetBox="[245,1343,196,1806]" captionTargetId="figure-19@12.[237,1350,190,1807]" captionTargetPageId="12" captionText="FIGURE 6. Cryptantha pondii. A. Close-up of single plant of presumed lectotype specimen (US41006), confirmed by Johnston (1928) as being correctly identified. Note linear, pustulate leaves and bracts at base of cymules. B,C. Images from Gentry 8684, 13 March 1949 (HCIB21969). B. Four nutlets removed from fruit, showing smooth surface and slight size heteromorphism. C. Close-up of cymules, showing linear bracts and relatively large corollas. D–F. Images from Philbrick B74-84, 25 March 1974 (SBBG49613). D. Fruit with four, smooth nutlets removed. Note finely hirsute calyx marginal region with ascending trichomes, as well as hispid calyx midrib of stouter trichomes. E. Close-up of four nutlets attached to gynobase. Note slight heteromorphism in size and slight extension of style tip/stigma from nutlet apices. F. Cymules, showing appressed-strigose vestiture of axes, scattered linear flower bracts, and relatively large corolla." figureDoi="http://doi.org/10.5281/zenodo.5426148" httpUri="https://zenodo.org/record/5426148/files/figure.png" pageId="11" pageNumber="196">Fig. 6A, C, F</figureCitation>
|
||
). The calyces are somewhat finely hirsute with ascending trichomes, described as “silky-hirsute” by
|
||
<bibRefCitation id="74379FACA523FFFEFC94FC2AFC227397" author="Johnston, I. M." box="[795,1021,914,940]" pageId="11" pageNumber="196" pagination="73 - 83" refId="ref13085" refString="Johnston, I. M. (1928) Studies in the Boraginaceae - VII. 2. Notes on various Boraginoideae. Contributions from the Gray Herbarium of Harvard University 81: 73 - 83." type="journal article" year="1928">Johnston (1928: 74)</bibRefCitation>
|
||
in addition to hispid along the midrib (
|
||
<figureCitation id="889DFED8A523FFFEFF1FFC0EFED873EA" box="[144,263,950,977]" captionStart="FIGURE 6" captionStartId="12.[136,229,1831,1853]" captionTargetBox="[245,1343,196,1806]" captionTargetId="figure-19@12.[237,1350,190,1807]" captionTargetPageId="12" captionText="FIGURE 6. Cryptantha pondii. A. Close-up of single plant of presumed lectotype specimen (US41006), confirmed by Johnston (1928) as being correctly identified. Note linear, pustulate leaves and bracts at base of cymules. B,C. Images from Gentry 8684, 13 March 1949 (HCIB21969). B. Four nutlets removed from fruit, showing smooth surface and slight size heteromorphism. C. Close-up of cymules, showing linear bracts and relatively large corollas. D–F. Images from Philbrick B74-84, 25 March 1974 (SBBG49613). D. Fruit with four, smooth nutlets removed. Note finely hirsute calyx marginal region with ascending trichomes, as well as hispid calyx midrib of stouter trichomes. E. Close-up of four nutlets attached to gynobase. Note slight heteromorphism in size and slight extension of style tip/stigma from nutlet apices. F. Cymules, showing appressed-strigose vestiture of axes, scattered linear flower bracts, and relatively large corolla." figureDoi="http://doi.org/10.5281/zenodo.5426148" httpUri="https://zenodo.org/record/5426148/files/figure.png" pageId="11" pageNumber="196">Fig. 6D, F</figureCitation>
|
||
). And, we confirm the observations of
|
||
<bibRefCitation id="74379FACA523FFFEFD39FC0EFC4A73EB" author="Johnston, I. M." box="[694,917,950,976]" pageId="11" pageNumber="196" pagination="73 - 83" refId="ref13085" refString="Johnston, I. M. (1928) Studies in the Boraginaceae - VII. 2. Notes on various Boraginoideae. Contributions from the Gray Herbarium of Harvard University 81: 73 - 83." type="journal article" year="1928">Johnston (1928: 74)</bibRefCitation>
|
||
that the corollas are relatively large (
|
||
<figureCitation id="889DFED8A523FFFEFABDFC0EFA7873EA" box="[1330,1447,950,977]" captionStart="FIGURE 6" captionStartId="12.[136,229,1831,1853]" captionTargetBox="[245,1343,196,1806]" captionTargetId="figure-19@12.[237,1350,190,1807]" captionTargetPageId="12" captionText="FIGURE 6. Cryptantha pondii. A. Close-up of single plant of presumed lectotype specimen (US41006), confirmed by Johnston (1928) as being correctly identified. Note linear, pustulate leaves and bracts at base of cymules. B,C. Images from Gentry 8684, 13 March 1949 (HCIB21969). B. Four nutlets removed from fruit, showing smooth surface and slight size heteromorphism. C. Close-up of cymules, showing linear bracts and relatively large corollas. D–F. Images from Philbrick B74-84, 25 March 1974 (SBBG49613). D. Fruit with four, smooth nutlets removed. Note finely hirsute calyx marginal region with ascending trichomes, as well as hispid calyx midrib of stouter trichomes. E. Close-up of four nutlets attached to gynobase. Note slight heteromorphism in size and slight extension of style tip/stigma from nutlet apices. F. Cymules, showing appressed-strigose vestiture of axes, scattered linear flower bracts, and relatively large corolla." figureDoi="http://doi.org/10.5281/zenodo.5426148" httpUri="https://zenodo.org/record/5426148/files/figure.png" pageId="11" pageNumber="196">Fig. 6C, F</figureCitation>
|
||
; described as
|
||
<quantity id="D75E4FB8A523FFFEFE97FC62FEAA73CE" box="[280,373,986,1013]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="11" pageNumber="196" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
|
||
broad by him, which we confirm here) and that there are four nutlets per fruit, these smooth and shiny and heteromorphic by size, with the odd nutlet axial (closest to the inflorescence axis) slightly larger than the other three consimilar nutlets (
|
||
<figureCitation id="889DFED8A523FFFEFE50FB9AFDA67406" box="[479,633,1058,1085]" captionStart="FIGURE 6" captionStartId="12.[136,229,1831,1853]" captionTargetBox="[245,1343,196,1806]" captionTargetId="figure-19@12.[237,1350,190,1807]" captionTargetPageId="12" captionText="FIGURE 6. Cryptantha pondii. A. Close-up of single plant of presumed lectotype specimen (US41006), confirmed by Johnston (1928) as being correctly identified. Note linear, pustulate leaves and bracts at base of cymules. B,C. Images from Gentry 8684, 13 March 1949 (HCIB21969). B. Four nutlets removed from fruit, showing smooth surface and slight size heteromorphism. C. Close-up of cymules, showing linear bracts and relatively large corollas. D–F. Images from Philbrick B74-84, 25 March 1974 (SBBG49613). D. Fruit with four, smooth nutlets removed. Note finely hirsute calyx marginal region with ascending trichomes, as well as hispid calyx midrib of stouter trichomes. E. Close-up of four nutlets attached to gynobase. Note slight heteromorphism in size and slight extension of style tip/stigma from nutlet apices. F. Cymules, showing appressed-strigose vestiture of axes, scattered linear flower bracts, and relatively large corolla." figureDoi="http://doi.org/10.5281/zenodo.5426148" httpUri="https://zenodo.org/record/5426148/files/figure.png" pageId="11" pageNumber="196">Fig. 6B, D, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1019E25DA523FFFEFF32FBFFFC2B7567" blockId="11.[136,1452,230,1373]" pageId="11" pageNumber="196">
|
||
We note that although
|
||
<bibRefCitation id="74379FACA523FFFEFE45FBFEFD5C745B" author="Johnston, I. M." box="[458,643,1094,1120]" pageId="11" pageNumber="196" pagination="73 - 83" refId="ref13085" refString="Johnston, I. M. (1928) Studies in the Boraginaceae - VII. 2. Notes on various Boraginoideae. Contributions from the Gray Herbarium of Harvard University 81: 73 - 83." type="journal article" year="1928">Johnston (1928)</bibRefCitation>
|
||
described the nutlets of
|
||
<taxonomicName id="D7A699DEA523FFFEFC26FBFFFBC9745B" authorityName=", Johnston" authorityYear="1928" box="[937,1046,1094,1121]" class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" kingdom="Plantae" order="Boraginales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="species" species="pondii">
|
||
<emphasis id="22D23E4FA523FFFEFC26FBFFFBC9745B" box="[937,1046,1094,1121]" italics="true" pageId="11" pageNumber="196">C. pondii</emphasis>
|
||
</taxonomicName>
|
||
as “smooth or very obscurely and minutely tuberculate above the middle”, we did not observe the latter sculpturing; all nutlets we observed were smooth and shiny. In addition, we were unable to observe specimens of
|
||
<taxonomicName id="D7A699DEA523FFFEFC5CFB37FBA07493" authorityName="Greene" authorityYear="1892" box="[979,1151,1166,1193]" class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" kingdom="Plantae" order="Boraginales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="species" species="bartolomaei">
|
||
<emphasis id="22D23E4FA523FFFEFC5CFB37FBA07493" box="[979,1151,1166,1193]" italics="true" pageId="11" pageNumber="196">C. bartolomaei</emphasis>
|
||
</taxonomicName>
|
||
, which Johnston believed to be synonymous with
|
||
<taxonomicName id="D7A699DEA523FFFEFE12FB0BFDD774F7" authorityName=", Johnston" authorityYear="1928" box="[413,520,1202,1229]" class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" kingdom="Plantae" order="Boraginales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="species" species="pondii">
|
||
<emphasis id="22D23E4FA523FFFEFE12FB0BFDD774F7" box="[413,520,1202,1229]" italics="true" pageId="11" pageNumber="196">C. pondii</emphasis>
|
||
</taxonomicName>
|
||
. In fact, the type of
|
||
<taxonomicName id="D7A699DEA523FFFEFD77FB0BFC7B74F7" authorityName="Greene" authorityYear="1892" box="[760,932,1202,1229]" class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" kingdom="Plantae" order="Boraginales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="species" species="bartolomaei">
|
||
<emphasis id="22D23E4FA523FFFEFD77FB0BFC7B74F7" box="[760,932,1202,1229]" italics="true" pageId="11" pageNumber="196">C. bartolomaei</emphasis>
|
||
</taxonomicName>
|
||
he believed was derived from the very same collection as that of
|
||
<taxonomicName id="D7A699DEA523FFFEFEFFFB6FFE0574CB" authorityName=", Johnston" authorityYear="1928" box="[368,474,1238,1265]" class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" kingdom="Plantae" order="Boraginales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="species" species="pondii">
|
||
<emphasis id="22D23E4FA523FFFEFEFFFB6FFE0574CB" box="[368,474,1238,1265]" italics="true" pageId="11" pageNumber="196">C. pondii</emphasis>
|
||
</taxonomicName>
|
||
. Online images of
|
||
<taxonomicName id="D7A699DEA523FFFEFD3AFB6FFCBF74CB" authorityName="Greene" authorityYear="1892" box="[693,864,1238,1265]" class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" kingdom="Plantae" order="Boraginales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="species" species="bartolomaei">
|
||
<emphasis id="22D23E4FA523FFFEFD3AFB6FFCBF74CB" box="[693,864,1238,1265]" italics="true" pageId="11" pageNumber="196">C. bartolomaei</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="CF1D5CFFA523FFFEFCFFFB6EFC0774CB" box="[880,984,1238,1264]" pageId="11" pageNumber="196" type="holotype">holotype</typeStatus>
|
||
:
|
||
<collectionCode id="76B77A98A523FFFEFC6BFB6EFBFC74CB" box="[996,1059,1238,1264]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15196" lsid="urn:lsid:biocol.org:col:15196" name="University of Notre Dame" pageId="11" pageNumber="196" type="Herbarium">NDG</collectionCode>
|
||
40169; isotypes:
|
||
<collectionCode id="76B77A98A523FFFEFB6DFB6EFAD574CB" box="[1250,1290,1238,1264]" country="USA" lsid="urn:lsid:biocol.org:col:15631" name="Harvard University - Gray Herbarium" pageId="11" pageNumber="196" type="Herbarium">GH</collectionCode>
|
||
00096299 and
|
||
<collectionCode id="76B77A98A523FFFEFF07FB42FF6E752F" box="[136,177,1274,1300]" country="USA" lsid="urn:lsid:biocol.org:col:15631" name="Harvard University - Gray Herbarium" pageId="11" pageNumber="196" type="Herbarium">GH</collectionCode>
|
||
00096293, the last apparently a fragment from the
|
||
<typeStatus id="CF1D5CFFA523FFFEFD77FB42FC83752E" box="[760,860,1274,1301]" pageId="11" pageNumber="196" type="holotype">holotype</typeStatus>
|
||
) do fit our concept of
|
||
<taxonomicName id="D7A699DEA523FFFEFBEEFB43FB14752F" authorityName=", Johnston" authorityYear="1928" box="[1121,1227,1274,1301]" class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" kingdom="Plantae" order="Boraginales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="species" species="pondii">
|
||
<emphasis id="22D23E4FA523FFFEFBEEFB43FB14752F" box="[1121,1227,1274,1301]" italics="true" pageId="11" pageNumber="196">C. pondii</emphasis>
|
||
</taxonomicName>
|
||
. However, the type specimens of
|
||
<taxonomicName id="D7A699DEA523FFFEFEADFAA7FE137503" authorityName="Greene" authorityYear="1892" box="[290,460,1310,1337]" class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" kingdom="Plantae" order="Boraginales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="species" species="bartolomaei">
|
||
<emphasis id="22D23E4FA523FFFEFEADFAA7FE137503" box="[290,460,1310,1337]" italics="true" pageId="11" pageNumber="196">C. bartolomaei</emphasis>
|
||
</taxonomicName>
|
||
need to be examined and verified as equivalent to
|
||
<taxonomicName id="D7A699DEA523FFFEFB8AFAA7FBB17503" authorityName=", Johnston" authorityYear="1928" box="[1029,1134,1310,1337]" class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" kingdom="Plantae" order="Boraginales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="species" species="pondii">
|
||
<emphasis id="22D23E4FA523FFFEFB8AFAA7FBB17503" box="[1029,1134,1310,1337]" italics="true" pageId="11" pageNumber="196">C. pondii</emphasis>
|
||
</taxonomicName>
|
||
, something we plan to do in the future. Note that
|
||
<taxonomicName id="D7A699DEA523FFFEFEFEFAFBFE067567" authorityName=", Johnston" authorityYear="1928" box="[369,473,1346,1373]" class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" kingdom="Plantae" order="Boraginales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="species" species="pondii">
|
||
<emphasis id="22D23E4FA523FFFEFEFEFAFBFE067567" box="[369,473,1346,1373]" italics="true" pageId="11" pageNumber="196">C. pondii</emphasis>
|
||
</taxonomicName>
|
||
has priority of publication over
|
||
<taxonomicName id="D7A699DEA523FFFEFCCCFAFBFC2E7567" authorityName="Greene" authorityYear="1892" box="[835,1009,1346,1373]" class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" kingdom="Plantae" order="Boraginales" pageId="11" pageNumber="196" phylum="Tracheophyta" rank="species" species="bartolomaei">
|
||
<emphasis id="22D23E4FA523FFFEFCCCFAFBFC897566" box="[835,854,1347,1373]" italics="true" pageId="11" pageNumber="196">C</emphasis>
|
||
.
|
||
<emphasis id="22D23E4FA523FFFEFCEBFAFAFC2E7567" box="[868,1009,1346,1372]" italics="true" pageId="11" pageNumber="196">bartolomaei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="22D23E4FA523FFFEFC7FFAFAFC2B7567" box="[1008,1012,1346,1372]" italics="true" pageId="11" pageNumber="196">.</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |